Scarabaeoidea – Scirtoidea – Dascilloidea – Buprestoidea – Byrrhoidea

Catalogue of Palaearctic Coleoptera
Volume 3

The titles published in this series are listed at brill.com/cpc

Odontotrypes (Odontotrypes) emei Král, Malý & Schneider, 2001

Scarabaeoidea – Scirtoidea –
Dascilloidea – Buprestoidea –
Byrrhoidea
Revised and Updated Edition
Volume 3

Edited by

I. Löbl
D. Löbl

LEIDEN | BOSTON

Cover illustration: Odontotrypes (Odontotrypes) emei Král, Malý & Schneider, 2001
Date of publication: 22 June 2016.
The index of the species-group names is available on site of the Koninklijke Brill NV, Leiden (http://dx.doi.org/10.6084/
m9.figshare.3187063).
The publication of the work was supported by the “Muséum de la Ville de Genève”, Geneva, Switzerland.

Library of Congress Cataloging-in-Publication Data
Names: Löbl, Ivan, editor. | Löbl, D. (Daniel), editor. Title: Catalogue of Palaearctic Coleoptera. Volume 3, Scarabaeoidea,
Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea / edited by I. Löbl, D. Löbl. Other titles: Scarabaeoidea, Scirtoidea,
Dascilloidea, Buprestoidea, Byrrhoidea Description: Revised and updated edition. | Leiden ; Boston : Brill, [2016] |
Includes bibliographical references and index. | Description based on print version record and CIP data provided by
publisher; resource not viewed.
Identifiers: LCCN 2016017450 (print) | LCCN 2016015134 (ebook) | ISBN 9789004309142 (E-book) | ISBN 9789004309135
(hardback : alk. paper) Subjects: LCSH: Beetles--Eurasia. | Beetles--Africa, North. Classification: LCC QL591.A1 (print) |
LCC QL591.A1 C38 2016 (ebook) | DDC 595.76--dc23
LC record available at https://lccn.loc.gov/2016017450

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Contents
Preface ix
Taxonomic Information xi
Distributional Information xiii
Bibliographic Information xix
Acknowledgements xxi
Editors and Authors xxii
New Nomenclatural and Taxonomic Acts, and Comments
Catalogue 33
Scarabaeoidea 33
Geotrupidae 33
Passalidae 52
Trogidae 53
Glaresidae 58
Lucanidae 58
Ochodaeidae 84
Hybosoridae 85
Glaphyridae 87
Scarabaeidae 97
Aegialiinae 97
Eremazinae 98
Aphodiinae 98
Aulonocneminae 166
Chironinae 166
Scarabaeinae 166
Dynamopodinae 208
Orphninae 209
Melolonthinae 210
Rutelinae 317
Dynastinae 358
Cetoniinae 367
Scirtoidea 412
Scirtidae 412
Decliniidae 425
Eucinetidae 425
Clambidae 426

1

viii

Contents
Dascilloidea 428
Dascillidae 428
Rhipiceridae 431
Buprestoidea 432
Buprestidae 432
Julodinae 432
Polycestinae 438
Galbellinae 456
Chrysochroinae 456
Buprestinae 494
Agrilinae 524
Byrrhoidea 574
Byrrhidae 574
Elmidae 591
Dryopidae 603
Limnichidae 607
Heteroceridae 610
Psephenidae 616
Ptilodactylidae 621
Chelonariidae 623
Eulichalidae 623
Callirhipidae 624

References 627
Index to family-group names 962
Index to genus-group names 965

Preface
An assessment of data is particularly challenging in popular groups that include high number of spectacular
taxa and many economically important species having also a major impact in ecosystems. This may be the
main reason why the previous edition of the Volume 3 of this Catalogue series that deals with such groups
has not fully responded to all expectations. Not all primary sources have been verified, and in a number of
cases the provided data were incomplete. Nevertheless, it was still a major advancement compared to currently available data sets that, no doubt, fulfilled its role and promoted new research. So, fourteen years since
the fixed limits for data entries in the first edition of this Catalogue volume, the present Revised and Updated
edition is expected to fill the gaps. It provides thousands of new data obtained by taxonomists and naturalists
all the world over.
As a by-product, the Catalogue demonstrates that taxonomy remains dynamic, and is certainly not a
dying discipline as outsiders sometimes believe. Taxonomists continue to describe species, build classifications, and specify knowledge about occurrences of taxa; and why taxonomists persist may be as mysterious
and complex as the origin of science. At this time the reality is that natural history research and biological
conservation are affected by policies and financial strategies just as much as social welfare. The science is in
flux, tracking changes in technology and paradigm shifts. Nowadays professional taxonomic jobs became
more fragile and can be lost as easily as jobs in basic industries. Many countries have experienced the contraction in organismal biology and if there is not a support of the taxonomic/natural history enterprise, then
jobs may go.
One way to survive the present boat ride is hope in a future systematics made possible by educating the
public about the importance of taxonomy as a centre for culture and conservation that directly helps our
species persist in a clean environment. There is a stronger advantage for supporting the need for taxonomic
research in places where beetle diversity has yet to be fully studied and where many widespread pests have
not reached. In many countries, students in taxonomy and other fields are integrating public education as
part of their routine activity, and this will promote taxonomic research.
Another way to overcome downturns is by sheer stoicism. In spite of some current trends, possibly also
due to inadequately informed decision makers and eventually resulting in quite Kafkaesque situations, we
should be resilient and recognize that taxonomy has survived some very hard blows, especially ideological
and sometimes from within their own ranks. One of them was the focus on population epiphenomena as
definers of species advanced by the New Synthesis, more recent are the barcoding movement and the
absurdist proposal to demolish the Linnaean Nomenclature by phylocoders.
Describing species is not necessarily high-tech (though can be) and can be done with basic equipment
and storage spaces. But, there are exciting new imaging techniques that may actually promote taxonomy,
especially for amateurs and in places like developing countries where researchers have little opportunity to
study type material because institutions will not loan material due to lending policies, shipping concerns or
lack of sufficient funds for postage (or, in the future, a curator to package loans). While high quality images
of whole bodies and external structures are commonplace, tomography imaging could be used for accurate
in situ rendering internal structures of type specimens available through cyber space. This type of technology
would facilitate the great taxonomic boom in Asia where there are researchers who have no access to foreign

x

Preface

type material, and the process reversed, where types are fully digitized, inside and out and available in the
Cloud.
Training the next generation of taxonomists should continue to balance the loss of expertise. The taxonomic impediment has been written about ad nauseam, a concern that began in earnest long ago with the
Species 2000 movement which led to an influx of funding and training grants in the United States. There are
also some steps in the EU, such as the Copenhagen Big4 project. This project is focused on student training
and by EU design it is a cooperative that involves commercial organizations, research institutions and universities. Thumbs up for this kind of activity and others such as the SYNTHESYS fund, also for EU nationals,
and those across political boundaries like the endowed Ernst Mayr Fund and the Mini-PEET Awards to
Enhance Transfer of Taxonomic Knowledge for students offered by the Society for Systematic Biology. We
need more of these initiatives, such as special resources through funding agencies that cross political
borders.
In summary, taxonomy as an essential field of natural history research will persist as long as there are
species to describe, synonyms to be established, and a bit of countryside remaining as a home to the beetles
which we adore and must be preserved for world’s health and for our own benefits.
Ivan Löbl & Rich Leschen

Taxonomic Information
The present Catalogue includes all available names, both valid and invalid (the synonyms of the family group
names excepted), of extant beetle taxa described before January 1, 2016 and known to occur in the Palaearctic
Region, as it is defined below. The classification is subservient to phylogeny, but in taxonomic praxis cannot
show all uncertitudes resulting from phylogenetic analyses. The higher classification largely follows that in
Bouchard & al. (2011) and Ślipiński & al. (2011). Notable divergences in Scarabaeoidea concern the Lucanidae
arranged following Huang & Chen (2010, 2013b), the Melolonthini, Rhizotrogini, Schizonychini, Leucopholini
and Heptophyllini being tribes within the Melolonthinae, and not subtribes of Melolonthini, and the
Osmodermatini which are likely basal within the Cetoniinae, unrelated to Trichiini (e.g., Micó & al. 2008).
As to other groups, the Scirtoidea are under Scirtiformia, as in Ślipiński & al. (2011).
With respect to present facilities in access to information, the editors adopted the policy that the gap
between publication date of the Catalogue and the date of the corresponding deadline entry should be as
short as possible. Therefore, also data published in 2015 are included though some may have remained
unknown to the respective authors. The present volume includes 22,520 valid taxa and their synonyms
(about 20,400 were listed in the 2006 edition).
Extinct taxa, names rejected by the ICZN (International Code of Zoological Nomenclature, Fourth
Edition. London: International Trust for Zoological Nomenclature, 1999), misspellings, misidentifications
and other nomina nuda are not included in the body of the Catalogue. However, concepts that are important
for nomenclatural purpose may be included, and if necessary, unavailable name may be discussed in the
chapter New Nomenclatural and Taxonomic acts, and Comments. Similarly, infrasubspecific names, such as
those established as “morpha”, “natio”, “race”, “subvariety” and “aberration”, or proposed as variety and
form of a subspecies or another variety, and names published, e.g., as A-us b-us c-us, but specified in the text
that they are actually proposed for a “natio” or “race”, etc., are not considered subspecific and are therefore
excluded from the Catalogue. Names proposed as varieties and forms before 1961 are included, if
deemed ­subspecific under the provisions of the ICZN, Article 45.6. Unjustified emendations are available
and included.
The currently valid names of the family-group taxa include the name of the author and the year of the
publication.
The names of the genus-group taxa are given with the name of the author, and the year and page of
publication. The page given is the page where the name and the actual description of the taxon is printed.
The type species of all genus-group names are given in their original combination. If the type species is
­currently regarded as a junior synonym, the valid senior synonym is given in brackets in its original
combination.
The names of the species-group taxa are given with the name of the author, and the year and page of
publication. The page given is the page where the name and the actual description of the taxon are printed.
In bi-languages Chinese/English and Japanese/English publications both respective pages of the actual
description may be given. For species-group taxa subsequently transferred to another genus, the name of the
original genus is given in parentheses, following the page of publication.

xii

Taxonomic Information

The following symbols, all given in square brackets following the page of publication, or the original
combination when applicable, are used for taxonomic information: HN for homonyms, eventually PHN for
primary homonyms and SHN for secondary homonyms, RN for replacement names, SN for substitute names,
NO for nomina oblita, NP for nomina protecta, DA for doubtful assignment, and EA for erroneous
assignment.
Taxa considered incertae sedis and nomina dubia are listed separately at the end of the nearest applicable
taxon.

Distributional Information
The limits of the Palaearctic region, as those of other biogeographical regions, are arbitrarily defined (Map 1).
For practical reasons, the boundaries of the Palaearctic Region, as they were established for the Catalogue
(see above), usually follow national boundaries. The region includes Europe, Africa north of the Sahara, and
Asia except for the part that is arbitrarily defined as belonging to the Oriental Region.
For each species and subspecies an outline of its present distribution is given. Fossil records are not considered. The information is given by means of symbols, presented at three levels.
The first level is the subdivision of the Palaearctic Region into three main parts, Europe (letter E, bold),
North Africa (letter N, bold) and Asia (letter A, bold).
Europe includes the Azores, Iceland and Turkey west of the Bosporus. The eastern boundaries are a matter of controversy. In the Catalogue, Europe includes Russia west of the main ridge of the Ural Mountains, the
Permsk Oblast, Bashkortostan Republic and Orenburskaya Oblast, and the small part of Kazakhstan west of
the Ural River. It includes the Caucasian republics of Georgia, Armenia and Azerbaijan. The south-eastern
boundaries are the political boundaries of the Asian part of Turkey, Iran, Kazakhstan, and the Caspian and
Black seas. Taxa known from “Russia” are in absence of detailed information given under the symbol RU. The
limits of Ukraine are as in the previous volumes of the Catalogue.
Former Yugoslavia (Serbia and Montenegro from 2003 to 2006) has been split into Serbia and Montenegro,
and Kosovo. The term Yugoslavia, listed in the body of the Catalogue under the symbol YU, is still used in
absence of more exact knowledge of the distribution of the respective taxa.
North Africa includes Morocco (incl. Western Sahara), Algeria, Tunisia, Libya and Egypt west of the Suez
Canal, and the Canary and Madeira islands.
Asia includes Sinai and the Arabian Peninsula (including Suqutra), Turkey east of the Bosporus, the
Middle East and Central Asian countries, Russia east of the main ridge of the Ural mountains, Korea, Japan
(including Ryukyu [= Nansei] Islands and the Japanese Pacific Islands), the entire People’s Republic of China,
Taiwan, Bhutan, Nepal, North India along the base of the Himalaya (Arunachal Pradesh, Uttarakhand
(= Uttaranchal), northwestern area of former Uttar Pradesh), Himachal Pradesh), Jammu & Kashmir and all
of Pakistan. Thus, India is the only state for which the strict political boundaries are not respected. Large
parts of Uttar Pradesh south of Nepal are overpopulated plains. Information on Coleoptera from this North
Indian state is based almost exclusively on its Himalayan districts lying west of Nepal, which are in the
­present state Uttarakhand.
The second level of the geographic information is provided by two-letter symbols for countries, major
areas of Russia and North Indian states, and by three-letter symbols for provinces of mainland China and for
Taiwan (see Table 1, Map 3).
The symbols are arranged in alphabetical order within the first-level subdivision. Some, usually older,
distributional records cannot be accommodated within the structure of the Catalogue (e.g., Arabia, Caucasus,
North India, Siberia). Such information is given in quotation marks (e.g., “Caucasus”) behind the last symbol
of the respective first-level symbol. Russia is subdivided into six major sub-regions (Map 2), each of which
has its own two-letter symbol. These, as well as the symbol RU for Russia, are used only when more detailed
information is not available. Similarly, the symbol CH for the People’s Republic of China, as well as the seven

xiv

Distributional Information

two-letter symbols for China’s major regions, are used only in the absence of more detailed geographical
information.
In general, the published distributional information is based on both identified material in collections
and on published records, scattered in an enormous number of taxonomic and faunistic papers that are
­virtually impossible to review in their entirety. Revisions of collections reveal a high proportion of misidentifications, attaining 40% of specimens in some large museums. The degree of identification reliability and
of the records derived from identifications is correlated to faunal diversity and quality of systematic revisions. Thus, the reliability in general increases from south to north and from poorly studied groups to “popular” groups. At present, a number of modern catalogues or check-lists, covering the beetle faunas of many
European and some extra-European countries or archipelagos, are available. The use of data contained in
these and other faunistic works is left to the discretion of the authors, who may also add unpublished information available to them. The second-level geographic information is not necessarily exhaustive, it should
rather be considered as a base for future faunal research.
The third-level geographic information concerns species and subspecies with restricted distribution.
Taxa of this category may be strict endemites, or taxa comparatively widely distributed in one area but
restricted in another area. For example, the distributional record of a species widely distributed in North
Africa with isolated occurrence on Pantelleria would appear as follows: E: IT (Pantelleria) N: AG MO TU. The
third-level information is facultative. It is given in parentheses after the respective second-level symbol. The
official language of the respective state is used for records in languages using the Latin alphabet, or it is
transliterated from the Cyrillic alphabet. Records in languages using non-Latin or Cyrillic characters (e.g.,
Chinese or Japanese pictographs) are translated into English, and the translated geographical terms are
spelled as closely as possible to those used in the Times Atlas®, or in other well-known sources. Detailed
geographical information may refer to natural geographical features such as islands, mountains, lakes, valleys, caves, or to administrative entities, such as districts.
The extralimital distribution of some Palaearctic species is indicated by three letter symbols in bold,
cosmopolitan species by the symbol COS, all located at the end of the respective geographical information
(see Table 1). Introductions are indicated by the letter “i” (e.g., Ei: GB). The extralimital regions for the needs
of the Catalogue are defined as follows (see Map 1): Nearctic (NAR): north of Mexico; Neotropical (NTR):
south of the United States; Afrotropical (AFR): south of the North African states included in the Palaearctic
Region; Oriental (ORR): areas south of the People’s Republic of China and Taiwan, areas south of the
Himalaya in India, the Philippines, Malaysia and Indonesia south to the Lydekker line; Australian (AUS):
south of the Lydekker line, and Pacific (PAC)

Distributional Information

map 1

The limits of the geographic Regions as defined for the purpose of this Catalogue

map 2

Subdivisions of Russia

xv

xvi

map 3

Table 1
E
AB
AL
AN
AR
AU
AZ
BE
BH
BU
BY
CR
CT
CZ
DE
EN
FA
FI

Distributional Information

Subdivisions and provinces of the People’s Republic of China

Geographical Symbols
Europe
Azerbaijan
Albania
Andorra
Armenia
Austria
Azores
Belgium
Bosnia Herzegovina
Bulgaria
Belarus
Croatia
Russia: Central European Territory
Czech Republic
Denmark
Estonia
Faeroe Islands
Finland

FR
GB
GE
GG
GR
HU
IC
IR
IT
KZ
LA
LS
LT
LU
MA
MC
MD
ME

France (incl. Corsica, Monaco)
Great Britain (incl. Channel Islands)
Germany
Georgia
Greece (incl. Crete)
Hungary
Iceland
Ireland
Italy (incl. Sardinia, Sicily, San Marino)
Kazakhstan
Latvia
Liechtenstein
Lithuania
Luxembourg
Malta
Macedonia
Moldova (Moldavia)
Montenegro

xvii

Distributional Information
NL
NR
NT
PL
PT
RO
RU
SB
SK
SL

The Netherlands
Norway
Russia: North European Territory
Poland
Portugal
Romania
Russia
Serbia
Slovakia
Slovenia

SP
SR
ST
SV
SZ
TR
UK
YU
KO

Spain (incl. Gibraltar)
Svalbard (Spitzbergen)
Russia: South European Territory
Sweden
Switzerland
Turkey
Ukraine
Serbia and Montenegro
Kosovo

N
AG
CI
EG

North Africa
Algeria
Canary Islands
Egypt

LB
MO
MR
TU

Libya
Morocco (incl. Western Sahara)
Madeira Archipelago
Tunisia

A
AE
AF
AP
BA
BT
CE
CH
CY
ES
FE
HP
IN
IQ
IS
JA
JO
KA
KI
KU
KZ
LE
MG
NE
NC

Asia
Arab Emirates
Afghanistan
India: Arunachal Pradesh
Bahrain
Bhutan
China: Central Territory
China
Cyprus
Russia: East Siberia
Russia: Far East
India: Himachal Pradesh
Iran
Iraq
Israel
Japan
Jordan
India: Kashmir
Kyrgyzstan
Kuwait
Kazakhstan
Lebanon
Mongolia
China: Northeast Territory
North Korea

NO
NP
NW
OM
PA
QA
RU
SA
SC
SD
SE

China: Northern Territory
Nepal
China: Northwest Territory
Oman
Pakistan
Qatar
Russia
Saudi Arabia
South Korea
India: Sikkim, Darjeeling District
China: Southeastern Territory (incl.
Macao, Hong Kong)
Egypt: Sinai
China: Southwestern Territory
Syria
Tajikistan
Turkmenistan
Turkey
India: Uttarakhand (Uttaranchal,
former part of Uttar Pradesh)
Uzbekistan
China: Western Plateau
Russia: West Siberia
Yemen (incl. Suqutra)

SI
SW
SY
TD
TM
TR
UP
UZ
WP
WS
YE

xviii
Table 1

Distributional Information
Geographical Symbols (cont.)

China: Provinces, Autonomous Regions or Municipalities, and Taiwan
ANH
BEI
CHQ
FUJ
GAN
GUA
GUI
GUX
HAI
HEB
HEI
HEN
HKG
HUB
HUN
JIA
JIL

Anhui (Anhwei)
Beijing (Peking or Peiping)
Chongqing
Fujian (Fukien)
Gansu (Kansu)
Guangdong (Kwantung)
Guizhou (Kweichow)
Guangxi (Kwangsi)
Hainan
Hebei (Hopeh)
Heilongjiang (Heilungkiang)
Henan (Honan)
Hong Kong
Hubei (Hupeh)
Hunan
Jiangsu (Kiangsu)
Jilin (Kirin)

JIX
LIA
MAC
NIN
NMO
QIN
SCH
SHA
SHG
SHN
SHX
TAI
TIA
XIN
XIZ
YUN
ZHE

Jiangxi (Kiangsi)
Liaoning
Macao
Ningxia (Ningsia)
Nei Mongol (Inner Mongolia)
Qinghai (Tsinghai)
Sichuan (Szechwan)
Shaanxi (Shensi)
Shanghai
Shandong (Shantung)
Shanxi (Shansi)
Taiwan (Formosa)
Tianjin (Tsientsin)
Xinjiang (Sinkiang)
Xizang (Tibet)
Yunnan
Zhejiang (Chekiang)

World Zoogeographical Regions
AFR
AUR
NAR
NTR
ORR

Afrotropical Region
Australian Region
Nearctic Region
Neotropical Region
Oriental Region

Unknown distribution

Several species described in absence of distributional information considered as possibly Palaearctic are
listed with appropriate note in square brackets.

Bibliographic Information
The total number of references in the present volume is 6,592. The references are given to primary sources of
the genus-group and species-group names included in the Catalogue, and for works supporting the New Acts
published in the present volume. No bibliographical reference is given to family-group names. Names of all
authors, if applicable, are given for each reference. An effort was put forth to unify the spellings of the names.
For names that appeared in publications in two or more different orthographies, the one used in the
Catalogue is that which was considered to be correct after thorough verification (e.g., the spelling Josef
Müller for the author who also published as Giuseppe Müller). In such cases the alternative spelling or initial
follows the original one in brackets. The original name is used for authors that subsequently adopted additional names, or honorary epithets (e.g., A. Semenov, and not A. Semenov-Tian-Shanskyi). Initials are given
for all authors, even if they are missing in the original publication. Names published in two or more alternative spellings, that seem to be equally correct, are spelled following Schmitt, Hübner & Gaedike (1998):
Nomina Auctorum. Auflösung von Abkürzungen taxonomischer Autoren-Namen. Nova Supplementa
Entomologica 11: 3–189. Names in Cyrillic are transliterated as for any other Cyrillic text, unless an alternative
spelling was preferred by the author, or the alternative name is generally used in the literature. It should be
noted that lower case prepositions such as French “de” and German “von” are not given with the author’s
name in the list of the taxa.
Titles of references are given in full, i.e. not abbreviated, with subtitles, as originally published. Text written in Cyrillic is transliterated. Titles in languages using non-Latin or non-Cyrillic characters are translated
into English and are presented in square brackets, followed by the indication of the original language in
parentheses.
Titles of periodicals are given in full, but without eventual subtitles and in single language if bilingual.
Series numbers are given in parentheses, and always precede the boldface volume number. The numbers of
separate issues are provided when they are paginated separately, in which case they are in parentheses following the volume number. In other cases they are facultative. Unnumbered pages are indicated in square
brackets. If descriptions appeared bilingual, the respective pages are separated by a slash. The titles of some
periodicals vary over the years. For these periodicals the titles are given as they appeared for the respective
items referred to (e.g., Coleopterologische Rundschau / Koleopterologische Rundschau). Contrarily, the journal
Entomologische Blätter had several titles and subtitles during its existence. Since there is no danger of confusion with any other periodical, the journal is cited simply as Entomologische Blätter until it changed to
Entomologische Blätter und Coleoptera. If the year of the volume differs from the actual year of publication,
the former follows the volume number and is given is square brackets. If periodicals have no volume number,
the year of the volume is given.
The Bulletin de la Société Entomologique de France is one of the more important periodicals in the field of
systematics. The title changed for more than 60 years, appearing usually as Bulletin des séances, Bulletin trimestriel or Bulletin entomologique. The first numbered volume, 37, was published 100 years after the creation
of the Annales and the Bulletin of the French Entomological Society in 1832. To avoid confusion, this periodical is consistently referred to as Bulletin de la Société Entomologique de France, and, from 1832 to 1931, is

xx

Bibliographic Information

i­ dentified by the originally given publication year, often preceding the true year of publication. From the year
1932, the volume number is given.
For ease of access, references of the same author and year were placed in order based on the first page
number of the paper, with Roman numerals being placed last, with respect to the number of the respective
issue. Books were placed first for a given year, unless a page range was cited, in which case the first-page rule
was followed. Papers of the same year and with the same first page numbers were placed in order of their last
page number. Thus the references are not arranged chronologically within each year.

Acknowledgements
The Editors and Authors of the present volume are indebted to following individuals who commented on
different parts of the Catalogue, pointed to nomenclatural problems, added distributional information,
­completed references, and helped in various ways: Miguel Alonso-Zarazaga, Madrid, Spain; Ming Bai,
Beijing, China; Daniele Baiocchi, Roma, Italy; Max Barclay, London, UK; Jan Bezděk, Brno, Czech
Republic, Svatopluk Bílý, Praha, Czech Republic; Patrice Bouchard, Ottawa, Canada; Yves Bousquet,
Ottawa, Canada; Hans Fery, Berlin, Germany; David Frank, Praha, Czech Republic; Stanislav Jákl, Praha,
Czech Republic; Manfred A. Jäch, Wien, Austria; Marek Kafka, Neratovice, Czech Republic; Tom Kwast,
Dresden, Germany; John F. Lawrence, Australia; Oxford, UK; Darren Mann, Oxford, UK; Richard A.B.
Leschen, Auckland, New Zealand; Kimio Masumoto, Tokyo, Japan; Hans Mühle, München, Germany;
Volker Puthz, Schlitz, Germany; Antonio Rey, Genova, Italy; Jian-Yue Qiu, Chongqing, China; Paul
Schoolmeesters, Herent, Belgium; Andrew B.T. Smith, Ottawa, Canada; Marco Uliana, Venezia, Italy;
Jiří C. Vávra, Ostrava, Czech Republic; Oscar Vorst, Utrecht, The Netherlands; Gao Xu, Chongqing, China;
Xing-Ke Yang, Beijing, China and Peter Zwick, Schlitz, Germany.
Richard Sehnal, Velenice, Czech Republic provided the illustration of Odontotrypes (Odontotrypes) emei
Král, Malý & Schneider, 2001.
Special thanks are due to Jacques Ayer, Muséum d’histoire naturelle, Geneva, Switzerland, who provided
support for the publication, and to Pascal Möschler, from the same institute, for encouragements.
The work of Jiří Hájek and Vítězslav Kubáň was partly supported by the Ministry of Culture of the Czech
Republic (DKRVO 2016/14, National Museum, 00023272).

Editors and Authors
The Editors
Ivan Löbl
Muséum d’histoire naturelle
Genève, Switzerland

Daniel Löbl
Veyrier, Switzerland

Coordinators of this Volume
Aleš Bezděk, Biology Centre CAS, Institute of Entomology, České Budějovice, Czech Republic: Scarabaeoidea
Vítĕzslav Kubáň, Department of Entomology, National Museum, Prague, Czech Republic: Buprestoidea
Ivan Löbl, Muséum d’histoire naturelle, Geneva, Switzerland: Scirtoidea, Dascilloidea, Byrrhoidea

Authors
superfamily SCARABAEOIDEA Latreille, 1802
family GEOTRUPIDAE Latreille, 1802
G. V. NIKOLAJEV, D. KRÁL, A. BEZDĚK
family PASSALIDAE Leach, 1815
M. KON, A. BEZDĚK
family TROGIDAE Macleay, 1819
R. PITTINO, A. BEZDĚK
family GLARESIDAE H.J. Kolbe, 1905
D. KRÁL, A. BEZDĚK
family LUCANIDAE Latreille, 1804	L. BARTOLOZZI, E. SPRECHER-UEBERSAX,
A. BEZDĚK
family OCHODAEIDAE Mulsant & Rey, 1871
G. V. NIKOLAJEV, A. BEZDĚK
family HYBOSORIDAE Erichson, 1847
A. BALLERIO, A. BEZDĚK
family GLAPHYRIDAE Macleay, 1819
M. NIKODÝM, A. BEZDĚK
family SCARABAEIDAE Latreille, 1802
subfamily Aegialiinae Laporte, 1840
A. BEZDĚK
subfamily Eremazinae Iablokoff-Khnzorian, 1977 A. BEZDĚK
subfamily Aphodiinae Leach, 1815
tribe Aphodiini Leach, 1815
M. DELLACASA, G. DELLACASA, D. KRÁL, A. BEZDĚK
tribe Corythoderini A. Schmidt, 1910
A. BEZDĚK
tribe Didactyliini Pittino, 1984
A. BEZDĚK
tribe Eupariini Schmidt, 1910
A. BEZDĚK
tribe Odochilini Rakovič, 1987
A. BEZDĚK
tribe Psammodiini Mulsant, 1942
M. RAKOVIČ, D. KRÁL, A. BEZDĚK

xxiii

Editors And Authors
tribe Rhyparini A. Schmidt, 1910
subfamily Aulonocneminae A. Janssens, 1946
subfamily Chironinae C.E. Blanchard, 1845
subfamily Scarabaeinae Latreille, 1802
tribe Ateuchini Laporte, 1840
tribe Canthonini van Lansberge, 1874
tribe Coprini Leach, 1815
tribe Gymnopleurini Lacordaire, 1856
tribe Oniticellini H. J. Kolbe, 1905
tribe Onitini Laporte, 1840
tribe Onthophagini Burmeister, 1846
tribe Scarabaeini Latreille, 1802
tribe Sisyphini Mulsant, 1842
subfamily Dynamopodinae Arrow, 1911
subfamily Orphninae Erichson, 1847
subfamily Melolonthinae Macleay, 1819
tribe Ablaberini Blanchard, 1850
tribe Chasmatopterini Lacordaire, 1856
tribe Diphycerini S. I. Medvedev, 1952
tribe Diplotaxini Burmeister, 1855
tribe Euchirini Hope, 1840
tribe Heptophyllini S.I. Medvedev, 1951
tribe Hopliini Latreille, 1829
tribe Leucopholini Burmeister, 1855
tribe Melolonthini Macleay, 1819
tribe Pachydemini Reitter, 1902
tribe Pachypodini Erichson, 1840
tribe Rhizotrogini Burmeister, 1855
tribe Schizonychini Burmeister, 1855
tribe Sericini Kirby, 1837
subfamily Rutelinae Macleay, 1819
subfamily Dynastinae Macleay, 1819
subfamily Cetoniinae Leach, 1815

A. BEZDĚK
D. KRÁL, A. BEZDĚK
D. KRÁL, A. BEZDĚK
D. KRÁL, A. BEZDĚK
D. KRÁL A. BEZDĚK
D. KRÁL, A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
S. ZIANI, A. BEZDĚK
D. KRÁL, A. BEZDĚK
A. BEZDĚK
D. KRÁL, A. BEZDĚK
A. BEZDĚK
D. AHRENS, A. BEZDĚK
T. BRANCO, A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
D. KRÁL, A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
D. AHRENS, A. BEZDĚK
C. ZORN, A. BEZDĚK
F.-T. KRELL, A. BEZDĚK
A. BEZDĚK

superfamily SCIRTOIDEA Fleming, 1821
family DECLINIIDAE Nikitsky et al., 1994
I. LÖBL
family EUCINETIDAE Lacordaire, 1857
I. LÖBL
family CLAMBIDAE Fischer von Waldheim, 1821
I. LÖBL
family SCIRTIDAE Fleming, 1821
B. KLAUSNITZER

xxiv

Editors And Authors

superfamily DASCILLOIDEA Guérin-Méneville, 1843
family DASCILLIDAE Guérin-Méneville, 1843
J. HÁJEK
family RHIPICERIDAE Latreille, 1834
J. HÁJEK
superfamily BUPRESTOIDEA Leach, 1815
family BUPRESTIDAE Leach, 1815
subfamily Julodinae Lacordaire, 1857
subfamily Polycestinae Lacordaire, 1857
subfamily Galbellinae Reitter, 1911
subfamily Chrysochroinae Laporte, 1835
tribe Chrysochroini Laporte, 1835
tribe Chalcophorini Lacordaire, 1857
tribe Dicercini Gistel, 1848
tribe Poecilonotini, Jakobson, 1913
tribe Paratassini Bílý & Volkovitsh, 1996
tribe Sphenopterini Lacordaire, 1857
subfamily Buprestinae Lacordaire, 1857
subfamily Agrilinae, except Agrilus and Aphanisticini
subfamily Agrilinae, genus Agrilus Curtis, 1825
tribe Aphanisticini Jacquelin du Val, 1859

V. KUBÁŇ
M.G. VOLKOVITSH
M.G. VOLKOVITSH
V. KUBÁŇ
V. KUBÁŇ
V. KUBÁŇ
V. KUBÁŇ
V. KUBÁŇ
M.Ju. KALASHIAN
V. KUBÁŇ
V. KUBÁŇ
E. JENDEK
M.Ju. KALASHIAN, V. KUBÁŇ

superfamily BYRRHOIDEA Latreille, 1804
family BYRRHIDAE Latreille, 1804
O. JÄGER, A. PÜTZ
family ELMIDAE Curtis, 1830
M.A. JÄCH, J. KODADA
family DRYOPIDAE Billberg, 1820
J. KODADA, M.A. JÄCH
family LIMNICHIDAE Erichson, 1846
C. HERNANDO, I. RIBERA
family HETEROCERIDAE Macleay, 1825
A. MASCAGNI
family PSEPHENIDAE Lacordaire, 1854
C.-F. LEE
family PTILODACTYLIDAE Laporte, 1836
J. HÁJEK
family CHELONARIIDAE C. E. Blanchard, 1845
J. HÁJEK
family EULICHADIDAE Crowson, 1973
J. HÁJEK
family CALLIRHIPIDAE Emden, 1924
J. HÁJEK

Addresses of the Authors
AHRENS Dirk, Museum A. Koenig, Adenaueralle 160, D-53113 Bonn, Germany. E-mail: d.ahrens.zfmk@unibonn.de
BALLERIO Alberto, Viale Venezia 45, I-25123 Brescia (BS), Italy. E-mail: alberto.ballerio.bs@numerica.it

Editors And Authors

xxv

BARTOLOZZI Luca, Museo Zoologico de “La Specola”, via Romana 17, I-50125 Firenze, Italy. E-mail:
luca.bartolozzi@unifi.it
BEZDĚK Aleš, Biology Centre ASCR, Institute of Entomology, Branišovská 31, CZ-370 05 České Budějovice,
Czech Republic. E-mail: bezdek@entu.cas.cz
BRANCO Tristao Valente, Rua de Camões 788 2 Dto, P-4000-142 Porto, Portugal. E-mail: tristao.branco
@gmail.com
DELLACASA Giovanni, Casella Postale 921, I-16100 Genova, Italy E-mail: dellacasag@alice.it
DELLACASA Marco, Università di Pisa. Centro interdipartimentale, Museo di Storia naturale e del Territorio,
Via Roma 79, I-56011 CALCI (PI) Italy. E-mail: dellacasa@museo.unipi.it
HÁJEK Jiří, National Museum, Department of Entomology, Cirkusová 1740, Praha 9 – Horní Počernice,
CZ-19300, Czech Republic. E-mail: jiri_hajek@nm. cz
HERNANDO Carles, P.O. Box 118, 08911 Badalona, Catalonia, Spain. E-mail: leptotyphlus@gmail.com
JÄCH Manfred A., 2. Zoologische Abteilung, Naturhistorisches Museum, Burgring 7, A-1014 Wien, Austria.
E-mail: manfred.jaech@nhm-wien.ac
JÄGER Olaf, Senckenberg Museum für Tierkunde, Abt. Entomologie, Königsbrücker Landstr. 159, D-01109
Dresden, Germany. E-mail: olaf.jaeger@senckenberg.de
JENDEK Eduard, Klincova 9, 82108 Bratislava, Slovakia; E-mail: jendeke@gmail.com
KALASHIAN Mark Jurievitsh, ul. Komitassa 42, kv. 48, 375051 Jerevan, Armenia. E-mail: mkalashian1
@gmail.com
KLAUSNITZER Bernhard, Lannerstrasse 5, D-01219 Dresden, Germany. E-mail: klausnitzer.col@t-online.de
KODADA Jan, Faculty of Natural Sciences, Comenius University, Ilkovičova 6, Mlynská dolina B-1, SK-842 15
Bratislava, Slovakia. E-mail: jankodada@hotmail.com
KON Masahiro, Pressamce Kyodai-Higashi 406, 116–3 Nishida-cho, Jodoji, Sakyo, Kyoto, 606–8417, Japan.
E-mail: dkblm500@kyoto.zaq.ne.jp
KRÁL David, Department of Zoology, Charles University, Viničná 7, 128 44 Praha 2, Czech Republic. E-mail:
kral@natur.cuni.cz
KRELL Frank-Thorsten, Department of Zoology, Denver Museum of Nature & Science, 2001 Colorado
Boulevard, Denver, CO 80205–5798 USA. E-mail: frank.krell@dmns.com
KUBÁŇ Vítĕzslav, National Museum, Department of Entomology, Cirkusová 1740, Praha 9 – Horní Počernice,
CZ-19300, Czech Republic. E-mails: vkuban@nm.cz, vkuban@volny.cz
LEE Chi-Feng, Applied Zoology Division, Taiwan Agricultural Research Institute, 189 Chung-Cheng Road,
Wufeng, Taichung 413, Taiwan. E-mail: chifeng38@yahoo.com.tw
LÖBL Ivan, Muséum d’histoire naturelle, Case postale 6434, CH 1211 Genève 6, Switzerland. E-mail:
ivan.lobl@ville-ge.ch
MASCAGNI Alessandro, Museo Zoologico de “La Specola”, via Romana 17, Via Romana 17, I-50125 Firenze,
Italy. E-mail: pcfmas@tin.it
NIKODÝM Milan, Wolkerova 1261, CZ-25263 Roztoky u Prahy, Czech Republic. E-mail: nikodym@cbox.cz
NIKOLAJEV Georgej V., Auezova 64/4 m.87, Almaty 050008, Kazakhstan. E-mail: nikolajevg@yahoo.com
PITTINO Riccardo, Via Zenon 10, I-20124 Milano, Italy. E-mail: riccardo.pittino@gmail.com
PÜTZ Andreas, Pohlitzer Strasse 1a, D-15890 Eisenhütenstadt, Germany. E-mail: byrrhus@aol.com

xxvi

Editors And Authors

RAKOVIČ Miloslav, 252 29 Dobřichovice, Czech Republic. E-mail: mrakovic@volny.cz
RIBERA Ignacio, Departamento de Biodiversidad y Biología Evolutiva, Museo Nacional de Ciencias Naturales,
José Gutiérrez Abascal 2, 28006 Madrid, Spain. E-mail: deronectes@gmx.net; i.ribera@mncn.csic.es
SPRECHER-UEBERSAX Eva, Naturhistorisches Museum, Augustinergasse 2, CH-4001 Basel, Switzerland.
E-mail: eva.sprecher@bs.ch
VOLKOVITSH Mark Gabrielevich, Zoological Institute, Russian Academy of Sciences, Universitetskaya
naberezhnaya 1, 199034 St. 199034 St. Petersburg, Russia. E-mails: polycest@zin.ru, acmaeodera@mail.ru
ZIANI Stefano, GeoLab, Via Case di Dozza 22, I-40026 Imola (BO), Italy. E-mail: stefanoziani@alice.it
ZORN Carsten, Suelzer Str. 52, D-17179 Gnoien, Germany. E-mail: czorn70@hotmail.com

The present work is based upon data compiled by the following authors of
the first edition of the Catalogue:
family LUCANIDAE Latreille, 1804
family PASSALIDAE Leach, 1815
family TROGIDAE Macleay, 1819
family GLARESIDAE H. J. Kolbe, 1905
family BOLBOCERATIDAE Mulsant, 1842
family GEOTRUPIDAE Latreille, 1802
subfamily Geotrupinae Latreille, 1802
subfamily Lethrinae Mulsant & Rey, 1871
family OCHODAEIDAE Mulsant & Rey, 1871
family CERATOCANTHIDAE Martínez, 1968
family HYBOSORIDAE Erichson, 1847
family GLAPHYRIDAE Macleay, 1819
family SCARABAEIDAE Latreille, 1802
subfamily Aegialiinae Laporte, 1840
subfamily Eremazinae Iablokoff-Khnzorian, 1977
subfamily Aphodiinae Leach, 1815
tribe Aphodiini Leach, 1815
tribe Corythoderini A. Schmidt, 1910
tribe Didactyliini Pittino, 1984
tribe Eupariini Schmidt, 1910
tribe Odochilini Rakovič, 1987
tribe Psammodiini Mulsant, 1942
tribe Rhyparini A. Schmidt, 1910
subfamily Aulonocneminae A. Janssens, 1946
subfamily Chironinae C. E. Blanchard, 1845
subfamily Scarabaeinae Latreille, 1802
tribe Ateuchini Laporte, 1840

L. BARTOLOZZI & E. SPRECHER
M. KON
R. PITTINO
D. KRÁL, I. LÖBL
D. KRÁL, I. LÖBL, G. V. NIKOLAJEV
I. LÖBL, G.V. NIKOLAJEV, D. KRÁL
D. KRÁL, G.V. NIKOLAJEV
J.I. LÓPEZ-COLÓN, I. LÖBL, G.V. NIKOLAJEV
A. BALLERIO
D. KRÁL, I. LÖBL
M. NIKODÝM, A. BEZDĚK
Z. T. STEBNICKA
Z. T. STEBNICKA
M. DELLACASA, G. DELLACASA
I. LÖBL
Z.T. STEBNICKA
Z.T. STEBNICKA
I. LÖBL
M. RAKOVIČ, D. KRÁL, I. LÖBL
I. LÖBL
D. KRÁL
D. KRÁL, I. LÖBL
D. KRÁL, I. LÖBL

xxvii

Editors And Authors
tribe Canthonini van Lansberge, 1874
tribe Coprini Leach, 1815
tribe Gymnopleurini Lacordaire, 1856
tribe Oniticellini H. J. Kolbe, 1905
tribe Onitiini Laporte, 1840
tribe Onthophagini Burmeister, 1846
tribe Scarabaeini Latreille, 1802
tribe Sisyphini Mulsant, 1842
subfamily Dynamopodinae Arrow, 1911
tribe Dynamopodini Arrow, 1911
tribe Thinorycterini Semenov & Reichardt, 1925
subfamily Orphninae Erichson, 1847
subfamily Pachypodinae Erichson, 1840
subfamily Eucheirinae Burmeister, 1842
subfamily Melolonthinae Macleay, 1819
tribe Chasmatopterini Lacordaire, 1856
tribe Diphycerini S. I. Medvedev, 1952
tribe Diplotaxini Burmeister, 1855
tribe Heptophyllini S. I. Medvedev, 1951
tribe Hopliini Latreille, 1829
tribe Leucopholini Burmeister, 1855
tribe Macrodactylini Burmeister, 1855
tribe Melolonthini Macleay, 1819
tribe Pachydemini Reitter, 1902
tribe Rhizotrogini Burmeister, 1855
tribe Schizonychini Burmeister, 1855
subfamily Sericinae Kirby, 1837
subfamily Rutelinae Macleay, 1819
tribe Adoretini Ohaus, 1912
tribe Anomalini C. E. Blanchard, 1851
tribe Rutelini Macleay, 1819
subfamily Dynastinae Macleay, 1819
subfamily Cetoniinae Leach, 1815
family DECLINIIDAE Nikitsky et al., 1994
family EUCINETIDAE Lacordaire, 1857
family CLAMBIDAE Fischer von Waldheim, 1821
family SCIRTIDAE Fleming, 1821
family DASCILLIDAE Guérin-Méneville, 1843
family RHIPICERIDAE Latreille, 1834
family BUPRESTIDAE Leach, 1815
subfamily Julodinae Lacordaire, 1857

D. KRÁL, I. LÖBL, F.-T. KRELL
I. LÖBL, F.-T. KRELL, D. KRÁL
A. BEZDĚK
A. BEZDĚK, F.-T. KRELL
A. BEZDĚK, F.-T. KRELL
I. LÖBL, F.-T. KRELL, S. ZIANI, D. KRÁL
I. LÖBL, F.-T. KRELL, D. KRÁL
A. BEZDĚK
D. KRÁL
Z.T. STEBNICKA
J.I. LÓPEZ-COLÓN
D. KRÁL, I. LÖBL
A. BEZDĚK
T. BRANCO
A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
A. SMETANA
A. BEZDĚK
A. BEZDĚK
A. BEZDĚK
D. KRÁL, A. SMETANA
A. SMETANA, D. KRÁL
A. BEZDĚK
D. AHRENS
D. KRÁL, A. SMETANA
C. ZORN
C. ZORN
F.-T. KRELL
A. SMETANA
I. LÖBL
S. VÍT
I. LÖBL
B. KLAUSNITZER
I. LÖBL
C. WURST
V. KUBÁŇ, M.G. VOLKOVITSH

xxviii

Editors And Authors

subfamily Polycestinae Lacordaire, 1857
M.G. VOLKOVITSH
subfamily Chrysochroinae Laporte, 1835
tribe Chrysochroini Laporte, 1835
V. KUBÁŇ
tribe Chalcophorini Lacordaire, 1857
V. KUBÁŇ
tribe Dicercini Gistel, 1848
V. KUBÁŇ
tribe Poecilonotini, Jakobson, 1913
V. KUBÁŇ
tribe Paratassini Bílý & Volkovitsh, 1996
S. BÍLÝ
tribe Sphenopterini Lacordaire, 1857	M.G. VOLKOVITSH, M.Ju.
KALASHIAN
subfamily Galbellinae Reitter, 1911	S. BÍLÝ, M.G. VOLKOVITSH,
V. KUBÁŇ
subfamily Buprestinae Lacordaire, 1857 (except Anthaxiini)
V. KUBÁŇ
tribe Anthaxiini Gory & Laporte, 1837
S. BÍLÝ
subfamily Agrilinae Laporte, 1835 (except Agrilus and Aphanisticina) V. KUBÁŇ
subfamily Agrilinae, genus Agrilus Curtis, 1825
E. JENDEK
subtribe Aphanisticina Jacquelin du Val, 1859
V. KUBÁŇ, M.Ju. KALASHIAN
family BYRRHIDAE Latreille, 1804
O. JÄGER, A. PÜTZ
family ELMIDAE Curtis, 1830	M.A. JÄCH, J. KODADA,
F. ČIAMPOR
family DRYOPIDAE Billberg, 1820
J. KODADA, M.A. JÄCH
family LIMNICHIDAE Erichson, 1846
C. HERNANDO, I. RIBERA
family HETEROCERIDAE Macleay, 1825
A. MASCAGNI
family PSEPHENIDAE Lacordaire, 1854	M.A. JÄCH, M.-L. JENG, C.-F. LEE,
M. SATÔ
family PTILODACTYLIDAE Laporte, 1836
M. SATÔ
family CHELONARIIDAE C. E. Blanchard, 1845
M. SATÔ
family EULICHADIDAE Crowson, 1973
M. JÄCH, J. HÁJEK
family CALLIRHIPIDAE Emden, 1924
J. HÁJEK

New Nomenclatural and Taxonomic Acts,
and Comments
Geotrupidae: Geotrupinae
D. Král
Resurrection and change of rank
Odontotrypes (Thorectomimus) semenowi nikodymi Král, Malý & Schneider, 2001 is resurrected from synonymy of O. (T.) semenowi (Reitter, 1887) and considered a valid subspecies of O. (T.) semenowi based on
study of the types and numerous additional material of both taxa.
Trogidae: Troginae
R. Pittino
New synonymy
Trox alpigenus Y.-W. Zhang, 1988, syn. nov. of Trox lama Pittino, 1985 based on illustrations in the original
description of T. alpigenus and study of the type and additional material of T. lama.
New subgeneric assignments
According to Scholtz (1986), almost all Afrotropical species of Trox species, including the two which also
occur in the Palaearctic region, are to be placed in Phoberus Macleay, 1819:
Trox (Phoberus) puncticollis Haaf, 1953, from Trox (Trox)
Trox (Phoberus) squamiger Rooth, 1851, from Trox (Trox)
Lucanidae: Lucaninae
L. Bartolozzi, E. Sprecher-Uebersax, A. Bezděk
New combinations
Dorcus apatani (Okuda & Maeda, 2015), comb. nov. from Digonophorus
Dorcus dungkharpai (Okuda & Maeda, 2015), comb. nov. from Digonophorus
Dorcus imrani (Okuda & Maeda, 2015), comb. nov. from Digonophorus
Eolucanus gracilis sharchokpai (Okuda & Maeda, 2014), comb. nov. from Lucanus
Hybosoridae: Hybosorinae
A. Ballerio
Hybosorus roei Westwood, 1846 and H. illigeri Reiche, 1853
Due the unsatisfactory taxonomy involving Hybosorus illigeri Reiche, 1853 and H. roei Westwood, 1846 (having
compelled the ICZN (2009: Opinion 2230) to reject an application purported to conserve the name
H. illigeri), the nomenclatural situation is still unclear. The names have not been added to the Official List
of Specific Names in Zoology or on the Official Index of Rejected or Invalid Specific Names in Zoology, so
the matter is still open to discussion. I have opted for using H. illigeri, since that name had been by far in
prevailing usage (I am not aware of any usage of H. roei as a valid name published after the Opinion).
A new application is, however, in preparation in order to clarify the matter once for all.
© koninklijke brill nv, leiden, ���6 | doi 10.1163/9789004309142_002

2

New Nomenclatural And Taxonomic Acts, And Comments
Scarabaeidae: Aegialiinae
A. Bezděk

New combinations
Psammoporus matalini (Gusakov, 2003), comb. nov. from Aegialia
Psammoporus sibiricus (Stebnicka, 1977), comb. nov. from Aegialia
Scarabaeidae: Aphodiinae: Aphodiini
M. Dellacasa, G. Dellacasa, D. Král & A. Bezděk
Changes in rank and new combinations
According to the generic treatment proposed for Aphodiini by G. Dellacasa et al. (2001), Chittius Tarasov,
2008, and Nipponoagoliinus Ochi & Kawahara, 2001 are e­ levated to genus rank and following new combinations are established:
Acanthobodilus languidulus (A. Schmidt, 1916), comb. nov. from Aphodius
Acrossus angustiarum (Balthasar, 1967), comb. nov. from Aphodius
Acrossus atratus (C. O. Waterhouse, 1875), comb. nov. from Aphodius
Acrossus belousovi (Frolov, 2001), comb. nov. from Aphodius
Acrossus binaevulus (Heyden, 1887), comb. nov. from Aphodius
Acrossus carpetanus (Graëlls, 1847), comb. nov. from Aphodius
Acrossus delavayi (W. Koshantschikov, 1916), comb. nov. from Aphodius
Acrossus devabhumi (Mittal, 1993), comb. nov. from Aphodius
Acrossus eberti (Balthasar, 1965), comb. nov. from Aphodius
Acrossus emodus (Petrovitz, 1976), comb. nov. from Aphodius
Acrossus formosanus (Nomura & Nakane, 1951), comb. nov. from Aphodius
Acrossus haliki (Balthasar, 1933), comb. nov. from Aphodius
Acrossus histrio (Balthasar, 1932), comb. nov. from Aphodius
Acrossus hrubanti (Balthasar, 1961), comb. nov. from Aphodius
Acrossus humerospinosus (Petrovitz, 1958), comb. nov. from Aphodius
Acrossus igai (Nakane, 1956), comb. nov. from Aphodius
Acrossus japonicus (Nomura & Nakane, 1951), comb. nov. from Aphodius
Acrossus jedlickai (Balthasar, 1932), comb. nov. from Aphodius
Acrossus jubingensis (Balthasar, 1967), comb. nov. from Aphodius
Acrossus klickai (Balthasar, 1932), comb. nov. from Aphodius
Acrossus koreanensis (J. I. Kim, 1986), comb. nov. from Aphodius
Acrossus kwanhsiensis (Balthasar, 1945), comb. nov. from Aphodius
Acrossus opacipennis (A. Schmidt, 1910), comb. nov. from Aphodius
Acrossus planicollis (Reitter, 1890), comb. nov. from Aphodius
Acrossus ritsemae (A. Schmidt, 1909), comb. nov. from Aphodius
Acrossus semiopacus (Reitter, 1887), comb. nov. from Aphodius
Acrossus superatratus (Nomura & Nakane, 1951), comb. nov. from Aphodius
Acrossus unifasciatus (Nomura & Nakane, 1951), comb. nov. from Aphodius
Acrossus ustulatus (Harold, 1862), comb. nov. from Aphodius

New Nomenclatural And Taxonomic Acts, And Comments
Aganocrossus pallidicornis (Walker, 1858), comb. nov. from Aphodius
Aganocrossus vejdovskyi (Balthasar, 1945), comb. nov. from Aphodius
Agoliinus amurensis (Iablokoff-Khnzorian, 1972), comb. nov. from Aphodius
Agoliinus kiuchii (Masumoto, 1984), comb. nov. from Aphodius
Agoliinus morii (Nakane, 1983), comb. nov. from Aphodius
Agoliinus pittinoi (Carpaneto, 1986), comb. nov. from Aphodius
Agoliinus setchan (Masumoto, 1984), comb. nov. from Aphodius
Agoliinus shibatai (Nakane, 1983), comb. nov. from Aphodius
Agoliinus tanakai (Masumoto, 1981), comb. nov. from Aphodius
Agolius bonvouloiri bonvouloiri (Harold, 1860), comb. nov. from Aphodius
Agolius bonvouloiri cantabricus (Pittino, 1981), comb. nov. from Aphodius
Agrilinus bardus (Balthasar, 1946), comb. nov. from Aphodius
Agrilinus bollowi (Balthasar, 1941), comb. nov. from Aphodius
Agrilinus breviusculus (Motschulsky, 1866), comb. nov. from Calamosternus
Agrilinus hasegawai (Nomura & Nakane, 1951), comb. nov. from Aphodius
Agrilinus ishidai (Masumoto & Kiuchi, 1987), comb. nov. from Aphodius
Agrilinus lindbergi (Petrovitz, 1959), comb. nov. from Aphodius
Agrilinus madara (Nakane, 1960), comb. nov. from Aphodius
Agrilinus monicae (Stebnicka, 1982), comb. nov. from Aphodius
Agrilinus montisamator (Balthasar, 1965), comb. nov. from Aphodius
Agrilinus obliviosus (Reitter, 1892), comb. nov. from Aphodius
Agrilinus punctator (Reitter, 1892), comb. nov. from Aphodius
Agrilinus ritsukoae (Kawai, 2004), comb. nov. from Aphodius
Agrilinus shukronajevi (Nikolajev, 1998), comb. nov. from Aphodius
Agrilinus spinulosus (A. Schmidt, 1910), comb. nov. from Aphodius
Agrilinus striatus (A. Schmidt, 1910), comb. nov. from Aphodius
Agrilinus tashigaonae (Emberson & Stebnicka, 2001), comb. nov. from Aphodius
Agrilinus tenax (Balthasar, 1932), comb. nov. from Aphodius
Agrilinus uniformis (C. O. Waterhouse, 1875), comb. nov. from Aphodius
Alocoderus elongatulus (Fabricius, 1801), comb. nov. from Aphodius
Alocoderus holdereri (Reitter, 1900), comb. nov. from Aphodius
Alocoderus machatschkei (Balthasar, 1960), comb. nov. from Aphodius
Alocoderus mineti (Clément, 1981), comb. nov. from Aphodius
Alocoderus strigimargo (Reitter, 1892), comb. nov. from Aphodius
Alocoderus teyrovskyi (Balthasar, 1935), comb. nov. from Aphodius
Alocoderus volubilis (Balthasar, 1961), comb. nov. from Aphodius
Alocoderus wolfgangi (Stebnicka, 1990), comb. nov. from Aphodius
Aparammoecius annapurnae (Stebnicka, 1986), comb. nov. from Aphodius
Aparammoecius bagmatiensis (Stebnicka, 1983), comb. nov. from Aphodius
Aparammoecius balangensis (Petrovitz, 1958), comb. nov. from Aphodius
Aparammoecius gansuensis (Stebnicka, 1997), comb. nov. from Aphodius

3

4

New Nomenclatural And Taxonomic Acts, And Comments

Aparammoecius heinzi (Pittino, 1997), comb. nov. from Aphodius
Aparammoecius holzschuhi (Pittino, 1997), comb. nov. from Aphodius
Aparammoecius ivani (Stebnicka, 1989), comb. nov. from Aphodius
Aparammoecius kanglae (Stebnicka, 1990), comb. nov. from Aphodius
Aparammoecius kaskiensis (D. Ahrens & Stebnicka, 1997), comb. nov. from Aphodius
Aparammoecius lamjungi (D. Ahrens & Stebnicka, 1997), comb. nov. from Aphodius
Aparammoecius mahriensis (Stebnicka, 1983), comb. nov. from Aphodius
Aparammoecius mizo (Nakane, 1967), comb. nov. from Aphodius
Aparammoecius mudukensis (Stebnicka, 1989), comb. nov. from Aphodius
Aparammoecius muzaffarabadensis (Pittino, 1997), comb. nov. from Aphodius
Aparammoecius pallidiligonis (C. O. Waterhouse, 1875), comb. nov. from Aphodius
Aparammoecius phulcokiensis (Stebnicka, 1986), comb. nov. from Aphodius
Aparammoecius sabhae (Stebnicka, 1990), comb. nov. from Aphodius
Aparammoecius schawalleri (Stebnicka, 1989), comb. nov. from Aphodius
Aparammoecius shogranicus (Pittino, 1997), comb. nov. from Aphodius
Aparammoecius sulcatus (Balthasar, 1953), comb. nov. from Aphodius
Aparammoecius tarokensis (Stebnicka, 1994), comb. nov. from Aphodius
Aparammoecius topali (Pittino, 1997), comb. nov. from Aphodius
Aparammoecius waniaramensis (Pittino, 1997), comb. nov. from Aphodius
Aparammoecius yangricus (Stebnicka, 1983), comb. nov. from Aphodius
Aparammoecius zojilae (Stebnicka, 1981), comb. nov. from Aphodius
Aphodaulacus koltzei (Reitter, 1892), comb. nov. from Aphodius
Aphodaulacus saraevi (Nikolajev, 2004), comb. nov. from Aphodius
Aphodobius amplipennis (Balthasar, 1945), comb. nov. from Aphodius
Bodilopsis sordida changajica (Endrődi, 1965), comb. nov. from Aphodius
Bodilus apiceopacus (Petrovitz, 1959), comb. nov. from Aphodius
Bodilus ardescens (A. Schmidt, 1916), comb. nov. from Aphodius
Bodilus arsissaensis (Petrovitz, 1968), comb. nov. from Aphodius
Bodilus crassus (A. Schmidt, 1916), comb. nov. from Aphodius
Bodilus furvus (A. Schmidt, 1916), comb. nov. from Aphodius
Bodilus hastatus (Reitter, 1892), comb. nov. from Aphodius
Bodilus insperatus (Petrovitz, 1967), comb. nov. from Aphodius
Bodilus kaznakovi (Frolov, 2001), comb. nov. from Aphodius
Bodilus kermanschahensis (Petrovitz, 1959), comb. nov. from Aphodius
Bodilus liesenfeldti (Petrovitz, 1958), comb. nov. from Aphodius
Bodilus longipennis (Rakovič, 1984), comb. nov. from Aphodius
Bodilus marani (Balthasar, 1929), comb. nov. from Aphodius
Bodilus nigriventris (Reitter, 1892), comb. nov. from Aphodius
Bodilus noxius (A. Schmidt, 1916), comb. nov. from Aphodius
Bodilus saoudi (R. Paulian, 1980), comb. nov. from Aphodius
Bodilus terminassianae (Kolov & Nikolajev, 2011), comb. nov. from Aphodius

New Nomenclatural And Taxonomic Acts, And Comments
Bodilus vartiani (Petrovitz, 1965), comb. nov. from Aphodius
Bodilus veselyi (Tesař, 1969), comb. nov. from Aphodius
Bodilus zarudnyi (Frolov, 2001), comb. nov. from Aphodius
Bodilus zunbajanicus (Stebnicka, 1975), comb. nov. from Aphodius
Boucomontiellus umbrinus (W. Koshantschikov, 1913), comb. nov. from Aphodius
Brachiaphodius ahrensi (Král, 2000), comb. nov. from Aphodius
Brachiaphodius babori (Balthasar, 1938), comb. nov. from Aphodius
Brachiaphodius eccoptus (Bates, 1889), comb. nov. from Aphodius
Brachiaphodius jendeki (Král, 2000), comb. nov. from Aphodius
Brachiaphodius nainiensis (Petrovitz, 1963), comb. nov. from Aphodius
Brachiaphodius pilifer (R. Paulian, 1934), comb. nov. from Aphodius
Brachiaphodius ruubpai (Masumoto, 1991), comb. nov. from Aphodius
Brachiaphodius taiwanicus (Petrovitz, 1976), comb. nov. from Aphodius
Brachiaphodius weibao (Král, 2000), comb. nov. from Aphodius
Brachiaphodius yuan (Král, 2000), comb. nov. from Aphodius
Calamosternus boreosinicus (Červenka, 1994), comb. nov. from Aphodius
Calamosternus machulkai (Balthasar, 1935), comb. nov. from Aphodius
Calamosternus prusai (Tesař, 1945), comb. nov. from Aphodius
Calamosternus pseudolucidus (Rakovič, 1977), comb. nov. from Aphodius
Calamosternus sauteri (Petrovitz, 1961), comb. nov. from Aphodius
Calamosternus spinosus (D. Koshantschikov, 1894), comb. nov. from Aphodius
Calamosternus srinagarensis (Pittino, 2001), comb. nov. from Aphodius
Calamosternus stercorarius (Mulsant & Rey, 1870), comb. nov. from Aphodius
Calamosternus vexator (Balthasar, 1933), comb. nov. from Aphodius
Calamosternus zmoskev (Červenka, 1994), comb. nov. from Aphodius
Carinaulus armandidavidi (Červenka, 2000), comb. nov. from Aphodius
Carinaulus coreensis (J. I. Kim, 1986), comb. nov. from Aphodius
Carinaulus dierli (Balthasar, 1967), comb. nov. from Aphodius
Carinaulus dundarum (Červenka, 2000), comb. nov. from Aphodius
Carinaulus gansuicus (Červenka, 2000), comb. nov. from Aphodius
Carinaulus imitabilis (Červenka, 2000), comb. nov. from Aphodius
Carinaulus inexspectatus (Balthasar, 1935), comb. nov. from Aphodius
Carinaulus jakli (Červenka, 2000), comb. nov. from Aphodius
Carinaulus krali (Červenka, 2000), comb. nov. from Aphodius
Carinaulus meghalayensis (Červenka, 2000), comb. nov. from Aphodius
Carinaulus milaninikodymi (Červenka, 2000), comb. nov. from Aphodius
Carinaulus pseudovseteckai (Červenka, 2003), comb. nov. from Aphodius
Carinaulus pseudozhanglaensis (Červenka, 2003), comb. nov. from Aphodius
Carinaulus pucholti (Balthasar, 1961), comb. nov. from Aphodius
Carinaulus tryznai (Červenka, 2000), comb. nov. from Aphodius
Carinaulus wrzecionkoi (Červenka, 2000), comb. nov. from Aphodius

5

6

New Nomenclatural And Taxonomic Acts, And Comments

Carinaulus zhanglaensis (Červenka, 2000), comb. nov. from Aphodius
Chilothorax albosetosus (Pittino, 1995), comb. nov. from Aphodius
Chilothorax alexis (Frolov, 2001), comb. nov. from Aphodius
Chilothorax aljibei (Blanco Villero, 1986), comb. nov. from Aphodius
Chilothorax altaicus (Nikolajev, 1984), comb. nov. from Aphodius
Chilothorax auliensis (Balthasar, 1938), comb. nov. from Aphodius
Chilothorax baghlanicus (Frolov, 1996), comb. nov. from Aphodius
Chilothorax bistriga (Reitter, 1900), comb. nov. from Aphodius
Chilothorax brancoi (Baraud, 1981), comb. nov. from Aphodius
Chilothorax chandmanicus (Frolov, 2001), comb. nov. from Aphodius
Chilothorax clausula (W. Koshantschikov, 1910), comb. nov. from Aphodius
Chilothorax discedens (A. Schmidt, 1907), comb. nov. from Aphodius
Chilothorax distinctus sevanicus (Rakovič, 1991), comb. nov. from Aphodius
Chilothorax dobrovljanskyi (W. Koshantschikov, 1913), comb. nov. from Aphodius
Chilothorax dzongensis (Petrovitz, 1976), comb. nov. from Aphodius
Chilothorax equitis (W. Koshantschikov, 1912), comb. nov. from Aphodius
Chilothorax figuratus (A. Schmidt, 1906), comb. nov. from Aphodius
Chilothorax fritschi (Balthasar, 1933), comb. nov. from Aphodius
Chilothorax glebi (Frolov, 2006), comb. nov. from Aphodius
Chilothorax grafi (Reitter, 1901), comb. nov. from Aphodius
Chilothorax huangyuanensis (Červenka, 1994), comb. nov. from Aphodius
Chilothorax hucklesbyi (R. Paulian, 1942), comb. nov. from Aphodius
Chilothorax jacobsoni (W. Koshantschikov, 1911), comb. nov. from Aphodius
Chilothorax kandaharicus (Balthasar, 1961), comb. nov. from Aphodius
Chilothorax kardonensis (R. Huang & Du, 1999), comb. nov. from Aphodius
Chilothorax kerzhneri (Nikolajev, 1984), comb. nov. from Aphodius
Chilothorax kukunorensis (Semenov, 1898), comb. nov. from Aphodius
Chilothorax lieni (Masumoto, Kiuchi & T.-C. Wang, 2014), comb. nov. from Aphodius
Chilothorax logunovi (Zinchenko, 2003), comb. nov. from Aphodius
Chilothorax mongolaltaicus (Nikolajev, 1984), comb. nov. from Aphodius
Chilothorax mossulensis (Mulsant & Godart, 1879), comb. nov. from Aphodius
Chilothorax naevuliger (Reitter, 1894), comb. nov. from Aphodius
Chilothorax ohishii (Masumoto, 1975), comb. nov. from Aphodius
Chilothorax okadai kiritachiensis (Ochi, Kawahara & Ohshio, 2012), comb. nov. from Aphodius
Chilothorax okadai okadai (Nakane, 1951), comb. nov. from Aphodius
Chilothorax pamirensis (S. I. Medvedev, 1930), comb. nov. from Aphodius
Chilothorax potanini (Frolov, 2001), comb. nov. from Aphodius
Chilothorax propola (Balthasar, 1946), comb. nov. from Aphodius
Chilothorax punctatus (C. O. Waterhouse, 1875), comb. nov. from Aphodius
Chilothorax shansianus (Nakane & Shirahata, 1957), comb. nov. from Aphodius
Chilothorax sinicus (Červenka, 1994), comb. nov. from Aphodius

New Nomenclatural And Taxonomic Acts, And Comments
Chilothorax subpolitus (Motschulsky, 1860), comb. nov. from Aphodius
Chilothorax tanhensis (Frolov, 2001), comb. nov. from Aphodius
Chilothorax tenuimanus (Sharp, 1878), comb. nov. from Aphodius
Chilothorax variipennis (A. Schmidt, 1916), comb. nov. from Aphodius
Chilothorax xanthellus (Frolov, 2002), comb. nov. from Aphodius
Chilothorax xanthopterus (Balthasar, 1938), comb. nov. from Aphodius
Chilothorax zaissanicus (Nikolajev, 1987), comb. nov. from Aphodius
Chittius anatolicus (Petrovitz, 1963), comb. nov. from Aphodius
Colobopterus indagator (Mannerheim, 1849), comb. nov. from Aphodius
Colobopterus notabilipennis (Petrovitz, 1972), comb. nov. from Aphodius
Colobopterus propraetor (Balthasar, 1932), comb. nov. from Aphodius
Colobopterus quadratus (Reiche, 1850), comb. nov. from Aphodius
Emadiellus jhelumensis (Rakovič, 1991), comb. nov. from Aphodius
Esymus alkani (Petrovitz, 1963), comb. nov. from Aphodius
Esymus filitarsis (Reitter, 1898), comb. nov. from Aphodius
Esymus finitimus (A. Schmidt, 1916), comb. nov. from Aphodius
Esymus helenaeliviae (G. Dellacasa & Pittino, 1985), comb. nov. from Aphodius
Esymus himalayanus (W. Koshantschikov, 1916), comb. nov. from Aphodius
Esymus hladili (Chromý, 1993), comb. nov. from Aphodius
Esymus hyalinus (Petrovitz, 1961), comb. nov. from Aphodius
Esymus indulgens (Balthasar, 1960), comb. nov. from Aphodius
Esymus ornatulus (Harold, 1866), comb. nov. from Aphodius
Esymus pokornyi (Chromý, 1993), comb. nov. from Aphodius
Esymus pyreti (Penecke, 1911), comb. nov. from Aphodius
Esymus sicardi (Reitter, 1892), comb. nov. from Aphodius
Esymus stribrnyi (Tesař, 1945), comb. nov. from Aphodius
Euorodalus longevittatus (A. Schmidt, 1916), comb. nov. from Aphodius
Euorodalus lucifer (D. Koshantschikov, 1894), comb. nov. from Aphodius
Euorodalus subtilitarsis (Petrovitz, 1963), comb. nov. from Aphodius
Eupleurus antiquus (Faldermann, 1835), comb. nov. from Aphodius
Gilletianus costatulus (A. Schmidt, 1908), comb. nov. from Aphodius
Koshantschikovius andrewesi (A. Schmidt, 1908), comb. nov. from Aphodius
Koshantschikovius chovdgolensis (Endrődi, 1983), comb. nov. from Aphodius
Koshantschikovius peculator (Balthasar, 1971), comb. nov. from Aphodius
Koshantschikovius schusteri (Balthasar, 1935), comb. nov. from Aphodius
Labarrus digitatus (Harold, 1871), comb. nov. from Aphodius
Labarrus hoabinhensis (Balthasar, 1946), comb. nov. from Aphodius
Labarrus kasyi (Petrovitz, 1965), comb. nov. from Aphodius
Labarrus nabeleki (Balthasar, 1936), comb. nov. from Aphodius
Labarrus nefandus (Reitter, 1907), comb. nov. from Aphodius
Labarrus paludani (Petrovitz, 1955), comb. nov. from Aphodius

7

8

New Nomenclatural And Taxonomic Acts, And Comments

Labarrus rugosocapita (Mittal, 1993), comb. nov. from Aphodius
Labarrus sublimbatus (Motschulsky, 1860), comb. nov. from Aphodius
Limarus hirtipennis (P. H. Lucas, 1846), comb. nov. from Aphodius
Loboparius drumonti (Rakovič & Mencl, 2010), comb. nov. from Aphodius
Loboparius dunchensis (Stebnicka, 1982), comb. nov. from Aphodius
Loboparius hazarae (Stebnicka, 1985), comb. nov. from Aphodius
Loboparius immarginatus cordieri (Boucomont, 1921), comb. nov. from Aphodius
Loboparius immarginatus immarginatus (A. Schmidt, 1907), comb. nov. from Aphodius
Loboparius immarginatus kumaoensis (Petrovitz, 1962), comb. nov. from Aphodius
Loboparius kotoensis (Miwa, 1930), comb. nov. from Aphodius
Loboparius mirificus (Balthasar, 1933), comb. nov. from Aphodius
Loboparius scheibei (Balthasar, 1956), comb. nov. from Aphodius
Loboparius schereri (Petrovitz, 1963), comb. nov. from Aphodius
Loraphodius corrugatus (Petrovitz, 1954), comb. nov. from Aphodius
Lunaphodius edgardi (Solsky, 1876), comb. nov. from Aphodius
Mecynodes alaiensis (Reitter, 1892), comb. nov. from Aphodius
Mecynodes anemurensis (Petrovitz, 1968), comb. nov. from Aphodius
Mecynodes angulosus (Harold, 1869), comb. nov. from Aphodius
Mecynodes asmaricus (Balthasar, 1956), comb. nov. from Aphodius
Mecynodes becvari (Červenka, 1995), comb. nov. from Aphodius
Mecynodes gilvipennis (Balthasar, 1946), comb. nov. from Aphodius
Mecynodes trochilus (Reitter, 1892), comb. nov. from Aphodius
Melinopterus abeillei (Sietti, 1903), comb. nov. from Aphodius
Melinopterus edithae (Reitter, 1906), comb. nov. from Aphodius
Melinopterus gissaricus (Akmetova & Frolov, 2012), comb. nov. from Aphodius
Melinopterus imamae (Stebnicka, 1988), comb. nov. from Aphodius
Melinopterus meuseli (Reitter, 1906), comb. nov. from Aphodius
Melinopterus scuticollis (Semenov, 1898), comb. nov. from Aphodius
Melinopterus sertavulensis (Pittino, 1988), comb. nov. from Aphodius
Melinopterus villarreali (Baraud, 1975), comb. nov. from Aphodius
Melinopterus wittmeri (Petrovitz, 1975), comb. nov. from Aphodius
Mendidaphodius angustatus (Klug, 1845), comb. nov. from Aphodius
Mendidaphodius fischeri (Petrovitz, 1962), comb. nov. from Aphodius
Mendidaphodius nodulifer (D. Koshantschikov, 1894), comb. nov. from Aphodius
Mendidaphodius palaestinensis (Petrovitz, 1963), comb. nov. from Aphodius
Mendidaphodius pseudoadustus (Petrovitz, 1958), comb. nov. from Aphodius
Mendidaphodius steinbergi (Nikritin, 1973), comb. nov. from Aphodius
Mendidius assectator (Balthasar, 1961), comb. nov. from Aphodius
Mendidius atricolor (Reitter, 1892), comb. nov. from Aphodius
Mendidius beluchistanicus (Petrovitz, 1962), comb. nov. from Aphodius
Mendidius bidens (Solsky, 1876), comb. nov. from Aphodius

New Nomenclatural And Taxonomic Acts, And Comments
Mendidius bivittatus (A. Schmidt, 1922), comb. nov. from Aphodius
Mendidius calliger (J. R. Sahlberg, 1908), comb. nov. from Aphodius
Mendidius diffidens (Reitter, 1892), comb. nov. from Aphodius
Mendidius endroedii (Balthasar, 1967), comb. nov. from Aphodius
Mendidius gobiensis (Endrődi, 1964), comb. nov. from Aphodius
Mendidius granulifrons (Fairmaire, 1883), comb. nov. from Aphodius
Mendidius heikertingeri (Balthasar, 1938), comb. nov. from Aphodius
Mendidius kabakovi (Nikritin, 1979), comb. nov. from Aphodius
Mendidius laevicollis (Harold, 1866), comb. nov. from Aphodius
Mendidius mamajevi (S. I. Medvedev, 1968), comb. nov. from Aphodius
Mendidius olexai (Král, 1996), comb. nov. from Aphodius
Mendidius osiris (W. Koshantschikov, 1916), comb. nov. from Aphodius
Mendidius palmetincolus (Karsch, 1881), comb. nov. from Aphodius
Mendidius pseudobidens (Balthasar, 1941), comb. nov. from Aphodius
Mendidius repetekensis (Nikritin, 1971), comb. nov. from Aphodius
Mendidius rutilinus (Reitter, 1892), comb. nov. from Aphodius
Mendidius samarcandicus (Nikritin, 1979), comb. nov. from Aphodius
Mendidius xerses (Petrovitz, 1963), comb. nov. from Aphodius
Mendidius zhantievi (Gusakov, 1997), comb. nov. from Aphodius
Mesontoplatys parvulus (Harold, 1871), comb. nov. from Aphodius
Neagolius dzhungaricus (Nikolajev & Frolov, 1997), comb. nov. from Aphodius
Neagolius haroldi (D. Koshantschikov, 1894), comb. nov. from Aphodius
Neagolius hengduanensis (Král, 1997), comb. nov. from Aphodius
Neagolius heydeni (Harold, 1871), comb. nov. from Aphodius
Neagolius interstitialis (D. Koshantschikov, 1894), comb. nov. from Aphodius
Neagolius montisjuldi (Pittino, 1988), comb. nov. from Aphodius
Neagolius moronensis (Frolov, 2009), comb. nov. from Aphodius
Neagolius orinus (W. Koshantschikov, 1912), comb. nov. from Aphodius
Neagolius ovitensis ovitensis (Pittino & Ballerio, 1994), comb. nov. from Aphodius
Neagolius petri (Frolov, 2014), comb. nov. from Aphodius
Neagolius savitskiyi (Gusakov, 2004), comb. nov. from Aphodius
Neagolius suschkini (W. Koshantschikov, 1913), comb. nov. from Aphodius
Neagolius takin (Král, 1995), comb. nov. from Aphodius
Neagolius zangbo (Král, 1997), comb. nov. from Aphodius
Nialus politus (Mulsant & Rey, 1870), comb. nov. from Aphodius
Nimbus harpagonis (Reitter, 1890), comb. nov. from Aphodius
Nimbus makowskyi (D. Koshantschikov, 1891), comb. nov. from Aphodius
Nipponoagoliinus yasutakai (Ochi & Kawahara, 2001), comb. nov. from Aphodius
Nobius bosaki (Tesař, 1945), comb. nov. from Aphodius
Nobius costatellus (A. Schmidt, 1916), comb. nov. from Aphodius
Nobius rhodiensis (Baraud, 1976), comb. nov. from Aphodius

9

10

New Nomenclatural And Taxonomic Acts, And Comments

Orodaliscus angulatulus (A. Schmidt, 1916), comb. nov. from Aphodius
Orodaliscus transaralicus (Nikolajev, 1983), comb. nov. from Aphodius
Paracrossidius kozlovi (W. Koshantschikov, 1916), comb. nov. from Aphodius
Paracrossidius laticapitus (Y.-W. Zhang, 1992), comb. nov. from Aphodius
Paracrossidius melanodiscus (Y.-W. Zhang, 1992), comb. nov. from Aphodius
Paracrossidius planipennis (Petrovitz, 1961), comb. nov. from Aphodius
Paracrossidius sichuanensis (Y.-W. Zhang, 1992), comb. nov. from Aphodius
Paracrossidius suvorovi (Kabakov, 1996), comb. nov. from Aphodius
Paracrossidius truculentus (Balthasar, 1963), comb. nov. from Aphodius
Paracrossidius transmontanus (Balthasar, 1932), comb. nov. from Aphodius
Paradidactylia biseriata (A. Schmidt, 1908), comb. nov. from Aphodius
Paradidactylia brivioi (Petrovitz, 1961), comb. nov. from Aphodius
Paradidactylia carinulata (Motschulsky, 1864), comb. nov. from Aphodius
Paradidactylia fastigata (A. Schmidt, 1908), comb. nov. from Aphodius
Paradidactylia haafi (Petrovitz, 1961), comb. nov. from Aphodius
Paradidactylia kathmandui (Stebnicka, 1985), comb. nov. from Aphodius
Paradidactylia shantungensis (Balthasar, 1941), comb. nov. from Aphodius
Paradidactylia wichei (Petrovitz, 1961), comb. nov. from Aphodius
Parammoecius amanicus (Stebnicka, 1978), comb. nov. from Aphodius
Parammoecius yamato (Nakane, 1960), comb. nov. from Aphodius
Paulianellus decoctor (Balthasar, 1965), comb. nov. from Aphodius
Paulianellus jacksoni (Petrovitz, 1975), comb. nov. from Aphodius
Paulianellus khandui (Stebnicka, 1980), comb. nov. from Aphodius
Paulianellus nepalensis (Balthasar, 1965), comb. nov. from Aphodius
Phaeaphodius fusculus (Reitter, 1892), comb. nov. from Aphodius
Phaeaphodius himalorectus (D. Ahrens & Stebnicka, 1997), comb. nov. from Aphodius
Phaeaphodius jouravliowi (Reitter, 1907), comb. nov. from Aphodius
Phaeaphodius kiulungensis (Balthasar, 1932), comb. nov. from Aphodius
Phaeaphodius pupillus (Balthasar, 1961), comb. nov. from Aphodius
Phaeaphodius roddi (W. Koshantschikov, 1911), comb. nov. from Aphodius
Phalacronothus ambulans (Petrovitz, 1971), comb. nov. from Aphodius
Phalacronothus botulus (Balthasar, 1945), comb. nov. from Aphodius
Phalacronothus flavolimbatus (Petrovitz, 1963), comb. nov. from Aphodius
Phalacronothus hilaris (Harold, 1869), comb. nov. from Aphodius
Phalacronothus pseudoavunculus (Masumoto, 1977), comb. nov. from Aphodius
Phalacronothus putoni (Reitter, 1894), comb. nov. from Aphodius
Phalacronothus semiangulus (Balthasar, 1952), comb. nov. from Aphodius
Pharaphodius aegyptiacus (Petrovitz, 1964), comb. nov. from Aphodius
Pharaphodius attritus (Balthasar, 1933), comb. nov. from Aphodius
Pharaphodius calo (Balthasar, 1971), comb. nov. from Aphodius
Pharaphodius campestris (Mittal, 1993), comb. nov. from Aphodius

New Nomenclatural And Taxonomic Acts, And Comments
Pharaphodius crenatus (Harold, 1862), comb. nov. from Aphodius
Pharaphodius jodhpurensis (Petrovitz, 1958), comb. nov. from Aphodius
Pharaphodius orientalis (Harold, 1862), comb. nov. from Aphodius
Pharaphodius parvatia (Mittal, 1993), comb. nov. from Aphodius
Pharaphodius pereirai (Balthasar, 1956), comb. nov. from Aphodius
Pharaphodius priscus (Motschulsky, 1858), comb. nov. from Aphodius
Pharaphodius putearius (Reitter, 1895), comb. nov. from Aphodius
Pharaphodius redargutus (Balthasar, 1971), comb. nov. from Aphodius
Pharaphodius robustus (Walker, 1858), comb. nov. from Aphodius
Pharaphodius rugosostriatus (C. O. Waterhouse, 1875), comb. nov. from Aphodius
Pharaphodius tenerus (Petrovitz, 1959), comb. nov. from Aphodius
Pharaphodius vernilis (A. Schmidt, 1911), comb. nov. from Aphodius
Plagiogonus bortkevitschi (W. Koshantschikov, 1916), comb. nov. from Aphodius
Plagiogonus burgaltaicus (Csiki, 1901), comb. nov. from Aphodius
Plagiogonus chan (Endrődi, 1967), comb. nov. from Aphodius
Plagiogonus culminarius (Reitter, 1900), comb. nov. from Aphodius
Plagiogonus farai (Balthasar, 1956), comb. nov. from Aphodius
Plagiogonus inflatus (Balthasar, 1938), comb. nov. from Aphodius
Plagiogonus martensi (Stebnicka, 1986), comb. nov. from Aphodius
Plagiogonus nanoides (Balthasar, 1961), comb. nov. from Aphodius
Plagiogonus oberthueri (R. Paulian, 1936), comb. nov. from Aphodius
Plagiogonus palea (Balthasar, 1967), comb. nov. from Aphodius
Plagiogonus quadriclypealis (Červenka, 1995), comb. nov. from Aphodius
Plagiogonus ramamiensis (Stebnicka, 1982), comb. nov. from Aphodius
Plagiogonus reitteri (D. Koshantschikov, 1894), comb. nov. from Aphodius
Plagiogonus shirahatai (Nakane, 1951), comb. nov. from Aphodius
Plagiogonus zobeideae (Petrovitz, 1980), comb. nov. from Aphodius
Planolinus nikolajevi (Berlov, 1989), comb. nov. from Aphodius
Planolinus rufoanalis (Petrovitz, 1961), comb. nov. from Aphodius
Platyderides jengi (Masumoto, Kiuchi & T.-C. Wang, 2014), comb. nov. from Aphodius
Platyderides klapperichi (Balthasar, 1942), comb. nov. from Aphodius
Pleuraphodius ilamensis (Stebnicka, 1990), comb. nov. from Aphodius
Pleuraphodius lewisi (C. O. Waterhouse, 1875), comb. nov. from Aphodius
Pseudacrossus absconditus (Balthasar, 1932), comb. nov. from Aphodius
Pseudacrossus caspius (Ménétriés, 1832), comb. nov. from Aphodius
Pseudacrossus grebenscikovi (Balthasar, 1961), comb. nov. from Aphodius
Pseudacrossus juxtus (Petrovitz, 1972), comb. nov. from Aphodius
Pseudacrossus kalabi (Král, 1997), comb. nov. from Aphodius
Pseudacrossus koshantschikovi (Jakobson, 1911), comb. nov. from Aphodius
Pseudacrossus kuskai (Stebnicka, 1982), comb. nov. from Aphodius
Pseudacrossus nasutus (Reitter, 1887), comb. nov. from Aphodius

11

12

New Nomenclatural And Taxonomic Acts, And Comments

Pseudacrossus przewalskyi (Reitter, 1887), comb. nov. from Aphodius
Pseudacrossus qinghaiensis (Král, 1997), comb. nov. from Aphodius
Pseudacrossus serrimargo (W. Koshantschikov, 1913), comb. nov. from Aphodius
Pseudacrossus smetanai (Král, 2011), comb. nov. from Aphodius
Pseudacrossus spalacophilus (Novikov, 1996), comb. nov. from Aphodius
Pseudacrossus subsericeus (Ballion, 1878), comb. nov. from Aphodius
Pseudacrossus wewalkai (Petrovitz, 1971), comb. nov. from Aphodius
Pseudacrossus zuercheri (Reitter, 1908), comb. nov. from Aphodius
Sinodiapterna gorodinskiyi (Gusakov, 2006), comb. nov. from Aphodius
Sinodiapterna hammondi (G. Dellacasa, 1986), comb. nov. from Aphodius
Sinodiapterna songrini (Stebnicka & Galante, 1992), comb. nov. from Aphodius
Sinodiapterna zeni (Ochi, 1991), comb. nov. from Aphodius
Sitiphus ammodites (Peyerimhoff, 1939), comb. nov. from Aphodius
Sitiphus sefrensis (Petrovitz, 1958), comb. nov. from Aphodius
Stenotothorax hibernalis hibernalis (Nakane & Tsukamoto, 1956), comb. nov. from Aphodius
Stenotothorax hibernalis saghalinensis (Nakane & Tsukamoto, 1956), comb. nov. from Aphodius
Subrinus clermonti (Reitter, 1907), comb. nov. from Aphodius
Subrinus feculentus (Fairmaire, 1892), comb. nov. from Aphodius
Subrinus motschulskyi (Nikritin, 1969), comb. nov. from Aphodius
Subrinus scintillius (Nikritin & Kabakov, 1979), comb. nov. from Aphodius
Subrinus subtilis (D. Koshantschikov, 1894), comb. nov. from Aphodius
Subrinus yemeniticus (Petrovitz, 1970), comb. nov. from Aphodius
Teuchestes analis (Fabricius, 1787), comb. nov. from Aphodius
Teuchestes brachysomus (Solsky, 1874), comb. nov. from Aphodius
Teuchestes sinofraternus (G. Dellacasa & Johnson, 1983), comb. nov. from Aphodius
Trichaphodius andreinii (Balthasar, 1939), comb. nov. from Aphodius
Trichaphodius hindustanicus (Balthasar, 1935), comb. nov. from Aphodius
Trichaphodius niloticus (Petrovitz, 1967), comb. nov. from Aphodius
Trichonotulus castetsi (R. Paulian, 1936), comb. nov. from Aphodius
Trichonotulus mongolicus (Mannerheim, 1852), comb. nov. from Aphodius
Scarabaeidae: Aphodiinae: Aphodiini
D. Král
New synonyms and combination
Aphodius virginalis Reitter, 1900, syn. nov. of Agrilinus obliviosus (Reitter, 1892), based on study of the type
material.
Aphodius inylchekensis Frolov, 2001, syn. nov. of Bodilus longipennis (Rakovič, 1984), based on study of the
type material.
Aphodius viturati Reitter, 1907, syn. nov. of Paracrossidius impressiusculus (Fairmaire, 1888), comb. nov. from
Aphodius, based on study of the type material.

New Nomenclatural And Taxonomic Acts, And Comments

13

Scarabaeidae: Aphodiinae: Aphodiini
H. Fery
Notes on the Aphodius fimetarius complex
Two species of the Aphodius fimetarius complex are treated in this catalogue under the names Aphodius
fimetarius (Linnaeus, 1758) and Aphodius pedellus (De Geer, 1774). The former has a mainly Mediterranean
distribution and does not occur in northern Europe and Russia. The latter species has a more northern
distribution, occurs among others in northern Europe and almost all parts of Russia. The nomenclature
of these species bases on results given in Wilson (2001) – which are incorrect in part – and on the
­succeeding Opinion 2345 of the ICZN (2014).
In contrast to Opinion 2345, Rössner (2012) and Fery & Rössner (2015) use and justify in detail another
­concept of the names of these two species: Aphodius fimetarius (Linnaeus, 1758) (=A. pedellus sensu
Wilson 2001) for the more northern species and Aphodius cardinalis Reitter, 1892 (=A. fimetarius sensu
Wilson 2001) for the more southern species. The users of this catalogue should feel free to make their own
decision about the names which they use in eventual publications or when labelling specimens, but do so
in an unambiguous naming of the two species.
Scarabaeidae: Aphodiinae: Psammodiini
M. Rakovič & D. Král
Resurrected name
Rhyssemodes nox Iablokoff-Khnzorian, 1971 is resurrected from synonymy with Granulopsammodius
t­ ranscaspicus (Petrovitz, 1961), based on the original description of Rhyssemodes nox and on the study of
the type material of both species.
Scarabaeidae: Aphodiinae: Psammodiini
A. Bezděk
Rhyssemina Pittino & Mariani, 1986 and Pleurophorina Mulsant, 1842
Usage of family-group name Rhyssemina Pittino & Mariani, 1986 (type genus Rhyssemus Mulsant, 1842) is
threatened by its senior subjective synonym Pleurophorina Mulsant, 1842 (type genus Pleurophorus
Mulsant, 1842). The name Pleurophorina has not been used since its proposal in 1842 until Ádám (1994a)
and Martynov & Shokhin (2014) mentioned it as valid. The name Pleurophorina was also listed by
Bouchard et al. (2011), but erroneously as subjective junior synonym of Psammodiina Mulsant, 1842.
A reversal of precedence (Art. 23.9) cannot be applied because the Article 23.9.1.1. of ICZN (1999) is met.
However, it is recommended that current usage (Rhyssemina as a valid name) should be maintained until
a case is made to the International Commission on Zoological Nomenclature and a ruling is published.
Scarabaeidae: Scarabaeinae: Oniticellini
A. Bezděk
Liatongus Reitter, 1892 and Onthosphaenus Motschulsky, 1860
Usage of the genus-group name Liatongus Reitter, 1892 (type species Onthophagus phanaeoides
Westwood, 1839 by subsequent designation of Arrow, 1931) is threatened by its senior subjective synonym
Onthosphaenus Motschulsky, 1860. The name Onthosphaenus has not been used since its proposal in 1860

14

New Nomenclatural And Taxonomic Acts, And Comments
until Paulian (1986: 214) designated Scarabaeus vertagus Fabricius, 1798 as the type species of
Onthosphaenus. Reversal of precedence (Art. 23.9) cannot be applied because Onthosphaenus was used as
valid after 1899 at least once by Paulian (1986). However, it is recommended that current usage (Liatongus
as a valid name) should be maintained until a case is made to the International Commission on Zoological
Nomenclature and a ruling is published.

Scarabaeidae: Scarabaeinae: Onthophagini
S. Ziani & A. Bezděk
Priority of Parentius Zunino, 1979 over Relictonthophagus Kabakov, 1979
The genus-group names Parentius Zunino 1979 and Relictonthophagus Kabakov, 1979, having the same type
species, are objective synonyms. Ziani (2001), appealing to the Art. 21 et seq of the International Code of
Zoological Nomenclature (1999), accorded the priority to Zunino’s name, published on July 5, over
Kabakov’s name, deemed as published on December 31. On the other hand, Kabakov & Shokhin (2014)
have considered the date printed at the end of the book where Kabakov’s paper is included, “подписано
к печати 16.VI.1979” [=signed to print 16.VI.1979] as its actual publication date and, for that reason, gave
priority to Kabakov’s name. Considering that the publication year for both papers is 1979, and that there
is a clear date printed on Zunino’s paper (July 5), that it is not possible to accept, as confirmed by other
Russian colleagues (e.g., V. K. Zinchenko, pers. comm.) that the lettering “подписано к печати” [=signed
to print] really means “published”, and that, for this reason, according to Art. 21.3.2 of the Code, the publication date to be adopted for Kabakov’s paper is December 31, we here confirm the priority of Parentius
Zunino, 1979 over Relictonthophagus Kabakov, 1979, as already stated by Ziani (2001).
New combination
Onthophagus (Parascatonomus) impar (Kabakov, 2006), comb. nov. from Pseudonthophagus
Scarabaeidae: Scarabaeinae: Onthophagini
M. Bai & X.-K. Yang
New replacement name
Onthophagus pseudoconvexicollis Bai & Yang, nom. nov. for Onthophagus sinicus Zhang & Wang, 1997 (non
Onthophagus sinicus Hope, 1842). This species epithet refers to the morphological similarity of
Onthophagus sinicus Zhang & Wang, 1997 and Onthophagus convexicollis Boheman, 1858.
Scarabaeidae: Sericinae
D. Ahrens
New synonymy
Serica pilosa (Nomura, 1971), syn. nov. of Serica inexspectata (Kontkanen, 1956)
After the examination of Kontkanen’s type specimens and a series from the type locality, Ahrens (2005)
synonymised S. inexspectata with S. nitidiceps (Nomura, 1971). Since Kobayashi (2009b), recognized the
synonymy of S. pilosa with S. nitidiceps, and given the earlier recognised synonymy of the latter with
S. inexspectata, S. pilosa has to be regarded a junior synonym of S. inexspectata too.

New Nomenclatural And Taxonomic Acts, And Comments

15

Scarabaeidae: Melolonthinae: Diphycerini
A. Bezděk
New tribal placement
In agreement with redefinition of melolonthine tribe Macrodactylini (Katovich, 2008), following genera are
transferred from Macrodactylini to Diphycerini: Dichelomorpha Burmeister, 1855; Metaceraspis Frey, 1962
and Xenoceraspis Arrow, 1920. Tribal placement of Hyperius Fairmaire, 1878 and Photyna Brenske, 1897 in
Diphycerini remains doubtful.
Scarabaeidae: Melolonthinae: Melolonthini
A. Bezděk
New replacement name
Carlschoenherria Bezděk, nom. nov. for Schoenherria Burmeister, 1855 (non Schoenherria Blanchard, 1853)
Scarabaeidae: Melolonthinae
A. Bezděk
Pachydemini Burmeister, 1855 versus Tanyproctini Erichson, 1847
The name used for this tribe of chafers has been inconsistent over the past few years. Pachydemini Burmeister,
1855 was in stable usage until Bouchard et al. (2011) resurrected the older but unused name Tanyproctini
Erichson, 1847 as a senior synonym (based on an inadvertent usage as a valid name by Smith (2006), and
not respecting the ICZN Article 35.5). Subsequent papers used either Tanyproctini (e.g., Král et al., 2012;
Mondaca & Ocampo, 2012; Neita & Ocampo, 2012; Sehnal, 2013a, 2014) or Pachydemini (e.g., Harrison,
2014; Chavanon & Labrique, 2013; Lacroix & Montreuil, 2014; Montreuil & Keith, 2013; Ziani et al., 2015).
Lacroix & Montreuil (2014) attempted to conserve the priority of Pachydemini over Tanyproctini using
Article 23.9 (ICZN, 1999). However, they overlooked the usage of Tanyproctini as a valid name by Smith
(2006), which invalids their action under Article 23.9.1.1.
The status of these names now falls under Article 23.10 (erroneous reversal of precedence), which
requests that the matter be referred to the International Commission on Zoological Nomenclature for
a ruling. With regard to long-term and conventional usage of Pachydemini, the name is used in the present Catalogue. It is also recommended that usage of Pachydemini as a valid name should be maintained
until a case is made to the International Commission on Zoological Nomenclature and a ruling is
published.
Scarabaeidae: Melolonthinae: Rhizotrogini
A. Bezděk
New combinations
Geotrogus cludtsi (Keith, 2005), comb. nov. from Pseudoapterogyna
Geotrogus euphytus lamantiai (Sparacio, 2014), comb. nov. from Pseudoapterogyna
Geotrogus michaelis (Sparacio, 2014), comb. nov. from Pseudoapterogyna
Miridiba (Miridiba) wangi (Y.-W. Zhang & Y. Li, 1997), comb. nov. from Holotrichia

16

New Nomenclatural And Taxonomic Acts, And Comments

Scarabaeidae: Dynastinae
Y. Bousquet, P. Bouchard & F.-T. Krell
Type species of Heteronychus Dejean, 1833
The correct type species of Heteronychus Dejean, 1833 is Scarabaeus syrichtus Fabricius, 1775, by subsequent
designation in Duponchel (1845), which had been overlooked until recently. Krell (2002) invalidly designated Geotrupes cricetus Hausmann, 1807 as the type species of Heteronychus and used the same species
as the type in the Catalogue of Palaearctic Coleoptera (Krell, 2006). Unfortunately, the valid type species
for Heteronychus, S. syrichtus, is currently placed in the genus Syrictes Prell, 1936 (see e.g., Endrődi, 1985).
Currently, a case is prepared for the International Commission on Zoological Nomenclature proposing
the conservation of the current taxonomic concepts of Heteronychus (with the type species G. cricetus)
and Syrictes. It is recommended to maintain this established usage until the International Commission on
Zoological Nomenclature has issued a ruling on this case.
Scarabaeidae: Dynastinae
F.-T. Krell
Availability of Pachypus truncatifrons Laporte, 1832, senior synonym of Calicnemis obesa (Erichson,
1841), syn. nov.
With the description of Calicnemis latreillii, Laporte (1832) introduced the name Pachypus truncatifrons as
a synonym. The latter name had been published by Dejean (1821) as a nomen nudum. According to
ICZN Art. 11.6. rules that a “name which when first published in […] was treated as a junior synonym
of a name then used as valid is not thereby made available.” But Art. 11.6.1. restricts this ruling:
“However, if such a name published as a junior synonym had been treated before 1961 as an available
name and either adopted as the name of a taxon of treated as a senior homonym, it is made available
thereby but dates from its first publication as a synonym.” Pachypus truncatifrons had been used as
valid by Guérin (1838d), Erichson (1841b), Ghiliani (1842), and, as Callicnemis truncatifrons, by Dufour
(1850). Verdugo & Drumont (2015) err in their assumption that Pachypus truncatifrons Laporte is a
nomen nudum on the grounds of Laporte having failed to provide a description. By listing Dejean’s
name as a synonym of the well described Calicnemis latreillii, the description of the latter refers to both
names. This description, combined with Guérin’s and Dufour’s usage, make Pachypus truncatifrons
Laporte, 1832, available.
Verdugo & Drumont (2015) assumed that the name was available from Dufour’s (1850) use of Callicnemis
truncatifrons and designated a lectotype for Callicnemis truncatifrons Dufour, 1850. This is a specimen
with a label from Dejean’s hand, therefore to be considered to be the species that Dejean had unavailably
named and that Laporte (1832) had cited as a synonym of his Calicnemis latreillii. Verdugo & Drumont
(2015) have identified this lectotype as Calicnemis obesa (Erichson, 1841), a name older than Dufour’s
Callicnemis truncatifrons, but younger than Laporte’s Pachypus truncatifrons. Therefore, the oldest available and valid name for Erichson’s species Colorhinus obesus is Pachypus truncatifrons Laporte, 1832, currently Calicnemis truncatifrons. Erichson himself (1841b) had stated this synonymy later in the very year
of the description of his species.

New Nomenclatural And Taxonomic Acts, And Comments

17

Scarabaeidae: Cetoniinae: Cetoniini
S. Jákl & A. Bezděk
Type species of Potosia Mulsant & Rey, 1870
The validly designated type species of Potosia Mulsant & Rey, 1870 is Scarabaeus speciosissimus Scopoli,
1786 – see Branco (2008), for details. However, it is recommended that current usage, i.e., type species
Cetonia floricola Herbst, 1790, should be maintained until a case is made to the International Commission
on Zoological Nomenclature and a ruling is published.
“Sinoliocola” Alexis & Delpont, 1998
Alexis & Delpont (1998) attempted to establish the genus Sinoliocola for four species: Cetonia brevitarsis
Lewis, 1879; Cetonia cathaica Bates, 1890; Cetonia speculifera Swartz, 1817; and Protaetia himalayana
Mikšić, 1987; without designating the type species (see also Smetana & Smith, 2006). Since no type species
was designated in the original publication, this name is unavailable. Later on, Devecis (2004a, b) described
two new species as Sinoliocola hainanensis Devecis, 2004 (from China: Hainan) and Sinoliocola pulverisata Devecis, 2004 (from China: Qinghai). These species are listed in the present Catalogue under Protaetia
(Liocola).
Comments to Cetoniinae described or recorded from China and based on material originating from
Li Jingke
Several cetoniine species were recently described or recorded on the material originating from Chinese
insect dealer Li Jingke and being labelled as collected in China. Unfortunately, part of this material was
mislabelled and in fact collected in Laos or in Thailand. Such newly described species based on Li Jingke’s
material should be very carefully revised in the future. Such problematic taxa are, e.g.: Campsiura
(Eucampsiura) lijingkei Legrand & Flutsch, 2007; Protaetia (Chrysoprotaetia) flutschi Legrand, 2005;
Protaetia (Dicranobia) potanini hainanensis Legrand, 2005.
Resurrected names
Eumimimetica Kraatz, 1881 (type species Cetonia terrosa Gory & Percheron, 1833) is resurrected from synonymy with Protaetia (subgenus Potosia Mulsant & Rey, 1870) and established as valid subgenus of Protaetia
Burmeister, 1842.
Protaetiophana Heller, 1896 (type species Protaetia excisithorax Heller, 1896) is resurrected from synonymy
with Protaetia (subgenus Potosia Mulsant & Rey, 1870) and established as valid subgenus of Protaetia
Burmeister, 1842.
New combinations
Protaetia (Chrysopotosia) mandschuriensis hainanensis (Devecis & Flutsch, 1915), comb. nov. from
Chrysopotosia
Protaetia (Netocia) subpilosa dorchini (Legrand & Drumont, 2006), comb. nov. from Netocia
Protaetia (Netocia) subpilosa iranica (Montreuil & Legrand, 2010), comb. nov. from Netocia

18

New Nomenclatural And Taxonomic Acts, And Comments

New synonymy
Oxythyrea cinctella orientalis Iablokoff-Khnzorian, 1967, syn. nov. of Oxythyrea cinctella (Schaum, 1841),
based on preliminary results of molecular phylogenetic study of the genus Oxythyrea (Šípek, pers.
comm.).
Scarabaeidae: Cetoniinae: Cetoniini
S. Jákl
New synonyms
Protaetia (Potosia) degrevei (Alexis & Delpont, 1998), syn. nov. of Protaetia (Potosia) multifoveolata (Reitter,
1899), based on study of type material and numerous additional material.
Glycyphana (Glycyphana) horsfieldii aurulenta Arrow, 1910; Glycyphana (Glycyphana) horsfieldii chinensis
Mikšić, 1967 and Glycyphana (Glycyphana) horsfieldii biargentata (Thomson, 1878), syn. nov. of
Glycyphana (Glycyphana) horsfieldii (Hope, 1831), based on study of part of type material and numerous
additional specimens across its geographic range.
Glycyphana (Glycyphaniola) nepalensis roquesi Bourgoin, 1916 and Glycyphana (Glycyphaniola) nepalensis fujianensis Krajčík, 2012, syn. nov. of Glycyphana (Glycyphaniola) nepalensis Kraatz, 1894, based
on study of part of type material and numerous additional specimens across its geographic range.
Change of rank
Glycyphana (Glycyphaniola) laotica Mikšić, 1968 is considered a valid species, based on study of numerous
specimens from continental Asia and the type material of Glycyphana nicobarica Janson, 1877.
Scarabaeidae: Cetoniinae: Cremastocheilini, Goliathini
S. Jákl
New synonyms
Clinterocera donckieri (Bourgoin, 1924), syn. nov. of Clinterocera davidis davidis (Fairmaire, 1878), based on
study of type material.
Clinterocera mandarina (Westwood, 1874), Clinterocera obsoleta (Fairmaire, 1878) and Clinterocera helleri
(Fairmaire, 1897), syn. nov. of Clinterocera scabrosa (Motschulsky, 1853), based on study of a part of the
type material and of numerous additional specimens across its distributional range.
Rhomborhina (Rhomborhina) bousqueti bousqueti Alexis & Delpont, 1998 and Rhomborhina (Rhomborhina)
bousqueti vietnamica Alexis & Delpont, 1998, syn. nov. of Rhomborhina (Rhomborhina) castanea Sakai,
1997, based on the study of type material.
Scarabaeidae: Cetoniinae: Trichiini
A. Bezděk
New combination
Indotrichius gorodinskii (Tauzin, 2013), comb. nov. from Paratrichius.

New Nomenclatural And Taxonomic Acts, And Comments

19

Dascillidae
J. Hájek
Comments
Jin et al. (2013) placed Dascillus aprutianus Depoli, 1936 as a doubtful synonym of Dascillus sicanus Fairmaire,
1861. This synonymy it taken over in the present Catalogue, though it was not formally established.
Scirtidae
B. Klausnitzer
Resurrection and new synonymy
Cyphon sasagawai Yoshitomi & Klausnitzer, 2002, was proposed as a replacement name for the homonymous
Cyphon intermedius Nakane, 1963. The latter has, however, two senior synonyms, C. ishiharai Sasagawa,
1985 and C. satoi Sasagawa, 1985. All are at present members of Herthania Klausnitzer, 2006; Herthania
satoi (Sasagawa, 1985) is resurrected from synonymy, while H. sasagawai (Yoshitomi & Klausnitzer, 2002)
is to be placed in its synonymy – syn. nov.
Comments
Elodes Latreille is traditionally treated as a feminine word. It is latinized from Greek ἑλώδης, −ες, meaning
marshy or frequenting marshes. Following the ICZN Art. 30.1.4.2, it is to be treated as masculine, since the
gender was originally not stated, either directly or by combination with an adjectival species name.
Buprestidae
V. Kubáň
New replacement name
Xanthocata nom. nov. [anagram of Catoxantha] for Catoxantha Solier, 1833 (nec Dejean, 1833); the type species is Buprestis opulenta Gory, 1832. Catoxantha Dejean, 1833 [issued January, 19] and Catoxantha Solier,
1833 [issued in July] are primary homonyms, Catoxantha Dejean, 1833 has priority, Catoxantha Solier, 1833
is permanently invalid.
Gender
Perotis Dejean, 1833: its gender is feminine, not masculine as in Bellamy (2006, 2008).
Authorships, assignments and type species
Anthaxia divina Reiche & Saulcy, 1856: 415 [not Reiche, 1860: 415] is a synonym of A. (Cratomerus) diadema
diadema (Fischer von Waldheim, 1824) and not of A. (C.) sponsa Kiesenwetter, 1857; based on the original
description of A. (C.) divina (see also Richter, 1949).
Anthaxia funerula var. hesperica Obenberger 1913 is subspecific (ICZN Article 45.6.4), and not infrasubspecific as in Bílý (2006b: 10, 32).
Buprestis cyanicornis is a synonym of Anthaxia (Cratomerus) hungarica hungarica (Scopoli, 1772). It was
established by Olivier (1790a: no 32: 71) and incorrectly credited to Fabricius (1781: 287).

20

New Nomenclatural And Taxonomic Acts, And Comments

Buprestis laeta is a synonym of Anthaxia (s.str.) nitidula (Linnaeus, 1758). It was established by Fabricius
(1775: 223) and incorrectly credited to Schaeffer (1766: 67).
Callanthaxia Richter is a synonym of Anthaxia (s.str.) Eschscholtz, 1829. Correct data for Callanthaxia Richter
are 1945c: 157 and not 1949: 168. The type species is Buprestis candens Panzer, 1792, by monotypy, and not
Buprestis passerini Pecchioli, 1837.
Cryptanthaxia Richter is a synonym of Anthaxia subgen. Haplanthaxia Reitter, 1911. Correct data for
Cryptanthaxia Richter are 1945c: 153 and not 1949: 77. The type species is Anthaxia rossica K. Daniel, 1903,
by monotypy.
Homalanthaxia Richter is a synonym of Anthaxia (s.str.) Eschscholtz, 1829. The correct data for Homalanthaxia
Richter are 1945c: 154, and not 1949: 86. The type species is Buprestis funerula Illiger, 1803, by monotypy.
Trichocratomerus Richter is a synonym of Anthaxia (s.str.) Eschscholtz, 1829. Correct data for Trichocratomerus
Richter are 1945a: 62 and not 1949: 161. Type species is Anthaxia intermedia Obenberger, 1913 (=Anthaxia
mancatula Abeille de Perrin, 1900), by monotypy, and not Buprestis manca Linnaeus, 1767.
Resurrected names
Anthaxia (Haplanthaxia) croesus (Villers, 1789) [sensu Kiesenwetter (1857)], resurrected from synonymy
with A. (s.str.) salicis salicis (Fabricius, 1777). See below, under New and reconfirmed synonyms.
Anthaxia (Haplanthaxia) millefolii polychloros Abeille de Perrin, 1894, resurrected from synonymy with
A. (H.) millefolii millefolii (Fabricius, 1801); “polychloros” is not a replacement name (see Mühle, 2015) but
was established for a new taxon subsequently erroneously identified as “Anthaxia millefolii (Fabricius,
1801)” (ICZN, Article 49).
Anthaxia (Melanthaxia) frankenbergeri Obenberger, 1933, resurrected from synonymy with A. (M.) godeti
Gory & Laporte, 1839; based on the original description and extensive material of A. (M.) godeti and on
study of the type and extensive recent material of A. (M.) frankenbergeri from the Balkans. Anthaxia
quadripunctata godeti var. frankenbergeri Obenberger 1914 is infrasubspecific. Anthaxia submontana var.
frankenbergeri Obenberger 1933 was made available by subsequent usage as valid subspecies by
Obenberger (1938). Thus, the correct authorships and data for A. (M.) frankenbergeri are Obenberger
1933b: 115, and not Obenberger 1914b: 14.
Anthaxia (Melanthaxia) venerabilis Marseul, 1869, resurrected from synonymy with A. (M.) istriana
Rosenhauer, 1847; synonymy with A. (M.) istriana is unfounded.
Capnodis miliaris metallica Ballion, 1871, resurrected from synonymy with C. miliaris (Klug, 1829) (see Mühle,
2015); type material studied.
Chrysobothris (s.str.) affinis adusta Abeille de Perrin, 1901 [replacement name for the homonymous C. (s.str.)
tristis Abeille de Perrin, 1900], resurrected from synonymy with C. (s.str.) affinis affinis (Fabricius, 1794);
based on the original description of C. (s.str.) tristis Abeille de Perrin, 1900, on study of the type material
of C. (s.str) carinithorax Obenberger, 1917, recent material of C. (s.str.) affinis adusta from southern Greece
and on the study of the extensive material of C. (s.str.) affinis affinis.
Coraebus nelsoni Akiyama & Ohmomo, 1989, resurrected from synonymy with C. hoscheki Gebhardt, 1929;
based on study of type and extensive recent material.
Julodis interpunctata J. Thomson, 1878, resurrected from synonymy with J. whitehillii whitehillii (G. R. Gray,
1832) (see Hołyński, 2014b); type material studied.

New Nomenclatural And Taxonomic Acts, And Comments

21

Poecilonota variolosa populialbae Richard, 1889, resurrected from synonymy with P. variolosa variolosa
(Paykull, 1799) (see Mühle, 2015); types and other extensive material studied.
Strigopteroides hellenica Cobos, 1990, resurrected from synonymy with S. depressa depressa (Fabricius, 1775);
recent material from continental Greece studied.
New and reconfirmed combinations, and new assignments
Amorphosoma aculeatum (Ganglbauer, 1890), comb. nov. from Coraebus.
Amorphosoma afghanicum (Alexeev & Volkovitsh, 1992), reconfirmed comb. from Vanroonia (see Bellamy,
2007, 2008).
Amorphosoma bispinosum (Wiedemann, 1823), reconfirmed comb. from Vanroonia (see Bellamy, 2007,
2008).
Amorphosoma cochinchinae Descarpentries & Villiers, 1967, reconfirmed comb. from Vanroonia (see Bellamy,
2007, 2008).
Amorphosoma himalayanum (Obenberger, 1958), reconfirmed comb. from Vanroonia (see Bellamy, 2007,
2008).
Amorphosoma pectorale Kerremans, 1892, reconfirmed comb. from Vanroonia (see Bellamy, 2007, 2008).
Amorphosoma perroti Descarpentries & Villiers, 1967, reconfirmed comb. from Vanroonia (see Bellamy, 2007,
2008).
Amorphosoma rambouseki (Obenberger, 1934), comb. nov. from Coraebus.
Anthaxia (Haplanthaxia) berytensis Abeille de Perrin, 1895, assign. nov. from Anthaxia s.str.
Anthaxia (Haplanthaxia) hypomelaena (Illiger, 1803), assign. nov. from Anthaxia s.str.
Anthaxia (Haplanthaxia) novickii Obenberger, 1938, assign. nov. from Anthaxia s.str.
Anthaxia (Haplanthaxia) olivieri Gory & Laporte, 1839, assign. nov. from Anthaxia s.str.
Anthaxia (Haplanthaxia) turcomanica Obenberger, 1937, assign. nov. from Anthaxia s.str.
Demochroa subgenus Demoxantha Hołyński, 2009, assign. nov. from Chrysochroa subgenus Demoxantha
Hołyński, 2009.
Julodis euphratica beludjistana Obenberger, 1924, reconfirmed assign. from J. whitehillii beludjistana
Obenberger, 1924 (see Hołyński, 2014b and Kubáň, 2006); type material studied.
Julodis interpunctata kabakovi Alexeev, 1990, assign. nov. from J. kabakovi Alexeev, 1990; not a synonym of
J. whitehillii klapperichi Cobos, 1966 (see Hołyński 2014b); type material studied.
New and reconfirmed synonyms
Anthaxia (s.str.) midas muelleri Obenberger, 1925, syn. nov. of A. (s.str.) spathuligera Obenberger, 1924 (see
Sakalian, 2007); type and recent extensive material from the Balkans studied.
Anthaxia (Haplanthaxia) millefolii millefolii (Fabricius, 1801) [junior homonym], syn. nov. of A. (H.) millefolii
millefolii (Herbst, 1801); based on the original description and illustration of A. (H.) millefolii millefolii
(Herbst, 1801), on the study of type and other extensive material of A. (H.) millefolii millefolii (Fabricius,
1801) from Europe and Turkey. Buprestis millefolii Herbst, 1801 [issued December, 5] and Buprestis millefolii
(Fabricius, 1801) [issued December, 31] are primary homonyms.
Anthaxia (Cratomerus) violacea Bílý, 1977, syn. nov. of A. (C.) sponsa Kiesenwetter, 1857; type and other extensive material from the Middle East studied.

22

New Nomenclatural And Taxonomic Acts, And Comments

Anthaxia (Haplanthaxia) berytensis alauita Obenberger, 1925, syn. nov. of A. (H.) berytensis Abeille de Perrin,
1895; type and other material from the Middle East studied.
Anthaxia (Haplanthaxia) nickerli Obenberger, 1923, reconfirmed syn. of A. (H.) pumila (Klug, 1829); type
material studied.
Anthaxia (Haplanthaxia) scutellaris Gené, 1839, reconfirmed syn. of A. (H.) croesus (Villers, 1789). The type
material of A. (H.) croesus is missing. Therefore a taxonomic consensus was adopted by V. Kubáň,
D. Baiocchi, S. Bílý, T. Kwast and M. Kafka, based on Kiesenwetter’s (1857) concept of A. croesus, the original descriptions and a study of extensive Mediterranean material.
Anthaxia (Haplanthaxia) scutellaris semireducta Pic, 1911, syn. nov. of A. (H.) croesus (Villers, 1789); based on
the original descriptions and on study of extensive Mediterranean material.
Anthaxia (Haplanthaxia) syrdarjensis Obenberger, 1934, syn. nov. of A. (H.) kuldjensis Obenberger, 1913; type
material studied.
Anthaxia (Haplanthaxia) viminalis Laporte, 1840 [junior homonym], syn. nov. of A. (H.) croesus (Villers,
1789); based on the original descriptions and on the study of extensive material from the Mediterranean.
Anthaxia (Melanthaxia) muehlei Niehuis, 1983, syn. nov. of A. (M.) frankenbergeri Obenberger, 1933; type
material studied (see under Resurrected names).
Bourgoinia Obenberger, 1926, reconfirmed syn. of Meliboeus Deyrolle, 1864 (see Bellamy 2007, 2008); junior
subjective synonym (see Kubáň, 2006: 45, 52); type material of both type species studied.
Brachycoraebus yajimai (Ohmomo, 2011), syn. nov. of B. punctatus (Baudon, 1968); based on the original
description and illustrations of B. yajimai, on the study of type and other extensive material of B. punctatus from China, India, Laos and Thailand.
Capnodis cariosa iranica Bogatchev, 1947, syn. nov. of C. cariosa hauseri Obenberger, 1928; based on the original description of C. cariosa iranica, on the study of type and other recent material of C. cariosa hauseri
from Iran, Iraq and Turkey.
Chrysobothris (s.str.) affinis carinithorax Obenberger, 1917, syn. nov. of C. (s.str..) affinis adusta Abeille de
Perrin, 1901 [replacement name]; based on the original description of C. (s.str.) tristis Abeille de Perrin,
1900 [junior homonym], on the study of type material of C. (s.str.) affinis carinithorax and recent material
of C. (s.str.) affinis adusta from southern Greece.
Chrysobothris (s.str.) chalcophana mediterranea G. Novak, 1986, syn. nov. of C. (s.str.) spuria Marseul, 1866;
based on the original description of C. (s.str.) spuria, on the study of a paratype of C. (s.str.) chalcophana
mediterranea from Israel and recent material of C. (s.str.) spuria from Egypt and Yemen.
Chrysodema subgenus Mitshekia Hołyński, 1994, syn. nov. of C. subgenus Cyalithoides Fisher, 1922; subjective
synonyms.
Coraebus elatus repletus Abeille de Perrin, 1893, syn. nov. of C. elatus (Fabricius, 1787); type and extensive
recent material studied.
Coraebus guangxiensis Peng, 1998, syn. nov. of C. sauteri Kerremans, 1912; type material studied.
Coraebus jiangxiensis Peng, 1990, syn. nov. of C. gorkai Gebhardt, 1929; type material studied.
Coraebus obscurus Peng, 1991, syn. nov. of C. quadriundulatus Motschulsky, 1866; type material studied.
Cylindromorphus acus Abeille de Perrin, 1897, syn. nov. of C. bifrons Rey, 1889; type material studied.
Cylindromorphus caspicus Obenberger, 1934, syn. nov. of C. pyrethri Stierlin, 1864; type material studied.

New Nomenclatural And Taxonomic Acts, And Comments

23

Cylindromorphus opacus Abeille de Perrin, 1897, syn. nov. of C. filum (Gyllenhal, 1817); type material studied.
Julodella fairmairei Théry, 1895, syn. nov. of J. dilaticollis Semenov, 1893; type and other material from
Azerbaijan and Iran studied.
Julodis armishi Makhan, 2012, syn. nov. of J. partha Obenberger, 1923; based on the original description and
holotype illustrations of J. armishi and on the study of extensive material of J. partha, including the type
material.
Julodis eoa Obenberger, 1923, syn. nov. of J. variolaris indica J. Thomson, 1878; type material studied.
Julodis euphratica bushirensis Obenberger, 1928, syn. nov. of J. consobrina consobrina Kerremans, 1914; type
material studied.
Julodis euphratica nusskina Obenberger, 1924, syn. nov. of J. euphratica scenica Kerremans, 1905; not a
­synonym of J. whitehillii beludjistana Obenberger, 1924 (see Hołyński 2014b and Mühle, 2015); based on the
description of J. iris var. scenica Kerremans, 1905 and on the study of the type material of J. euphratica
beludjistana and J. euphratica nusskina.
Julodis monstrosa Abeille de Perrin, 1904, syn. nov. of J. speculifer speculifer Laporte, 1835; type material
studied.
Julodis oberthuri Kerremans, 1898, reconfirmed syn. of J. variolaris variolaris (Pallas, 1771) (see Kubáň 2006);
not a subspecies of J. whitehillii (G. R. Gray, 1832) (see Hołyński 2014b); type material of J. oberthuri and
J. whitehillii studied.
Julodis ormarensis Obenberger, 1924 (lectotype designaton by Hołyński 2014b), reconfirmed syn. of J. euphratica scenica Kerremans, 1905 (see Kubáň 2006); not a subspecies of J. whitehillii (G. R. Gray, 1832) (see
Hołyński 2014b); type material of J. ormarensis and J. whitehillii studied.
Julodis palmyrensis Obenberger, 1923, syn. nov. of J. syriaca distincta Gory, 1840; type material studied.
Julodis variolaris kokandensis Obenberger, 1924, syn. nov. of J. variolaris variolaris (Pallas, 1771); based on the
original description of J. variolaris and on study of the type material of J. variolaris kokandensis.
Julodis zablodskii Motschulsky, 1845, syn. nov. of J. variolaris variolaris (Pallas, 1771); based on the original
descriptions of J. zablodskii and J. variolaris.
Kisanthobia ariasi curta Pic, 1898, reconfirmed syn. of K. ariasi (Robert, 1858); type and extensive recent
material from Cyprus, Turkey, Syria and Europe studied.
Lakhonia Descarpentries & Villiers 1967, reconfirmed syn. of Meliboeus Deyrolle, 1864 (see Bellamy 2007,
2008). Lakhonia Descarpentries & Villiers is a junior subjective synonym (see Kubáň 2006: 48, 52) permanent invalid because of homonymy with Lakhonia Yang, 1936 (Hemiptera).
Lampetis (Spinthoptera) interrupta (Laporte & Gory, 1837), syn. nov. of L. (S.) mimosae (Klug, 1829); type
material studied.
Lampetis (Spinthoptera) mimosae cyanea (Alfieri, 1976) [homonym] and L. (S.) mimosae kubani Bellamy, 2007
[replacement name], synn. nov. of L. (S.) mimosae (Klug, 1829); type material studied.
Latipalpis (Palpilatis) plasoni tauricola (Pic, 1916), syn. nov. of L. (P.) plasoni (Reitter, 1888); type material
studied.
Megaloxantha Kerremans, 1903, syn. nov. of Catoxantha Dejean, 1833; objective synonyms.
Meliacanthus Théry, 1942, reconfirmed syn. of Meliboeus Deyrolle, 1864 (see Bellamy 2007, 2008); junior subjective synonym (see Kubáň 2006: 48, 52); type material of both type species studied.

24

New Nomenclatural And Taxonomic Acts, And Comments

Meliboeus (Meliboeoides) parvulus ponticus Obenberger, 1924, syn. nov. of M. (s.str.) parvulus (Küster, 1852);
type and extensive recent material studied.
Meliboeus (s.str.) cupreomarginatus isanensis (Ohmomo, 2011), syn. nov. of M. (s.str.) cupreomarginatus
(Saunders, 1866); based on the original description and illistration of M. (s.str.) cupreomarginatus
i­ sanensis, on the study of type and other extensive material of M. (s.str.) cupreomarginatus from China,
Laos, Myanmar, Thailand and Vietnam.
Meliboeus (s.str.) fokienicus Obenberger, 1944, syn. nov. of M. (s.str.) wenigi Obenberger, 1927; type material
studied.
Meliboeus (s.str.) graminis (Herbst, 1801), syn. nov. of M. (s.str.) graminis graminis (Panzer, 1799); based on the
original descriptions and illustrations. Buprestis graminis Herbst, 1801 is a junior primary homonym of
B. graminis Panzer, 1799.
Meliboeus (s.str.) shirozui Kurosawa, 1954, syn. nov. of M. (s.str..) chinensis Obenberger, 1927; type material
studied.
Meliboeus (s.str.) staneki Obenberger, 1935, syn. nov. of M. (s.str.) heydeni (Abeille de Perrin, 1897); type material studied.
Melixes Schaefer, 1949, reconfirmed syn. of Meliboeus Deyrolle, 1864 (see Bellamy 2007, 2008); junior subjective synonym (see Kubáň 2006: 49, 52); type material of both type species studied.
Nalanda Théry, 1904, reconfirmed syn. of Meliboeus Deyrolle, 1864 (see Bellamy 2007, 2008); junior subjective
synonym (see Kubáň 2006: 49, 52); type material of both type species studied.
Neocoraebus Kerremans, 1903, syn. nov. of Brachycoraebus Kerremans, 1903; the type species of Neocoraebus
is Amorphosoma abyssinicum Kerremans, 1899 is very similar to the Asian B. klapperichi (Obenberger,
1959), B. herychi (Obenberger, 1940) and B. luzonicus (Obenberger, 1959). Thus, Ethiopia was obviously
given by error as the type locality of B. abyssinicus and the species is considered Oriental.
Neotoxoscelus shirahatai Kurosawa, 1977, syn. nov. of N. kerzhneri (Alexeev, 1975); type material studied.
Perotis planidorsis (Liskenne, 1994), syn. nov. of P. unicolor (A.G. Olivier, 1790); based on the original description and illustrations of P. planidorsis and on the study of extensive material of P. unicolor from France,
Spain and northern Africa.
Perotis striata Spinola, 1837, syn. nov. of P. unicolor (A.G. Olivier, 1790); based on the original description
of P. striata and on the study of extensive material of P. unicolor from France, Spain and northern
Africa.
Perotis unicolor igniventris Escalera, 1914, syn. nov. of P. unicolor (A.G. Olivier, 1790); type material of
P. unicolor igniventris and other extensive material of P. unicolor from France, Spain and northern Africa
studied.
Steraspis squamosa var. kindermanni Marseul, 1865 [made available by subsequent usage as valid subspecies], syn. nov. of S. squamosa (Klug, 1829); type material of S. squamosa and other extensive material
studied.
Tonkinula Obenberger, 1923, reconfirmed syn. of Meliboeus Deyrolle, 1864 (see Bellamy, 2007, 2008); junior
subjective synonym (see Kubáň, 2006: 50, 52); type material of both type species studied.
Trachys subglaber Rey, 1891, syn. nov. of T. quercicola Marseul, 1871; type material studied. Type locality is
“Hyères” (Rey, 1891c).

New Nomenclatural And Taxonomic Acts, And Comments

25

New lectotype designation
Trachys subglaber Rey, 1891: Lectotype (by present designation), female, in Muséum dʼHistoire Naturelle,
Lyon [MNHL] is labelled: [round black label = “Provence”, H. Labrique (MNHL) pers. comm.] “LECTOTYPE
Trachys marseuli[sic!] var. subglabra Rey, 1891 ♀ V. Kubáň design. 2010 [printed, red label]” “Trachys
­quercicola Marseul, 1871 Vít. Kubáň det. 2010 [printed]”. Designated in order to preserve stability of
nomenclature.
New assignments of invalid names
Acmaeodera feisthameli Gory, 1840 is a synonym of A. (Acmaeotethya) octodecimguttata quattuordecimpunctata (Villers, 1789) and not of A. (A.) degener degener (Scopoli, 1763); based on the original description and
illustration of A. (A.) feisthameli.
Anthaxia nitidula lichacevi Obenberger, 1934 (originally introduced as Anthaxia nitidula signaticollis
ab. lichacevi Obenberger, 1930c: 512, subsequently validated by Obenberger, 1934m: 139) is a synonym of
A. (s.str.) signaticollis Krynicki, 1832 and not of A. (s.str.) nitidula (Linnaeus, 1758); based on the original
descriptions.
Epacmene Gistel, 1848, replacement name, is a synonym of Catoxantha Dejean, 1833 and not of Catoxantha
Solier, 1833.
Euanthaxia Richter, 1945 is a synonym of Anthaxia (s.str.) Eschscholtz, 1829 and not of Anthaxia subgenus
Haplanthaxia Reitter, 1911. The correct data for Euanthaxia Richter are 1945a: 67 and not 1949: 86. The type
species is Anthaxia discicollis Gory & Laporte, 1839, by monotypy, and not Buprestis millefolii Fabricius,
1801.
Julodis eoa aberrans Obenberger, 1924 is a synonym of J. variolaris indica J. Thomson, 1878 and not of
J. v­ ariolaris eoa Obenberger, 1923 (see Hołyński 2014b); type material studied.
Julodis euphratica mrazi Obenberger, 1924 is a synonym of J. euphratica karelinii Mannerheim, 1837 and not
of J. whitehillii beludjistana Obenberger, 1924 (see Hołyński 2014b); type material studied.
Julodis klapaleki Obenberger, 1924 (lectotype designaton by Hołyński 2014b) is a synonym of J. euphratica
scenica Kerremans, 1905 and not of J. whitehillii beludjistana Obenberger, 1924 (see Hołyński 2014b),
­neither of J. atkinsoni Kerremans, 1896 (see Kubáň 2006); type material of J. atkinsoni, J. euphratica beludjistana and J. klapaleki studied.
Lampetis sancta Théry, 1922 is a synonym of Lampetis (Spinthoptera) arabica Gahan, 1895 and not of L. (S.)
nigrita Fairmaire, 1882; based on the original description of L. (S.) sancta and on the study of the type
material of L. (S.) arabica and L. (S.) nigrita.
Meliboeus (s.str.) hoberlandti Obenberger, 1944 is a synonym of M. (s.str.) alfierianus Théry, 1935 and not of
M. (s.str.) impressithorax Pic, 1924; type material studied.
Poecilonota variolosa setulosa Fleischer, 1896 is a synonym of P. variolosa populialbae Richard, 1889 and not
of P. variolosa variolosa (Paykull, 1799) (see Mühle, 2015); type and other extensive material studied.
Changes in rank
Anthaxia (Haplanthaxia) kuldjensis Obenberger, 1913, stat. nov. of A. (H.) flammifrons var. kuldjensis
Obenberger, 1913 [described as a subspecific name]; type material studied.

26

New Nomenclatural And Taxonomic Acts, And Comments

Buprestis (Ancylocheira) haemorrhoidalis rifensis Kocher, 1964, stat. nov. from B. (A.) rifensis Kocher, 1964;
based on the original description.
Chrysobothris (s.str.) spuria Marseul, 1866, stat. nov., from C. (s.str.) chalcophana spuria Marseul, 1866; based
on the original description and recent material of C. (s.str.) spuria from Egypt and Yemen, and on the
study of a paratype of C. (s.str.) chalcophana mediterranea G. Novak, 1986 from Israel.
Cylindromorphus carpetanus Perris, 1864, stat. nov., from C. filum carpetanus Perris, 1864; type material
studied.
Julodis atkinsoni Kerremans, 1896, reconfirmed stat. from J. whitehillii atkinsoni Kerremans, 1896 (see
Hołyński 2014b and Kubáň 2006); type material studied.
Julodis klapperichi Cobos, 1966, reconfirmed stat. from J. whitehillii klapperichi Cobos, 1966 (see Hołyński
2014b and Kubáň 2006); type material studied.
Julodis proxima Gory, 1840, reconfirmed stat. from J. whitehillii proxima Gory, 1840 (see Hołyński 2014b and
Kubáň 2006); type material studied.
Lampetis (Spinthoptera) svobodai Krajčík, 2009, stat. nov., from L. (S.) catenulata svobodai Krajčík, 2009; type
material studied.
Meliboeus (s.str.) primoriensis Alexeev, 1979, stat. nov., from M. (s.str.) ohbayashii primoriensis Alexeev, 1979;
type and other material from China and Far East studied.
Perotis lugubris aereiventris (Reiche, 1861), stat. nov., from P. aereiventris (Reiche, 1861); based on the original
description of P. aereiventris.
Trachys compressus Abeille de Perrin 1891, stat. nov., from T. troglodytes compressus Abeille de Perrin 1891;
type material studied.
Available names
[The following names were previously either considered unavailable, or overlooked.]
Acmaeodera octodecimguttata var. friwaldskyi Marseul, 1965: 278 (syn. of A. (Acmaeotethya) octodecimguttata
octodecimguttata (Piller & Mitterpacher, 1783)) was described as a subspecific name.
Anthaxia canifrons var. bucharica Obenberger, 1913 (syn. of A. (Melanthaxia) conradti Semenov, 1891) was
described as a subspecific name.
Anthaxia bivulnerata var. splichali Obenberger 1914 (syn. of A. (s.str.) baconis J. Thomson, 1879) was described
as a subspecific name.
Anthaxia candens var. marchii Obenberger 1925 (syn. of A. (s.str.) candens candens (Panzer, 1792)) was
described as a subspecific name.
Anthaxia lucens var. phoenica Ganglbauer, 1882 (syn. of A. (s.str.) urens Abeille de Perrin, 1882) was described
as a subspecific name.
Anthaxia bonvouloiri var. amabilis Obenberger 1913 (syn. of A. (Cratomerus) bonvouloiri Abeille de Perrin,
1869) was described as a subspecific name.
Anthaxia scorzonerae var. juno Obenberger 1913 (syn. of A. (C.) diadema diadema (Fischer von Waldheim,
1824)) was described as a subspecific name.
Anthaxia hungarica var. juvenilis Abeille de Perrin 1895 (syn. of A. (C.) eugeniae eugeniae Ganglbauer, 1885)
was described as a subspecific name.
Anthaxia sponsa var. cyanescens Marseul, 1865 (syn. of A. (C.) sponsa Kiesenwetter, 1857) was described as a
subspecific name.

New Nomenclatural And Taxonomic Acts, And Comments

27

Anthaxia exsul var. satanella Obenberger 1914 (syn. of A. (Haplanthaxia) exsul Obenberger, 1914) was
described as a subspecific name.
Anthaxia flammifrons var. ignea Obenberger, 1913 (syn. of A. (H.) flammifrons Semenov, 1891) was described
as a subspecific name.
Anthaxia flavicomes var. erivana Obenberger 1913 (syn. of A. (H.) flavicomes Abeille de Perrin, 1900) was
described as a subspecific name.
Anthaxia ignipennis var. flammeola Abeille de Perrin 1907 (syn. of A. (H.) ignipennis Abeille de Perrin, 1882)
was described as a subspecific name.
Anthaxia pleuralis var. robustior Obenberger 1913 (syn. of A. (H.) umbellatarum domina Abeille de Perrin,
1900) was described as a subspecific name.
Anthaxia inculta var. aerea Rey, 1891 (syn. of A. (H.) umbellatarum umbellatarum (Fabricius, 1787)) was
described as subspecific name.
Anthaxia godeti var. cupricolor Abeille de Perrin 1909 (syn. of A. (Melanthaxia) godeti Gory & Laporte, 1839)
was described as a subspecific name.
Anthaxia helvetica varr. bulgarica Obenberger 1913, pecirkai Obenberger 1913 and teriolensis Obenberger, 1913
(synn. of A. (M.) helvetica helvetica Stierlin, 1868) were described as subspecific names.
Anthaxia sepuchralis var. nigrocyanea Rey, 1891 (syn. of A. (M.) helvetica helvetica Stierlin, 1868) was described
as a subspecific name.
Anthaxia morio f. meridionalis Obenberger 1913 (syn. of A. (M.) morio (Fabricius, 1792)) was described as
subspecific name.
Anthaxia tenella var. insularis Obenberger, 1914 (syn. of A. (M.) tenella tenella Kiesenwetter, 1858) was
described as a subspecific name.
Chalcophorella stigmatica var. biimpressa Pic, 1916 (syn. of C. (s.str.) stigmatica (Dalman, 1817)) was described
as a subspecific name.
Chalcophorella fabricii var. semiviridis Pic, 1916 (syn. of Chalcophorella (Rossiella) fabricii (P. Rossi, 1794)) was
described as a subspecific name.
Psiloptera pisna var. gallica Abeille de Perrin, 1891 (syn. of Latipalpis (s.str.) plana plana (A.G. Olivier, 1790))
was described as a subspecific name.
Comments
Clema elegans Obenberger, 1923, was described from “Turkestan”; it is synonym with Pseudoclema
t­ ransvaalense (Kerremans, 1911) from southern Africa and therefore excluded from the Catalogue.
Coraebus perpulcher Obenberger, 1916, was described from “China”; it is synonym with Asymydes transvalensis Kerremans, 1893 from southern Africa and therefore excluded from the Catalogue.
Changes proposed by Hołyński (2006, 2009, 2010, 2011, 2014b) based on his MICSEQ method of phylogenetic
analysis (Hołyński, 2001b) are not followed in this catalogue.
Buprestidae: Polycestinae: Acmaeoderini
M.G. Volkovitsh
Authorships
Acmaeodera flavipennis ab. dispersemaculata Lotte, 1943: 145 is unavailable. Acmaeodera flavipennis dispersemaculata was made available by subsequent usage as valid subspecies by Alfieri (1976). Correct

28

New Nomenclatural And Taxonomic Acts, And Comments

­authorships and date for A. flavipennis dispersemaculata is Alfieri, 1976 (see also under New combinations
and New and reconfirmed synonyms).
Acmaeodera flavipennis ab. brunneicostata Lotte, 1943: 145 is unavailable. Acmaeodera flavipennis brunneicostata was made available by subsequent usage as valid subspecies by Alfieri (1976). Correct authorships
and date for A. flavipennis brunneicostata is Alfieri, 1976 (see also under New combinations and assignments, and New and reconfirmed synonyms).
Acmaeodera philistina var. immaculata Andres, 1920 was made available by subsequent usage as valid subspecies by Alfieri (1976) (as Acmaeodera convoluta). Correct authorships and date for A. philistina immaculata is Andres, 1920 (see also under New combinations and assignments, and New and reconfirmed
synonyms).
Acmaeodera (Ptychomus) polita var. hedjazensis Théry, 1931 was made available by subsequent usage as valid
subspecies by Cobos (1958). Correct authorships and date for A. (P.) polita hedjazensis is Théry, 1931 (see
also under Resurrected names).
New combinations and assignments
Acmaeodera (Acmaeotethya) octodecimguttata quattuordecimpunctata (Villers, 1789), new assign. for A. (A.)
degener quattuordecimpunctata (Villers, 1789); based on the original descriptions and on the study of the
extensive material of both subspecies of A. (A.) octodecimguttata (Piller & Mitterpacher, 1783).
Acmaeodera (Palaeotethya) lugubris fracta Wollaston, 1864, new assign. from A. (P.) rubromaculata P.H. Lucas,
1844.
Acmaeodera (Palaeotethya) lugubris grosseseriata Cobos, 1960, new assign. from A. (P.) rubromaculata
P.H. Lucas, 1844.
Acmaeodera (Palaeotethya) lugubris lugubris Spinola, 1838, new assign. from A. (P.) rubromaculata
r­ ubromaculata P.H. Lucas, 1844.
Acmaeodera (Palaeotethya) lugubris malacensis Cobos, 1954, new assign. from A. (P.) rubromaculata
P.H. Lucas, 1844.
Acmaeodera (Palaeotethya) lugubris oertzeni Ganglbauer, 1889, new assign. from A. (P.) rubromaculata
P.H. Lucas, 1844.
Acmaeodera (Palaeotethya) lugubris segurensis Escalera, 1904, new assign. from A. (P.) rubromaculata
P.H. Lucas, 1844.
Acmaeodera (Rugacmaeodera) mirei dispersemaculata Alfieri, 1976, new assign. from A. flavipennis (Klug,
1829) (see also under Authoships and New and reconfirmed synonyms).
Acmaeoderella (Euacmaeoderella) alfierii (Théry, 1929), new assign. from A. subgenus Acmaeoderella.
Acmaeoderella (Euacmaeoderella) nivetecta Volkovitsh, 1976, new assign. from A. subgenus Acmaeoderella.
Acmaeoderella (Euacmaeoderella) pseudonivetecta Volkovitsh, 2011, new assign. from A. subgenus
Acmaeoderella.
Xantheremia (s.str.) flavipennis brunneicostata (Alfieri, 1976), new comb. from Acmaeodera (see also under
Authoships and New and reconfirmed synonyms).
Xantheremia (s.str.) philistina immaculata (Andres, 1920), new comb. from Acmaeodera (see also under
Authoships and New and reconfirmed synonyms).

New Nomenclatural And Taxonomic Acts, And Comments

29

Resurrected names
Acmaeodera (Acmaeotethya) octodecimguttata (Piller & Mitterpacher, 1783), resurrected from synonymy
with A. (A.) degener degener (Scopoli, 1763); based on the original descriptions and on the study of the
extensive material of both species.
Acmaeodera (Palaeotethya) ceylonensis Obenberger, 1932, resurrected from synonymy with A. (P.) bipunctata
bipunctata (A.G. Olivier, 1790a). Type and other extensive material studied.
Acmaeodera (Palaeotethya) lugubris Spinola, 1838 (as A. vaillant var. α), resurrected from synonymy with
A. (P.) rubromaculata rubromaculata P.H. Lucas, 1844. Type material studied.
Acmaeodera (Ptychomus) polita hedjazensis Théry, 1931, resurrected from synonymy with A. (P.) polita (Klug,
1829) (Bellamy, 2008) (see also chapter Authoships). Type material studied.
Acmaeoderella (Euacmaeoderella) olivacea (Abeille de Perrin, 1904, resurrected from synonymy with A. (E.)
vetusta (Ménétriés, 1832). Syntype (male) in coll. A. Cobos in the Museo Nacional de Ciencias Naturales,
Madrid and other recent material from Iran studied.
New lectotype designations
Acmaeodera (Palaeotethya) ceylonensis Obenberger, 1932: Lectotype (by present designtion) male in National
Museum, Prague is labelled: “Ceylon” [false locality]. Designated in order to preserve stability of nomenclature. This species is known from Cyprus exclusively, it occurs together with A. (P.) bipunctata guillebeaui Abeille de Perrin, 1891.
Acmaeodera (Palaeotethya) lugubris Spinola, 1838: Lectotype (by present designation) sex not examined, in
Museo Regionale di Scienze Naturali, Turin, labelled [bottom label]: “Acmaeodera Vaillant m. lugubris
Icon D. Vaillant Algeria”. Designated in order to preserve stability of nomenclature.
Acmaeodera (Ptychomus) polita hedjazensis Théry, 1931: Lectotype (by present designation), sex not
­examined, in the Museum National d’Histoire Naturelle, Paris is labelled: “Meyer-Darcis Patria [printed]
Hedjaz Arabia [handwritten]” “Type [printed, red label]” “polita var. hedjazensis Thery [handwritten]
Type [printed, red font]” “Muséum Paris 1936 Coll. A. Théry [printed]” “hedjazensis Théry [handwritten, E. Holm, red font] Holotype [printed, red font]”. Designated in order to preserve stability of
nomenclature.
Xantheremia (Xantheremia) philistina immaculata (Andres, 1920): Lectotype (by present designation), sex
not examined, in the Naturhistorisches Museum, Basel is labelled: “Sinai (Sudwest), 22–24.6.1904, des
chasses de Kneucker (rear de A. Andres) [handwritten]” “Acmaeodera philistina var. nov. immaculata
Andres Type [handwritten]” “Acmaeodera flavipennis Klug. var. immaculata Andres (Thery det. 1929)
[handwritten]” “729 [handwritten]” “Coll. Alfieri Egypte [printed]”.
New synonyms
Acmaeodera (s.str.) revelierii Mulsant & Rey, 1861, syn. nov. of A. (s.str.) revelierii Mulsant & Rey, 1859; based
on the original descriptions.
Acmaeodera (Palaeotethya) rubromaculata P.H. Lucas, 1844, syn. nov. of A. (P.) lugubris lugubris Spinola, 1838;
based on the original description of A. (P.) rubromaculata and on the study of the lectotype of A. (P.)
lugubris lugubris (see Resurrected names).

30

New Nomenclatural And Taxonomic Acts, And Comments

Acmaeodera (Ptychomus) omana Krajčík, 2015, syn. nov. of A. (P.) arabica Gory, 1840 based on original
description and male holotype illustrations of A. (P.) omana and on the study of the type material of
A. (P.) arabica.
Acmaeodera (Rugacmaeodera) mirei dispersemaculata Alfieri, 1976, syn. nov. of A. (R.) mirei Descarpentries &
Mateu, 1965 (see also Authorships and new combinations). Holotype (sex not examined) studied (Coll. Frey
in Naturhistorisches Museum, Basel): “Eclos au Caire, d’Acacia tortilis en avril 1937” “Bois ramassé au Wadi
Feiran, Sinai, le 29.5.35” “A. flavipennis ab. dispersemaculata mihi [Lotte] Type” “727” “Coll. Alfieri Egypte”).
Acmaeoderella (s.str.) normanna Sparacio, 2006, syn. nov. of A. (s.str.) circassica (Reitter, 1890); type material
studied.
Acmaeoderella (Carininota) flavofasciata veselyi (Obenberger, 1934), syn. nov. of A. (C.) flavofasciata flavofasciata (Piller & Mitterpacher, 1783); type material of A. (C.) flavofasciata veselyi studied.
Acmaeoderella (Euacmaeoderella) corsica (Obenberger, 1922), syn. nov. of A. (E.) gibbulosa (Ménétriés, 1832);
type material studied.
Acmaeoderella (Liogastria) alesi (Obenberger, 1936), syn. nov. of A. (L.) virgulata (Illiger, 1803); type material
studied.
Xantheremia (s.str.) flavipennis brunneicostata (Alfieri, 1976), syn. nov. of X. (s.str.) flavipennis (Klug, 1829)
(see also under Authorships). Holotype (sex not examined) studied (Coll. Frey in Naturhistorisches
Museum, Basel): “Eclos au Caire, de Acacia tortilis 24.6.1935” “Wadi Feiran Sud Sinai 29–30.5.1935
A. Rabinovitch cep.” “A. flavipennis ab. brunneicostata mihi [Lotte] Type” “726” “Coll. Alfieri Egypte”.
Xantheremia (s.str.) philistina immaculata (Andres, 1920), syn. nov. of X. (s.str.) philistina (Marseul, 1865).
Type material studied.
New assignments of invalid names
Acmaeodera insolita Théry, 1908 is a synonym of A. (Acmaeotethya) octodecimguttata quattuordecimpunctata
(Villers, 1789) and not of A. (A.) degener quattuordecimpunctata (Villers, 1789); type material of A. (A.)
insolita studied.
Acmaeodera multipunctata P.H. Lucas, 1844 is a synonym of A. (Acmaeotethya) octodecimguttata quattuordecimpunctata (Villers, 1789) and not of A. (A.) degener quattuordecimpunctata (Villers, 1789); type material of A. multipunctata studied.
Acmaeodera quattuordecimguttata (Fabricius, 1798) is a synonym of A. (Acmaeotethya) octodecimguttata
quattuordecimpunctata (Villers, 1789) and not of A. (A.) degener quattuordecimpunctata (Villers, 1789);
based on the original description of A. quattuordecimguttata.
Acmaeodera sedecimpunctata (Schrank, 1789) is a synonym of A. (Acmaeotethya) octodecimguttata octodecimguttata (Piller & Mitterpacher, 1783) and not of A. (A.) degener degener (Scopoli, 1763); based on the
original description of A. sedecimpunctata.
Comments
Acmaeodera (Palaeotethya) vaillant Spinola, 1838 is unnecessary new replacement name, and a objective
synonym of A. (Palaeotethya) bipunctata bipunctata (A.G. Olivier, 1790).
Acmaeodera (Palaeotethya) vaillanti Marseul, 1865 is unjustified subsequent emendation for A. vaillant
Spinola, 1838 and a synonym of A. (Palaeotethya) bipunctata bipunctata (A.G. Olivier, 1790).

New Nomenclatural And Taxonomic Acts, And Comments

31

Buprestidae: Chrysochroinae: Sphenopterini
V. Kubáň & M. Yu. Kalashian
Resurrected names
Rethia Théry, 1901, resurrected from synonymy with S. subgen. Tropeopeltis; it is a monotypical Ethiopian
taxon excluded from the Catalogue.
New and reconfirmed synonyms
Sphenoptera (Deudora) gemmata laevis Rey, 1891, reconfirmed syn. of S. (D.) gemmata gemmata (A. G.
Olivier, 1790); based on the original descriptions and on the study of extensive material of S. (D.) gemmata
gemmata from western Europe and northern Africa.
Sphenoptera (s.str.) barbarica sicula Jakovlev, 1886, syn. nov. of S. (s.str.) barbarica barbarica (Gmelin, 1790);
type material of S. (s.str.) barbarica sicula and other extensive material of S. (s.str.) barbarica barbarica
from western Europe and northern Africa studied.
Sphenoptera (s.str.) barbarica silvestrii Leoni, 1911, reconfirmed syn. of S. (s.str.) barbarica barbarica (Gmelin,
1790); based on the original descriptions and on the study of extensive material of S. (s.str.) barbarica
barbarica from western Europe and northern Africa.
Buprestidae: Agrilinae: Aphanisticini
M.Yu. Kalashian
New synonymy
Aphanisticus klapperichi Obenberger, 1944 syn. nov. of Aphanisticus peninsulae Obenberger, 1924; type material studied.
Change in rank and lectotype designation
Endelus continentalis Obenberger, 1944, stat. nov. from Endelus collaris continentalis Obenberger, 1944. Type
material studied: Lectotype (by present designation), female, in the National Museum, Prague is labelled:
“Tainingli, Ch[ina], Fukien [handwritten]” “Typus [printed, red paper]” “Endelus collaris ssp. continentalis
m. [handwritten] Dr. Obenberger det. [printed]”. Designated in order to preserve stability of nomenclature.
Buprestidae: Agrilinae: genus Agrilus
E. Jendek
New assignments of invalid names
Agrilus rubi Kaszab, 1940 is a synonym of A. roscidus Kiesenwetter, 1857 and not of A. viridicaerulans rubi
Schaefer, 1937; type material studied.
Comments
A number of subgenera have been established in Agrilus, and most of them are still valid, though not applicable to the majority of species. The concepts of subgenera are confusing in current state, as the case of
the common A. pratensis (Ratzeburg, 1837) shows: this species was placed in four different subgenera
(Agrilus s.str., Anambus, Arquagrilus, Robertius). With respect to the current situation, all species are
listed alphabetically in the present work.

32

New Nomenclatural And Taxonomic Acts, And Comments

Agrilus auricollis Kiesenwetter, 1857 is a junior primary homonym. It will be declared nomen protectum in a
subsequent work.
Limnichidae
C. Hernando & I. Ribera
Comments
Thaumastodus and Thaumastodinae were described by Champion, 1924a: 25. The genus was subsequently
placed by Champion (1924b) in synonymy with Pseudeucinetus Heller, 1921. Csiki (1924) established for
Pseudeucinetus the new tribe Pseudeucinetini, placed within the Melandryidae. Csiki’s paper was issued
on February, 24 while the day of the publication of Thaumastodinae is unknown and therefore deemed to
be the last day of February 1924. However, according to the ICZN Article 35.5, the older name
(Pseudeucinetini) is not to displace the younger name (i.e. Thaumastodinae) in prevailing usage.
Psephenidae
I. Löbl
Eubria Latreille, 1829 and Eubria Germar, 1818
The name Eubria is often credited to Germar, 1818: 239 (e.g., Bouchard et al., 2011; Löbl & Smetana, 2011), likely
based on overlooked statement or misunderstanding of the original German text. In fact, Germar (1818:
238, 289) noted under Cyphon palustris “… dürfte zu einer besondern Gattung sich eignen, und Dahl verwendet es auch unter dem Namen Eubria palustris, doch ich finde diese Gattung nicht hinreichend nach
allen Merkmales durchgeführt, und bis dahin mag es unter Cyphon stehen. Meinen Freuden habe ich es
früher als Cyphon striatus mihi gesand. … ” Thus, Germar mentioned, but clearly refused the use of the
Dahl’s unavailable Name Eubria, and considered specimens identified by Dahl as Eubria palustris the
same as his Cyphon striatus. Latreille (1829) was the first who established the name Eubria.

Catalogue
series SCARABAEIFORMIA Crowson, 1960
superfamily SCARABAEOIDEA Latreille, 1802
family GEOTRUPIDAE Latreille, 1802
subfamily Athyreinae Lynch Arribálzaga, 1878

genus Pseudoathyreus Howden & Martínez, 1963: 348 type species Athyreus porcatus Laporte, 1840
flavohirtus Walker, 1871: 12 (Athyreus) A: AE “Arabia”
orientalis Laporte, 1840: 103 (Athyreus) A: XIZ ORR

subfamily Bolboceratinae Mulsant, 1842
tribe Bolbelasmini Nikolajev, 1996

genus Bolbelasmus Boucomont, 1911: 335 type species Bolboceras gallicum Mulsant, 1842
subgenus Bolbelasmus Boucomont, 1911: 335 type species Bolboceras gallicum Mulsant, 1842
bocchus bocchus Erichson, 1841a: 170 (Bolboceras) E: PT SP N: AG MO A: IS
fissicornis Mulsant, 1843: 280 (Bolboceras)

bocchus vaulogeri Abeille de Perrin, 1898: 254 (Bolboceras) N: EG LB TU
gallicus Mulsant, 1842: 350 (Bolboceras) E: FR PT SP
conjunctus Mulsant, 1842: 351 (Bolboceras)
provincialis Mulsant, 1842: 351 (Bolboceras)

keithi Miessen & Trichas, 2011: 183 E: GR (Kríti, Kássos)
makrisi Miessen, 2011: 111 A: CY
nireus Reitter, 1895b: 81 (Bolboceras) A: IN IQ SY TR
orientalis Petrovitz, 1968d: 185 A: FE (Vladivostok)
romanorum Arnone & Massa, 2010: 403 E: IT (Sicilia)
shibatai Masumoto, 1984a: 75 A: JA (Ryukyus)
tauricus Petrovitz, 1973d: 338 A: TR
unicornis Schrank, 1789: 61 (Scarabaeus) E: AU BH BU CR CZ FR GE GR HU IT MD PL RO SK SL SZ TR UK
A: TR
aeneas Panzer, 1793a: pl. 5 (Scarabaeus)
quadridens Panzer, 1795: 34 (Scarabaeus)

subgenus Kolbeus Boucomont, 1911: 335 type species Bolboceras coreanus H.J. Kolbe, 1886
coreanus H.J. Kolbe, 1886: 188 (Bolboceras) A: ANH FUJ GAN GUI NC SC SCH SD SHA TAI YUN ZAI ZHE ORR
conicifrons Fairmaire, 1896a: 82 (Bolboceras)
kurosawai Masumoto, 1984a: 76

krikkeni Nikolajev, 1979f: 225 A: SD
meridionalis Krikken, 1977c: 285 A: JIX ORR
minutus C.-L. Li & Masumoto, 2008: 482 A: TAI
nativus ishigakiensis Masumoto, 1984a: 73 A: JA (Ryukyus)
nativus nativus Krikken, 1977c: 287 A: TAI
© koninklijke brill nv, leiden, ���6 | doi 10.1163/9789004309142_003

34

GEOTRUPIDAE

genus Bolbogonium Boucomont, 1911: 340 type species Bolboceras triangulus Westwood, 1852
davatchii Baraud, 1973a: 14 (Bolboceras) A: IN
howdeni Krikken, 1977a: 97 A: AF PA ORR
impressum Wiedemann, 1823: 6 (Scarabaeus) A: HP UP ORR
kabakovi Nikolajev, 1976d: 693 A: AF
kabulicum Nikolajev & Kabakov, 1977: 646 A: AF
pseudopunctatissimum Krikken, 1977a: 90 A: HP UP ORR
triangulum Westwood, 1852a: 26 (Bolboceras) A: HP PA UP ORR
genus Bolbotrypes Olsoufieff, 1907a: 21 type species Bolboceras davidis Fairmaire, 1891
davidis Fairmaire, 1891a: 6 (Bolboceras) A: BEI FE HEB HEI NE NO TAI ORR
chaoyanum Niijima & Kinoshita, 1937: 4 (Bolboceras)

tribe Bolboceratini Mulsant, 1842

genus Bolbaffroides Nikolajev, 1979e: 189 type species Bolboceras validus Klug, 1845
carenicollis Laporte, 1840: 104 (Bolboceras) A: PA ORR
capitatus Westwood, 1848b: 386 (Bolboceras)

kubaricus Krikken, 1978b: 308 (Bolboceroides) A: YE
scotti R. Paulian, 1948: 144 (Bolboceras) A: YE
validus Klug, 1845: 47 (Bolboceras) N: EG A: OM SA YE
genus Bolboceras Kirby, 1819b: 459 type species Scarabaeus quadridens Fabricius, 1781
Indobolbus Nikolajev, 1979e: 190 type species Scarabaeus quadridens Fabricius, 1781

berytense Petrovitz, 1963c: 239 A: LE
extraneum Krikken, 2013: 48 A: PA ORR
perpunctatum Krikken, 2013: 54 A: SD
schulzei Krikken, 2013: 44 A: UP ORR

tribe Bolbochromini Nikolajev, 1970

genus Bolbocerodema Nikolajev, 1973: 856 type species Bolboceras nigroplagiatus C.O. Waterhouse, 1875
apicatum Fairmaire, 1891a: 6 (Bolboceras) A: HUB
dierli Krikken, 1979: 47 (Bolbocerosoma) A: NP
garritor Krikken, 1979: 45 (Bolbocerosoma) A: JIA SCH TAI
kiyoyamai Nomura, 1973b: 38 (Bolbocerosoma) A: TAI
nigroplagiatum C.O. Waterhouse, 1875a: 96 (Bolboceras) A: JA
sikkimense Krikken, 1979: 48 (Bolbocerosoma) A: SD
zonatum Nikolajev, 1973: 859 A: FE LIA NC SC
genus Bolbochromus Boucomont, 1909b: 117 type species Bolboceras laetus Westwood, 1852
subgenus Bolbochromus Boucomont, 1909b: 117 type species Bolboceras laetus Westwood, 1852
laetus Westwood, 1852a: 27 (Bolboceras) A: GUI ORR

35

GEOTRUPIDAE
nomurai C.-L. Li & Krikken, 2013: 47 A: YUN ORR
plagiatus Westwood, 1852a: 27 (Bolboceras) A: NP UP ORR
ryukyuensis Masumoto, 1984b: 168 A: JA (Ryukyus) TAI
sinensis Krikken & C.-L. Li, 2013: 514 A: FUJ

tribe Eubolbitini Nikolajev, 1970

genus Bolboceratops Krikken, 1978: 72 type species Bolboceras indicus Westwood, 1848
indicus Westwood, 1848b: 356 (Bolboceras) A: PA SD XIZ AFR ORR
bidentatus Boucomont, 1923b: 99 (Bolboceras)

genus Bolbohamatum Krikken, 1980: 5 type species Scarabaeus cyclops A.G. Olivier, 1789
calanus Weswood, 1848: 385 (Bolboceras) A: UP ORR
tumidulus Westwood, 1852a: 21 (Bolboceras)

cyclops A.G. Olivier, 1789: no 3: 60 (Scarabaeus) A: HP NP UP ORR
subglobosus Westwood, 1852a: 20 (Bolboceras)
veter Fabricius, 1792a: 33 (Scarabaeus)

drescheri drescheri Krikken, 1980: 34 A: HKG
laterale Westwood, 1848b: 385 (Bolboceras) A: KA SD
laevicolle Westwood, 1848b: 385 (Bolboceras) A: “Himalaya” ORR
grandis Westwood, 1848b: 385 (Bolboceras)

marginale Krikken, 1980: 30 A: PA
robustum Krikken, 1980: 25 A: “Himalaya”
genus Eubolbitus Reitter, 1892a: 125 [=1893a: 4] type species Bolboceras radoszkowskii Solsky, 1876
radoszkowskii Solsky, 1876a: 352 (Bolboceras) A: AF HP IN KI KZ TD UZ
gagarinei Fairmaire, 1892c: cxxii (Bolboceras)

sicardi Reitter, 1896g: 270 N: MO LB TU
zarudnyi Semenov & S.I. Medvedev, 1929b: 167 A: IN IQ SY
petrovitzi Baraud, 1973a: 13

genus Socotrabolbus Cambefort, 1998: 443 type species Socotrabolbus canui Cambefort, 1998
canui Cambefort, 1998: 444 A: YE (Suqutra)

tribe Odonteini Shokhin, 2007

genus Odonteus Samouelle, 1819: 189 type species Scarabaeus mobilicornis Fabricius, 1775 (=Scarabaeus
armiger Scopoli, 1772)
Odontaeus Dejean, 1821: 56 type species Scarabaeus mobilicornis Fabricius, 1775 (=Scarabaeus armiger Scopoli, 1772)

armiger Scopoli, 1772: 78 (Scarabaeus) E: AB AR AU BE BH BU BY CR CT CZ DE EN FR GB GE GR HU IT KZ
LA LT LU MC MD NL PL RO SK SB SL SP ST SV SZ TR UK A: TR
bicolor Fabricius, 1775: 15 (Scarabaeus)
fulvus Mulsant, 1842: 349 (Bolboceras)

36

GEOTRUPIDAE
mobilicornis Fabricius, 1775: 11 (Scarabaeus)
obliteratus Mulsant, 1842: 349 (Bolboceras)
recticornis Mulsant, 1842: 348 (Bolboceras)
rufescens Ponza, 1805: 33 (Scarabaeus)
testaceus Fabricius, 1775: 15 (Scarabaeus)

gandhara Carpaneto & Mignani, 2005: 2 A: PA
orientalis Mittal, 1998: 385 (Odontaeus) A: PA UP
subfamily Bolboceratidae, species incertae sedis
Bolboceras bicarinatum Westwood, 1852a: 24 A: “East India”

subfamily Geotrupinae Latreille, 1802
tribe Enoplotrupini Paulian, 1945

genus Chelotrupes Jekel, 1866: 549 type species Scarabaeus momus Fabricius, 1792 (=Scarabaeus momus
A.G. Olivier, 1789)
algarvicus Hillert, Král & Schneider, 2012: 3 E: PT
annamariae Byk, 2012: 110

brancoi Hillert, Král & Schneider, 2012: 6 E: PT
feryi Hillert, Král & Schneider, 2012: 8 E: PT
hendrichi Hillert, Král & Schneider, 2012: 10 E: SP
hiostius Gené, 1836: 187 (Geotrupes) E: IT (Sardegna)
kyliesi Hillert, Král & Schneider, 2012: 16 E: SP
laevipennis Mulsant & Godart, 1855a: 1 [=1855e: 1] (Ceratophyus) E: SP
andalusiacus Fairmaire, 1869: 10 (Geotrupes)

matutinalis Baudi di Selve, 1870: 70 (Geotrupes) E: IT (Sardegna)
momus A.G. Olivier, 1789: no 3: 60 (Scarabaeus) E: SP
momoides Reitter, 1892a: 130 [=1893a: 9] (Ceratophyus)
momus Fabricius, 1792a: 13 (Scarabaeus)

genus Enoplotrupes P.H. Lucas, 1869b: xiii type species Enoplotrupes sinensis P.H. Lucas, 1869
subgenus Enoplotrupes P.H. Lucas, 1869b: xiii type species Enoplotrupes sinensis P.H. Lucas, 1869
Enoplotrypes Fairmaire, 1886: 320 type species Enoplotrupes sinensis P.H. Lucas, 1869

barmanicus Gestro, 1888b: 111 A: YUN ORR
chaslii Fairmaire, 1886: 320 (Enoplotrypes) A: FUJ GUI JIX ZHE
crassicornis Boucomont, 1905: 212 A: GAN HUB JIX SCH
gaoligong Král, Malý & Schneider, 2015: 324 A: YUN
kubani Král, Malý & Schneider, 2015: 327 A: SCH YUN
largeteaui R. Oberthür, 1883a: 58 A: GUI JIX SCH YUN ORR
variicolor Fairmaire, 1886: 321 (Enoplotrypes)

monachus Král, Malý & Schneider, 2015: 239 A: YUN
rhinoceros Král, Malý & Schneider 2012: 3 A: AP ORR
sinensis P.H. Lucas, 1869b: ciii A: CHQ GAN HUN SCH SHA YUN “Thibet”
kumei Masumoto, 1991c: 179

GEOTRUPIDAE
subgenus Gynoplotrupes R. Oberthür, 1883a: 55 type species Enoplotrupes bieti R. Oberthür, 1883
Gynaecoplotrupes Boucomont, 1912a: 19 [unjustified emendation]

bieti bieti R. Oberthür, 1883a: 56 (Gynoplotrupes) A: BT SCH SD SHX UP YUN XIZ
potanini Semenov, 1889a: 401
serricornis Bates, 1891b: 73 (Geotrupes)

bieti morishitai Miyake, 1996: 33 A: NP
gansuensis Král, Malý & Schneider, 2015: 332 A: GAN
latus Boucomont, 1909a: 16 A: SCH YUN
yunnanus Fairmaire, 1888a: 17 (Enoplotrypes) A: YUN
genus Phelotrupes Jekel, 1866: 575 type species Geotrupes orientalis Westwood, 1839
subgenus Chromogeotrupes Bovo & Zunino, 1983: 404 type species Geotrupes auratus Motschulsky, 1858
auratus auratus Motschulsky, 1858a: 31 (Geotrupes) A: FE HEI JA NC NO SC
japonicus Jekel, 1866: 585 (Geotrupes)
purpurascens C.O. Waterhouse, 1875a: 97 (Geotrupes)
ruri Nakane, 1952a: 24 (Geotrupes)
viridiaureus Nakane, 1952a: 24 (Geotrupes)

auratus yaku Tsukamoto, 1973: 43 (Geotrupes) A: JA (Yaku-shima)
bicolor Fairmaire, 1888a: 18 (Geotrypes) A: SCH SHA YUN
immarginatus Boucomont, 1905: 231 (Geotrupes) A: SCH YUN
scutellatus Fairmaire, 1887b: 103 (Geotrypes) A: YUN
subgenus Eogeotrupes Bovo & Zunino, 1983: 407 type species Geotrupes laevistriatus Motschulsky, 1858
amethystinus Jekel, 1866: 582 (Geotrupes) A: SD ORR
cambeforti Král, Malý & Schneider, 2001: 89 A: BT NP SD XIZ
deuvei Král, Malý & Schneider, 2001: 90 A: FUJ GUA
formosanus Miwa, 1930: 164 (Geotrupes) A: TAI
laevifrons Jekel, 1866: 164 (Geotrupes) A: ANH GAN GUI HUB JIX SCH YUN ORR
laevistriatus Motschulsky, 1858a: 32 (Geotrupes) A: FE JA NC NE NO SC ZHE
deyrollei Jekel, 1866: 586 (Geotrupes)

oshimanus Fairmaire, 1895e: ccxxvii (Geotrypes) A: JA (Ryukyus)
subaeneus Ochi, Kon & Bai, 2010: 146 A: FUJ
tenuestriatus Fairmaire, 1887b: 103 (Geotrypes) A: SCH YUN
armicrus Fairmaire, 1888a: 17 (Geotrypes)

subgenus Phelotrupes Jekel, 1866: 575 type species Geotrupes orientalis Westwood, 1839
businskyorum Král, Malý & Schneider, 2001: 100 A: XIZ ORR
davidis Deyrolle, 1878: 97 (Geotrupes) A: SCH
denticulatus Boucomont, 1905: 232 (Geotrupes) A: SCH
holzschuhi Král, Malý & Schneider, 2001: 105 A: BT NP SD YUN ORR
imurai Masumoto, 1995b: 383 (Geotrupes) A: SHA
indicus Boucomont, 1905: 226 (Geotrupes) A: NP
haroldi Boucomont, 1905: 250 (Geotrupes) [RN]
jekeli Harold, 1869b: 104 (Geotrupes)

37

38

GEOTRUPIDAE
tenebrosus Fairmaire, 1901b: 267 (Geotrypes)

kerzhneri Shokhin, 2008: 289 A: YUN
kubani Král, Malý & Schneider, 2001: 110 A: YUN
lama Červenka, 2009: 19 A: XIZ
metallescens Fairmaire, 1894a: 218 (Geotrypes) A: HUB SCH
obscuratus Fairmaire, 1889a: 14 (Geotrypes) A: YUN
orientalis Westwood, 1839b: liv (Geotrupes) A: BT HP KA NP PA SD UP XIZ ORR
henrici Jekel, 1866: 579 (Geotrupes)
sylheticus Jekel, 1866: 580 (Geotrupes)

wrzecionkoi Král, Malý & Schneider, 2001: 118 A: XIZ
yunnanensis Král, Malý & Schneider, 2001: 120 A: YUN
subgenus Sinogeotrupes Bovo & Zunino, 1983: 401 type species Geotrypes substriatellus Fairmaire, 1897
armatus Boucomont, 1905: 246 (Geotrupes) A: SCH
belousovi Nikolajev, 2007b: 8 A: SCH
bolm Král, Malý & Schneider, 2001: 124 A: HUB HUN ZHE
cheni Ochi, Kon & Bai, 2010: 141 A: ZHE
compressidens Fairmaire, 1891d: vi (Geotrypes) A: FUJ HUB HUN ZHE
hunanensis Král, Malý & Schneider, 2001: 127 A: HUN
insulanus Howden, 1965: 502 (Geotrupes) A: FUJ TAI
jendeki Král, Malý & Schneider, 2001: 129 A: GUI
oberthuri Boucomont, 1905: 232 (Geotrupes) A: SCH
substriatellus Fairmaire, 1897f: 242 (Geotrypes) A: HUN SCH
taiwanus Miyake & Yamaya, 1995: 32 (Sinogeotrupes) A: FUJ JIA TAI ZHE
yangi Ochi, Kon & Bai, 2010: 143 A: HUB YUN
zhangi Ochi, Kon & Bai, 2010: 145 A: JIX YUN
genus Phelotrupes, species incertae sedis
nikolajevi Král, Malý & Schneider, 2001: 138 A: YUN
smetanai Král, Malý & Schneider, 2001: 139 A: YUN
variolicollis Boucomont, 1905: 226 (Geotrupes) A: YUN
genus Typhaeus Leach, 1815: 97 type species Typhaeus vulgaris Leach, 1815 (=Scarabaeus typhoeus
Linnaeus, 1758)
Armideus A. Villa & G.B. Villa, 1833: 16 type species Scarabaeus typhoeus Linnaeus, 1758
Mesotrypes Jakobson, 1892: 256 type species Geotrupes lateridens Guérin-Méneville, 1838
Minotaurus Mulsant & Godart, 1855a: 4 [=1855e: 4] type species Scarabaeus typhoeus Linnaeus, 1758

fossor Waltl, 1838: 456 E: BU GR MC TR A: LE SY TR
subarmatus Fairmaire, 1848: 172 (Geotrupes)

lateridens Guérin-Méneville, 1838c: 48 (Geotrupes) E: AL CR GR ME
quadrigeminus Fairmaire, 1859: 48 (Geotrupes)
subarmatus Erichson, 1847a: 727 (Geotrupes)

typhaeoides Fairmaire, 1852: 85 (Geotrupes) N: AG MO TU

39

GEOTRUPIDAE

typhoeus Linnaeus, 1758: 346 (Scarabaeus) E: AN BE CT (Kaliningrad) CZ DE FI FR GB GE IR IT LA LS LU NL
PL PT SP SV SZ N: MO
brunneus Mulsant, 1842: 355 (Ceratophyus)
pumilus Marsham, 1802: 8 (Scarabaeus)
pusillus Mulsant, 1842: 355 (Ceratophyus)
vulgaris Leach, 1815: 97

tribe Geotrupini Latreille, 1802

genus Allotrypes François, 1904: 64 type species Geotrupes mandibularis Reitter, 1896
mandibularis Reitter, 1896g: 271 (Geotrupes) N: AG MO TU
genus Anoplotrupes Jekel, 1866: 525 type species Scarabaeus sylvaticus Panzer, 1797 (=Scarabaeus stercorosus
Scriba, 1791)
Melanotrupes F. Blanchard, 1888: 105 type species Melanotrupes hornii F. Blanchard, 1888

stercorosus Scriba, 1791: 251 (Scarabaeus) E: AL AN AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR
IT KO LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP SV ST SZ UK A: ES TR WS
amoethysticus Mulsant, 1842: 362 (Geotrupes)
fauconneti Pic, 1926b: 9 (Geotrupes)
inaequalis Faldermann, 1835a: 263 (Geotrupes)
juvenilis Mulsant, 1842: 362 (Geotrupes)
monticola Heer, 1841: 499 (Geotrupes)
nigrinus Mulsant, 1842: 362 (Geotrupes)
prussicus Czwalina, 1884: 264 (Geotrupes)
sylvaticus Panzer, 1797c: 3 (Scarabaeus)
violaceus Dalla Torre, 1879: 111 (Geotrupes)
viridis Dalla Torre, 1879: 111 (Geotrupes)
viturati Pic, 1926b: 9 (Geotrupes)

genus Baraudia López-Colón, 1996: 362 type species Geotrupes geminatus Gené, 1839
geminata Gené, 1839: 61 (Geotrupes) E: FR (Corse) IT (Sardegna) SP (Menorca)
genus Ceratophyus Fischer von Waldheim, 1824: 143 type species Scarabaeus dispar Fabricius, 1781
(=Scarabaeus polyceros Pallas, 1771)
dauricus Jekel, 1866: 538 A: ES (Transbaikalia) MG NMO
hoffmannseggi Fairmaire, 1856: 530 (Geotrupes) E: PT SP N: MO
olivieri Jekel, 1866: 539
rossii Rosenhauer, 1856: 126 [NO]

kabaki Nikolajev, 2007c: 55 A: SCH
maghrebinicus Hillert & Král, 2013: 174 N: AG LB
martinezi Lauffer, 1909: 275 E: SP
mesasiaticus S.I. Medvedev & Nikolajev, 1974: 119 A: KI KZ XIN

40

GEOTRUPIDAE

polyceros Pallas, 1771: 461 (Scarabaeus) E: CT KZ ST UK A: KI KZ UZ (Karakalpakiya)
ammon Pallas, 1776: 707 (Scarabaeus)
dispar Fabricius, 1781a: 5 (Scarabaeus)
fischeri Fischer von Waldheim, 1823: pl. XVIII [=1824: 148]
menetriesi Mulsant & Rey, 1871a: 422 [=1871c: 166]
nigricans Jekel, 1866: 537
pallasi Mulsant & Rey, 1871a: 422 [=1871c: 166]

rossii Jekel, 1866: 539 [NP] E: IT
monoceros Jekel, 1866: 539
perroudi Mulsant & Rey, 1871a: 423 [=1871c: 167]

schaffrathi Hillert & Král, 2013: 177 E: GR (Kós)
sinicus Zunino, 1973: 17 A: BEI GAN MG NMO QIN SHX XIN
sulcicornis Fairmaire, 1887f: cxcii (Geotrypes) A: AP NP SD ORR
genus Geotrupes Latreille, 1797: 6 type species Scarabaeus stercorarius Linnaeus, 1758
subgenus Geotrupes Latreille, 1797: 6 type species Scarabaeus stercorarius Linnaeus, 1758
Geotrypes Agassiz, 1846: 161 type species Scarabaeus stercorarius Linnaeus, 1758

baicalicus Reitter, 1892a: 135 [=1893a: 14] E: CT KZ A: ES KZ MG WS
corinthius Fairmaire, 1886: 322 A: GAN SCH QIN XIN YUN
semicupreus Reitter, 1892a: 136 [=1893a: 15]

ibericus Baraud, 1958: 119 E: PT SP
jakowlewi Semenov, 1891: 314 A: AF KA KI KZ TD UZ XIN
foveatus Solsky, 1876a: 358 [HN]

kashmirensis Sharp, 1878b: 171 A: KA
koltzei Reitter, 1892a: 135 [=1893a: 14] A: ES FE GAN HEI JA MG NC NE NO SC SCH QIN
amoenus Jakobson, 1893: 120 (Geotrypes)

lenardoi Petrovitz, 1973d: 341 A: SA YE
mutator Marsham, 1802: 22 (Scarabaeus) E: AB AL AR AU BE BH BY CR CT CZ FR GB GE GG GR HU IT KO KZ
LA LS LT LU MC MD ME NL PL PT RO SB SK SL SP ST SV SZ TR UK A: IN KZ TR
aenescens Depoli, 1924: 156
cyanescens Luigioni, 1920: 186
fulvopubens Reitter, 1909b: 77
politus Malinowsky, 1811: 10 (Scarabaeus)
impressicollis Ferrari, 1852: 307
incertus Tournier, 1886: cxxiii
virescens Mulsant, 1842: 358

olgae Olsoufieff, 1918: 73 (Geotrypes) E: AR GG ST UK (Krym) A: IN TR
montanus Olsoufieff, 1918: 74 (Geotrypes)

spiniger Marsham, 1802: 21 (Scarabaeus) [NP] E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FR GB GE GR
HU IR IT LA LS LT LU MC MD ME NL NR RO SB SK SL SP ST SV SZ UK A: IN TM TR AURi
mesoleius C.G. Thomson, 1868: 331

GEOTRUPIDAE

41

puncticollis Malinowsky, 1811: 10 (Scarabaeus)
sublaevigatus Stephens, 1830: 185

stercorarius Linnaeus, 1758: 349 (Scarabaeus) E: AN AU BE BY CT CZ DE EN FI FR GB GE HU IR IT LA LS LT
LU MD NL NR NT PL PT RO SK SL SP SV SZ UK NARi
chalybaeus Mulsant, 1842: 358
exaratus Mulsant, 1842: 358
fimicola Mulsant & Godart, 1855b: 177 [=1855d: 277]
foveatus Marsham, 1802: 21 (Scarabaeus)
intermedius Ferrari, 1852: 306
juvencus Mulsant, 1842: 358
minor Erichson, 1847a: 728
punctatostriatus Stephens, 1830: 183
putridarius Erichson, 1847a: 730
subrugulosus Mulsant, 1842: 358
subviolaceus Mulsant, 1842: 358

subgenus Glyptogeotrupes Nikolajev, 1979e: 190 type species Geotrupes impressus Gebler, 1841
Epigeotrupes Bovo & Zunino, 1983: 430 type species Geotrupes kuluensis Bates, 1891

castanipennis Reitter, 1887b: 228 A: GAN SCH QIN
infraopacus Fairmaire, 1901b: 267

crenulipennis Fairmaire, 1891e: cxxii (Geotrypes) A: SCH YUN
folwarcznyi Červenka, 2005: 21 A: XIZ
genestieri Boucomont, 1905: 220 (Geotrupes) A: SCH
kuluensis Bates, 1891a: 13 A: KA SD
impressus Gebler, 1841a: 372 A: AF KI TD UZ
murrayi Ballion, 1871a: 334
turkestanicus Boucomont, 1899: 64

molestus Faldermann, 1835a: 264 A: TR
micanticollis Faldermann, 1835a: 265
caucasicus Sharp, 1871a: 11

ovchinnikovi Nikolajev, 2004d: 98 A: PA
subgenus Stereopyge A. Costa, 1847: 86 type species Geotrupes siculus A. Costa, 1847
Canthothrupes Jekel, 1866: 616 type species Geotrupes douei Gory, 1841
Streopuge A. Costa, 1847: 86 [unused original spelling]

douei Gory, 1841: 3 E: IT (Sicilia) MA N: AG MO TU
dentifrons Mulsant, 1843: 281
siculus A. Costa, 1847: 82

genus Jekelius López-Colón, 1989b: 72 type species Scarabaeus intermedius O.G. Costa, 1839
subgenus Jekelius López-Colón, 1989b: 72 type species Scarabaeus intermedius O.G. Costa, 1839
albarracinus Wagner, 1928: 153 (Thorectes) E: SP
balearicus López-Colón, 1985a: 318 (Thorectes) E: SP (Mallorca)

42

GEOTRUPIDAE

brullei africanus Baraud, 1965b: 148 (Geotrupes) N: AG LB TU
brullei anatolicus Jekel, 1866: 556 (Geotrupes) E: GR TR A: TR
brullei brullei Jekel, 1866: 554 (Geotrupes) E: AL BH CR GR IT ME SL
brullei creticus Fairmaire, 1876a: 25 (Geotrupes) E: GR (Kríti)
brullei syriacus Jekel, 1866: 544 (Geotrupes) A: IS JO SY
castillanus López-Colón, 1985b: 222 (Thorectes) E: SP
catalonicus López-Colón, 1991: 403 (Thorectes) E: SP
chalconotus Chevrolat, 1840: 15 (Geotrupes) E: PT
rugosicollis Jekel, 1866: 557 (Geotrupes)

chersinus J.-J. Delabie, 1954: 133 (Thorectes) E: FR SP
hernandezi López-Colón, 1988a: 115 (Thorectes) E: SP
hispanus Reitter, 1892a: 144 [=1893a: 23] (Geotrupes) E: PT SP
ardoini Baraud, 1966c: 192 (Geotrupes)

intermedius O.G. Costa, 1839a: 5 (Scarabaeus) E: FR IT SP (Menorca) N: AG TU
areolatus Reitter, 1892a: 151 [=1893a: 30] (Geotrupes)
cheroni Croissandeau, 1892b: ccxxi (Thorectes)
desjardinii Mulsant, 1842: 368 (Geotrupes)
italicus Jekel, 1866: 561 (Geotrupes)
lineicollis Mulsant, 1842: 368 (Geotrupes)
romanus Reitter, 1892a: 151 [=1893a: 30] (Geotrupes)
simplicidens Mulsant, 1842: 368 (Geotrupes)
subgeminalis Mulsant, 1842: 368 (Geotrupes)
subrugulosus Mulsant, 1842: 368 (Geotrupes)

juengeri Romero-Samper, 1995: 33 (Thorectes) E: SP
nitidus Jekel, 1866: 559 (Geotrupes) E: PT SP
heydeni Reitter, 1892a: 152 [=1893a: 31] (Geotrupes)
reitteri François, 1904: 140 (Geotrypes) [RN]

punctatolineatus François, 1904: 141 (Geotrypes) E: SP
sardous Erichson, 1847a: 738 (Geotrupes) E: FR (Corse) IT (Sardegna)
sericeus Jekel, 1866: 558 (Geotrupes) E: FR
subgenus Reitterius López-Colón, 1996: 367 type species Geotrupes punctulatus Jekel, 1866
punctulatus Jekel, 1866: 565 (Geotrupes) E: AL BU BH GR KO MC SB
brancsiki Apfelbeck, 1890: 167 (Geotrupes)
rhilensis Tesař, 1935: 186 (Geotrupes)

subgenus Rudolfpetrovitzia A. Rey & López-Colón, 2003: 138 [RN] type species Scarabaeus marginatus
Poiret, 1787
Petrovitzia López-Colón, 1996: 366 [HN] type species Scarabaeus marginatus Poiret, 1787

marginatus Poiret, 1787: 111 (Scarabaeus) E: IT (Sicilia) N: AG MO TU
hemisphaericus A.G. Olivier, 1789: no 3: 66 (Scarabaeus)
rotundatus P.H. Lucas, 1846a: pl. 24 (Geotrupes)
siculus Jekel, 1866: 570 (Geotrupes)

GEOTRUPIDAE
genus Odontotrypes Fairmaire, 1887b: 102 type species Geotrypes impressiusculus Fairmaire, 1887
subgenus Odontotrupes Boucomont, 1905: 234 type species Geotrupes orichalceus Fairmaire, 1895
Bootrupes Boucomont, 1911: 349 [RN] type species Geotrupes orichalceus Fairmaire, 1895

bhutan Král, Malý & Schneider, 2001: 59 A: BT
biconiferus Fairmaire, 1887b: 103 (Geotrypes) A: SCH
gogona Král, Malý & Schneider, 2001: 62 A: BT
lassallei Král, Malý & Schneider, 2001: 63 A: SCH
orichalceus Fairmaire, 1895d: ccv (Geotrupes) A: SD
subgenus Odontotrypes Fairmaire, 1887b: 102 type species Geotrypes impressiusculus Fairmaire, 1887
baimaensis Nikolajev, 2014b: 150 A: YUN
balthasari Mikšić, 1958a: 3 (Geotrupes) A: NP XIZ
bimaculatus Červenka, 2005: 20 A: XIZ
cariosus Fairmaire, 1886: 322 (Geotrypes) A: YUN
caverniferus Nikolajev, 2014a: 223 A: YUN
cicatricosus Nikolajev, 1977: 380 (Geotrupes) A: YUN
davidiani Nikolajev, 2009a: 47 A: SCH
duplopunctatus Nikolalev & Kolov, 2014a: 224 A: YUN
emei Král, Malý & Schneider, 2001: 16 A: SCH
gnom Nikolajev, 2009a: 47 A: SCH
gongga Král, Malý & Schneider, 2001: 17 A: SCH
haba Král, Malý & Schneider, 2001: 19 A: YUN
hayekae adamiki Červenka, 2007: 22 A: XIZ
hayekae hayekae Mikšić, 1961: 271 (Geotrupes) A: NP XIZ
howdeni Král, Malý & Schneider, 2001: 21 A: XIZ
hubeicus Král, Malý & Schneider, 2014: 214 A: HUB
impressiusculus Fairmaire, 1887b: 101 (Geotrypes) A: YUN
jiuding Král, Malý & Schneider, 2001: 24 A: SCH
kabakovi Nikolajev, 2006a: 43 A: YUN
kalabi Král, Malý & Schneider, 2001: 25 A: SCH
karnali Král, Malý & Schneider, 2001: 26 A: NP
kryzhanovskii Nikolajev, 1998: 39 (Geotrupes) A: YUN
kucerai Král, Malý & Schneider, 2001: 29 A: YUN
lama Král, Malý & Schneider, 2001: 30 A: SCH
leptopterus Nikolajev, 2014b: 150 A: YUN
luguensis Nikolajev, 2013a: 243 A: YUN
medvedevi Nikolajev, 1998: 41 (Geotrupes) A: SCH
meigu Shokhin, 2008: 287 A: SCH
meyomintang Král, Malý & Schneider, 2001: 32 A: SCH
mirek Král, Malý & Schneider, 2001: 33 A: YUN
paulusi Král, Malý & Schneider, 2001: 35 A: XIZ
purpureiaeneus Nikolajev, 1977: 381 (Geotrupes) A: SCH

43

44

GEOTRUPIDAE

purpureipunctatus Boucomont, 1905: 239 (Geotrupes) A: SCH
qinling Král, Malý & Schneider, 2001: 39 A: SHA
radiosus Fairmaire, 1895e: ccxxviii (Geotrypes) A: SCH
pauma Král, Malý & Schneider, 2001: 34

roborowskyi Reitter, 1887b: 229 (Geotrupes) A: GAN QIN
corrugatus J. Frivaldszky, 1890: 198 (Geotrupes)
koiwayai Masumoto, 1995b: 384 (Geotrupes)

rufipes Boucomont, 1905: 242 (Geotrupes) A: SCH
rugososulcatus Nikolajev, 2009a: 48 A: SCH
sabde Král, Malý & Schneider, 2001: 45 A: SCH
satanas Král, Malý & Schneider, 2001: 46 A: SCH
semiscribrosus Fairmaire, 1891e: cxxii (Geotrypes) A: KA
sichuanensis Nikolajev, 2013a: 244 A: SCH
szetshwanus Nikolajev, 1977: 381 (Geotrupes) A: SCH
taurus Boucomont, 1905: 240 (Geotrupes) A: GAN SCH
paradoxus Mikšić, 1958a: 3 (Geotrupes)

tibetanus Nikolajev, 1977: 379 (Geotrupes) A: QIN
turnai Král, Malý & Schneider, 2001: 53 A: SCH
uenoi Masumoto, 1995b: 382 (Geotrupes) A: SHA
yulong Král, Malý & Schneider, 2001: 56 A: YUN
zhongdianensis cavazzutii Král, Malý & Schneider, 2001: 14 A: YUN
zhongdianensis zhongdianensis Král, Malý & Schneider, 2001: 57 A: YUN
subgenus Thorectomimus Nikolajev, 2009c: 58 type species Geotrupes semenowi Reitter, 1887
cribripennis Fairmaire, 1887b: 102 (Geotrypes) A: YUN
densopunctatus Nikolajev & Kolov, 2014b: 150 A: YUN
farkaci Král, Malý & Schneider, 2001: 68 A: YUN
glaber Nikolajev, 1998: 40 (Geotrupes) A: SCH
kabaki Nikolajev, 2005b: 29 A: SCH
korolevi Nikolajev, 2013b: 356 A: YUN
mursini Nikolajev, 1998: 40 (Geotrupes) A: YUN
semenowi nikodymi Král, Malý & Schneider, 2001: 70 A: GAN SCH
semenowi semenowi Reitter, 1887b: 230 (Geotrupes) A: QIN
semirugosus Fairmaire, 1887b: 102 (Geotrypes) A: YUN
shokhini Nikolajev, 2009c: 60 A: YUN
slavek Král, Malý & Schneider, 2001: 75 A: YUN
tryznai Král, Malý & Schneider, 2001: 76 A: XIZ
xue Král, Malý & Schneider, 2001: 78 A: YUN
genus Renaudtrupes López-Colón, 2006: 416 [RN] type species Thorectes distinctus Marseul, 1878
Renaudia López-Colón, 1996: 359 [HN] type species Thorectes distinctus Marseul, 1878

distinctus Marseul, 1878: 59 (Thorectes) N: AG MO

GEOTRUPIDAE
genus Sericotrupes Zunino, 1984: 66 type species Scarabaeus niger Marsham, 1802
niger Marsham, 1802: 22 (Scarabaeus) E: BE FR GE IT NL PT SP SZ N: AG MO TU
hypocrita Illiger, 1803: 209 (Scarabaeus)
laevicollis Mulsant, 1842: 361 (Geotrupes)
substriatus Mulsant, 1842: 361 (Geotrupes)
subvirescens Mulsant, 1842: 361 (Geotrupes)

genus Silphotrupes Jekel, 1866: 553 type species Geotrupes punctatissimus Chevrolat, 1840
escorialensis escorialensis Jekel, 1866: 572 (Geotrupes) E: PT SP
silphoides Jekel, 1866: 573 (Geotrupes)

escorialensis opaculus Reitter, 1892a: 154 [=1893a: 33] (Geotrupes) E: PT SP
petrovitzi Baraud, 1970a: 141 (Thorectes)

orocantabricus Verdú & Galante, 2000: 197 (Thorectes) E: SP
punctatissimus Chevrolat, 1840: 16 (Geotrupes) E: PT SP
genus Thorectes Mulsant, 1842: 367 type species Scarabaeus laevigatus Fabricius, 1798
armifrons Reitter, 1892a: 150 [=1893a: 29] (Geotrupes) N: MO
asperifrons Fairmaire, 1866b: 256 (Geotrupes) [DA] A: TR
banghaasi Reitter, 1892a: 146 [=1893a: 25] (Geotrupes) [DA] A: TD
bankhaasi Reitter, 1892a: 146 [incorrect original spelling]

baraudi López-Colón, 1981: 277 E: SP
coiffaiti Baraud, 1969: 359 [DA] N: MO
coloni Ruiz, 1998a: 62 [DA] N: MO
demoflysi Baraud, 1965b: 151 (Geotrupes) [DA] N: LB TU
ferreri López-Colón, 1983: 299 E: SP
laevigatus Fabricius, 1798: 23 (Scarabaeus) N: AG MO
truncaticornis Reitter, 1892a: 148 [=1893a: 27] (Geotrupes)

latus Sturm, 1826: 65 (Geotrupes) [DA] N: AG LB TU
inflatus Reitter, 1892a: 144 [=1893a: 23] (Geotrupes)

lusitanicus Jekel, 1866: 653 (Geotrupes) E: PT SP
cobosi Baraud, 1965b: 144 [=1966: 184] (Geotrupes)

martensi Krikken, 1981: 372 (Geotrupes) [DA] A: NP
nepalensis Baraud, 1974: 123 (Geotrupes) [DA] A: NP
puncticollis P.H. Lucas, 1846a: pl. 24 [DA] N: AG LB MO TU
reflexus Jekel, 1866: 568 (Geotrupes) [DA] N: TU
rugatulus Jekel, 1866: 562 (Geotrupes) [DA] N: AG EG MO TU
semisericeus Jekel, 1866: 564 (Geotrupes)

shankara Carpaneto & Mignani, 1999: 1005 [DA] A: HP
stellosus annapurnicus Krikken, 1981: 377 (Geotrupes) [DA] A: NP
stellosus stellosus Krikken, 1981: 374 (Geotrupes) [DA] A: NP
trituberculatus Reitter, 1892a: 150 [=1893a: 29] (Geotrupes) [DA] N: AG MO

45

46

GEOTRUPIDAE

valencianus Baraud, 1966c: 186 (Geotrupes) E: SP
variolipennis Marseul, 1876: ci [DA] N: MO
subcostatus Fairmaire, 1883d: cix (Geotrupes)

genus Trypocopris Motschulsky, 1860b: 160 type species Scarabaeus vernalis Linnaeus, 1758
subgenus Lethrotrypes Jakobson, 1892: 257 type species Geotrupes inermis Ménétriés, 1832
inermis Ménétriés, 1832: 183 (Geotrupes) E: GG ST (Kavkaz)
subgenus Pseudotrypocopris Mikšić, 1954d: 246 type species Geotrupes amedei Fairmaire, 1861
amedei Fairmaire, 1861: 582 (Geotrupes) E: BU GR TR A: LE SY TR
subgenus Trypocopris Motschulsky, 1860b: 160 type species Scarabaeus vernalis Linnaeus, 1758
Silotrupes Mulsant & Rey, 1871a: 450 [=1871c: 194] type species Silotrupes epistomalis Mulsant & Rey, 1871 (=Geotrupes
alpinus Sturm & Hagenbach, 1825)
Sternotrupes Jekel, 1866: 618 type species Scarabaeus vernalis Linnaeus, 1758

alpinus alpinus Sturm & Hagenbach, 1825: 487 (Geotrupes) E: AU GE IT SL SZ
auverti Croissandeau, 1892a: lix (Silotrupes)
epistomalis Mulsant & Rey, 1871a: 450 [=1871c: 194] (Silotrupes)
obscurus Mulsant, 1842: 365 (Geotrupes)

alpinus balcanicola Mikšić, 1954d: 244 (Geotrupes) E: AL BH IT ME
alpinus marianii Löbl, 2006: 30 [RN] E: FR IT
opacus Mariani, 1958: 38 (Geotrupes) [HN]

caspius Motschulsky, 1845a: 55 (Geotrupes) E: GG ST A: IN TR
caucasicus J. Weise, 1879: 193 (Geotrypes)
laticollis Motschulsky, 1845a: 55 (Geotrupes)

fausti Reitter, 1890j: 195 (Geotrupes) E: AB A: IN
fulgidus Motschulsky, 1845a: 55 (Geotrupes) E: BU TR A: TR
purpureus Küster, 1852: 46 (Geotrupes)

pyrenaeus bartolii G. Dellacasa, 1974: 9 E: IT
pyrenaeus cyanicolor Capra, 1930: 122 (Geotrupes) E: IT
pyrenaeus pyrenaeus T. Charpentier, 1825: 208 (Scarabaeus) E: BE FR GB PT SP
coruscans Chevrolat, 1840: 15 (Geotrupes)

pyrenaeus splendens Heer, 1841: 499 (Geotrupes) E: GE IT SL SZ
autumnalis Heer, 1841: 499 (Geotrupes)
erichsoni Boucomont, 1911: 348 (Geotrupes)
heyrovskyi Balthasar, 1929c: 112 (Geotrupes)

vernalis apenninicus Mariani, 1958: 31 (Geotrupes) E: IT
vernalis fauveli Bedel, 1911b: 379 (Geotrypes) [RN] E: FR IT SP
obscurus Fauvel, 1892: 57 (Geotrypes) [HN]

vernalis vernalis Linnaeus, 1758: 349 (Scarabaeus) E: AL AN AU BE BH BU BY CR CT CZ DE EN FI FR GB GE
GR HU IR IT KO LA LS LT LU MC MD ME NL NR NT PL RO SB SK SL SP SV ST SZ UK A: TR
autumnalis Erichson, 1847a: 737 (Geotrupes)
balcanicus Reitter, 1892a: 139 [=1893a: 18] (Geotrupes)

47

GEOTRUPIDAE
cyanescens Leoni, 1911c: 63
cyanicollis Depoli, 1938: 247 (Geotrupes)
insularis O. Schneider, 1898: 69 (Geotrupes)
laevis Haworth, 1807: 79 (Scarabaeus)
manifestus Reitter, 1892a: 141 [=1893a: 20] (Geotrupes) [DA]
politus Mulsant, 1842: 366 (Geotrupes)
rambouseki Tesař, 1935: 186 (Geotrupes)
splendens Erichson, 1847a: 737 (Geotrupes)
sulzeri Gistel, 1857b: 567 (Geotrupes)
thoracalis Depoli, 1938: 247 (Geotrupes)
timotheus Gistel, 1857b: 571 (Geotrupes)
varians Mulsant, 1842: 365 (Geotrupes)
violaceus Mulsant, 1842: 365 (Geotrupes)
viridissimus Mikšić, 1958a: 1 (Geotrupes)

zaitzevi Olsoufieff, 1918: 75 (Geotrypes) E: AB A: TR
genus Zuninoeus López-Colón, 1989b: 74 type species Geotrupes hoppei Sturm & Hagenbach, 1825
hoppei Sturm & Hagenbach, 1825: 486 (Geotrupes) E: CR IT SL
aurichalceus J. Müller, 1902: 450 (Geotrupes)
rugulosus T. Charpentier, 1825: 209 (Geotrupes)

subfamily Lethrinae Oken, 1843

genus Lethrus Scopoli, 1777: 439 type species Lucanus apterus Laxmann, 1770
subgenus Abrognathus Jakovlev, 1890: 563 type species Lethrus tuberculifrons Ballion, 1871
Microlethrus Semenov, 1894: 476 type specie Lethrus pygmaeus Ballion, 1871

majusculus Semenov, 1899c: 632 A: UZ
pygmaeus Ballion, 1871a: 337 A: KI KZ UZ
tuberculifrons Ballion, 1871a: 337 A: KI KZ
breviceps Ballion, 1871a: 338

subgenus Autolethrus Semenov, 1892a: 236 type species Lethrus laevigatus Ballion, 1871 (=Lethrus scoparius
Fischer von Waldheim, 1821)
anisodon Semenov & Gussakowski, 1935: 1401 A: UZ
appendiculatus Jakovlev, 1891a: 122 A: AF TD
afghanus Král & Olexa, 1996: 53
coptotis Semenov, 1894: 514
degener S.I. Medvedev, 1962: 598

arnoldii Nikolajev, 1987: 46 A: UZ
bajsuntavicus Nikolajev, 2003a: 183 A: UZ
chistjakovae Nikolajev, 2003a: 184 A: UZ
geminatus Kraatz, 1882i: 311 A: UZ
glaber S.I. Medvedev, 1962: 596 A: TD

48

GEOTRUPIDAE

karateghinicus Nikolajev, 1976c: 233 A: TD
marakandicus Nikolajev, 2003a: 178 A: UZ
mucronatus Semenov, 1894: 522 A: KI UZ
obliteratus Semenov, 1894: 522
taschkumyricus Protsenko, 1970: 16

politus Solsky, 1876a: 366 A: TD UZ
scoparius Fischer von Waldheim, 1821a: 11 A: KI KZ UZ
bradytus Fairmaire, 1892c: cxxii
laevigatus Ballion, 1871a: 334
marquardti Reitter, 1902c: 138

sogdianus Semenov, 1894: 516 A: UZ
splendidus Semenov & S.I. Medvedev, 1935: 284 A: UZ
submandibularis Lebedev, 1932: 86 A: UZ
sulcipennis Kraatz, 1883e: 340 A: KI UZ
dinotherium Wilkins, 1885: 292
litvinowi Semenov, 1900c: 308
spathulatus Lebedev, 1932: 88

superbus Kraatz, 1882i: 312 A: TM UZ
fedtschenkoi Semenov, 1894: 519

tshatkalensis Protsenko, 1965: 14 A: KI UZ
vachshianus Nikolajev & Schukronajev, 1989: 154 A: TD
zeravshanicus Nikolajev, 2003a: 179 A: TD
subgenus Ceratodirus Fischer von Waldheim, 1845: 340 type species Ceratodirus dispar Fischer von Waldheim,
1845 (=Lethrus longimanus Fischer von Waldheim, 1821)
aequidentatus Nikolajev, 1968: 536 A: KZ
aralicus Nikolajev, 2003a: 86 A: KZ UZ
borealis Nikolajev, 1973: 859 E: ST (Kumak) A: KZ
cephalotes Pallas, 1771: 461 (Scarabaeus) E: ST (Akbulak) A: KZ
dostojewskii Semenov, 1899c: 638 A: KZ
desertus Skopina, 1965: 109 (Ceratodirus)
miliaris Skopina, 1965: 108 (Ceratodirus)

gladiator Reitter, 1897h: 213 A: KZ
karelini Gebler, 1845: 330 A: KI KZ
subaurosus Reitter, 1909b: 78

lamellifer Nikolajev, 1976c: 233 A: KZ
longimanus Fischer von Waldheim, 1821a: 12 E: ST (Orenburg, Volgograd) A: KZ
dispar Fischer von Waldheim, 1845: 340
eversmannii Krynicki, 1832: 122

mugodzharicus Nikolajev, 1987: 44 A: KZ
serpentifer Semenov & S.I. Medvedev, 1935: 286 A: QIN
subgenus Furcilethrus Nikolajev, 1968: 533 type species Lethrus furcatus Jakovlev, 1890

GEOTRUPIDAE
antovae S.I. Medvedev, 1957: 695 A: TD
rugulosus S.I. Medvedev, 1959: 860

banghaasi Reitter, 1893e: 222 A: TD
inconspicuus S.I. Medvedev, 1959: 863

frantsevichi Nikolajev, 1979b: 65 A: TD
frater Nikolajev, 1975c: 1259 A: TD
furcatus Jakovlev, 1890: 568 A: TD
kiritschenkoi S.I. Medvedev, 1965b: 169 A: TD
legezini Nikolajev, 2001: 103 A: TD
michailovi Nikolajev & Schukronajev, 1977: 157 A: TD
mikitovae Nikolajev & Schukronajev, 1989: 153 A: TD
muskinabadensis Nikolajev & Ivanova, 2014: 65 A: TD
nasreddinovi Nikolajev, 1987: 43 A: TD
paki Ivanova, 2012a: 45 A: TD
rustam Nikolajev & Ivanova, 2014: 69 A: TD
saryhissoricus Nikolajev, 1987: 42 A: TD
sohrab Nikolajev, 1976c: 232 A: TD
tadzhikorum S.I. Medvedev, 1959: 862 A: TD
tenuidens obsoletus Semenov, 1894: 513 A: KI TD
tenuidens tenuidens Reitter, 1890n: 294 A: TD
staudingeri Reitter, 1893e: 222

subgenus Heteroplistodus Jakovlev, 1890: 563 type species Lethrus crenulatus Gebler, 1845
Lethrulus Semenov, 1892a: 235 type species Lethrus cicatricosus Reitter, 1889

bispinus Jakovlev, 1889b: 259 A: KI UZ “Fergana”
talassicola S.I. Medvedev, 1965b: 171

bulbocerus bulbocerus Fischer von Waldheim, 1845: 341 A: KZ
jakowlewi Semenov, 1892a: 238

bulbocerus tarbagataicus Nikolajev, 2001: 102 A: KZ
cicatricosus Reitter, 1889d: 104 [=1890n: 295] A: AF TM
ciskungesicus Nikolajev, 2001: 101 A: XIN
crenulatus Gebler, 1845: 334 A: KZ
kentauensis Král & Olexa, 1996: 50 A: KZ
korshinskii Semenov, 1899c: 635 A: TD UZ
kuldshensis Lebedev, 1926: 38 A: XIN
lebedevi Semenov & S.I. Medvedev, 1935: 287 A: KZ
petri Semenov & S.I. Medvedev, 1935: 287

lopatini S.I. Medvedev, 1959: 866 A: TD
medvedevi Semenov & Gussakowski, 1935: 1400 A: TD
obliquus Semenov, 1894: 510 A: AF TD
potanini Jakovlev, 1889b: 261 A: GAN MG NIN NMO
conformis J. Frivaldszky, 1890: 199

49

50

GEOTRUPIDAE

serridens Nikolajev, 1971: 128 A: TD
tekesicus Nikolajev, 2003a: 72 A: XIN
tschitscherini Semenov, 1894: 508 A: KZ XIN
uncidens Nikolajev, 1976c: 231 A: TD UZ
subgenus Lethrus Scopoli, 1777: 439 type species Lucanus apterus Laxmann, 1770
Bulbocerus Acharius, 1781: 246 type species Bulbocerus cephalotes Acharius, 1781 (=Lucanus apterus Laxmann, 1770)
Clunipes Hochentwarth, 1785: 347 type species Clunipes scarabaeoides Hochenwarth, 1785 (=Lucanus apterus
Laxmann, 1770)
Goniolethrus Semenov, 1894: 478 type species Lethrus brachiicollis Fairmaire, 1855

apterus Laxmann, 1770: 594 (Lucanus) E: AU BH BU CR CZ HU LA PL MD RO SB SK ST UK
cephalotes Acharius, 1781: 249 (Bulbocerus)
podolicus Fischer von Waldheim, 1822: 139
scarabaeoides Hochenwarth, 1785: 347 (Clunipes)
subaeneus Fairmaire, 1866b: 257

ares Král, Rejsek & Schneider, 2001: 254 E: GR
armeniacus Reitter, 1890n: 294 A: TR
brachiicollis Fairmaire, 1855b: 314 A: TR
elephas Reitter, 1890n: 290 E: AL GR MC
mandibularis Jakovlev, 1892: 182

fallax Nikolajev, 1975a: 147 E: GR TR A: TR
halkidikiensis Král & Hillert, 2013a: 222 E: GR
liviae Pittino, 2011b: 382 E: TR
macrognathus Fairmaire, 1866b: 255 A: TR
perun Král & Hillert, 2013a: 227 E: BU
raymondi Reitter, 1890n: 293 E: GR MC
rotundicollis Fairmaire, 1866b: 256 E: GR (Lesvós) A: TR
heydenii Fairmaire, 1866b: 256

schaumii Reitter, 1890n: 293 E: BU GR
forcipatus Jakovlev, 1892: 187

schneideri Král & Hillert, 2013b: 95 E: GR
strymonensis Král & Hillert, 2013a: 234 E: GR
subgenus Mesolethrus Nikolajev, 2003a: 122 type species Lethrus microbuccis Ballion, 1871
elisae Nikolajev, 2001: 102 A: UZ
inermis Reitter, 1897k: 235 A: KI UZ TD
microbuccis Ballion, 1871a: 336 A: KI UZ TD
puncticollis Kraatz, 1882i: 312
glasunowi D. Koshantschikov, 1894c: 118

nuratavicus Nikolajev, 1987: 41 A: UZ
shakhristanicus Nikolajev, 2003a: 129 A: TD
subgenus Neolethrus Nikolajev, 1987: 50 type species Lethrus arcanus S.I. Medvedev, 1971
arcanus S.I. Medvedev, 1971: 120 A: TM

GEOTRUPIDAE
subgenus Paralethrus Nikolajev, 2003a: 191 type species Lethrus turkestanicus Ballion, 1871
bituberculatus bituberculatus Ballion, 1871a: 338 A: KZ UZ
bituberculatus Lebedev, 1913: 6 [HN]
bituberculifrons Lebedev, 1926: 40 [RN]

bituberculatus impressifrons Ballion, 1871a: 336 A: KI KZ UZ
eous Semenov, 1894: 522
hauseri Reitter, 1894b: 44
jacobsoni Semenov & S.I. Medvedev, 1935: 285
nigroaeneus Skopin, 1955: 150

bituberculatus macrodon Nikolajev, 1969: 528 A: KI KZ
crassus Hillert, 2004: 824 A: UZ
kabaki Nikolajev, 1998: 41 A: KI UZ
karatavicus Nikolajev & Skopin, 1971: 27 A: KZ
turkestanicus Ballion, 1871a: 335 A: KZ UZ
aenescens Fairmaire, 1892c: cxxi
silus Reitter, 1894b: 43
zarudnyi Semenov & S.I. Medvedev, 1935: 284

subgenus Scelolethrus Semenov, 1892a: 236 type species Lethrus rosmarus Ballion, 1871
aktavicus Nikolajev, 2003c: 179 A: TD
andrejewae Nikolajev, 1987: 48 A: TD
auriculatus Semenov, 1894: 500 A: TD
bactrianus Semenov, 1894: 502 A: TD
baglanicus Nikolajev, 2003c: 189 A: AF
carinatus Nikolajev, 1976c: 234 A: AF TD
chorassanicus Semenov & S.I. Medvedev, 1935: 283 A: IN
costatus Semenov, 1894: 497 A: TD
crypticus Král & Olexa, 1996: 59 A: TM
gissaricus Nikolajev, 1976c: 237 A: TD UZ
kattaghanicus Nikolajev, 1976c: 236 A: AF
koshantschikowi Semenov, 1892a: 240 A: TD
bucharicus Semenov, 1894: 504

kryzhanovskii S.I. Medvedev, 1971: 119 A: AF TD
mithras Reitter, 1904d: 256 A: IN TM
zarudnianus Semenov & S.I. Medvedev, 1935: 282

nikolajevi S.I. Medvedev, 1971: 117 A: AF TD
obtritus Nikolajev, 1976c: 235 A: TD
percutens Gusakov & Klimenko, 2011: 3 A: IN
rosmarus Ballion, 1871a: 336 A: KZ UZ TD TM
spinimanus Jakovlev, 1892: 190 A: TM
substriatus Kraatz, 1883e: 341 A: TM
turcomanicus Semenov, 1894: 507

51

52

PASSALIDAE

sulcatus Kraatz, 1883e: 342 A: TM
komarowi Semenov, 1894: 491

tekkensis Král & Olexa, 1996: 62 A: TM
tenuepunctus Semenov & S.I. Medvedev, 1935: 283 A: TD
tenuestriatus Nikolajev, 1976c: 237 A: TD
valentinae Nikolajev, 2003a: 224 A: UZ
subgenus Teratolethrus Semenov, 1894: 477 type species Lethrus sieversi D. Koshantschikov, 1894
acutangulus Ballion, 1871a: 335 A: IN
persicus Reitter, 1890n: 291
baiocchii Hillert & Sechi, 2014: 706 A: IN
sieversi D. Koshantschikov, 1894: [149] A: TM
genus Lethrus, species incertae sedis
mediocris Fairmaire, 1892c: cxxii A: “Turkestan”
genus Lethrus, nomen dubium
semenowi D. Koshantschikov, 1893: 137 A: TD

family PASSALIDAE Leach, 1815
subfamily Aulacocyclinae Kaup, 1868

genus Ceracupes Kaup, 1871: 16 type species Passalus fronticornis Westwood, 1842
arrowi Heller, 1911: 256 A: TAI ORR
chingkini Okano, 1988: 2 A: GUX TAI ORR
fronticornis Westwood, 1842d: 124 (Passalus) A: AP BT HAI SD XIZ ORR
austeni Stoliczka, 1873: 151

yui Okano, 1988: 1 A: TAI
genus Cylindrocaulus Fairmaire, 1880a: 164 type species Passalus bucerus Fairmaire, 1880
Auritulus Zang, 1905b: 229 type species Aulacocyclus patalis Lewis, 1883

bucerus Fairmaire, 1880a: 164 A: SCH
davidi Boucher & Reyes-Catillo, 1996: 88 A: GAN SCH
mishmi Kon, Araya & Johki, 2015: 127 A: AP
patalis Lewis, 1883: 341 (Aulacocyclus) A: JA
genus Taeniocerus Kaup, 1871: 41 type species Passalus bicanthatus Percheron, 1841
bicuspis Kaup, 1868: 5 (Aulacocyclus) A: AP BT SD ORR

subfamily Passalinae Leach, 1815

genus Aceraius Kaup, 1868: 26 type species Passalus grandis Burmeister, 1847
grandis Burmeister, 1847: 463 (Passalus) A: AP BT SD SE SW TAI XIZ ORR
chinensis Kuwert, 1898: 344
hirsutus Kuwert, 1891: 163 (Acerajus)

53

TROGIDAE
helferi Kuwert, 1891: 163 (Acerajus) A: AP NC SD SE SW XIZ ORR
assamensis Kuwert, 1898: 347
himalayensis Gravely, 1914: 236

genus Leptaulax Kaup, 1868: 11 type species Passalus dentatus Fabricius, 1792
Leptaulacides Zang, 1905a: 106 type species Leptaulax eschscholtzi Kaup, 1868 (=Passalus bicolor Fabricius, 1801)

bicolor Fabricius, 1801b: 256 (Passalus) A: AP BT SD SE SW XIZ AUR ORR
aurivillii Kuwert, 1891: 189
eschscholtzi Kaup, 1868: 14
evidens Kuwert, 1898: 295

cyclotaenius Kuwert, 1891: 188 A: AP SD ORR
anipunctatus Zang, 1905c: 234 (Leptaulacides)
himalayae Kuwert, 1898: 285

dentatus Fabricius, 1792b: 241 (Passalus) A: AP BT HAI NP SD SE SW TAI XIZ AUR ORR
interponendus Kuwert, 1898: 299

formosanus van Doesburg, 1992: 525 A: TAI
koreanus Nomura, Kon, Johki & C.E. Lee, 1993: 52 A: SC
genus Macrolinus Kaup, 1868: 18 type species Passalus latipennis Percheron, 1841
foveolatus Ma, 1988: 268 A: XIZ
medogensis Y.-W. Zhang, 1981: 340 A: XIZ
sikkimensis Stoliczka, 1873: 161 (Basilianus) A: AP SD ORR
genus Ophrygonius Zang, 1904: 697 type species Ophrygonius quadrifer Zang, 1904
birmanicus Gravely, 1914: 226 A: AP BT ORR
cantori Percheron, 1844: 3 (Passalus) A: BT NP SD XIZ ORR
dunsiriensis Gravely, 1914: 225
interrogationis Kuwert, 1891: 164 (Basilianus)

chinensis Endrődi, 1955c: 232 A: CE SE
convexifrons Zang, 1904: 698 (Basilianus) A: BT SD ORR
genus Tiberioides Gravely, 1913: 405 type species Tiberius kuwerti Arrow, 1907
austeni Gravely, 1914: 216 A: AP BT SD
borealis Arrow, 1907c: 446 (Chilomazus) A: SD ORR
kuwerti Arrow, 1907c: 446 (Tiberius) A: AP BT NP SD ORR

family TROGIDAE W.S. Macleay, 1819
subfamily Omorginae Nikolajev, 2005

genus Afromorgus Scholtz, 1986: 362 type species Trox squalidus A.G. Olivier, 1789
chinensis Boheman, 1858: 52 (Trox) A: FUJ GUX HAI HKG HUB HUN JIX NO NE TAI JA ORR
obscurus C.O. Waterhouse, 1875: 98 (Trox)

54

TROGIDAE

frater Pittino, 2005: 74 (Omorgus) A: PA ORR
gemmatus A.G. Olivier, 1789: no 4: 7 (Trox) N: EG A: AE AF OM SA AFR
granulatus Herbst, 1783: 12 (Trox) A: AF HP PA SD ORR
guttalis Haaf, 1954: 336 (Trox) N: EG AFR
haagi Harold, 1872: 59 (Trox) A: PA ORR
inclusus Walker, 1858: 208 (Trox) A: HAI ORR
italicus Reiche, 1853b: 89 (Trox) Ei: IT A: CH HP KA PA SD ORR
maindroni Pittino, 2005: 78 (Omorgus) A: PA
melancholicus Fåhraeus, 1857: 378 (Trox) Ei: IT (Sicilia) A: YE AFR
larvatus Gerstäcker, 1873: 309 (Trox)
madagascariensis Fairmaire, 1871b: 34 (Trox)

niloticus desertorum Harold, 1872: 77 (Trox) N: EG A: AE SA YE AFR
philbyi R. Paulian, 1948: 143 (Trox)

niloticus niloticus Harold, 1872: 79 (Trox) N: EG AFR
erlangeri H.J. Kolbe, 1904b: 293 (Trox)
massaicus H.J. Kolbe, 1904b: 293 (Trox)

omacanthus Harold, 1872: 64 (Trox) A: SD UP ORR
pauliani Haaf, 1954: 385 (Trox) A: HEN HKG HUB NP SD SW TAI ORR
procerus Harold, 1872: 127 (Trox) N: EG A: AE SA IN PA AFR ORR
reiterorum Král & Kubáň, 2012: 148 A: YE (Suqutra)
squalidus A.G. Olivier, 1789: no 4: 12 (Trox) N: AG EG
incultus Fåhraeus, 1857: 377 (Trox)

testudo Arrow, 1927: 466 (Trox) A: PA ORR
verrucosus Reiche & Saulcy, 1856: 404 (Trox) N: EG A: AE IQ SY YE AFR
nomadicus Scholtz, 1980: 138 (Trox)

genus Omorgus Erichson, 1847b: 111 type species Trox suberosus Fabricius, 1775
Chesas Burmeister, 1876: 264 type species Trox pastillarius C.É. Blanchard, 1846
Lagopelas Burmeister, 1876: 265 type species Trox ciliatus C.É. Blanchard, 1846
Megalotrox Preudhomme de Borre, 1886a: 59 type species Trox gigas Harold, 1872

costatus Wiedemann, 1823: 30 (Trox) A: HAI HEN SW TAI AUR ORR
montalbanensis Schultze, 1915: 272 (Trox)
regularis Harold, 1868b: 85 (Trox)
velutinus Blackburn, 1892a: 40 (Trox)

nanningensis Pittino, 2005: 71 A: GUX
subcarinatus W.J. Macleay, 1864: 128 (Trox) Ei: SP AUR
fenestratus Harold, 1872: 97 (Trox)

suberosus Fabricius, 1775: 31 (Trox) Ei: BE CZ HU SP Ni: CI MO AFRi AURi NAR NTR
alternatus Say, 1835: 179 (Trox)
crenatus A.G. Olivier, 1789: no 4: 7 (Trox)
denticulatus Palisot de Beauvois, 1805: 176 (Trox) [HN]

55

TROGIDAE
gibbus A.G. Olivier, 1789: no 4: 13 (Trox)
manilensis Schultze, 1915: 271 (Trox)
nobilis Wollaston, 1867: 93 (Trox)
novaecaledoniae Balthasar, 1966b: 184 (Trox)
ovatus Palisot de Beauvois, 1805: 175 (Trox)
punctatus LeConte, 1854d: 215 (Trox)
torresalai Báguena Corella, 1959: 212 (Trox)
tricolor Blackburn, 1904b: 296 (Trox)

subfamily Troginae W.S. Macleay, 1819

genus Trox Fabricius, 1775: 31 type species Scarabaeus sabulosus Linnaeus, 1858
Pseudotrox Robinson, 1948: 1 type species Trox laticollis LeConte, 1854

subgenus Trox Fabricius, 1775: 31 type species Scarabaeus sabulosus Linnaeus, 1858
aproximans Escalera, 1914a: 158 N: MO
boucomonti R. Paulian, 1933c: 205 A: FUJ JIX YUN ORR
brahminus Pittino, 1985: 338 A: HP NP SD SW UP XIZ YUN ORR
cadaverinus cadaverinus Illiger, 1802b: 44 E: AU BE BY CR CT CZ GE HU PL SK ST SZ UK A: ES FE GAN HEI JIL
KI LIA MG TM WS
undulatus Zoubkov, 1829: 158

cadaverinus komareki Balthasar, 1931b: 130 A: FE HEI MG
camberforti Pittino, 1985: 334 A: SCH
clathratus Reiche, 1861b: 205 E: FR (Corse) IT (Sardegna)
confluens Wollaston, 1864: 193 N: AG CI LB MO TU
cotodognanensis Compte, 1986: 69 E: SP (Huelva, Cádiz)
cribrum Gené, 1836: 186 E: FR (Corse) IT (Sardegna)
cyrenaicus Pittino, 2011a: 374 N: LB
dhaulagiri Paulus, 1972a: 55 A: NP
elkantaraensis Pittino, 2011a: 373 N: AG
eversmannii Krynicki, 1832: 123 E: AB AR AU BU CR CZ GE HU MD PL RO SK ST UK A: IN KI KZ TM UZ XIN
concinnus Erichson, 1848a: 932

eximius Faldermann, 1835b: 368 A: MG NMO
fabricii Reiche, 1853b: 89 [RN] E: IT (Sicilia, Pelagie) MA PT SP N: AG EG LB MO TU
granulatus Fabricius, 1801a: 110 [HN]

formosanus Nomura, 1973b: 37 A: SC TAI
gansuensis Ren, 2003: 110 A: GAN
granulipennis Fairmaire, 1852: 83 E: ITi SP “Caucasus” N: AG CI EG LB MO TU A: IN LE SI SY
barbarus Harold, 1872: 169

hispanicus Harold, 1862a: 161 E: FR (Pyrénées) PT SP
hispidus Pontoppidan, 1763: 431 (Scarabaeus) E: AU BE BH CR CZ DE FR GE GR HU MC NL PL PT SB SK SL SP
SV SZ N: CI A: IN SY TR
arenarius Paykull, 1798: 80

56

TROGIDAE
cricetulus Ádám, 1994c: 37
insularis O. Schneider, 1898: 69
luridus P. Rossi, 1790: 17 [HN]
sabulosus Preyssler, 1790: 48 [HN]

horiguchii Ochi & Kawahara, 2002: 54 A: JA TAI
ineptus Balthasar, 1931b: 131 A: ES FE MG
inadai Ochi, Kawahara & Inagaki 2008: 135 A: JA
iranicus Petrovitz, 1980: 613 A: IN TM
klapperichi Pittino, 1983c: 280 E: GR (Ródos) UK (Krym) TR A: AE CY IN IQ IS JO SA SY TR
koreanus J.I. Kim, 1991: 136 A: FE SC
kyotensis Ochi & Kawahara, 2000: 53 A: JA
lama Pittino, 1985: 329 A: XIZ
alpigenus Y.-W. Zhang, 1988: 234

leonardii Pittino, 1983c: 277 E: PT SP N: AG EG LB MO TU A: IS SY
litoralis Pittino, 1991: 77 E: AL AU BH CR GR IT MA ME TR Ni: AG A: TR
mandli Balthasar, 1931b: 128 A: ES FE FUJ HEI JA SC
nishijimai Nakane & Tsukamoto, 1955: 90

martini Reitter, 1892e: 153 N: AG LB TU
matsudai Ochi & Hori, 1999: 37 A: JA (Amami-ôshima)
mitis Balthasar, 1933b: 81 A: BEI ES FE HEI JA SC
mixtus Harold, 1872: 167 A: CY IS LE SY TR
berytensis Petrovitz, 1980: 614

morticinii Pallas, 1781: 11 (Scarabaeus) E: ST A: ES KI KZ TM WS XIN
pinguis Faldermann, 1835b: 369

mutsuensis Nomura, 1937: 85 A: JA
niger P. Rossi, 1792: 9 E: AL AR BH BU CR CT CZ FR GE GG GR HU IT MC ME PL RO SB SK ST SZ TR UK A: ES
KI KZ TR WS
asiaticus Faldermann, 1835a: 261

niponensis Lewis, 1895c: 387 A: FE JA TAI ORR
nodulosus Harold, 1872: 165 E: FR (Corse) IT (Sardegna)
nohirai Nakane, 1954: 10 A: FE JA
opacotuberculatus Motschulsky, 1861a: 14 A: FE HEI NC TAI
parvisetosus Ochi, Kon & Bai, 2010a: 131 A: SCH
perlatus Goeze, 1777: 74 (Scarabaeus) E: BE FR GB GE IT NL SL SP SZ Ni: AG
chevrolati Harold, 1868b: 86
sabulosus A.G. Olivier, 1789: no 4: 8 [HN]
subterraneus Geoffroy, 1785: 8 (Scarabaeus)

perrisii Fairmaire, 1868a: 482 E: AU BE BY CR CT CZ FR GE HU IT PL PT SK SL SP ST SZ UK N: AG MO A: TR
haroldi Flach, 1879: 155
nidicola Bonnaire, 1881: 63

TROGIDAE

57

placosalinus Ren, 2003: 112 A: XIN
quadrimaculatus Ballion, 1871a: 338 A: IN IS KI KZ TD UZ
sabulosus fujiokai Ochi, 2000: 43 A: JA (Honshu)
sabulosus sabulosus Linnaeus, 1758: 350 (Scarabaeus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR
HU IT LA LT LU MD NL NR NT PL PT RO SB SK SL SP ST SV SZ UK A: ES KZ MG NC SC SY TR WS
femoratus De Geer, 1774: 269 (Scarabaeus)

scaber Linnaeus, 1767: 573 (Silpha) [NP] E: AL AU AZ BE BH BU BY CR CZ DE EN FI FR GB GE GR HU IT LA
LT LU MA NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: AG CI MO MR TU A: FE IS JA MG “Korea” AUR
NAR NTR
arenarius Fabricius, 1787a: 18
arenosus Gmelin, 1790: 1586
barbosus Laicharting, 1781: 31
hispidus Paykull, 1798: 81 [HN]
trisulcatus Curtis, 1845: 446

setifer C.O. Waterhouse, 1875a: 98 A: JA JIL LIA NC SC
aino Nakane & Tsukamoto, 1955: 92

sonorae LeConte, 1854d: 211 Ai: CH NAR
alternans LeConte, 1854d: 211
conjunctus Petrovitz, 1975a: 2
econtei Harold, 1872: 156 [RN]

sordidatus Balthasar, 1936d: 452 E: BU CR GR TR A: SY TR
strandi Balthasar, 1936d: 453 N: AB LB MO TU
sugayai Masumoto & Kiuchi, 1995: 206 A: JA
taiwanus Masumoto, Ochi & C.-L. Li, 2005: 233 A: TAI
tibialis Masumoto, Ochi & C.-L. Li, 2005: 235 A: TAI
transversus Reiche, 1856b: 405 E: GR
demaisoni Reitter, 1904c: 156
graecus Mulsant & Rey, 1871a: 479 [=1871c: 223]

uenoi matsumurai Miyake & Yamaya, 1995: 31 A: JA (Okinawa-shotō)
uenoi uenoi Nomura, 1961b: 70 A: JA
ussuriensis Balthasar, 1931b: 129 A: ES FE HEI JIL LIA MG NC SC
vimmeri Balthasar, 1931b: 130 A: BEI FE MG SC
yamayai Nakane, 1983b: 139 A: JA (Ryukyus)
yangi Masumoto, Ochi & C.-L. Li, 2005: 238 A: TAI
zoufali Balthasar, 1931b: 131 A: BEI FE HUB NC TAI ORR
subgenus Phoberus W.S. Macleay, 1819: 137 type species Trox horridus Fabricius, 1775
puncticollis Haaf, 1953: 330 A: AE SA
squamiger Roth, 1851: 133 A: AE SA YE AFR
angulatus Fåhraeus, 1857: 381
setulosus H.J. Kolbe, 1891: 23

58

GLARESIDAE

genus Trox, nomina dubia
lutosus Marsham, 1802: 25 (Scarabaeus) E: GB
maurus Herbst, 1790: 26 [distribution unknown]

family GLARESIDAE H.J. Kolbe, 1905

genus Glaresis Erichson, 1848a: 925 type species Glaresis rufa Erichson, 1848
Afroglaresis Petrovitz, 1968h: 270 type species Afroglaresis obscura Petrovitz, 1968
Eoglaresis Semenov & S.I. Medvedev, 1932b: 338 type species Glaresis oxiana Semenov, 1892
Eremoglaresis Semenov & S.I. Medvedev, 1932b: 341 type species Glaresis beckeri Solsky, 1870

alfierii Petrovitz, 1968h: 266 N: EG
arabica R. Paulian, 1980: 142 (Eoglaresis) A: OM QA SA
beckeri Solsky, 1870: 463 E: KZ ST A: IN KZ MG TD TM UZ
carthagensis Petrovitz, 1968h: 268 N: MO TU
ceballosi Pardo Alcaide, 1958: 163 N: MO
contrerasi Pardo Alcaide, 1958: 164 N: MO
gineri Pardo Alcaide, 1958: 166 N: AG MO
handlirschi Reitter, 1892a: 223 [=1893a: 102] N: AG EG LB A: SI
hispana Báguena Corella, 1959: 210 (Eoglaresis) E: SP
holzschuhi Petrovitz, 1968g: 468 A: TR
kocheri Petrovitz, 1968h: 263 N: MO
mandibularis Petrovitz, 1968h: 265 N: AG
maroccana Petrovitz, 1968h: 262 N: MO
orientalis S.I. Medvedev, 1976: 155 A: MG
oxiana armena Iablokoff-Khnzorian, 1967: 52 E: AR
oxiana oxiana Semenov, 1892b: 469 A: AF IQ KI KZ PA TD TM UZ
pardoi Petrovitz, 1968h: 267 N: TU
porrecta Semenov & S.I. Medvedev, 1932b: 341 (Eoglaresis) A: IN
quedenfeldti Petrovitz, 1968h: 264 N: MO
rufa Erichson, 1848a: 925 E: AB AU BU GR HU PL RO SB SK ST UK A: KZ
frivaldszkyi Westwood, 1852b: 72

thiniensis Verdú & Galante, 2001: 273 E: SP
tripolitana Petrovitz, 1968h: 268 N: MO LB TU
zarudniana Semenov & S.I. Medvedev, 1932b: 340 (Eoglaresis) A: IN

family LUCANIDAE Latreille, 1804
subfamily Aesalinae W.S. Macleay, 1819
tribe Aesalini W.S. Macleay, 1819

genus Aesalus Fabricius, 1801b: 254 type species Lucanus scarabaeoides Panzer, 1793
asiaticus asiaticus Lewis, 1883: 340 A: JA
asiaticus sawaii Fujita & Ichikawa, 1985: 4 A: JA
imanishii Inahara & Ratti, 1981: 191 A: TAI

59

LUCANIDAE

scarabaeoides meridionalis Bartolozzi, 1989: 105 E: IT
scarabaeoides scarabaeoides Panzer, 1793d: 15/16 (Lucanus) E: AU BH BU CR CZ FR GE HU IR IT PL RO SB SK
SL SP ST SV SZ UK A: TM
scarabaeoides siculus Baviera, 2008: 586 E: IT (Sicilia)
sichuanensis Araya, Tanaka & Tanikado, 1995: 210 A: CHQ
ulanowskii Ganglbauer, 1886d: 89 E: AB AR GG ST (Kavkaz) UK (Krym) A: IN
daghestanicus Didier & Séguy, 1953a: 59

genus Echinoaesalus Zelenka, 1993: 235 type species Echinoaesalus jaechi Zelenka, 1993
subgenus Echinoaesalus Zelenka, 1993: 235 type species Echinoaesalus jaechi Zelenka, 1993
chungi H. Huang & C.-C. Chen, 2015a: 164 A: TAI
genus Himaloaesalus H. Huang & C.-C. Chen, 2013b: 39 type species Aesalus himalayicus Kurosawa, 1985
himalayicus Kurosawa, 1985a: 49 (Aesalus) A: NP SD ORR
saburoi Araya, Tanaka & Bartolozzi, 1998: 408 (Aesalus) A: NP
zhejiangensis H. Huang & Bi, 2009: 20 (Aesalus) A: ZHE

tribe Nicagini LeConte, 1861

genus Nicagus LeConte, 1861: 130 type species Ochodaeus obscurus LeConte, 1848
japonicus Nagel, 1928a: 260 A: JA

subfamily Syndesinae W.S. Macleay, 1819
tribe Ceruchini LeConte, 1861

genus Ceruchus W.S. Macleay, 1819: 115 type species Lucanus tenebroides Fabricius, 1787 (=Lucanus chrysomelinus Hochenwarth, 1785)
Platycerus Latreille, 1806: 133 [HN] type species Lucanus tenebroides Fabricius, 1787 (=Lucanus chrysomelinus
Hochenwarth, 1785)
Tarandus Dejean, 1833: 174 [HN] type species Lucanus tenebroides Fabricius, 1787 (=Lucanus chrysomelinus
Hochenwarth, 1785)

atavus Fairmaire, 1891b: 88 A: KA PA
chrysomelinus Hochenwarth, 1785: 356 (Lucanus) E: AU BH BY CR CT CZ EN FI FR GE IT LA LS LT NT PL RO
SB SK SL SP SV SZ A: WS
piceus Bonsdorff, 1785: 222 (Lucanus)
sylvicola Mulsant, 1842: 598
tarandus Panzer, 1789: 25 (Lucanus)
tenebroides Fabricius, 1787a: 2 (Lucanus)

deuvei Boucher & Král, 1997: 43 A: SCH
katerinae gonjoicus J. Li, 2015b: 41 A: XIZ
katerinae katerinae Král, 1995a: 96 A: SCH
lignarius lignarius Lewis, 1883: 339 A: FE (Kuril’skie ostrova, Sakhalin) JA
lignarius monticola Nakane, 1978: 78 A: JA

60

LUCANIDAE

lignarius nodai Fujita, 1987: 6 A: JA
luojishanensis Okuda, 2008: 35 A: SCH
minor Tanikado & Okuda, 1994: 4 A: CHQ GAN SCH SHA SHX
niger Boucher & Král, 1997: 42 A: SCH YUN
reginae Boucher & Král, 1997: 44 A: YUN
sinensis Nagel, 1933: 226 A: SCH YUN ORR
tabanai Okuda, 2008: 33 A: SCH
yangi H. Huang, Imura & C.-C. Chen, 2011: 11 A: GUI SCH
yingqii H. Huang & C.-C. Chen, 2013b: 36 A: SCH

tribe Sinodendrini LeConte, 1861

genus Sinodendron Hellwig, 1792: 391 type species Scarabaeus cylindricus Linnaeus, 1758
Ligniperda Fabricius, 1790: 216 [HN] type species Scarabaeus cylindricus Linnaeus, 1758

cylindricum Linnaeus, 1758: 346 (Scarabaeus) E: AL AU BE BH BY CR CZ DE EN FI FR GB GE HU IT KO LA LS
LT LU MC MD ME NR NT PL RO SB SK SL SP ST SV SZ UK A: ES KZ MG WS XIN
americanum Palisot de Beauvois, 1805: 192
aspromontanum J. Müller, 1938: 58
juvenilis Mulsant, 1842: 602
tenebroides Scopoli, 1777: 75 (Lucanus)

persicum Reitter, 1902a: 81 E: AB A: IN
yunnanense Král, 1994: 48 A: SCH YUN

subfamily Lucaninae Latreille, 1804
tribe Aegini H. Huang & C.-C. Chen, 2013

genus Aegus W.S. Macleay, 1819: 112 type species Aegus chelifer W.S. Macleay, 1819
angustus Bomans, 1989a: 17 A: FUJ HUB SCH ZHE
bidens Möllenkamp, 1902b: 353 A: GUA GUI YUN ORR
imitator Nagel, 1941: 58

callosilatus Bomans, 1989a: 20 A: FUJ ORR
dispar Didier, 1931: 211 A: FUJ HUN SCH YUN ZHE
fukiensis Bomans, 1989a: 21 A: FUJ ORR
kuangtungensis Nagel, 1925: 170 A: FUJ GUA
kurosawai Okajima & Ichikawa, 1986: 181 A: TAI
laevicollis abei Ichikawa & Imanishi, 1976: 10 A: JA (Tokara-rettō)
laevicollis asaii A. Murayama & Shimizu, 2004: 33 A: JA (Tokara: Nakano-shima)
laevicollis doii Fujita, 2002: 22 A: JA (Okinawa: Iheyajima)
laevicollis formosae Bates, 1866: 347 A: TAI
laevicollis fujitai Ichikawa & Imanishi, 1976: 10 A: JA (Izu-shotō)
laevicollis ishigakiensis Nomura, 1960b: 43 A: JA (Okinawa: Ishigaki-jima)
laevicollis laevicollis W.W. Saunders, 1854: 54 A: CHQ FUJ GUA GUI HUB HUN JA JIX XIZ YUN ORR
labilis Westwood, 1864: 54

61

LUCANIDAE
punctiger W.W. Saunders, 1854: 55

laevicollis matsushitai Asai, 2001: 23 A: JA (Ryukyus)
laevicollis mizunumai Ichikawa & Fujita, 1985: 8 A: JA (Yaeyama-shotō)
laevicollis nakanei Ichikawa & Imanishi, 1976: 9 A: JA (Okinawa-shotō)
laevicollis subnitidus C.O. Waterhouse, 1873: 277 A: JA NC SC
laevicollis tamanukii Ichikawa & Imanishi, 1976: 10 A: JA (Okinoerabu-jima)
laevicollis taurulus Didier, 1928: 55 A: JA (Ôshima)
linealis Didier, 1928: 54 A: BT SD ORR
melli Nagel, 1925: 173 A: FUJ GUA
nakaneorum Ichikawa & Fujita, 1986: 139 A: TAI
ogasawarensis chichijimaensis Hosoguchi, 2000: 24 A: JA (Chichi-jima)
ogasawarensis ogasawarensis Okajima & Kobayashi, 1975: 119 A: JA (Ogasawara-shotō)
parallelus Hope & Westwood, 1845: 22 (Lucanus) A: HUB XIZ ORR
aequalis Westwood, 1845c: 102
capitatus Westwood, 1847c: 275 (Lucanus)
distinctus Westwood, 1845c: 102
malabaricus Westwood, 1845c: 102
platycepalus Westwood, 1845c: 101
sinister Hope & Westwood, 1845: 22 (Lucanus)

parvus Boileau, 1902c: 320 A: SCH
pichoni Didier, 1931: 210 A: ZHE
taurus Boileau, 1899d: 320 A: CH ORR

tribe Cyclommatini H. Huang & C.-C. Chen, 2013

genus Cyclommatus Parry, 1863: 449 type species Lucanus tarandus Thunberg, 1806
Cyclommatellus Nagel, 1936b: 293 type species Cyclommatellus bucephalus Nagel, 1936
Cyclommatinus Didier, 1927c: 103 type species Lucanus strigiceps Westwood, 1848
Cyclophthorus Arrow, 1950: 63 type species Lucanus tarandus Thunberg, 1806
Megaloprepes J. Thomson, 1862: 420 [HN] type species Lucanus tarandus Thunberg, 1806

albersii Kraatz, 1894d: 268 A: AP YUN ORR
asahinai Y. Kurosawa, 1974a: 103 A: TAI
mniszechii J. Thomson, 1856: 526 (Cyclophthalmus) A: HUN SHG TAI ORR
tonkinensis Didier, 1927d: 164

multidentatus Westwood, 1848a: 17 (Lucanus) A: AP BT NP SD ORR
scutellaris Möllenkamp, 1912a: 7 A: TAI
strigiceps Westwood, 1848a: 18 (Lucanus) A: CHQ GUI SD
elsae Kriesche, 1921a: 95

taiwanus Bomans, 1989c: 263 A: TAI

tribe Dorcini Parry, 1864

genus Aulacostethus C.O. Waterhouse, 1869: 13 type species Aulacosthetus archeri C.O. Waterhouse, 1869
Aegomorphus Houlbert, 1914b: 344 [HN] type species Aegomorphus ruditemporalis Houlbert, 1914

62

LUCANIDAE

archeri C.O. Waterhouse, 1869: 14 A: NP SD XIZ ORR
doani M. Baba, 2000: 3 A: YUN ORR
ruditemporalis Houlbert, 1914b: 344 (Aegomorphus) A: SCH
tianmuxing H. Huang & C.-C. Chen, 2013d: 313 A: ZHE
genus Dorcus W.S. Macleay, 1819: 111 type species Scarabaeus parallelipipedus Linnaeus, 1758
Digonophorus C.O. Waterhouse, 1895: 157 type species Digonophorus atkinsoni C.O. Waterhouse, 1895
Ditomoderus Parry, 1864: 45 type species Ditomoderus mirabilis Parry, 1864
Durelius Houlbert, 1915b: 92 type species Dorcus derelictus Parry, 1862
Dynodorcus Didier, 1937: 195 type species Dorcus antaeus Hope, 1842
Eurydorcus Didier, 1937: 196 type species Dorcus reichei Hope, 1842
Eurytrachellelus Didier, 1937: 185 [RN] type species Dorcus semirugosus J. Thomson, 1862
Eurytrachelus J. Thomson, 1862: 421 [HN] type species Dorcus semirugosus J. Thomson, 1862
Goniodorcus Didier, 1937: 196 type species Dorcus coranus Gestro, 1881
Hemisodorcus J. Thomson, 1862: 421 type species Lucanus nepalensis Hope, 1831
Macrodorcas Motschulsky, 1862a: 15 type species Psalidostomus rectus Motschulsky, 1857
Nipponodorcus Nomura & Kurosawa, 1960: 41 type species Eurytrachelus rubrofemoratus Snellen van Vollenhoven, 1865
Telodorcus Didier, 1937: 196 type species Lucanus saiga A.G. Olivier, 1789

alexisi Muret & Drumont, 1999: 485 A: CY
amamianus amamianus Nomura, 1964: 47 (Macrodorcas) A: JA
kubotai Fujita & Ichikawa, 1985: 9 (Macrodorcas)
nomurai Mizunuma, 1994: 263 [RN] (Macrodorcas)
okinawanus Nomura, 1964: 47 [HN] (Macrodorcas)

amamianus yaeyamaensis Hori, 1991: 75 (Macrodorcas) A: JA (Yaeyama-shotō)
antaeus antaeus Hope, 1842b: 83 [=1845b: 74] A: BT NP SD XIZ ORR
dispar De Lisle, 1970: 109
parryi Boileau, 1902a: 49 (Rhaetus)
scaritides Hope & Westwood, 1845: 24 (Lucanus)

antaeus miyashitai Fujita, 2010: 19 A: GUI GUX HAI YUN ORR
apatani Okuda & Maeda, 2015a: 31 (Digonophorus) A: AP
arrowi arrowi Boileau, 1911b: 263 (Hemisodorcus) A: NP YUN ORR
chudoraziensis Fujita, 2010: 247

arrowi magdaleinae J.P. Lacroix, 1972a: 67 (Macrodorcas) A: YUN ORR
nobuhiroi Fujita, 2010: 248

bashanus H. Huang & C.-C. Chen, 2013d: 445 A: CHQ
branaungi Nagai, 2000b: 104 A: YUN ORR
carinulatus koreanus Jang & Kawai, 2008: 103 A: SC
carinulatus carinulatus Nagel, 1941: 56 A: TAI
cervulus Boileau, 1901: 284 (Eurytrachelus) A: CHQ GUI GUX XIZ YUN ORR
chucheni H. Huang & C.-C. Chen, 2013d: 433 A: YUN

LUCANIDAE

63

cuonaensis H. Huang & C.-C. Chen, 2013d: 460 A: XIZ
curvidens curvidens Hope, 1840a: 78 (Lucanus) A: AP BT CHQ GUX HAI NP PA SD XIZ YUN ORR
dehaani Hope, 1842d: 247 [=1842c: 127] (Dorcas)
klugii J. Thomson, 1862: 424

cylindricus J. Thomson, 1862: 427 A: NP ORR
bobi Didier, 1927e: 191
rosti Zang, 1906: 184

daedalion Didier & Séguy, 1953c: 104 (Eurytrachellelus) A: CHQ FUJ GUA GUI GUX JIX SCH YUN ORR
tibetanus Schenk, 2006b: 7 (Serrognathus)

davidis Fairmaire, 1887d: 314 (Gnaphaloryx) A: BEI HEB MG NIN NMO SHX
davidianus Arrow, 1943: 138 [RN]
septentrionalis Bomans, 1990: 174

derelictus Parry, 1862: 112 A: BT NP SD XIZ
donckieri donckieri Boileau, 1898c: 227 (Hemisodorcus) A: NP SD XIZ YUN ORR
dungkharpai Okuda & Maeda, 2015b: 38 (Digonophorus) A: AP
elegans Parry, 1862: 110 (Cladognathus) A: BT NP SD XIZ ORR
atkinsoni C.O. Waterhouse, 1895: 157
rudis Westwood, 1864: 35
sulcatipennis Houlbert, 1915a: 51 (Prosopocoelus)

fukinukii Schenk, 2000: 79 (Hemisodorcus) A: AP ORR
fuliginosus Didier, 1929: 77 (Eurytrachelus) A: KA
gongshanus H. Huang & C.-C. Chen, 2013d: 446 A: YUN
haitschunus Didier & Séguy, 1952: 227 (Eurytrachellelus) A: FUJ GUI GUX NC SC SCH
hansi Schenk, 2008a: 3 A: GUA GUI GUX HAI
yoshinarii Tsukawaki, 2008: 41

hirticornis clypeatus Benesh, 1950a: 14 A: TAI
hirticornis hirticornis Jakovlev, 1897: 457 (Eurytrachelus) A: CHQ FUJ GUA GUI GUX HUB HUN JIX SCH ZHE
hopei formosanus Miwa, 1929b: 351 A: TAI
hopei hopei W.W. Saunders, 1854: 50 (Platyprosapus) A: ANH BEI CHQ FUJ GUA GUI GUX HEN HUB HUN JA
JIA JIX LIA NC SC SCH SHN TIA ZHE
binodulosus C.O. Waterhouse, 1874a: 6
striatopunctatus W.W. Saunders, 1854: 51
striatus W.W. Saunders, 1854: 53

hyperion hyperion Boileau, 1899b: 177 A: YUN ORR
imrani Okuda & Maeda, 2015b: 36 (Digonophorus) A: AP
intricatus J.P. Lacroix, 1981: 68 (Hemisodorcus) A: HUB SCH
japonicus Nakane & Makino, 1985: 21 A: JA
katsurai Ikeda, 2000b: 29 A: CHQ FUJ GUI JIX SCH YUN ZHE ORR
kikunoae Hosoguchi, 2004: 30 A: SD XIZ
kyawi Nagai & Maeda, 2009: 46 A: YUN ORR

64

LUCANIDAE

laevidorsis Fairmaire, 1888a: 15 A: YUN ORR
affinis Pouillaude, 1913d: 336 (Eurytrachelus)
ohtai Ikeda, 1998: 24

lhoba H. Huang & C.-C. Chen, 2013d: 452 A: XIZ
lineatopunctatus gaoligongshanus H. Huang & C.-C. Chen, 2013d: 520 A: YUN
lineatopunctatus lineatopunctatus Hope, 1831: 22 (Lucanus) A: AP BT NP SD XIZ ORR
linwenhsini H. Huang & C.-C. Chen, 2013d: 450 A: YUN
liyingbingi H. Huang & C.-C. Chen, 2013d: 467 A: YUN
lvbu H. Huang & C.-C. Chen, 2013d: 442 A: CHQ GUI
macleayii Hope & Westwood, 1845: 19 (Lucanus) A: SD XIZ YUN ORR
menba H. Huang & C.-C. Chen, 2013d: 455 A: XIZ
mencius Kriesche, 1935: 172 (Eurytrachelus) A: GUI SCH YUN
miwai Benesh, 1936: 12 A: TAI
monpa Okuda & Maeda, 2014: 14 A: AP
montivagus adachii Fujita & Ichikawa, 1987: 20 (Nipponodorcus) A: JA (Kyūshū)
montivagus montivagus Lewis, 1883: 337 (Macrodorcas) A: JA ORR
yezoensis Miwa, 1927: 27 (Eurytrachelus)

motuensis H. Huang & C.-C. Chen, 2013d: 429 A: XIZ
musimon Gené, 1836: 192 E: IT (Sardegna) N: AG TU
lineatipennis Normand, 1936: 205
semisulcatus Mulsant & Rey, 1874: 52

myinti Nagai & Maeda, 2009: 48 A: YUN ORR
nepalensis Hope, 1831: 22 (Lucanus) A: AP BT NP PA SD XIZ
chevrolatii Chenu, 1840: pl. 44 (Lucanus)
parryi Hope, 1843b: 94 (Lucanus)
rafflesii Hope, 1840a: 77 (Lucanus)
similis Hope, 1831: 22 (Lucanus)

nosei Nagai, 2000b: 106 A: XIZ ORR
parallelipipedus Linnaeus, 1758: 354 (Scarabaeus) E: AB AL AR AU BE BU BY CR CT CZ DE EN FR GB GE GG
GR HU IT KZ LA LS LT LU MC ME NT PL PT RO SK SL SP ST SV SZ UK N: MO A: IN IS SY TR WS
bipunctatus Pontoppidan, 1765: 199 (Scarabaeus)
capra Panzer, 1798: 12 (Lucanus)
dama O.F. Müller, 1776: 52 (Lucanus)
immaturus Mulsant, 1842: 591
infractus Bergsträsser, 1778: 40 (Lucanus)
leuthneri Ganglbauer, 1886a: 140 [RN]
mexicanus Benesh, 1944: 45
minor Seabra, 1905: 9
reichei Ganglbauer, 1886a: 81
truquii Mulsant, 1855a: 14 [=1855c: 14]
tuberculatus W.S. Macleay, 1819: 112

LUCANIDAE

65

peyronis Reiche & Saulcy, 1856: 407 E: AR BU GR MC A: IN IS SY TR
vavrai Nonfried, 1905a: 12

prochazkai Schenk, 2003: 17 A: IN
ratiocinativus Westwood, 1871: 356 A: BT NP SD XIZ
rectus kobayashii Fujita & Ichikawa, 1985: 9 (Macrodorcas) A: JA (Tokara-rettō)
rectus miekoae Yoshida, 1991: 40 (Macrodorcas) A: JA (Izu-shotō)
rectus mishimaensis Tsuchiya, 2003: 50 A: JA (Ôsumi-shotō)
rectus rectus Motschulsky, 1858a: 29 (Psalidostomus) A: CHQ FE HEI JA LIA NC SC TAI ORR
binervis Motschulsky, 1861a: 16
diabolicus J. Thomson, 1862: 423
niponensis Snellen van Vollenhoven, 1861: 113
rugipennis Motschulsky, 1862a: 16 (Macrodorcas)

rectus yakushimaensis Tsuchiya, 2003: 50 A: JA (Ôsumi-shotō)
reichei Hope, 1842b: 83 [=1845b: 75] A: BT NP SD XIZ YUN ORR
bengalensis Hope & Westwood, 1845: 22 (Lucanus)
blanchardi Hope, 1842b: 84
cognatus Hope, 1842b: 84 [=1845: 75]
castelnaudii Deyrolle, 1865: 31 (Eurytrachelus)
exaratus J. Thomson, 1862: 426
glabripennis Westwood, 1871: 359
praecellens Möllenkamp, 1902a: 283 (Eurytrachelus)
punctatostriatus Kollar & L. Redtenbacher, 1844: 532

rubrofemoratus chenpengi J. K. Li, 1992: 67 (Nipponodorcus) A: BEI HEB HEI HEN HUB LIA NC SC ZHE
rubrofemoratus miyamai Nagai & Okazaki, 2005: 24 A: CHQ GAN SCH
rubrofemoratus rubrofemoratus Snellen van Vollenhoven, 1865: 152 (Eurytrachelus) A: FE JA TAI ORR
schenklingi Möllenkamp, 1913: 20 (Eurytrachelus) A: TAI
semenowi Jakovlev, 1900a: 36 (Hemisodorcus) A: CHQ GAN GUI SCH YUN
barbatus Nagel, 1939: 327 (Hemisodorcus)

sewertzowi Semenov, 1891: 309 A: AF KA PA TD UZ ORR
rugatus Didier, 1927e: 193

sinensis concolor Bomans, 1971a: 66 (Macrodorcas) A: YUN
sinensis kentai Tsukawaki, 1999: 7 A: YUN ORR
sinensis sinensis Boileau, 1899b: 176 (Hemisodorcus) A: YUN
castaneus Bomans, 1971a: 69 (Macrodorcas)

striatipennis davidi Séguy, 1954a: 187 (Hemisodorcus) A: CHQ GUX HUB JIX SCH SHX ZHE
continentalis Sakaino, 1997: 12
emikoae Ikeda, 2001: 31

striatipennis koyamai Nakane, 1978: 77 (Macrodorcas) A: JA (Yaku-shima)
striatipennis striatipennis Motschulsky, 1862a: 17 (Macrodorcas) A: FE (Kuril’skie ostrova, Sakhalin) JA ORR
cribellatus Motschulsky, 1862a: 17 (Macrodorcas)
opacus C.O. Waterhouse, 1870: 208

66

LUCANIDAE
vanvolxemi Lewis, 1879b: 462

striatipennis yushiroi Sakaino, 1997: 11 A: TAI
submolaris Hope & Westwood, 1845: 23 (Lucanus) A: KA NP PA UP ORR
brachycerus Boileau, 1904a: 27 (Eurytrachelus)
kucerai Baba, 2008: 42

sukkiti Fukinuki, 2004: 29 A: YUN ORR
suturalis Westwood, 1871: 358 A: KA NP SD ORR
opacipennis Zang, 1906: 184
rotundopunctatus Nagel, 1936a: 209

taiwanicus Nakane & Makino, 1985: 21 A: TAI
tanakai tanakai Nagai, 2002a: 12 A: FUJ GUA GUI GUX HAI
tenuihirsutus S.I. Kim & J.I. Kim, 2010: 66 A: BEI GAN HEN SC
tityus Hope, 1842b: 83 [=1845b: 74] A: BT SD XIZ YUN ORR
punctilabris Hope, 1842b: 84 [=1845b: 75]
semirugosus J. Thomson, 1862: 422
tethys Didier, 1929: 115 (Eurytrachelus)

ursulae Schenk, 1996a: 440 (Hemisodorcus) A: CHQ HUB ZHE
ursulus Arrow, 1938a: 55 A: BT SD XIZ YUN ORR
velutinus J. Thomson, 1862: 426 A: AP BT FUJ GUA GUI GUX HAI HEB HUB HUN NP SCH SD XIZ YUN ZHE
ORR
cinereus Boileau, 1902c: 321 (Gnaphaloryx)

vicinus W.W. Saunders, 1854: 51 A: FUJ HUN ZHE
wemckeni Schenk, 2008b: 176 (Digonophorus) A: AP XIZ
wardi Arrow, 1943: 136 A: XIZ ORR
wui H. Huang & C.-C. Chen, 2013d: 378 A: SHA
yaksha gracilicornis Benesh, 1950a: 11 A: CHQ FUJ GUA GUI GUX SCH TAI ZHE
yaksha yaksha Gravely, 1915: 422 A: SD XIZ YUN ORR
tonkinensis Fujita, 2010: 254

yamadai Miwa, 1937: 166 (Macrodorcus) A: TAI
zhangmuensis H. Huang & C.-C. Chen, 2013d: 456 A: XIZ
genus Epidorcus Séguy, 1954a: 192 type species Cladognathus piceipennis Westwood, 1855
bidentatus Bomans, 1978: 162 (Prosopocoilus) A: YUN ORR
denticulatus Boileau, 1901: 284 (Prosopocoilus) A: GUX YUN ORR
katsurai Fujita, 2010: 200 (Prosopocoilus)

gracilis W.W. Saunders, 1854: 47 (Cladognathus) A: CHQ FUJ GUA GUI GUX HUB HUN JIA JIX SCH SHG YUN
ZHE ORR
piceipennis Westwood, 1855: 202 (Cladognathus) A: GUI GUX YUN
andreasi Schenk, 2009b: 1 (Prosopocoilus)
similis Schenk, 2009b: 2 (Prosopocoilus)

rondoni Bomans & J.P. Lacroix, 1970: 118 (Prosopocoilus) A: YUN ORR
tonkinensis Pouillaude, 1913d: 335 (Prosopocoilus) A: GUA GUX HAI ORR

LUCANIDAE
genus Falcicornis Planet, 1894: 44 type species Falcicornis groulti Planet, 1894
Pogonodorcus Séguy, 1954a: 189 type species Prosopocoilus elegantulus Albers, 1891

bisignatus Parry, 1862: 111 (Cladognathus) A: AP BT CHQ SD XIZ YUN ORR
dierli Endrődi, 1968a: 51 (Dorcus) A: NP
fulvonotatus Parry, 1862: 111 (Cladognathus) A: NP SD XIZ ORR
bilineatus Bomans, 1971b: 76 (Hemisodorcus)

heyangi H. Huang & C.-C. Chen, 2013d: 280 A: AP XIZ
himalayae Schenk, 2009c: 3 (Macrodorcas) A: AP XIZ
humilis Arrow, 1935: 111 (Dorcus) A: AP BT SD XIZ ORR
itoi Bomans, 1993: 400 (Macrodorcas) A: GUX YUN ORR
kusakabei Fujita, 2010: 233 (Dorcus) A: XIZ ORR
mellianus Kriesche, 1921a: 98 (Hemisodorcus) A: CHQ FUJ GUA GUI GUX HUN ORR
lasiodontus De Lisle, 1964: 44 (Macrodorcas)
perroti J.P. Lacroix, 1972a: 64 (Hemisodorcus)

moellenkampi moellenkampi Nagel, 1924b: 402 (Hemisodorcus) A: YUN
moellenkampi bomansi J.P. Lacroix, 1981: 69 (Hemisodorcus) A: YUN ORR
neolucanoides H. Huang & C.-C. Chen, 2013d: 261 A: YUN ORR
pseudaxis Didier, 1926c: 178 (Hemisodorcus) A: YUN ORR
ruficrus De Lisle, 1970: 108 (Macrodorcas) A: YUN ORR
rufonotatus giselae Bomans, 1991: 142 (Hemisodorcus) A: YUN ORR
seguyi De Lisle, 1955: 6 (Macrodorcas) A: FUJ GUA GUX HAI YUN ZHE ORR
songianus Didier & Séguy, 1953a: 46 (Hemisodorcus) A: CHQ FUJ GUA GUI GUX SCH YUN
elsiledis Séguy, 1954a: 188 (Hemisodorcus)

taibaishanensis Schenk, 2008a: 9 (Macrodorcas) A: FUJ GUA GUI GUX SHA ZHE
tenuecostatus mochizukii Miwa, 1937: 164 (Macrodorcas) A: TAI
tenuecostatus pieli Didier & Séguy, 1953a: 45 (Hemisodorcus) A: ANH CHQ FUJ GUA GUI JIX SCH ZHE
sancha Fujita, 2010: 18 (Dorcus)

tenuecostatus tenuecostatus Fairmaire, 1888b: 116 (Dorcus) A: BEI HEN TIA
genus Gnaphaloryx Burmeister, 1847: 396 type species Gnaphaloryx opacus Burmeister, 1847
opacus Burmeister, 1847: 397 A: HAI TAI YUN ORR
andamanus Kriesche, 1921a: 100
burmeisteri Nagel, 1926: 119 [RN]
rugosus Albers, 1885: 236
tonkinensis Kriesche, 1921a: 101

genus Hexarthrius Hope, 1842d: 247 [=1842c: 127] type species Hexarthrius parryi Hope, 1842
aduncus aduncus Jordan, 1884: 484 A: XIZ ORR
forsteri forsteri Hope, 1840: 77 (Lucanus) A: XIZ ORR
serricollis Hope, 1840: 78 (Lucanus)

melchioritis Séguy, 1954b: 269 A: YUN ORR
mniszechii J. Thomson, 1857c: 396 (Lucanus) A: NP ORR

67

68

LUCANIDAE

parryi deyrollei Parry, 1864: 11 A: YUN ORR
vitalisi tsukamotoi Nagai, 1998a: 8 A: GUX
vitalisi vitalisi Didier, 1925b: 202 A: YUN ORR
genus Kirchnerius Schenk, 2009a: 2 type species Kirchnerius guangxii Schenk, 2009
boreli Boileau, 1904c: 284 (Prosopocoilus) A: XIZ ORR
bulbosus Hope, 1840a: 78 (Lucanus) A: SD ORR
punctiger Hope, 1840a: 78 (Lucanus)

guangxii Schenk, 2009a: 2 A: GUX
parryi Boileau, 1913: 233 (Prosopocoilus) A: BT NP SD ORR
angustus Didier, 1929: 101 (Prosopocoilus)
latus Didier, 1929: 100 (Prosopocilus)
polymorphus Arrow, 1943: 138 (Dorcus) [RN]

simianshanus H. Huang & C.-C. Chen, 2011: 41 (Prosopocoilus) A: CHQ
spencei mandibularis Möllenkamp, 1902a: 283 (Prosopocoilus) A: GUX SCH YUN ORR
spencei spencei Hope, 1840a: 78 (Lucanus) A: AP NP SD XIZ ORR
crenicollis J. Thomson, 1862: 418 (Prosopocoilus)
laticeps Möllenkamp, 1904: 403 (Prosopocoilus)
mordax Boileau, 1904c: 285 (Prosopocoilus)

yangi Fukinuki, 2004: 31 (Prosopocoilus) A: CHQ GUI HUB FUJ
genus Metallactulus Ritsema, 1885: 54 type species Lucanus parvulus Hope & Westwood, 1845
parvulus Hope & Westwood, 1845: 25 (Lucanus) A: TAI ORR
genus Miwanus H. Huang & C.-C. Chen, 2013c: 77 type species Leptinopterus formosanus Miwa, 1929
formosanus capricornus Didier, 1931: 173 (Prosopocoilus) A: GUX ORR
formosanus formosanus Miwa, 1929a: 74 (Leptinopterus) A: TAI
formosae Arrow, 1943: 138 (Dorcus) [RN]

formosanus kishidai Fujita, 2010: 17 (Dorcus) A: GUA GUX
genus Prosopocoilus Hope & Westwood, 1845: 4 type species Lucanus cavifrons Hope & Westwood, 1845
Cladognathinus Didier & Séguy, 1952: 225 type species Cladognathus decipiens Parry, 1864
Cladognathus Burmeister, 1847: 364 [RN] type species Lucanus girafa A.G. Olivier, 1789
Cyclotropus R. Oberthür & Houlbert, 1913b: 449 type species Lucanus occipitalis Hope & Westwood, 1845
Eulucanus Didier, 1927b: 87 type species Eulcanus spectabilis Didier, 1927
Homoderinus Kriesche, 1926: 384 type species Homoderus variegatus Boileau, 1904
Hoplitocranum Jakovlev, 1896: 172 type species Metopodontus calcaratus Jakovlev, 1896
Macrodorcinus Maes, 1990: 8 type species Lucanus passaloides Hope & Westwood, 1845
Macrognathus Hope & Westwood, 1845: 5 [HN] type species Lucanus girafa A.G. Olivier, 1789
Metopodontus Hope & Westwood, 1845: 4 type species Lucanus downesii Hope, 1835
Metopotropus R. Oberthür & Houlbert, 1913a: 416 type species Prosopocoilus mohniki Parry, 1873

LUCANIDAE

69

Pelecognathus Houlbert, 1915a: 52 type species Pelecognathus prosopocoeloides Houlbert, 1915
Prosopocoelinus Maes, 1990: 8 type species Lucanus curvipes Hope & Westwood, 1845
Psalidognathus Motschulsky, 1862a: 13 [HN] type species Lucanus inclinatus Motschulsky, 1858
Psalidoremus Motschulsky, 1862b: 55 [RN] type species Lucanus inclinatus Motschulsky, 1858
Pseudodontolabis Maes, 1990: 6 type species Prosopocoilus lumawigi De Lisle, 1977

approximatus Parry, 1864: 33 (Cladognathus) A: YUN ORR
astacoides astacoides Hope, 1840a: 78 (Lucanus) A: AP BT SD ORR
foveatus Hope, 1840a: 78 (Lucanus)
omissus Hope, 1840a: 78 (Lucanus)

astacoides blanchardi Parry, 1873: 337 (Metopodontus) A: ANH BEI CHQ GAN GUI HEB HUB HUN JIA MG
NMO SHA SHN SC SCH SHX TIA TAI XIZ YUN ZHE
thibetanus Planet, 1899f: 385 (Metopodontus)

astacoides castaneus Hope, 1845: 12 (Lucanus) A: BT SD NP XIZ ORR
croceus Didier, 1929: 121 (Metopodontus)
poultoni Boileau, 1911a: 63 (Metopodontus)

astacoides dubernardi Planet, 1899d: 35 (Metopodontus) A: GUA GUI SCH YUN ORR
astacoides fraternus Hope & Westwood, 1845: 12 (Lucanus) A: GUX YUN ORR
birmanicus Gravely, 1915: 417 (Metopodontus)

astacoides kachinensis Bomans & Miyashita, 1997: 2 A: XIZ YUN ORR
biplagiatus biplagiatus Westwood, 1855: 200 (Lucanus) A: BT FUJ GUA GUI GUX NP XIZ YUN ORR
nigripes Boileau, 1905a: 17 (Metopodontus)

buddha Hope, 1842c: 127 (Lucanus) A: AP SD UP ORR
cardoni Didier, 1927f: 220 A: SD ORR
confucius Hope, 1842a: 60 [=1843a: 62, 1845: 5] (Lucanus) A: AP FUJ GUA GUI GUX HAI JIX SD YUN ZHE ORR
arrowi Gravely, 1915: 416 (Cladognathus)

dentifer Deyrolle, 1865: 29 (Cladognathus) A: NP ORR
dissimilis dissimilis Boileau, 1898b: 95 (Psalidoremus) A: JA
loochooensis Maes, 1981: 157 (Psalidoremus)

dissimilis elegans Inahara, 1958: 12 (Psalidoremus) A: JA (Takarajima)
dissimilis hayashii Fujita, 2009: 38 A: JA (Iheyajima)
dissimilis kumejimaensis Ichikawa & Fujita, 1985: 7 (Psalidoremus) A: JA (Kumejima)
dissimilis makinoi Ichikawa & Fujita, 1985: 7 (Psalidoremus) A: JA (Tokuno-shima)
dissimilis okinawanus Nomura, 1962: 37 (Psalidoremus) A: JA (Okinawa-shotō)
dissimilis okinoerabuanus Ichikawa & Fujita, 1985: 7 (Psalidoremus) A: JA (Okinoerabujima)
doris Kriesche, 1921a: 94 A: GUX HAI ORR
semifuscus Didier, 1929: 113 (Metopodontus)

forficula austerus De Lisle, 1967: 527 (Psalidoremus) A: TAI
forficula forficula J. Thomson, 1856: 527 (Dorcus) A: ANH FUJ GUA GUI GUX HUN SCH YUN ZHE
forficula hainanensis Schenk, 1999: 116 A: HAI
fulgens Didier, 1927d: 158 (Metopodontus) A: HAI XIZ YUN ORR
fuscocinctus De Lisle, 1973: 140 (Prosopocoelus) A: NP

70

LUCANIDAE

girafa girafa A.G. Olivier, 1789: no 1: 21 (Lucanus) A: BT NP PA YUN ORR
braminus Hope, 1842d: 247 [=1842c: 127] (Lucanus)
bouvieri Didier, 1936: 191 (Cladognathus)

hachjioensis Nomura, 1960b: 40 A: JA (Hachijojima)
inclinatus inclinatus Motschulsky, 1858a: 29 (Lucanus) A: JA LIA NC SC
inflexus Harold, 1875e: 288 (Psalidoremus)
koreanus Li, 2002: 98
mandibularis J. Thomson, 1862: 417 (Cladognathus)

inclinatus kuchinoerabuensis Shimizu & Murayama, 1998: 24 A: JA (Kochinoerabujima)
inclinatus kuroshimaensis Shimizu & Murayama, 2004: 12 A: JA (Kuro-shima)
inclinatus miyakejimaensis Adachi, 2009: 34 A: JA (Izu-shotō)
mikuraensis Matsuoka & Takatoji, 2010: 20

inclinatus mishimaiouensis Shimizu & Murayama, 1998: 22 A: JA (Kagoshima)
inclinatus yakushimaensis Adachi, 2014: 1 A: JA (Yaku-shima)
impressus C.O. Waterhouse, 1869: 17 (Cladognathus) A: NP ORR
jenkinsi hiekei De Lisle, 1975: 269 A: YUN ORR
sukkiti Nagai, 2005b: 24

jenkinsi jenkinsi Westwood, 1848a: 21 (Lucanus) A: BT PA ORR
calcaratus Jakovlev, 1896: 172 (Metopodontus)

laterotarsus maedaorum Nagai, 2005b: 24 A: XIZ ORR
maclellandi maclellandi Hope, 1842b: 83 (Lucanus) A: SD YUN ORR
quadrinodosus Parry, 1862: 109 (Cladognathus)

maclellandi miyashitai Nagai, 2005b: 24 A: GUX ORR
motschulskyii C.O. Waterhouse, 1869: 16 (Cladognathus) A: TAI
occipitalis Hope & Westwood, 1845: 13 (Lucanus) A: AP JA TAI ORR
anoa Kriesche, 1921c: 121
anoella Kriesche, 1921c: 121
anoides Kriesche, 1921c: 121
astericus J. Thomson, 1862: 417 (Cladognathus)
marginatus Burmeister, 1847: 369 (Cladognathus)
preangerensis R. Oberthür & Houlbert, 1913b: 450
roepstorffi C.O. Waterhouse, 1890a: 35 (Metopodontus)

oweni melli Kriesche, 1922: 123 A: GUA GUI GUX FUJ
duplodentatus Benesh, 1943: 39

oweni oweni Hope & Westwood, 1845: 14 (Lucanus) A: AP BT SD ORR
subangulatus Hope & Westwood, 1845: 24 (Lucanus)

politus Parry, 1862: 110 (Cladognathus) A: BT SD XIZ ORR
porrectus Bomans, 1978: 156 A: GUA GUX HAI ORR
pouillaudei Houlbert, 1915a: 50 A: SD
prosopocoeloides Houlbert, 1915a: 53 (Pelecognathus) A: BT
pseudodissimilis Kurosawa, 1976c: 190 (Pseudoremus) A: JA (Ishigaki-jima)

LUCANIDAE
reni H. Huang & C.-C. Chen, 2011: 49 A: HAI
spineus Didier, 1927a: 38 (Metopodontus) A: GUA GUX YUN ORR
suturalis A.G. Olivier, 1789: no 1: 16 (Lucanus) A: FUJ GUA GUI HAI HUN NP SD YUN ORR
thibeticus Westwood, 1855: 199 (Lucanus) A: YUN ORR
elegans Bomans, 1978: 157

valgipes Kriesche, 1940: 40 A: KA
yazakii Nagai, 2005b: 24 A: XIZ ORR
genus Pseudorhaetus Planet, 1899e: 174 type species Pseudorhaetus oberthuri Planet, 1899
Weinreichius J.P. Lacroix, 1978: 255 type species Weinreichius perroti J.P. Lacroix, 1978

oberthuri Planet, 1899e: 175 A: YUN ORR
sinicus concolor Benesh, 1960: 110 A: TAI
sinicus sinicus Boileau, 1899a: 111 (Rhaetulus) A: FUJ GUI GUX JIX ZHE
genus Rhaetulus Westwood, 1871: 353 type species Rhaetulus crenatus Westwood, 1871
crenatus boileaui Didier, 1925a: 154 A: YUN ORR
crenatus crenatus Westwood, 1871: 353 A: TAI
minor Kriesche, 1921a: 96
sauteri Möllenkamp, 1912a: 6

crenatus rubrifemoratus Nagai, 2000a: 5 A: FUJ GUA GUI GUX HAI HUN JIX
recticornis Kurosawa, 1964b: 444 A: JA
genus Rhaetus Parry, 1864: 10 type species Hexarthrius westwoodi Parry, 1862
westwoodii westwoodi Parry, 1862: 108 (Hexarthrius) A: SD XIZ ORR
arrowi Gravely, 1915: 416 (Cladognathus)

genus Serrognathus Motschulsky, 1862a: 11 type species Serrognathus castanicolor Motschulsky, 1861
Brontodorcus Didier, 1937: 196 type species Brontodorcus eurycephalus Burmeister, 1847
Lasiodorcus Didier, 1937: 196 type species Lucanus gypaetus Laporte, 1840
Platyprosopus Hope & Westwood, 1845: 6 [HN] type species Lucanus titanus Boisduval, 1835

consentaneus akahorii Tsukawaki, 1998: 82 (Dorcus) A: CHQ HUB
consentaneus consentaneus Albers, 1886: 28 (Eurytrachelus) A: BEI HUN JA JIA JIX LIA NC SC SHN ZHE
costatus Boileau, 1898a: 45 (Eurytrachelus) A: JA (Ryukyus)
metacostatus Kikuta, 1985: 25 (Dorcus) [RN]

damoiseaui Maes, 1982: 7 (Eurytrachellelus) A: JA
kyanrauensis Miwa, 1934: 324 (Eurytrachelus) A: TAI
coussementi Maes, 1982: 5 (Eurytrachellelus)

titanus castanicolor Motschulsky, 1862a: 12 A: JA JIL LIA NC SC
fasolt Kriesche, 1921b: 118 (Eurytrachelus)

titanus daitoensis Fujita & Ichikawa, 1986: 28 A: JA
titanus elegans Boileau, 1899b: 177 (Eurytrachelus) A: JA

71

72

LUCANIDAE

titanus fafner Kriesche, 1921b: 116 (Eurytrachelus) A: GUX HAI YUN ORR
titanus hachijoensis Fujita & Okuda, 1989: 24 A: JA
titanus karasuyamai M. Baba, 1999: 19 (Dorcus) A: JA (Fuke-jima, Hisaka-jima)
titanus okinawanus Kriesche, 1922: 124 (Eurytrachelus) A: JA
titanus okinoerabuensis Fujita & Ichikawa, 1985: 10 A: JA
titanus pilifer Snellen van Vollenhoven, 1861: 112 (Dorcus) A: JA
titanus platymelus W.W. Saunders, 1854: 50 (Platyprosopus) A: ANH CHQ FUJ GUA GUI GUX HEI HEN HUB
HUN JIA JIX SCH SHA ZHE
hangul Kriesche, 1921b: 117 (Eurytrachelus)
hymir Kriesche, 1935: 173 (Eurytrachelus)
marginalis W.W. Saunders, 1854: 53 (Dorcus)
obscurus W.W. Saunders, 1854: 52 (Dorcus)

titanus sakishimanus Nomura, 1964: 62 (Dorcus) A: JA
titanus sika Kriesche, 1921b: 118 (Eurytrachelus) A: TAI
titanus takaraensis Fujita & Ichikawa, 1985: 9 A: JA
titanus tatsutai Shiokawa, 2001: 6 (Dorcus) A: JA
titanus tokunoshimaensis Fujita & Ichikawa, 1985: 9 A: JA
titanus typhoniformis Nagel, 1924a: 72 (Eurytrachelus) A: CHQ GUI YUN
prometheus Didier, 1925d: 251 (Eurytrachelus)

genus Sinodorcus H. Huang & C.-C. Chen, 2013c: 78 type species Dorcus sawaii Tsukawaki, 1999
sawaii sawaii Tsukawaki, 1999: 6 A: FUJ ZHE

tribe Figulini Burmeister, 1847

genus Figulus W.S. Macleay, 1819: 109 type species Lucanus striatus A.G. Olivier, 1789
binodulus C.O. Waterhouse, 1873: 227 A: CH JA NC SC TAI ORR
boninensis Nakane & Y. Kurosawa, 1953: 110 A: JA (Ogasawara)
caviceps Boileau, 1902b: 205 A: NP XIZ ORR
curvicornis Benesh, 1950b: 56 A: TAI ORR
daitoensis Fujita & Ichikawa, 1986: 24 A: JA
deletus Bomans, 1989: 18 A: GUX ORR
fissicollis Fairmaire, 1849: 414 A: TAI ORR
insolitus Bomans, 1987: 10 A: BT
napu Kriesche, 1922: 130 A: YUN ORR
praecipuus Bomans, 1987: 13 A: CH
punctatus C.O. Waterhouse, 1873: 278 A: JA SC TAI
venustus Bomans, 1989b: 16 A: SC
wittmeri Bomans & J.P. Lacroix, 1989: 24 A: NP
yuji Fujita, 1994: 15 A: JA
yunnanensis Nagel, 1928b: 295 A: FUI YUN ORR

73

LUCANIDAE
genus Nigidionus Kriesche, 1926: 385 type species Nigidius parryi Bates, 1866
parryi Bates, 1866: 347 (Nigidius) A: FUJ GUA HAI HUN YUN TAI ORR
gigas Möllenkamp, 1901: 363 (Nigidius)

genus Nigidius W.S. Macleay, 1819: 108 type species Nigidius cornutus W.S. Macleay, 1819
Eudora Laporte, 1840: 174 type species Eudora midas Laporte, 1840 (=Lucanus bubalus Swederus, 1787)
Hadroginidius Kraatz, 1896a: 65 type species Hadroginidius bennigseni Kraatz, 1896
Platycerus Gory, 1833c: 109 type species Platycerus auriculatus Gory, 1833

acutangulus Heller, 1917: 173 A: TAI YUN ORR
baeri Boileau, 1905a: 18 A: TAI ORR
elongatus Boileau, 1902b: 204 A: YUN ORR
formosanus Bates, 1866: 347 A: HAI TAI
himalayae Gravely, 1915: 429 A: NP SD ORR
lewisi Boileau, 1905c: 61 A: HAI JA TAI
miwai Nagel, 1941: 66 A: SC
sinicus Schenk, 2011: 4 A: GUA
sticheri Okuda & Fukinuki, 2000: 8 A: BT
genus Penichrolucanus Deyrolle, 1863: 485 type species Penichrolucanus copricephalus Deyrolle, 1863
Xizangia Y.-W. Zhang, 1988: 233 type species Xizangia cryptonychus Y.-W. Zhang, 1988

cryptonychus Y.-W. Zhang, 1988: 234 (Xizangia) A: XIZ

tribe Lucanini Latreille, 1804

genus Cladophyllus Houlbert, 1912a: 138 type species Cladophyllus oberthuri Houlbert, 1912
bousqueti Boucher, 1996a: 77 A: YUN
nobuhikoi Ikeda, 2000a: 42 A: YUN ORR
oberthuri Houlbert, 1912a: 139 A: YUN
flavus Didier, 1930: 126

genus Cyclorasis J. Thomson, 1862: 397 type species Lucanus platycephalus Hope, 1842
Cyclophthalmus Hope & Westwood, 1845: 4 [HN] type species Lucanus platycephalus Hope, 1842

platycephalus Hope, 1842b: 83 [=1845b: 73] (Lucanus) A: BT NP SD SCH YUN ORR
branczicki Nonfried, 1905a: 11 (Prismognathus)

subnitens Parry, 1862: 112 (Cyclorasis) A: BT SD ORR
parvus Didier, 1929: 79 (Prismognathus)

genus Eolucanus Kurosawa, 1970: 159 type species Lucanus gracilis Albers, 1889
davidis Deyrolle, 1878: 93 (Pseudolucanus) A: CHQ GAN SCH
gracilis gracilis Albers, 1889: 319 (Lucanus) A: BT NP SD XIZ ORR
gracilis sharchokpai Okuda & Maeda, 2014: 18 (Lucanus) A: AP

74

LUCANIDAE

kerleyi Boucher, 1995: 512 (Lucanus) A: NP UP
lesnei Planet, 1905: 212 (Lucanus) A: SCH XIZ YUN ORR
mingyiae H. Huang, 2006: 14 (Lucanus) A: YUN ORR
oberthuri Planet, 1896c: 279 (Lucanus) A: AP NP SD XIZ
pani H. Huang, 2006: 16 (Lucanus) A: XIZ
prometheus Boucher & J. Huang, 1991: 32 (Lucanus) A: XIZ
genus Lucanus Scopoli, 1763: 1 type species Scarabaeus cervus Linnaeus, 1758
Hexaphyllus Mulsant, 1839: 119 type species Hexaphyllus pontbrianti Mulsant, 1839 (=Scarabaeus cervus Linnaeus, 1758)
Pseudolucanus Hope & Westwood, 1845: 30 type species Scarabaeus capreolus Linnaeus, 1764

angusticornis inclinatus Schenk, 2008a: 5 A: GUX YUN
atratus Hope, 1831: 22 A: BT NP SD XIZ ORR
barbarossa Fabricius, 1801b: 251 E: FR PT SP N: MO
boileaui Planet, 1897b: 205 A: SCH
brivioi Zilioli, 2003: 266 A: FUJ
bruanti J.P. Lacroix, 1973: 138 A: BT ORR
busignyi Planet, 1909: 173 (Pseudolucanus) A: TR
cantori Hope, 1842b: 83 [=1845b: 73] A: BT NP SD XIZ ORR
cervus akbesianus Planet, 1896b: 256 A: SY TR
cervus cervus Linnaeus, 1758: 353 (Scarabaeus) E: AL AN AU BE BH BU BY CR CT CZ DE EN FR GB GE GG GR
HU IT LA LS LT LU MD ME NL NT PL PT RO SB SK SL SP ST SV SZ UK A: KZ TR
americanus Hope & Westwood, 1845: 10
antennatus Rey, 1890a: 164
armiger Herbst, 1790: 301
brevicollis Motschulsky, 1870: 34
capitalis Möllenkamp, 1909c: 242
capra A.G. Olivier, 1789: no 1: 11
capreolus Fuessly, 1775: 3 [HN]
dorcas O.F. Müller, 1776: 52
europaeus Motschulsky, 1870: 30
grandis Haworth, 1807: 77
heptaphyllus Abeille de Perrin, 1867: 70
hircus Herbst, 1790: 299
inermis Marsham, 1802: 48
longipennis Rothenburg, 1900: 99
lusitanicus Hope & Westwood, 1845: 9
major Burmeister, 1847: 350
maxillaris Motschulsky, 1845a: 60
mediadonta J.P. Lacroix, 1978: 253
microcephalus Mulsant, 1842: 586
motschulskyi Benesh, 1960: 140 [RN]

LUCANIDAE
nigrinus Marcu, 1931: 214
poujadei Planet, 1897d: 267
smyrniacus Didier & Séguy, 1953b: 29
syriacus Planet, 1897a: 65 [HN]
tauricus Motschulsky, 1845a: 60
turcicus Sturm, 1843: 347
validus Möllenkamp, 1912b: 128
vicinus Hope & Westwood, 1845: 10

cervus judaicus Planet, 1902: 21 A: IS SY TR
cheni H. Huang, 2011: 56 A: XIZ
choui H. Huang & C.-C. Chen, 2013a: 30 A: XIZ
confusus Boucher, 1995: 507 A: BT NP SD
continentalis Zilioli, 1998: 145 A: FUJ ZHE
datunensis Hashimoto, 1984: 25 A: TAI
ritae J.P. Lacroix, 1984b: 10

delavayi Fairmaire, 1887e: xxvii A: SCH YUN
deuvei deuvei J.P. Lacroix, 1988: 7 A: KA
deuvei pakistanicus Schenk, 2009c: 8 A: PA
deuveianus Boucher, 1998: 182 A: GUX
guangxii Schenk, 2009a: 4

dirki Schenk, 2002: 11 A: YUN
didieri Planet, 1927: 106 A: YUN
dybowski dybowski Parry, 1873: 335 A: ANH BEI CHQ FE GAN HEN HUB JIL LIA NC SC SHA TIA
jilinensis J. K. Li, 1992: 68

dybowski lhasaensis Schenk, 2006b: 5 A: SCH
dybowski taiwanus Miwa, 1936: 2 A: TAI
fairmairei Planet, 1897d: 265 A: GUI SCH
jietuii Nagai & Maeda, 2010: 36

fanjingshanus H. Huang & C.-C. Chen, 2010: 123 A: GUI
feglini J.P. Lacroix & Bomans, 1973: 148 A: AF
ferriei Planet, 1898: 251 A: JA (Amami-Ôshima)
fonti Zilioli, 2005: 150 A: FUJ ZHE
formosanus Planet, 1899c: 36 A: TAI
formosus Didier, 1925c: 220 A: YUN ORR
cyclommatoides Didier, 1928: 51

fortunei W.W. Saunders, 1854: 46 A: ANH FUJ GUA HUN JIX ZHE
laevigatus Didier, 1931: 225

fryi fryi Boileau, 1911a: 431 A: XIZ YUN ORR
fryi schepanskii Schenk, 2008a: 6 A: AP
fujianensis Schenk, 2008a: 7 A: FUJ GUA GUX
suzumurai Fujita, 2010: 12

75

76

LUCANIDAE

fujitai Katsura & Giang, 2002: 14 A: YUN ORR
fukinukiae Katsura & Giang, 2002: 14 A: YUN ORR
gamunus Sawada & Watanabe, 1960: 99 A: JA (Izu-shotō)
groulti Planet, 1897c: 227 A: CH NP
hayashii Nagai, 2000b: 88 A: YUN ORR
hansi Zilioli, 2000b: 41

hermani De Lisle, 1973: 137 A: FUJ GUX HAI HUN ZHE
hewenjiae H. Huang & C.-C. Chen, 2013a: 33 A: GUX
ibericus ibericus Motschulsky, 1845a: 60 E: AB AL AR CT GG GR TR ST A: IN
caspica Semenov, 1901: 105
ravasinii Schatzmayr, 1923: 8
syriacus Motschulsky, 1870: 45

ibericus subvelutinus Motschulsky, 1870: 46 E: AR GG A: TR IN TM
imitator Boucher & J. Huang, 1991: 36 (Pseudolucanus) A: XIZ
bicolor Schenk, 1996b: 509 (Pseudolucanus)

jibii Okuda & Maeda, 2015a: 29 A: AP
kanoi chuyunshanus Sakaino & C. K. Yu, 1993: 15 A: TAI
kanoi kanoi Kurosawa, 1966: 339 A: TAI
kanoi ogakii Imanishi, 1990: 15 A: TAI
masumotoi Hirasawa & Akiyama, 1990: 55

kanoi piceus Kurosawa, 1966: 342 A: TAI
kirchneri Zilioli, 1999: 84 A: FUJ
klapperichi Bomans, 1989a: 9 A: FUJ GUA HUN JIX ZHE
koyamai Akiyama & Hirasawa, 1990: 53 A: YUN ORR
kraatzi kraatzi Nagel, 1926: 116 A: FUJ GUI SCH YUN
kurosawai Sakaino, 1995: 5 A: TAI
laminifer laminifer C.O. Waterhouse, 1890a: 33 A: SD YUN ORR
langi H. Huang, He & Shi, 2011: 59 A: XIZ
laticornis Deyrolle, 1864b: 312 A: TR
liuyei liuyei H. Huang & C.-C. Chen, 2010: 93 A: CHQ GUI
ludivinae Boucher, 1998: 178 A: YUN ORR
lunifer Westwood, 1839b: lv A: AP BT KA NP SD XIZ ORR
rugifrons Hope & Westwood, 1845: 4

macrophyllus Kraatz, 1860: 271 A: TR
maculifemoratus adachii Tsukawaki, 1995: 12 A: JA
maculifemoratus maculifemoratus Motschulsky, 1862a: 9 A: FE JA
balachowskyi J.P. Lacroix, 1968: 180
elegans Planet, 1898: 19
hopei Parry, 1862: 108
montivagus Möllenkamp, 1912b: 128

LUCANIDAE
mearesii Hope, 1842b: 83 A: AP BT NP SD XIZ ORR
nigripes Hope & Westwood, 1845: 10

mishmi Okuda & Maeda, 2014: 16 A: AP
miwai Kurosawa, 1966: 342 A: TAI
nobilis Didier, 1925c: 221 A: YUN ORR
nyishwini bretschneideri Schenk, 2008a: 8 A: AP XIZ
orientalis Kraatz, 1860: 269 A: IN IQ LE SY TR
intermedius Motschulsky, 1870: 41
piger Motschulsky, 1870: 37
tenebrosus Motschulsky, 1870: 47

parryi laetus Arrow, 1943: 134 [RN] A: CHQ GUI HEN HUB SCH SHA
oberthuri Planet, 1896a: 179

parryi parryi Boileau, 1899a: 111 A: FUJ JIX ZHE
aterrimus Didier, 1929: 95
thoracicus Didier, 1929: 96

planeti dayaoshanensis Schenk, 2011: 6 [RN] A: GUA GUX
guangxii Schenk, 2009c: 5 [HN]

planeti planeti Planet, 1899c: 72 A: YUN ORR
pontbrianti Mulsant, 1839: 120 (Hexaphyllus) E: FR SP
fabiani Mulsant & Godart, 1855c: 250
pentaphyllus Reiche, 1853a: 71
reichei Motschulsky, 1870: 31

prossi Zilioli, 2000a: 53 A: YUN ORR
smithii Parry, 1862: 108 A: BT NP SD XIZ ORR
swinhoei swinhoeyi Parry, 1874: 370 A: TAI
szetschuanicus Hanuš, 1932: 101 A: CHQ HUB HUN GUI SHX
hildegardae Zilioli, 2002a: 131

tetraodon provincialis Colas, 1949: 128 E: FR SP (Pirineos)
tetraodon serraticornis Fairmaire, 1859a: 275 E: FR (Corse)
corsicus Gautier des Cottes, 1860: liii

tetraodon sicilianus Planet, 1899a: 98 E: IT (Sicilia)
tetraodon tetraodon Thunberg, 1806b: 188 E: AL GR IT
bidens Thunberg, 1806b: 197
impressus Thunberg, 1806b: 197

thibetanus furcifer Arrow, 1950: 46 A: SCH YUN ORR
katsurai Mizunuma, 1994: 7
pseudosingularis Didier & Séguy, 1953b: 31
singularis Planet, 1903: 12 [HN]

thibetanus isaki Nagai, 2000b: 90 A: YUN ORR
thibetanus singularis Planet, 1900a: 11 A: YUN
gennestieri J.P. Lacroix, 1971: 565

77

78

LUCANIDAE

thibetanus thibetanus Planet, 1898: 277 A: SCH
victorius Zilioli, 2002b: 210 A: YUN
villosus Hope, 1831: 22 A: NP
wemckeni Schenk, 2006a: 271 A: AP
westermanni Hope & Westwood, 1845: 10 A: NP SD XIZ ORR
mniszechi Planet, 1900b: 96 (Pseudolucanus)

wittmeri J.P. Lacroix, 1984b: 7 (Pseudolucanus) A: NP PA
wuyishanensis Schenk, 1999: 114 A: FUJ JIX ZHE
xerxes Král, 2004: 184 A: IN
genus Lucanus, species incertae sedis
curtulus Motschulsky, 1845a: 60 E: “Caucasus”
genus Prismognathus Motschulsky, 1860e: 138 type species Prismognathus subaeneus Motschulsky, 1860
Eligmodontus Houlbert, 1915a: 17 type species Eligmodontus arcuatus Houlbert, 1915
Gonometopus Houlbert, 1915a: 19 type species Gonometopus triapicalis Houlbert, 1915

alessandrae Bartolozzi, 2003: 18 A: SCH YUN
angularis angularis C.O. Waterhouse, 1874a: 6 A: FE (Kuril’skie ostrova, Sakhalin) JA
angularis morimotoi Kurosawa, 1975b: 157 A: JA
angularis tokui Kurosawa, 1975b: 156 A: JA (Yaku-shima)
arcuatus Houlbert, 1915a: 17 (Eligmodontus) A: SCH
bretschneideri Schenk, 2008b: 178 A: AP
dauricus Motschulsky, 1860e: 137 A: BEI ES FE GUA HEI HUN JIX MG NC SC YUN ORR
subaeneus Motschulsky, 1860e: 138
jekelii Parry, 1864: 41 (Cyclorasis)

davidis cheni Bomans & Ratti, 1973: 283 A: TAI
nigerrimus Sakaino & C. K. Yu, 1993: 14

davidis davidis Deyrolle, 1878: 94 A: ANH BEI CHQ FUJ GAN GUA GUI HEB HEN HUB JIA NP SHA SCH XIZ
YUN ZHE
formosanus Nagel, 1928b: 274 A: TAI
fukinukii Nagai, 2000a: 7 A: XIZ ORR
haojiani H. Huang & C.-C. Chen, 2012: 16 A: GUI
kanghianus kanghianus Didier & Séguy, 1953a: 44 (Eligmodontus) A: YUN ORR
klapperichi Bomans, 1989a: 15 A: CHQ FUJ GUA GUI GUX SCH ZHE
kucerai Baba, 2004: 27 A: UP
mekolaorum Okuda & Maeda, 2015a: 33 A: AP
miyashitai miyashitai Ikeda, 1997: 29 A: AP ORR
nigricolor Boucher, 1996a: 70 A: YUN ORR
passangi Okuda & Maeda, 2015b: 38 A: AP
piluensis Sakaino, 1992: 11 A: TAI
prossi Bartolozzi & Wan, 2006: 61 A: GUI GUX SCH XIZ
shani H. Huang & C.-C. Chen, 2012: 9 A: SCH

79

LUCANIDAE
sinensis Bomans, 1989a: 16 A: FUJ
siniaevi siniaevi Ikeda, 1997: 29 A: YUN ORR
triapicalis Houlbert, 1915a: 19 (Gonometopus) A: CHQ GUA GUI GUX HUB HUN SCH YUN ZHE
zhangi H. Huang & C.-C. Chen, 2013b: 35 A: YUN
genus Tetrarthrius Didier, 1926a: 28 type species Tetrarthrius castaneus Didier, 1926
castaneus Didier, 1926a: 29 A: SD ORR
castaneicolor Arrow, 1950: 167 (Dorcus)
rufobrunneus Arrow, 1943: 136 (Dorcus) [RN]

delislei Endrődi, 1971: 10 (Prismognathus) A: NP
lucidus Boileau, 1904b: 278 (Prismognathus) A: BT SD
ruficephalus J.P. Lacroix, 1978: 281 (Prismognathus) A: SD

tribe Noseolucanini H. Huang & C.-C. Chen, 2013

genus Noseolucanus Araya & Tanaka, 1998: 334 type species Noseolucanus rugosus Araya & Tanaka, 1998
(=Pseudolucanus denticulus Boucher, 1996)
denticulus Boucher, 1996b: 441 (Pseudolucanus) A: XIZ YUN ORR
rugosus Araya & Tanaka, 1998: 335

zhengi H. Huang, 2006: 11 A: XIZ

tribe Platycerini Mulsant, 1842

genus Platycerus Geoffroy, 1762: 62 type species Scarabaeus caraboides Linnaeus, 1758
Systenocerus J. Weise, 1883: 93 type species Scarabaeus caraboides Linnaeus, 1758

acuticollis Kurosawa, 1969b: 478 A: JA
akitaorum Imura, 2007b: 483 A: JA
albisomni albisomni K. Kubota, N. Kubota & Otobe, 2008: 84 A: JA
albisomni chichibuensis K. Kubota, N. Kubota & Otobe, 2008: 85 A: JA
bashanicus Imura & Tanikado, 1998: 93 A: SCH
businskyi Imura, 1996: 42 A: SCH SHX
canae Imura, 2010: 227 A: HEN
caprea De Geer, 1774: 335 (Lucanus) E: AU BH BU BY CR CZ EN FI FR GE HU IT LA LS LT MC MD ME NR NT
PL RO SK SL SP SV SZ TR A: ES FE (Skovorodino) KZ WS
caraboides Linnaeus, 1758: 354 (Scarabaeus) E: AL AU BE BH BY CR CZ DE EN FI FR GB GE GG GR HU IT KO
LA LS LT LU MC MD ME NT PL PT RO SB SK SL SP ST SV SZ UK N: AG MO TU A: ES IN KZ NE SY TR WS
azureus Dalla Torre, 1879: 104
caerulosus Didier & Séguy, 1953a: 55
chalybaeus W.S. Macleay, 1819: 117
chlorizans Dalla Torre, 1879: 104
coeruleus Dalla Torre, 1879: 104
cribratus Mulsant & Rey, 1863: 7
diabolus Dalla Torre, 1879: 104

80

LUCANIDAE
nigripes W.S. Macleay, 1819: 117
pseudocaprea Paulus, 1970a: 113
rufipes Herbst, 1790: 311 (Lucanus)
trivialis Dalla Torre, 1879: 104
virescens Mulsant, 1842: 595
viridiaeneus Mulsant, 1842: 595
viridicollis Depoli, 1917: 190 (Systenocerus)
viridis Dalla Torre, 1879: 104

caucasicus Parry, 1864: 60 E: AB AR GG ST A: TR
consimilis consimilis Tanikado & Tabana, 1998b: 13 A: SCH
consimilis phagophilus Imura, 2005c: 260 A: SCH
cupreimicans Imura, 2006: 121 A: YUN
cyanidraconis Imura, 2008a: 122 A: SCH
delagrangei Fairmaire, 1892b: 145 A: SY TR
delicatulus delicatulus Lewis, 1883: 338 A: FE (Kuril’skie ostrova) JA NC SC YUN
delicatulus unzendakensis Fujita & Ichikawa, 1982b: 9 A: JA
diluvialis Imura, 2012: 61 A: SCH
dundai dundai Imura & Bartolozzi, 1994: 142 A: SCH
benesi Imura & Bartolozzi, 1994: 140

dundai miyatakei Tanikado & Tabana, 1998a: 26 A: SCH
feminatus Tanikado & Tabana, 1997: 4 A: SCH
hiurai hiurai Tanikado & Tabana, 1997: 6 A: SCH
hiurai tanikadoi Imura, 2002: 20 A: SCH
hongwonpyoi dabashanensis Okuda, 1997: 12 A: SCH
hongwonpyoi hongwonpyoi Imura & Choe, 1989: 20 A: SC
hongwonpyoi funiuensis Imura, 2005a: 498 A: HEN
hongwonpyoi merkli Imura & Choe, 1989: 21 A: NC
hongwonpyoi mongolicus Imura & Bartolozzi, 2006: 136 A: NMO
hongwonpyoi qinlingensis Imura, 1993: 12 A: SHA
hongwonpyoi quanbaoshanus Imura, 2011: 139 A: HEN
hongwonpyoi shennongjianus Imura, 2008a: 126 A: HUB
hongwonpyoi tianmushanus Imura & Wan, 2006: 294 A: ZHE
kawadai Fujita & Ichikawa, 1982a: 1 A: JA
kitawakii Imura & Tanikado, 1998: 95 A: CHQ SCH SHX
ladyae Imura, 2005b: 507 A: SCH
mandibularis Imura, 2009a: 77 A: GUI
masumotoi Imura, 2011: 131 A: SCH
nagahatai Imura, 2008a: 109 A: SHA
perplexus Gusakov, 2003a: 26 E: AB ST
primigenius E. Weise, 1960: 139 E: AB AR ST A: IN TR
rugosus jaroslavi Imura, 2009b: 277 A: HUB

81

LUCANIDAE
rugosus rugosus Okuda, 1997: 9 A: CHQ SCH
senguni Schweiger, 1966: 30 A: TR (West Anatolia)
spinifer Schaufuss, 1862: 198 E: FR PT SP
sue Imura, 2007c: 492 A: JA
sugitai Okuda & Fujita, 1987: 4 A: JA
tabanai baotianmanus Imura, 2005a: 498 A: HEN
tabanai tabanai Okuda & Tanikado, 1994: 6 A: SCH SHX
tabanai taibashanensis Okuda, 2003: 23 A: SHA (Taibai Shan)
takakuwai akitai Fujita, 1987: 5 A: JA
takakuwai namedai Fujita, 1987: 5 A: JA
takakuwai takakuwai Fujita, 1987: 5 A: JA
tangi Imura, 2008a: 1177 A: SCH
tieguanzi Imura, 2007a: 319 A: SCH
turnai sichuanus Imura, 2008b: 297 A: SCH
turnai turnai Imura, 2001: 27 A: HUB
urushiyamai Imura, 2007b: 480 A: JA (Kyūshū)
vicinus Gusakov, 2003a: 26: E: AB AR GG
viridicuprus kanadai K. Kubota, N. Kubota & Otobe, 2008: 95 A: JA
viridicuprus viridicuprus K. Kubota, N. Kubota & Otobe, 2008: 94 A: JA
xiongmao Imura, 2008b: 291 A: SCH
yangi H. Huang, C.-C. Chen & Imura, 2010: 234 A: GAN
yeren Imura, 2008a: 113 A: HUB
yingqii H. Huang & C.-C. Chen, 2009: 27 A: SHA

tribe Odontolabini Parry, 1870

genus Neolucanus J. Thomson, 1862: 415 type species Odontolabis baladena Hope, 1842
angulatus Hope & Westwood, 1845: 17 (Lucanus) A: BT SD XIZ ORR
celebensis Möllenkamp, 1900: 46
lama Leuthner, 1885: 430
ollenbachi Didier, 1930: 131

atratus atratus Didier, 1926d: 210 A: GUX HAI ORR
atratus donckieri Didier, 1926d: 211 A: YUN ORR
baladeva Hope, 1842d: 247 [=1842c: 127] (Odontolabis) A: AP BT FUJ GUA GUI GUX HUB NP SD TAI XIZ
YUN ORR
benoiti Schenk, 2009a: 5 A: GUA GUX
bimaculatus Schenk, 2013b: 7 A: YUN
brevis Boileau, 1899c: 197 A: YUN ORR
birmaensis Möllenkamp, 1900: 46

brochieri Bomans & Miyashita, 1997: 3 A: YUN ORR
castanopterus castanopterus Hope, 1831: 22 (Lucanus) A: AP BT FUJ GUA GUI GUX HAI NP SCH SD UP XIZ
YUN ORR

82

LUCANIDAE
bicolor Burmeister, 1847: 360 (Anoplocnemus)
melas Didier, 1930: 146
parvus Nagel, 1941: 54

castanopterus elongatulus Möllenkamp, 1907: 109 A: YUN ORR
pallidus Boileau, 1914a: 73

championi Parry, 1864: 20 A: FUJ GUA GUI GUX HKG TAI ZHE
confucius J.P. Lacroix, 1972b: 114 A: BT SD ORR
fiedleri Schenk, 2006b: 9 A: GUX HAI YUN ORR
fuliginatus Mizunuma, 1994: 8 A: YUN
fuscus fuscus Didier, 1926d: 213 A: GUX HAI YUN ORR
armatus J.P. Lacroix, 1972b: 111
sianoukei J.P. Lacroix, 1978: 269

fuscus zhongguo Schenk, 2012: 3 A: GUX
giganteus Pouillaude, 1914b: 37 A: GUA GUI GUX YUN ORR
guangxii Schenk, 2009c: 11 A: GUX
guizhoui Schenk, 2011: 2 A: GUI
imitator Kriesche, 1935: 170 A: HUN
hengshanensis Ichikawa & Fujita, 1987: 12

insularis Miwa, 1929a: 73 A: JA (Ishigaki-jima)
insulicola donan Mizunuma, 1985: 18 A: JA (Yayeyama-shotō)
insulicola insulicola Kurosawa, 1976c: 190 A: JA
kachinensis Schenk, 2014: 2 A: YUN ORR
latissimus Schenk, 2013a: 6 A: FUJ GUA ZHE
maedai maedai Nagai, 2001a: 9 A: YUN ORR
marginatus dohertyi Houlbert, 1914a: 281 A: YUN ORR
marginatus marginatus C.O. Waterhouse, 1872: 53 A: BT SD XIZ ORR
maximus Houlbert, 1912b: 193 A: GUI XIZ YUN ORR
montanus Kriesche, 1935: 169 A: SCH XIZ
aterrimus Weinreich, 1959: 757

nitidus lividus Didier, 1930: 142 A: ANH FUJ GUA GUX HUB JIA JIX ZHE
nitidus nitidus W.W. Saunders, 1854: 47 (Odontolabis) A: SCH YUN
oberthueri bisignatus Houlbert, 1914a: 252 A: YUN ORR
oberthueri oberthueri Leuthner, 1885: 429 A: GUI GUX YUN ORR
okinawanus Sakaino, 1984: 10 A: JA
pallescens diffusus Bomans, 1989: 12 A: FUJ
pallescens pallescens Leuthner, 1885: 426 A: HUN JIX YUN ZHE
pallescens rutilans Bomans, 1989: 14 A: FUJ
parryi parryi Leuthner, 1885: 424 A: GUA GUI GUX YUN
perarmatus goral Kriesche, 1926: 382 A: FUI GUA GUX HAI JIX ZHE
perarmatus perarmatus Didier, 1925e: 262 A: YUN ORR
protogenetivus hamaii Mizunuma, 1994: 10 A: JA

LUCANIDAE

83

protogenetivus protogenetivus Kurosawa, 1976c: 190 A: JA
pseudopacus intermedius Houlbert, 1914a: 257 A: GUA GUI
curvidens J.P. Lacroix, 1978: 267

robustus lemeei Houlbert, 1914a: 276 A: GUA GUX ORR
robustus robustus Boileau, 1914b: 133 A: YUN ORR
rudolphi Schenk, 2008a: 10 A: AP XIZ
saundersi Parry, 1864: 20 A: BT SD ORR
shaanxiensis Schenk, 2008a: 11 A: SCH SHA YUN
sinicus opacus Boileau, 1899b: 176 A: FUJ GUA GUX HAI HUB HUN JIX
nosei Mizunuma, 1994: 8

sinicus sinicus W.W. Saunders, 1854: 48 (Odontolabris) A: FUJ GUA GUI GUX HUN JIX ZHE SCH SHA SHX
YUN ZHE
extremus Kriesche, 1940: 40

spicatus Didier, 1930: 141 A: FUJ GUA HUB JIX ZHE
fujitai Mizunuma, 1994: 10

svenjae Schenk, 2003: 12 A: XIZ
swinhoei Bates, 1866: 346 A: TAI
bifoveolatus Möllenkamp, 1912b: 128
cephalotes Möllenkamp, 1909b: 58
doro Mizunuma, 1994: 8
eugeniae Bomans, 1992: 363
horaguchii Nagai, 2001b: 13

taiwanus Mizunuma, 1994: 8 A: TAI
tanakai Mizunuma, 1994: 10 A: HAI
tao Kriesche, 1935: 170 A: GUI GUX
hainanensis Mizunuma, 1994: 218

tibetanus Schenk, 2003: 13 A: XIZ
vendli Dudich, 1923: 17 A: TAI
waterhousei Boileau, 1899b: 178 A: SD ORR
zebra J.P. Lacroix, 1988: 5 A: TAI
genus Odontolabis Hope, 1842d: 247 type species Odontolabis cuvera Hope, 1842
Anoplocnemus Hope, 1843d: 279 type species Lucanus burmeisteri Hope, 1841

cuvera bouloixi J.P. Lacroix, 1984a: 136 A: YUN
cuvera cuvera Hope, 1842d: 247 [=1842c: 127] A: BT NP SD ORR
gestroi Boileau, 1902b: 204
mandibularis Möllenkamp, 1909a: 4
prinseppii Hope & Westwood, 1845: 16 (Lucanus)

cuvera fallaciosa Boileau, 1901: 284 A: GUA GUI ORR
fruhstorferi Meyer-Darcis, 1901: 355
salvazae Pouillaude, 1913d: 334

84

OCHODAEIDAE

cuvera rattii J.P. Lacroix, 1981: 60 A: BT
cuvera sinensis Westwood, 1848a: 54 (Lucanus) A: FUJ GUI HAI HKG HUN SCH SHG YUN ORR
macrocephala J.P. Lacroix, 1984a: 80 A: GUA GUX ORR
platynota Hope & Westwood, 1845: 18 (Lucanus) A: FUJ GUA GUI HUN SCH YUN ZHE ORR
coomani Didier, 1927g: 271
emarginatus W.W. Saunders, 1854: 49 (Odontolabris)
evansii Westwood, 1855: 201 (Odontolabris)

siva Hope & Westwood, 1845: 16 (Lucanus) A: AP BT FUJ GUA GUI GUX HAI HEN HUB HUN JIX NP SCH SD
TAI YUN ORR
bellicosa Reiche, 1853a: 72
carinatus Parry, 1864: 76
chinensis Arrow, 1943: 134 (Calcodes) [RN]
mingianus Didier & Séguy, 1953a: 37
parryi Boileau, 1905b: 47

family OCHODAEIDAE Mulsant & Rey, 1871
subfamily Ochodaeinae Mulsant & Rey, 1871
tribe Ochodaeini Mulsant & Rey, 1871

genus Codocera Eschscholtz, 1821: 398 type species Lethrus ferrugineus Eschscholtz, 1818
Stomphax Fischer von Wladheim, 1823: 158 type species Stomphax crucirostris Fischer von Waldheim, 1824

ferruginea chinensis Balthasar, 1936b: 98 A: HEB HEI
ferruginea ferruginea Eschscholtz, 1818: 451 (Lethrus) E: AB AR AU BH BU CT CZ GG HU KZ MD RO SB SK ST
UK A: ES FE KI KZ MG NC NE NMO TD TM UZ WS XIN
crucirostris Fischer von Waldheim, 1824: 160 (Stomphax)

tuberculata S.I. Medvedev & Nikolajev, 1972: 445 A: MG
genus Ochodaeus Dejean, 1821: 56 type species Melolontha chrysomelina Fabricius, 1792 (=Scarabaeus chrysomeloides Schrank, 1781)
Psephus Kirby & Spence, 1826: 678 type species Scarabaeus chrysomeloides Schrank, 1781

berytensis Petrovitz, 1965d: 681 A: IS LE TR
chrysomeloides Schrank, 1781: 16 (Scarabaeus) E: AB AL AR AU BH BY CR CT CZ FR GE GG GR HU IT MC MD
PL RO SB SK SL SP ST SZ UK N: AG
chrysomelina Fabricius, 1792b: 175 (Melolontha)
clypeatus Motschulsky, 1860d: 133
scymnoides Mulsant, 1842: 342

coomani Paulian, 1945: 185 A: SCH ORR
cornifrons Solsky, 1876a: 394 E: AB AR BU KZ RO ST A: IN IS KZ SY TD TM UZ
alleonis Fairmaire, 1884f: cxli
nigricollis Semenov, 1891: 313

cychramoides Reitter, 1892e: 255 E: IT

85

HYBOSORIDAE
gigas Marseul, 1878: 58 N: AG EG LB MO TU A: IS
graecus Petrovitz, 1968b: 15 E: GR
holzschuhi Petrovitz, 1971e: 566 A: TR
inermis Reitter, 1892e: 256 E: SP
inscutellaris Pic, 1900a: 6 A: IS
integriceps Semenov, 1891: 313 E: AB AR AU BU CR CZ GG HU MC SB SK ST UK
euxinus Semenov, 1900a: 93

mongolicus Petrovitz, 1967d: 430 A: HEB
montanus Fuente, 1912: 366 E: SP
nurestanicus Nikolajev, 1995: 77 A: AF (Nurestan)
pocadioides Motschulsky, 1860d: 132 E: PT SP
mariateresae Luna de Carvaldo, 1982: 10

seleuciensis Petrovitz, 1963c: 245 A: TR
solskyi Semenov, 1891: 311 A: AF IN KZ TD TM UZ XIN
pallens Petrovitz, 1965d: 682
zarudnyi Semenov & S.I. Medvedev, 1931: 394

thalycroides Reitter, 1892e: 256 E: BU GR MC TR A: TR
tuberculifrons Reitter, 1908b: 47 N: EG

tribe Nothochodaeini Nikolajev, 2015

genus Nothochodaeus Nikolajev, 2005a: 219 type species Ochodaeus maculatus C.O. Waterhouse, 1875
Mimochodaeus Nikolajev, 2009 type species Ochodaeus grandiceps Fairmaire, 1897

asahinai Kurosawa, 1968b: 240 (Ochodaeus) A: JA (Ryukyus)
formosanus Kurosawa, 1968b: 241 (Ochodaeus) A: TAI
grandiceps Fairmaire, 1897f: 241 (Ochodaeus) A: SCH
interruptus horii Ochi, Kawahara & Inagaki, 2006a: 141 (Ochodaeus) A: JA (Amami-Ôshima)
interruptus interruptus Kurosawa, 1968b: 243 (Ochodaeus) A: JA (Ishigaki-jima) TAI
interruprus kurosawai Ochi & Kawai, 2002: 51 [RN] (Ochodaeus) A: JA (Okinawa-shotō)
carinatus Kurosawa, 1968b: 243 (Ochodaeus) [HN]

jengi Huchet & C.-L. Li, 2015: 182 A: TAI
lanyuensis Ochi, Masumoto & C.-L. Li, 2006: 304 (Ochodaeus) A: TAI
maculatus koreanus J.I. Kim, 1990: 154 (Ochodaeus) A: SC
maculatus maculatus C.O. Waterhouse, 1875a: 95 (Ochodaeus) A: JA
sakaii Ochi, Masumoto & C.-L. Li, 2006: 302 (Ochodaeus) A: TAI
tonkineus Balthasar, 1952: 222 (Ochodaeus) A: CH TAI ORR

family HYBOSORIDAE Erichson, 1847
subfamily Ceratocanthinae Martínez, 1968

genus Madrasostes R. Paulian, 1975: 610 type species Madrasostes nigrum R. Paulian, 1975
deharvengi Gao, 2009: 155 A: GUX
feae Gestro, 1898: 486 (Pterorthochaetes) A: AP NP SD UP ORR

86

HYBOSORIDAE

hisamatsui Ochi, 1990: 114 A: JA
kazumai Ochi, Johki & Nakata, 1990: 31 A: JA
suzukii Ochi, Tsai & Tsukamoto, 2005: 494 A: TAI
taiwanense Ochi, Tsai & Masumoto, 2005: 492 A: TAI
genus Pterorthochaetes Gestro, 1898: 479 type species Synarmostes gestroi Harold, 1874
dembickyi Ballerio, 2014b: 522 A: AP
insularis Gestro, 1898: 482 A: NP YUN ORR
septemtrionalis Ballerio, 1999: 223 A: NP
yunnanensis Ballerio, 2014a: 277 A: YUN

subfamily Hybosorinae Erichson, 1847

genus Celaenochrous Kuijten, 1984: 47 type species Celaenochrous sinensis Kuijten, 1984
sinensis Kuijten, 1984: 47 A: SCH
genus Hybosorus W.S. Macleay, 1819: 120 type species Geotrupes arator Illiger, 1803 (=Hybosorus illigeri
Reiche, 1853)
illigeri Reiche, 1853b: 87 E: AB AR BU CR FR GG GR IT MA MC PT RO SB SP ST UK (Krym) N: AG EG LB MO
TU A: AE AF CH CY HP IN IQ IS JO PA QA SA SI SD SY TD TM TR UZ YE AFR NARi NTRi ORR
arator Illiger, 1803: 210 (Geotrupes) [HN]
carolinus LeConte, 1848: 84
nitidus van Lansberge, 1882: xxiii
nossibianus Fairmaire, 1895a: 17
pinguis Westwood, 1846: 159
palearcticus Endrődi, 1957: 46
roei Westwood, 1846: 159

laportei Westwood, 1846: 159 Ei: FR N: EG AFR
orientalis Westwood, 1846: 159 A: AF HP NP PA SD UP XIZ ORR
genus Phaeochroops Candèze, 1876: lxiii type species Phaeochroops lansbergei Candèze, 1876
taiwanus Nomura, 1973b: 39 A: TAI
genus Phaeochrous Laporte, 1840: 108 type species Phaeochrous emarginatus Laporte, 1840
Silphodes Westwood, 1841b: 41 type species Silphodes sumatrensis Westwood, 1841

beccarii Harold, 1871a: 26 A: YE AFR
arabicus Arrow, 1909b: 497
elgonensis Schouteden, 1918: 199
gonsalvezi Petrovitz, 1975d: 621
niloticus Burgeon, 1928: 184
stercorarius Kolbe, 1895b: 342
tangensis Schouteden, 1918: 199

borealis Kuijten, 1984: 46 A: SCH

87

GLAPHYRIDAE

dissimilis dissimilis Arrow, 1909b: 496 A: YUN ORR
dissimilis vietnamicola Kuijten, 1978: 33 A: GUI ORR
emarginatus davidis Fairmaire, 1886: 323 A: YUN ORR
emarginatus emarginatus Laporte, 1840: 109 A: BT CHQ GUI GUX HP JA (Ryukyus) NP PA SC TAI UP YUN
“Thibet” AUR ORR
alternatus Fairmaire, 1879b: 112
asiaticus Lewis, 1896: 332
celebensis Pic, 1928b: 5
gracilis Petrovitz, 1975: 222
hirtipes W.J. Macleay, 1864: 125
pallidus Arrow, 1909b: 496
perroudi Pic, 1943: 15
sumatrensis Westwood, 1841b: 41

emarginatus suturalis van Lansberge, 1885b: 394 A: XIZ ORR
emarginatus thilliezi Keith, 1999: 12 A: PA
enigmaticus enigmaticus Kuijten, 1978: 32 A: YUN ORR
enigmaticus ovchinnikovi Nikolajev, 2008a: 202 A: PA
hainanensis Y.-W. Zhang, 1990a: 145 A: HAI HKG
intermedius intermedius Pic, 1928b: 5 A: GUI ORR
intermedius occidentalis Kuijten, 1978: 29 A: BT NP UP ORR
pseudointermedius Kuijten, 1978: 29 A: GUI ORR
rufus Pic, 1928b: 5 A: JIX SCH
separabilis Y.-W. Zhang, 1990a: 146 A: CH
tokaraensis Nomura, 1961b: 71 A: JA (Ryukyus) SC
genus Phaeocroides Péringuey, 1908: 647 type species Phaeocroides damarinus Péringuey, 1908
nanus Arrow, 1942: 924 (Phaeochrous) A: NP ORR
genus Seleucosorus Kuijten, 1983: 36 type species Hybosorus punctatissimus Reiche, 1861
punctatissimus Reiche, 1861a: 1 (Hybosorus) A: SY TR

subfamily Pachyplectrinae Ocampo, 2006

genus Brenskea Reitter, 1891g: 254 type species Brenskea coronata Reitter, 1891
Spermohybosorus Pic, 1922b: 21 type species Spermohybosorus testaceus Pic, 1922 (=Brenskea coronata Reitter, 1891)

coronata Reitter, 1891g: 255 E: AB N: AG EG LB MO TU A: AF IN IS KZ OM PA SI TD TM UZ
chudeaui Reitter, 1909d: 526
testaceus Pic, 1922b: 21 (Spermohybosorus)
varentzowi Semenov, 1896b: 45

family GLAPHYRIDAE W.S. Macleay, 1819
subfamily Amphicominae C.É. Blanchard, 1845

genus Amphicoma Latreille, 1806: 118 type species Melolontha abdominalis Fabricius, 1781

88

GLAPHYRIDAE
Amphitriche Gistel, 1848a: 162 [RN] type species Melolontha abdominalis Fabricius, 1781
Anthypnoides Yawata, 1942: 34 type species Anthypnoides splendens Yawata, 1942
Arrhephora Fairmaire, 1891a: 8 type species Arrhephora chalcochrysea Fairmaire, 1891
Toxocerus Fairmaire, 1891a: 7 type species Toxocerus rothschildii Fairmaire, 1891
Thypnia Endrődi, 1952a: 4 type species Anthypna carceli Laporte, 1832

abdominalis abdominalis Fabricius, 1792b: 171 (Melolontha) E: IT SL SZ
alpina A.G. Olivier, 1789: no 5: 83 (Melolontha)
cyanipennis Reitter, 1903a: 154 (Anthypna)

abdominalis aemiliana Ghidini, 1956: 61 (Anthypna) E: IT
abdominalis binaghii Ghidini, 1956: 60 (Anthypna) E: IT (Liguria)
amaliae Nikodým, 2005: 17 A: ANH
aurata Yawata, 1942: 36 (Anthypnoides) A: TAI
brittoni Endrődi, 1952a: 17 (Anthypna) A: SE
businskyi Nikodým, 2005: 11 A: HUB
carceli Laporte, 1832b: 411 (Anthypna) E: IT
duponcheli Luigioni, 1903: 22 (Anthypna)
romana Duponchel, 1833: 254 (Anthypna)

carinata Endrődi, 1952a: 34 (Anthypna) A: FUJ GAN HUB JIX
cervenkai Nikodým, 2005: 16 A: YUN
corinthia Fairmaire, 1891a: 9 (Arrhephora) A: FUJ HKG
damiani Nikodým, 2005: 7 A: YUN
davidis Fairmaire, 1886: 323 (Anthypna) A: YUN
dolorosa Fairmaire, 1891a: 8 (Arrhephora) A: JIX
dubia Semenov, 1891: 330 (Toxocerus) A: GAN
dundai Nikodým, 2005: 19 A: SHX
emeia Nikodým, 2005: 10 A: SCH
fairmairei Semenov, 1891: 330 (Toxocerus) A: GAN
florentini Fairmaire, 1893d: 287 (Toxocerus) A: GUI YUN ORR
floriani Nikodým, 2009a: 25 A: SCH
formosana Miyake, 1982b: 173 A: TAI
fruhstorferi Endrődi, 1952a: 23 (Anthypna) A: YUN
graeca Nikodým, 2009b: 15 E: AL GR
iberica Drioli, 1980: 345 (Anthypna) E: SP
klapperichi Endrődi, 1952a: 31 (Anthypna) A: FUJ
lalashanensis C.-L. Li, 2011: 25 A: TAI
latouchei Fairmaire, 1900b: 617 (Toxocerus) A: FUJ SCH
magdalenae Nikodým, 2007: 21 A: HEN
marginata Nikodým, 2009b: 16 A: FUJ
nikodymi Keith, 2008c: 23 A: YUN
pectinata Lewis, 1895c: 388 (Anthypna) A: JA
purpuripennis Petrovitz, 1972a: 28 (Anthypna) A: HUB
regalis Arrow, 1938b: 289 (Anthypna) A: FUJ GUA GUX ORR

GLAPHYRIDAE

89

riaultii Fairmaire, 1897f: 243 (Toxocerus) A: GUI
rothschildii Fairmaire, 1891a: 7 (Toxocerus) A: CHQ JIX
chalcochrysea Fairmaire, 1891a: 8 (Arrhephora)

schneideri Nikodým, 2005: 18 A: SCH SHX
sichuana Nikodým, 2005: 8 A: SCH
splendens Yawata, 1942: 35 (Anthypnoides) A: JA (Ishigaki-jima, Yonaguni-jima)
tesari Endrődi, 1952a: 32 (Anthypna) A: GUI JIX
yunnanica Petrovitz, 1972a: 27 (Anthypna) A: YUN
genus Amphicoma, nomen dubium
metallica A. Villa & G.B. Villa, 1833: 34 E: IT
genus Anthypna Eschscholtz, 1818: 472 type species Melolontha cyanipennis Fabricius, 1801 (=Melolontha
meles Fabricius, 1792)
Psilodema C.É. Blanchard, 1845: 235 type species Melolontha meles Fabricius, 1792

ciliata Ménétriés, 1836b: 180 (Amphicoma) A: TR
mustela Waltl, 1838: 459 (Amphicoma)

meles Fabricius, 1792b: 183 (Melolontha) N: AG MO TU
barbara Reitter, 1890b: 55 (Amphicoma)
cyanipennis Fabricius, 1801b: 184 (Melolontha)
laufferi Reitter, 1903a: 140 (Amphicoma)

genus Eulasia Truqui, 1848: 16 type species Scarabaeus arctos Pallas, 1781
subgenus Eulasia Truqui, 1848: 16 type species Scarabaeus arctos Pallas, 1781
arctos anatolica Reitter, 1903a: 148 (Amphicoma) A: TR
arctos arctos Pallas, 1781: 16 (Scarabaeus) E: AR GG ST A: TR
chloridicollis Motschulsky, 1845a: 58 (Amphicoma)
faldermanni Reitter, 1903a: 148 (Amphicoma)
subfasciatula Reitter, 1903a: 147 (Amphicoma)

arctos armeniaca Reitter, 1890b: 61 (Amphicoma) E: AR A: TR
arctos martes I. Frivaldszky, 1845: 182 (Amphicoma) E: BU GR MC A: TR
aurantica Reitter, 1890b: 61 (Amphicoma) E: TR A: IN
azarbaijanica Petrovitz, 1980: 614 (Amphicoma) A: IN
bicolor bicolor Waltl, 1838: 458 (Amphicoma) E: TR A: TR
limbipennis Reitter, 1899b: 158 (Amphicoma)

bicolor dichroa Reitter, 1890b: 60 (Amphicoma) E: BU GR MC
bodemeyeri Petrovitz, 1965d: 683 (Amphicoma) A: IN
bombyliformis athenae Petrovitz, 1980: 615 (Amphicoma) E: GR
bombyliformis bombyliformis Pallas, 1781: 17 (Scarabaeus) E: AB AR GG KZ ST UK (Krym) A: IN KZ TM TR
erevanica Petrovitz, 1972a: 23 (Amphicoma)
ochraceipennis Ménétriés, 1832: 187 (Amphicoma)

bombyliformis pygidialis Petrovitz, 1963c: 261 (Amphicoma) A: TR

90

GLAPHYRIDAE

corniculata Reitter, 1903a: 147 (Amphicoma) A: TR
cornifrons Reitter, 1903a: 147 (Amphicoma) A: TR
chrysopyga Faldermann, 1835a: 284 (Amphicoma) E: AB AR GG ST A: IN TR
pallidopilosa Motschulsky, 1860g: 166 (Trichopleurus)

diadema Reitter, 1890b: 58 (Amphicoma) A: IN TR
diademata Reitter, 1903a: 143 (Amphicoma)

eichleri Zaitzev, 1923: 23 (Amphicoma) E: AR
ernae Petrovitz, 1962b: 118 (Amphicoma) A: TR
hyrax Truqui, 1848: 36 A: IS LE TR SY
truqui Reitter, 1890b: 59 (Amphicoma)

jordanica Mitter, 1992a: 31 (Amphicoma) A: JO
korbi Petrovitz, 1972a: 23 (Amphicoma) E: AR A: TR
kordestana Mitter, 2004: 305 A: IN
kuschakevitschi Ballion, 1871a: 339 (Amphicoma) A: AF KI KZ TD UZ
auricollis Reitter, 1890b: 60 (Amphicoma)
belia Reitter, 1900b: 53 (Amphicoma)
clypeata Solsky, 1876a: 389 (Amphicoma)
dubia Solsky, 1876a: 389 (Amphicoma)
schneideri Reitter, 1903a: 145 (Amphicoma)
viridicollis Reitter, 1890b: 60 (Amphicoma)

montana Reitter, 1890b: 61 (Amphicoma) A: TR
naviauxi Baraud, 1971c: 43 (Amphicoma) A: IN
nitidicollis Reiche, 1862b: 539 (Amphicoma) A: IS JO LE SY TR
nitidinatis Baraud, 1990a: 120 A: IN
sublineata Reitter, 1890b: 59 (Amphicoma)

persidis Baraud, 1990a: 121 A: IN
praeusta Champenois, 1896: 15 (Amphicoma) A: IN TR
clypealis Petrovitz, 1972a: 23 (Amphicoma)

subgenus Rudeulasia Baraud, 1990a: 109 type species Amphicoma papaveris Sturm, 1843
aegyptiaca Petrovitz, 1972a: 25 (Amphicoma) N: EG
anemurensis Petrovitz, 1964c: 103 (Amphicoma) A: TR
baumanni Mitter, 1996b: 363 A: IS LE
bocquilloni Zaitzev, 1923: 14 (Amphicoma) A: IN
carinata Baraud, 1990a: 124 A: IN IQ
chalybaea brenskei Reitter, 1890b: 62 (Amphicoma) E: AR A: TR
adusta Ganglbauer, 1905: 286 (Amphicoma)

chalybaea chalybaea Faldermann, 1835a: 287 (Amphicoma) E: AB AR A: IN TM
daccordii Uliana & Sabatinelli, 2013: 125 A: IS JO
dilutipennis Reitter, 1890b: 62 (Amphicoma) A: IS LE SY
eiselti Petrovitz, 1967b: 331 (Amphicoma) A: TR
fastuosa Reitter, 1890b: 63 (Amphicoma) E: GR A: IS LE SY TR
heydeni Reitter, 1890b: 63 (Amphicoma)

GLAPHYRIDAE
reitteri Ganglbauer, 1905: 284 (Amphicoma)
semicyanea Reitter, 1890b: 63 (Amphicoma)

genei Truqui, 1848: 28 A: IN IQ IS JO LE SY TR
harmonia Petrovitz, 1968g: 474 (Amphicoma) A: SY TR
hybrida agricola Reitter, 1890b: 64 (Amphicoma) A: TR
hybrida hybrida Reitter, 1890b: 64 (Amphicoma) A: TR
ithae Reitter, 1903a: 151 (Amphicoma)

papaveris Sturm, 1843: 342 (Amphicoma) A: IN IQ IS JO LE SY TR
cupripennis Kollar, 1843: 986 (Amphicoma)
dominula Reitter, 1890b: 63 (Amphicoma)
testaceipennis Champenois, 1896: 16 (Amphicoma)

pietschmanni Breit, 1920: 55 (Amphicoma) A: IQ JO SY
pretiosa Truqui, 1848: 30 A: IS
chaifensis Reitter, 1890b: 63 (Amphicoma)
japhoensis Petrovitz, 1972a: 24 (Amphicoma)

pulchra kurdistana Petrovitz, 1968g: 474 (Amphicoma) A: IN TR
pulchra pulchra Reitter, 1890b: 62 (Amphicoma) E: AR A: IN TR
rapillyi Baraud, 1988a: 206 A: IS JO LE SY TR
regeli Ballion, 1878: 286 (Amphicoma) A: KI TD UZ
bucharica Reitter, 1903a: 152 (Amphicoma)
iris Reitter, 1897e: 125 (Amphicoma)
theana Reitter, 1903a: 152 (Amphicoma)
violaceipennis Ballion, 1878: 288 (Amphicoma)

rittneri Uliana & Sabatinelli, 2010: 36 A: IS
rodionovi Zaitzev, 1923: 22 (Amphicoma) E: AB
saccai Petrovitz, 1972a: 25 (Amphicoma) A: IS JO LE SY
speciosa Champenois, 1900: 248 (Amphicoma) A: IN SY TR
straussi Ganglbauer, 1905: 285 (Amphicoma) A: IN IQ TR
subgenus Solskiola Semenov, 1903g: 391 type species Amphicoma analis Solsky, 1876
analis Solsky, 1876a: 387 (Amphicoma) A: AF IN TD TM UZ
subgenus Trichopleurus Motschulsky, 1860f: 162 type species Melolontha bombylius Fabricius, 1787
Vittateulasia Baraud, 1990a: 109 type species Melolontha vittata Fabricius, 1775

bombylius Fabricius, 1787a: 25 (Melolontha) E: SP N: AG MO TU
goudotii goudotii Laporte, 1840: 153 (Amphicoma) N: MO
goudotii pardoi Baraud, 1981c: 159 N: MO
goudotii lajonquierei Baraud, 1981c: 159 N: MO
palmyrensis Mitter, 1996c: 933 A: JO SY
pareyssei Brullé, 1832: 183 (Amphicoma) E: AL BU GR MC TR A: TR
lasserrei Germar, 1836: 4 (Amphicoma)

vittata lineata Faldermann, 1835a: 283 (Amphicoma) E: AR N: EG A: IN IQ IS JO LE SY TR
africana Reitter, 1890b: 57 (Amphicoma)
chrysura Reitter, 1890b: 57 (Amphicoma)

91

92

GLAPHYRIDAE
decorata Reitter, 1890b: 57 (Amphicoma)
lateritia Reitter, 1903a: 142 (Amphicoma)
syriaca Kollar, 1843: 986 (Amphicoma)

vittata persica Petrovitz, 1963c: 261 (Amphicoma) A: IN
vittata vittata Fabricius, 1775: 40 (Melolontha) E: BU GR TR A: TR
flavicans Truqui, 1848: 21
semifulva Reitter, 1890b: 57 (Amphicoma)
smyrnensis Laporte, 1840: 153 (Amphicoma)
strigada Waltl, 1838: 458 (Amphicoma)

genus Eulasia, species incertae sedis
zaitzevi Bogatchev, 1947: 16 (Amphicoma) A: IN
genus Pygopleurus Motschulsky, 1860f: 162 type species Scarabaeus syriacus Linnaeus, 1758
akbesianus Petrovitz, 1958b: 44 (Amphicoma) A: IS SY TR
cupreus Petrovitz, 1958b: 44 (Amphicoma)
viridis Petrovitz, 1958b: 44 (Amphicoma) [HN]

aleppensis Petrovitz, 1958b: 56 (Amphicoma) A: IS SY TR
palaestinensis Petrovitz, 1963c: 247 (Amphicoma)
violaceus Petrovitz, 1958b: 56 (Amphicoma) [HN]
viridis Petrovitz, 1958b: 56 (Amphicoma) [HN]

anahitae Mitter, 2001: 1041 A: IN
anemoninus Brullé, 1832: 180 (Amphicoma) E: GR TR A: LE TR
albopilosus Petrovitz, 1958b: 50 (Amphicoma)
chrysonatus Brullé, 1832: 181 (Amphicoma)
coeruleus Petrovitz, 1958b: 50 (Amphicoma) [HN]
cupreus Petrovitz, 1958b: 50 (Amphicoma) [HN]
pseudopsilotrichius Petrovitz, 1958b: 50 (Amphicoma)
purpuricollis Waltl, 1838: 458 (Amphicoma)
viridis Petrovitz, 1958b: 50 (Amphicoma) [HN]

angulatus Fairmaire, 1884b: 168 (Amphicoma) A: SY TR
apicalis Brullé, 1832: 182 (Amphicoma) E: BU GR MC
humerosus Reitter, 1890b: 55 (Amphicoma)

banghaasi Reitter, 1895b: 81 (Amphicoma) A: IN TM
coeruleus Petrovitz, 1958b: 46 (Amphicoma) [HN]
coeruleipennis Petrovitz, 1958b: 47 (Amphicoma)
cupreus Petrovitz, 1958b: 47 (Amphicoma) [HN]
mixtus Brancsik, 1899: 99 (Amphicoma)

basalis Reitter, 1890b: 53 (Amphicoma) A: IN IQ TR
purpureus Petrovitz, 1958b: 63 (Amphicoma)
viridis Petrovitz, 1958b: 61 (Amphicoma) [HN]

besucheti Baraud, 1989d: 348 A: IS JO

GLAPHYRIDAE
bimaculatus L. Redtenbacher, 1850: 48 (Amphicoma) A: SY
caesarae Baraud, 1989d: 348 A: IS SY
cirrius Petrovitz, 1958b: 60 (Amphicoma) A: IQ SA
costatus Petrovitz, 1958b: 61 (Amphicoma) A: SY TR
cyaneoviolaceus cyaneoviolaceus Motschulsky, 1860g: 166 E: AB GG A: IN
cyaneoviolaceus versicolor Baraud, 1989d: 351 A: TR
cyanescens Reitter, 1890b: 55 (Amphicoma) E: AB AR A: IN IQ SY TR
demelti Petrovitz, 1967b: 330 (Amphicoma) A: TR
despectus Petrovitz, 1958b: 45 (Amphicoma) A: TR SY
deuvei Montreuil & Serri, 2007: 2 A: IN
diffusus Petrovitz, 1958b: 45 (Amphicoma) E: AL GR MC TR A: TR
cupreus Petrovitz, 1958b: 46 (Amphicoma) [HN]
viridis Petrovitz, 1958b: 60 (Amphicoma) [HN]

distinctus Faldermann, 1835a: 286 (Amphicoma) E: AB AR ST A: IN IQ TR
persicus Ganglbauer, 1905: 289 (Amphicoma)
violaceus Petrovitz, 1958b: 59 (Amphicoma) [HN]
viridis Petrovitz, 1958b: 60 (Amphicoma) [HN]

distinguendus Muche, 1963a: 165 (Amphicoma) E: AR GG
foina Reitter, 1890b: 53 (Amphicoma) E: BU GR TR A: TR
cupreus Petrovitz, 1958b: 55 (Amphicoma) [HN]
viridis Petrovitz, 1958b: 55 (Amphicoma) [HN]

gordyenensis Petrovitz, 1971e: 571 (Amphicoma) E: AB AR A: IN TR
hirsutus Brullé, 1832: 182 (Amphicoma) E: BU GR MC A: TR
suturangulus Reitter, 1890b: 53 (Amphicoma)

humeralis Brullé, 1832: 183 (Amphicoma) E: BU GR MC TR A: TR
isikdagensis Petrovitz, 1963c: 248 (Amphicoma)
nigroapicalis Petrovitz, 1958b: 49 (Amphicoma)
petrowitzi Mikšić, 1959b: 40 (Amphicoma)
uniformis Petrovitz, 1958b: 49 (Amphicoma)
violaceus Petrovitz, 1958b: 49 (Amphicoma) [HN]
viridis Petrovitz, 1958b: 49 (Amphicoma) [HN]
viridisuturatus Reitter, 1903a: 138 (Amphicoma)

immundus Reitter, 1903a: 137 (Amphicoma) A: IN TR
distinguendus Muche, 1963a: 165 (Amphicoma)
violaceus Petrovitz, 1958b: 61 (Amphicoma) [HN]
viridis Petrovitz, 1958b: 61 (Amphicoma) [HN]

israelitus Muche, 1963a: 167 (Amphicoma) A: IS LE SY
kareli Petrovitz, 1962b: 116 (Amphicoma) E: GR (Lesvós) A: TR
katbehi Sabatinelli, 2008: 149 A: IS JO
keithi Sabatinelli & Uliana, 2009: 33 A: SY
koniae koniae Petrovitz, 1958b: 57 (Amphicoma) A: IN TR

93

94

GLAPHYRIDAE

koniae vulpoides Petrovitz, 1968b: 473 (Amphicoma) A: TR
labaumei Petrovitz, 1971e: 572 (Amphicoma) E: GR (Lesvós) A: TR
libanonensis Petrovitz, 1958b: 46 (Amphicoma) A: IS LE SY TR
rufus Petrovitz, 1958b: 46 (Amphicoma)
viridis Petrovitz, 1958b: 46 (Amphicoma) [HN]

lucarellii Piattella & Sabatinelli, 1992a: 93 (Amphicoma) E: GR
lyciensis Petrovitz, 1971e: 570 (Amphicoma) A: TR
madenensis Petrovitz, 1968b: 471 (Amphicoma) E: GG A: TR
mardiensis Baraud, 1989d: 363 A: IN
medius Petrovitz, 1958b: 57 (Amphicoma) E: GR (Lesvós) A: TR
nigropillosus Petrovitz, 1958b: 57 (Amphicoma)
violaceus Petrovitz, 1958b: 57 (Amphicoma) [HN]

mithridates Petrovitz, 1962b: 116 (Amphicoma) A: TR
monticola Petrovitz, 1964c: 102 (Amphicoma) A: SY TR
orientalis Petrovitz, 1958b: 47 (Amphicoma) A: IS JO LE SY
ottomanus Baraud, 1989d: 366 A: TR
ponticus Petrovitz, 1958b: 52 (Amphicoma) A: IS JO SY TR
cupreus Petrovitz, 1958b: 52 (Amphicoma) [HN]
viridis Petrovitz, 1958b: 52 (Amphicoma) [HN]

pseudomedius Mikšić, 1964: 286 (Amphicoma) A: TR
psilotrichius annemariae Mitter, 1992b: 170 (Amphicoma) A: TR
psilotrichius psilotrichius Faldermann, 1835a: 285 (Amphicoma) E: AB AR GG ST UK (Krym) A: IN JO SY TM TR
aurigaster Reitter, 1895b: 82 (Amphicoma)
cyanellus Zaitzev, 1923: 19
nigripennis Champenois, 1896: 13 (Amphicoma)
maculipennis Petrovitz, 1958b: 44 (Amphicoma)
violaceus Petrovitz, 1958b: 43 (Amphicoma)
viridis Petrovitz, 1958b: 43 (Amphicoma) [HN]

rapuzzii Keith & Uliana, 2008: 60 A: IN
ressli Petrovitz, 1963c: 246 (Amphicoma) A: LE TR SY
rufovillosus nigropubescens Petrovitz, 1963c: 249 (Amphicoma) A: TR
rufovillosus rufovillosus Reitter, 1907a: 28 (Amphicoma) E: AR GG A: IN IQ TR
pompa Petrovitz, 1963c: 249 (Amphicoma)

rufovillosus undofi Keith, 2000b: 568 A: SY
saltinii Uliana, 2010: 23 A: TR
samai Keith, 2001a: 5 A: LE
scutellatus Brullé, 1832: 181 (Amphicoma) E: GR
sexualis Petrovitz, 1968g: 472 (Amphicoma) A: TR
simplex Petrovitz, 1963c: 246 (Amphicoma) A: TR
sinaicus Petrovitz, 1958b: 56 (Amphicoma) A: SI

95

GLAPHYRIDAE
syriacus Linnaeus, 1758: 351 (Melolontha) A: IS JO LE SY
carinatus Muche, 1963a: 165 (Amphicoma)
nabataeensis Mitter, 1996a: 155 (Amphicoma)
piesbergeni Trappen, 1906: 124 (Amphicoma)

transcaucasicus Petrovitz, 1962b: 115 (Amphicoma) E: AB AR GG ST A: TM TR
tristis Petrovitz, 1968g: 469 (Amphicoma) A: IN TR
vulpes Fabricius, 1781a: 47 (Melolontha) E: AL BU CT GR KZ MC MD ME RO SB ST TR UK A: KZ TR
alopecias Pallas, 1781: 15 (Scarabaeus)
coeruleus Petrovitz, 1958b: 42 (Amphicoma)
griseovillosa Reitter, 1903a: 137 (Amphicoma)
hirtus Fabricius, 1781a: 47 (Melolontha)
pyrrothrix Dohrn, 1885: 80 (Amphicoma)
rufus Petrovitz, 1958b: 40 (Amphicoma)
viridis Petrovitz, 1958b: 40 (Amphicoma)
vulpeculus Truqui, 1848: 39 (Eulasia)

weilli Keith, 2008b: 40 A: SY
zagrosensis Nikodým & Král, 1998: 343 A: IN

subfamily Glaphyrinae W.S. Macleay, 1819

genus Glaphyrus Latreille, 1802a: 150 type species Scarabaeus maurus Linnaeus, 1758
subgenus Eoglaphyrus Semenov, 1926: 33 type species Glaphyrus turkestanicus Semenov, 1889
turkestanicus Semenov, 1889b: 195 A: AF TD UZ
sogdianus Semenov, 1892d: 477

subgenus Glaphyrus Latreille, 1802a: 150 type species Scarabaeus maurus Linnaeus, 1758
Pachymerus Faldermann, 1835a: 281 type species Glaphyrus micans Faldermann, 1835

aulicus Chevrolat, 1854c: 479 A: IS JO LE SY
syriacus Harold, 1896: 437

calvaster Zaitzev, 1923: 16 E: AR
cinnaberinus Schweiger, 1949: 115 A: IN
comosus Harold, 1869e: 433 A: IS JO LE SY
equestris Champenois, 1898: 248 N: EG
festivus Ménétriés, 1836a: 150 E: AB AR A: IN TR
rufipennis Gory, 1884: 103

haroldi G. Quedenfeldt, 1891: 131 N: LB
laufferi Reitter, 1903a: 130 A: IN
luristanus Reitter, 1903a: 128 A: IN
maurus Linnaeus, 1758: 348 (Scarabaeus) N: AG MO TU
cardui Fabricius, 1787a: 21 (Melolontha)
semibrunneus Reitter, 1903a: 131
viridipennis Pic, 1897b: 202

96

GLAPHYRIDAE

micans euphraticus Mitter, 1990: 111 A: TR
micans micans Faldermann, 1835a: 282 E: AB AR A: IN IQ SY TR
purpureus Pic, 1947b: 2
schelkownikowi Reitter, 1903a: 132

muticus Reitter, 1903a: 135 A: IN
olivieri Laporte, 1840: 153 N: EG
fulgidus Burmeister, 1844: 14

onopordi Reitter, 1903a: 13 A: IN IQ
opulentus mesatlanticus Kocher, 1953: 74 N: MO
opulentus opulentus Bedel, 1884: 249 N: AG MO
viridicollis Harold, 1869e: 440

orbachi Sabatinelli, Miessen & Rittner, 2012: 84 A: IS
ornatus Schweiger, 1949: 114 A: TR
oxypterus Pallas, 1771: 462 (Scarabaeus) E: AB KZ ST A: IN KZ NW
uralensis Gmelin, 1790: 1557 (Scarabaeus)
vittatus Herbst, 1790: 143 (Melolontha)

panousei Kocher, 1958: 259 N: AG MO
pubescens Schweiger, 1949: 114 A: IQ
reymondi Kocher, 1958: 259 N: MO
rothi Harold, 1869e: 441 A: LE SY
sequensi Reitter, 1903a: 131 E: AR A: IN TR
serratulae maroccanus Chobaut, 1923: 20 N: MO
serratulae serratulae Fabricius, 1792b: 168 (Melolontha) N: AG MO TU
cyaneus Pic, 1897b: 202
purpuripennis Heyden, 1887c: 438
maurus A.G. Olivier, 1789: no 5: 38 (Melolontha) [HN]

serratulae villosipennis M. Quedenfeldt, 1887: 322 N: MO
superbus straussi Reitter, 1903a: 134 E: AB AR A: IN TR
superbus superbus Champenois, 1898: 247 A: IN TM
incertus Pic, 1947b: 2

varians rufipennis Gory, 1844: 8 A: TR
varians turcicus Mitter, 1990: 108 A: TR
varians variabilis Reitter, 1903a: 131 A: TR
varians varians Ménétriés, 1836a: 150 A: IN TR
globulicollis Ménétriés, 1836a: 150

viridicollis P.H. Lucas, 1846d: pl. 25 N: AG MO TU
pulchellus P.H. Lucas, 1855a: 290

subgenus Hemiglaphyrus Champenois, 1903: 145 type species Glaphyrus caucasicus Kraatz, 1882
caucasicus Kraatz, 1882i: 312 E: AR A: IN TR
globulicollis Ménétriés, 1839: 30
persicus Pic, 1947a: 6

97

SCARABAEIDAE
modestus Kiesenwetter, 1858: 235 E: GR
nigripes Schaufuss, 1879: 478

family SCARABAEIDAE Latreille, 1802
subfamily Aegialiinae Laporte, 1840

genus Aegialia Latreille, 1806: 96 type species Scarabaeus globosus Kugelann, 1794 (=Scarabaeus arenarius
Fabricius, 1787)
arenaria Fabricius, 1787a: 11 (Scarabaeus) E: AZ BE CT (Kaliningrad) CZ DE EN FI FR GB GE IR ITi LA LT NL
NR PL PT SP SV SZi Ai: JA NARi
globosa Kugelann, 1794: 514 (Scarabaeus)

nitida C.O. Waterhouse, 1875a: 95 A: JA SC
koreana J.I. Kim, 1979: 9

genus Caelius Lewis, 1895c: 381 type species Caelius denticollis Lewis, 1895
Leptaegialia W.J. Brown, 1931: 12 type species Aegialia humeralis W.J. Brown, 1931

denticollis Lewis, 1895c: 382 A: JA
genus Psammoporus C.G. Thomson, 1859: 82 type species Scarabaeus sabuleti Paykull, 1798 (=Scarabaeus
sabuleti Panzer, 1797)
Dimalia Mulsant & Rey, 1870c: 646 [=1871a: 606] type species Scarabaeus sabuleti Panzer, 1797

abditus Nikritin, 1975: 108 A: ES MG WS
comis Lewis, 1895c: 384 (Psammobius) A: FE HEI JA NC SC
freyi Petrovitz, 1961d: 136

friebi Balthasar, 1935c: 121 (Aegialia) A: ES FE JA NO
hybridus Reitter, 1892e: 251 (Aegialia) A: ES FE MG NO
insularis Pittino, 2006d: 330 E: GB
kamtschaticus Motschulsky, 1860e: 132 (Aegialia) A: ES FE NC
latipunctus Gredler, 1866: 470 (Aphodius) E: AU GE IT SZ
matalini Gusakov, 2003a: 28 (Aegialia) A: WS (Altai)
mimicus Pittino, 2006d: 328 E: AU BE BU CZ FR GE GR LU NL PL RO SK SZ
nakanei horii Ochi & Kawahara, 2012: iii A: JA (Hokkaido)
nakanei nakanei Masumoto, 1986b: 144 A: JA
sabuleti Panzer, 1797a: 3 (Scarabaeus) E: BY CT DE EN FI GE LA LT NR NT PL SB SL SV UK
sibiricus Stebnicka, 1977b: 485 (Aegialia) A: FE
tsukamotoi Masumoto, 1986b: 146 A: JA
genus Rhysothorax Bedel, 1911a: 93 type species Scarabaeus rufus Fabricius, 1792
Anomalaegialia W.J. Brown, 1931: 15 type species Aegialia spissipes LeConte, 1878

rufus Fabricius, 1792a: 39 (Scarabaeus) E: BE BU CT DE FL FR GB GE KZ LA LT NI NR PL RO SV UK NARi
arnoldii S.I. Medvedev, 1951b: 546 (Mothon)
rufina Silfverberg, 1977: 91 (Aegialia) [RN]

98

SCARABAEIDAE
spissa A. Schmidt, 1912: 7 (Aegialia) [RN]
spissipes LeConte, 1878: 611 (Aegialia)

genus Silluvia Landin, 1949: 3 type species Silluvia elongata Landin, 1949
subgenus Silluvia Landin, 1949: 3 type species Silluvia elongata Landin, 1949
gansuensis Frolov, 2008: 44 (Aegialia) A: GAN SCH
gogona Stebnicka, 1977a: 269 (Aegialia) A: BT NP
gosainkundae Stebnicka, 1986a: 42 (Aegialia) A: NP
himalayana Petrovitz, 1968d: 186 (Psammoporus) A: SD
igori Frolov, 2008: 45 (Aegialia) A: SCH
kabaki Frolov, 2002c: 308 (Aegialia) A: SCH
petrovitzi Stebnicka, 1977a: 270 (Aegialia) A: HP
shashi Stebnicka, 1977b: 429 (Aegialia) A: SCH
simbuae Stebnicka, 1990: 13 (Aegialia) A: NP
sinica Stebnicka, 1977b: 428 (Aegialia) A: SCH
striatocrenata Fairmaire, 1889a: 14 (Oxyomus) A: SHN ORR
wassuensis Petrovitz, 1963a: 118 (Psammoporus) A: SCH
wittmeri Stebnicka, 1981b: 539 (Aegialia) A: NP
yunnanica Frolov, 2008: 43 (Aegialia) A: YUN

subfamily Eremazinae Iablokoff-Khnzorian, 1977

genus Eremazus Mulsant, 1851a: 139 type species Eremazus unistriatus Mulsant, 1851
Millingenia Sharp, 1874b: 124 type species Millingenia fossor Sharp, 1874 (=Aegialia punctatus Harold, 1869)
Tolisus Sharp, 1874b: 125 type species Tolisus aeneus Sharp, 1874 (=Eremazus unistriatus Mulsant, 1851)

cribratus Semenov, 1893b: 449 E: AB GG ST A: AF IN KI KZ PA TM UZ WS
marmottani Fairmaire, 1871a: 374 (Aegialia) N: AG MO TU
punctatus Harold, 1869b: 103 (Aegialia) N: EG TU
fossor Sharp, 1874b: 125 (Millingenia)

sefrensis Clouët des Pesruches, 1897: 266 N: AG LB MO
unistriatus Mulsant, 1851a: 140 E: AR GG TR N: AG CI EG IS LB MO TU A: AE AF BA IN IQ IS KU KZ OM PA SA
SY TD TM TR UP UZ AFR
aeneus Sharp, 1874b: 126 (Tolisus)
cloueti Semenov & S.I. Medvedev, 1938: 241
minor Sharp, 1874b: 126 (Tolisus)

subfamily Aphodiinae Leach, 1815
tribe Aphodiini Leach, 1815
subtribe Aphodiina Leach, 1815

genus Acanthobodilus G. Dellacasa, 1983a: 105 type species Aphodius immundus Creutzer, 1799
immundus Creutzer, 1799b: 57 (Aphodius) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FR GE GG GR HU
IT LA LT LU MC MD ME NT PL PT RO SB SK SL SP ST SV SZ TR UK N: EG MO A: AF CY ES FE IN IQ IS JA
KI KZ MG NMO SY TD TM TR UZ WS XIN

SCARABAEIDAE

99

arvernicus Hoffmann, 1928b: 118 (Aphodius)
fulvicollis Mulsant, 1842: 227 (Aphodius)
melinopleurus Mulsant, 1842: 227 (Aphodius)

languidulus A. Schmidt, 1916: 98 (Aphodius) [RN] A: BEI ES FE GAN HEI JA LIA NC SC SCH SHG SHN SHX TAI
obsoletus C.O. Waterhouse, 1875a: 88 (Aphodius) [HN]

genus Acrossus Mulsant, 1842: 269 type species Scarabaeus luridus Fabricius, 1775
Hemicyclorhinus Seabra, 1909: 22 type species Scarabaeus luridus Fabricius, 1775

angustiarum Balthasar, 1967b: 125 (Aphodius) A: NP SD UP ORR
hingstoni Petrovitz, 1976: 2 (Aphodius)
simlaicus Petrovitz, 1970b: 17 (Aphodius)

atratus C.O. Waterhouse, 1875a: 91 (Aphodius) A: JA
belousovi Frolov, 2001c: 139 (Aphodius) A: SCH
bimaculatus Laxmann, 1770: 593 (Scarabaeus) E: AU BY CR CT CZ EN GE HU LA LT NT PL RO SB SK SL ST SZ
UK A: ES KI KZ TM UZ WS
bipunctatus Lepechin, 1775: 201 (Scarabaeus)
coccinelloides Pallas, 1781: 12 (Scarabaeus)
obscurior Hochhuth, 1873b: 133 (Aphodius)

binaevulus Heyden, 1887b: 303 (Aphodius) A: ES FE NC NP SC SD
diaphanomaculatus Heyden, 1887b: 303 (Aphodius)

bolognai Carpaneto & Piattella, 1989: 183 (Aphodius) E: GG ST A: TR
prometheus Kabakov, 1996: 874 (Aphodius)

carpetanus Graëlls, 1847: 306 (Aphodius) E: SP
delavayi W. Koshantschikov, 1916: 199 (Aphodius) A: YUN
depressus Kugelann, 1792a: 262 (Scarabaeus) E: AB AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG GR
HU IR IT KZ LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK A: AF ES FE IN KI KZ
MG NC TD TR WS XIN NARi
atramentarius Erichson, 1848a: 897 (Aphodius)
biceps Dalla Torre, 1879: 109 (Aphodius)
caminarius Faldermann, 1835a: 251 (Aphodius)
circumlineatus A. Schmidt, 1916: 98 (Aphodius) [RN]
humeralis W. Kolbe, 1911: 11 (Aphodius) [HN]
kolbeanus A. Schmidt, 1916: 97 (Aphodius) [RN]
marginatus Dalla Torre, 1879: 109 (Aphodius) [HN]
nigripes Duftschmid, 1805: 117 (Aphodius)
nigripes Kriechbaumer, 1847: 21 (Aphodius) [HN]
nigripes Zetterstedt, 1828: 182 (Aphodius) [HN]
rufus Dalla Torre, 1879: 109 (Aphodius)
squalidissimus Gistel, 1857b: 591 (Aphodius)

devabhumi Mittal, 1993: 239 (Aphodius) A: UP ORR
eberti Balthasar, 1965a: 108 (Aphodius) A: NP UP XIZ
thibetanus Petrovitz, 1976: 1 (Aphodius)

100

SCARABAEIDAE

emodus Petrovitz, 1976: 4 (Aphodius) A: BT NP SD
formosanus Nomura & Nakane, 1951: 36 (Aphodius) A: TAI
gagatinus Ménétriés, 1832: 182 (Aphodius) E: AR GG ST A: IN KI KZ TM TR
haliki Balthasar, 1933a: 60 (Aphodius) A: SCH SHA
histrio Balthasar, 1932h: 205 (Aphodius) A: SCH
hrubanti Balthasar, 1961c: 366 (Aphodius) A: SCH
humerospinosus Petrovitz, 1958b: 136 (Aphodius) A: SCH
igai Nakane, 1956b: 120 (Aphodius) A: JA
miyakei Nakane, 1960: 3 (Aphodius)
pseudoigai Nakane & Masumoto, 1967: 38 (Aphodius)
sakuraii Nakane & Masumoto, 1967: 39 (Aphodius)
tsurugianus Nakane & Masumoto, 1967: 39 (Aphodius)

japonicus Nomura & Nakane, 1951: 37 (Aphodius) A: JA
jedlickai Balthasar, 1932h: 207 (Aphodius) A: SCH
jubingensis Balthasar, 1967b: 126 (Aphodius) A: NP
klickai Balthasar, 1932h: 204 (Aphodius) A: SCH
koreanensis J.I. Kim, 1986: 9 (Aphodius) A: SC
kwanhsiensis Balthasar, 1945c: 162 (Aphodius) A: SCH
laticollis Baudi di Selve, 1870: 68 (Aphodius) E: FR IT
luridus Fabricius, 1775: 19 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG GR
HU IR IT KO KZ LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: MO A: AF
CY ES FE IN IQ IS KI KZ SY TD TM TR UZ WS XIN
apicalis Mulsant, 1842: 276 (Aphodius)
arator Herbst, 1783: 9 (Scarabaeus)
aterunicolor Hochhuth, 1873b: 134 (Aphodius)
bipaginatus Mulsant, 1842: 277 (Aphodius)
connexus Mulsant, 1842: 275 (Aphodius)
deplanatus Ménétriés, 1832: 181 (Aphodius)
faecis Gistel, 1857b: 575 (Aphodius)
faillae Ragusa, 1883: 296 (Aphodius)
gagates O.F. Müller, 1776: 55 (Scarabaeus)
gagatinus Geoffroy, 1785: 10 (Scarabaeus)
hilleri Schilsky, 1888b: 317 (Aphodius)
humeralis Dalla Torre, 1879: 110 (Aphodius) [HN]
informis Mulsant, 1842: 275 (Aphodius)
interpunctatus Herbst, 1783: 8 (Scarabaeus)
intricarius Mulsant, 1842: 275 (Aphodius)
lateralis Mulsant, 1842: 276 (Aphodius) [HN]
lividibasis Reitter, 1898c: 342 (Aphodius)
lividus Walckenaer, 1802: 12 (Aphodius)
lutarius Fabricius, 1801a: 77 (Aphodius)

SCARABAEIDAE

101

nigripes Fabricius, 1792a: 35 (Scarabaeus)
nigrosulcatus Marsham, 1802: 27 (Scarabaeus)
pellitus A. Schmidt, 1916: 97 (Aphodius) [RN]
rufitarsis Latreille, 1806: 88 (Aphodius)
rufonotatus Dalla Torre, 1879: 110 (Aphodius)
strigosus Dalla Torre, 1879: 110 (Aphodius)
variegatus Herbst, 1783: 9 (Scarabaeus)
varius Gmelin, 1790: 1553 (Scarabaeus)

opacipennis A. Schmidt, 1910c: 356 (Aphodius) A: NP SCH SD
planicollis Reitter, 1890s: 390 (Aphodius) E: AB AR GG ST A: TR
ritsemae A. Schmidt, 1909d: 112 (Aphodius) A: NP SD
rufipes Linnaeus, 1758: 353 (Scarabaeus) E: AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG HU IR
IT KO KZ LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: CI TU A: ES FE JA JIL
KI KZ MG NC NP SC TM TR UZ WS XIN AFR NARi NTRi
boum Gistel, 1857b: 596 (Aphodius)
capicola Harold, 1862b: 381 (Aphodius)
capitatus De Geer, 1774: 263 (Scarabaeus)
juvenilis Mulsant, 1842: 272 (Aphodius)
matsuzawai Yawata, 1943: 7 (Aphodius)
muticus Stephens, 1830: 200 (Aphodius)
oblongus Scopoli, 1763: 8 (Scarabaeus)
rufotestaceus Dalla Torre, 1879: 109 (Aphodius) [HN]

semiopacus Reitter, 1887b: 227 (Aphodius) A: GAN SCH XIN XIZ
luteoirroratus Heyden, 1889c: 657 (Aphodius)

siculus buturensis G. Dellacasa, 1983a: 122 (Aphodius) E: IT (Basilicata, Calabria)
siculus siculus Harold, 1862b: 395 (Aphodius) E: IT (Sicilia)
superatratus Nomura & Nakane, 1951: 36 (Aphodius) A: ES FE JA NC SC
arsenjevi Berlov, 1989b: 394 (Aphodius)

tingitanus Reitter, 1892e: 246 (Aphodius) N: MO
unifasciatus Nomura & Nakane, 1951: 35 (Aphodius) A: JA
hayashii Nakane, 1955: 33 (Aphodius)
kiiensis Nakane, 1960: 3 (Aphodius)

ustulatus Harold, 1862b: 386 (Aphodius) A: HP UP ORR
genus Aganocrossus Reitter, 1895f: 208 type species Aphodius postpilosus Reitter, 1895
pallidicornis Walker, 1858: 207 (Aphodius) A: CH JA NP PA ORR
khuntianus Balthasar, 1940: 38 (Aphodius) [RN]
urostigmoides Balthasar, 1939a: 20 (Aphodius) [HN]

postpilosus Reitter, 1895f: 209 (Aphodius) A: JA NC SC SD SHG TAI ORR
urostigma Harold, 1862a: 170 (Aphodius) A: AF BEI BT FE FUJ HAI HKG HP JIA LIA NC NP PA SC SD SHG SHX
TAI XIZ YUN AFR ORR

102

SCARABAEIDAE

compacticollis Motschulsky, 1858d: 55 (Aphodius)
deplorandus A. Schmidt, 1907a: 202 (Aphodius) [RN]
mixtus Motschulsky, 1858d: 56 (Aphodius) [HN]
piceoniger Motschulsky, 1864: 461 (Aphodius)

vejdovskyi Balthasar, 1945c: 161 (Aphodius) [DA] N: AG
genus Agoliinus A. Schmidt, 1913b: 162 type species Aphodius congregatus Mannerheim, 1853
aleutus Eschscholtz, 1822: 27 (Aphodius) A: ES FE (Kamchatka, Kuril’skie ostrova) JA NAR
ursinus Motschulsky, 1845b: 365 (Aphodius)

amurensis Iablokoff-Khnzorian, 1972: 283 (Aphodius) A: FE
durrelli Rakovič & Mencl, 2011: 309 A: SD
guttatus Eschscholtz, 1823: 97 (Aphodius) A: FE (Kamchatka) NAR
isajevi Kabakov, 1994a: 33 (Aphodius) E: CT ST UK A: WS
kiuchii Masumoto, 1984b: 170 (Aphodius) A: JA
lapponum Gyllenhal, 1806: 72 (Aphodius) E: FA FI GB IC IR NR NT SV A: ES FE KI KZ MG TM WS XIN
anachoreta Bouskell, 1901: 22 (Aphodius) [HN]
axillaris Stephens, 1839: 162 (Aphodius)
humeratus Bouskell, 1901: 22 (Aphodius)
miniatulus Balthasar, 1931c: 218 (Aphodius)
maurus Gebler, 1833: 285 (Aphodius)
nigritus Bouskell, 1901: 22 (Aphodius)
rhenonum Zetterstedt, 1828: 179 (Aphodius)
sobrinus Harold, 1863: 384 (Aphodius)
subalpinus J. Hardy, 1847: 380 (Aphodius)

morii Nakane, 1983b: 140 (Aphodius) [DA] A: JA
nemoralis Erichson, 1848a: 816 (Aphodius) E: AU BE BU BY CT CZ DE EN FI FR GB GE HU IT LA LT NL NR NT
PL RO SK SL ST SV SZ UK
ardoei Landin, 1954: 27 (Aphodius)
ferrugineus Schilsky, 1888b: 309 (Aphodius) [HN]
ferruginus A. Schmidt, 1916: 97 (Aphodius) [RN]

parastorkani Červenka, 1995: 88 (Aphodius) A: YUN
piceus Gyllenhal, 1808: 21 (Aphodius) E: AB AU BE BY CR CT CZ DE EN FI FR GE GG HU IT LA LT NL NR NT
PL RO SB SK SL SP ST SV SZ UK A: ES FE KZ MG WS XIN YUN
alpicola Mulsant, 1842: 192 (Aphodius)
ambulans Gistel, 1857b: 596 (Aphodius)
melanarius Germar, 1824: 110 (Aphodius)
orobius Mulsant, 1842: 191 (Aphodius)

pittinoi Carpaneto, 1986: 146 (Aphodius) E: GR
pseudostorkani Stebnicka, 1982a: 75 (Aphodius) A: SCH
ragusae Reitter, 1892e: 216 (Aphodius) E: IT (Basilicata, Calabria, Sicilia)

SCARABAEIDAE

103

satyrus Reitter, 1892e: 198 (Aphodius) E: AL AU BU CR FR GE GG IT ME SB SL ST SZ
kofleri Petrovitz, 1956: 22 (Aphodius)

setchan Masumoto, 1984b: 171 (Aphodius) A: FE JA
shibatai Nakane, 1983b: 141 (Aphodius) [DA] A: JA
storkani Balthasar, 1932e: 123 (Aphodius) A: SCH XIZ
tanakai Masumoto, 1981: 73 (Aphodius) A: JA
genus Agolius Mulsant & Rey, 1870c: 472 [=1871a: 232] type species Aphodius mixtus A. Villa & G.B. Villa, 1833
(=Aphodius abdominalis Bonelli, 1812)
Calaphodius Reitter, 1892e: 228 type species Aphodius bonvouloiri Harold, 1860

abdominalis abdominalis Bonelli, 1812: 155 (Aphodius) E: AU BH CR FR (Alpes) GE GR IT (Alpi) MC SB SL SZ
bovinus Gistel, 1857b: 586 (Aphodius)
chimarius Gistel, 1857b: 581 (Aphodius)
discus W.L.E. Schmidt, 1840b: 127 (Aphodius) [HN]
commaculatus A. Schmidt, 1916: 96 (Aphodius) [RN]
conjunctus Schilsky, 1888b: 316 (Aphodius)
cyclocephalus Mulsant, 1842: 270 (Aphodius)
mixtus A. Villa & G.B. Villa, 1833: 34 (Aphodius)
oreophilus Gistel, 1857b: 587 (Aphodius)
protectus A. Schmidt, 1916: 99 (Aphodius) [RN]
unicolor Schilsky, 1888b: 316 (Aphodius) [HN]

abdominalis balazuci J.-L. Nicolas, 1971: 154 (Aphodius) E: FR (Massif Central)
abdominalis emilianus G. Dellacasa, 1983a: 136 (Aphodius) E: IT (Appennino Ligure e Tosco-Emiliano)
abdominalis pecoudi Hoffmann, 1954a: 81 (Aphodius) E: AN FR (Hautes-Pyrénées) SP
abdominalis tatrensis M. Dellacasa & G. Dellacasa, 2005: 65 E: CZ PL RO SK UK
bonvouloiri bonvouloiri Harold, 1860: 615 (Aphodius) E: PT SP
bonvouloiri cantabricus Pittino, 1981: 218 (Aphodius) E: SP (Picos de Europa)
genus Agrilinus Mulsant & Rey, 1870c: 419 [=1871a: 179] type species Scarabaeus ater De Geer, 1774
Pseudolimarus Balthasar, 1931c: 217 type species Aphodius jugurtha Balthasar, 1931

ater De Geer, 1774: 270 (Scarabaeus) E: AU BE BH BU BY CR CT CZ DE EN FA FI FR GB GE GG HU IR IT LA LT
LU MC MD ME NL NR NT PL RO SB SK SL SP ST SV SZ UK A: ES FE KI KZ MG TR UZ WS XIN
mediocris A. Schmidt, 1916: 98 (Aphodius) [RN]
obscurus Marsham, 1802: 18 (Scarabaeus) [HN]
terrenus Stephens, 1830: 195 (Aphodius) [HN]
terrestris Fabricius, 1775: 15 (Scarabaeus)

bardus Balthasar, 1946c: 63 (Aphodius) A: ES FE JIL
bollowi Balthasar, 1941a: 86 (Aphodius) A: FUJ
breviusculus Motschulsky, 1866: 170 (Calamosternus) A: FE HEI JA NC NMO SC
nigerrimus C.O. Waterhouse, 1875a: 83 (Aphodius)
nitidulus C.O. Waterhouse, 1875a: 93 (Aphodius) [HN]

104

SCARABAEIDAE

ovalis C.O. Waterhouse, 1875a: 89 (Aphodius)
trituberculatus A. Schmidt, 1907a: 201 (Aphodius) [RN]

constans Duftschmid, 1805: 94 (Aphodius) E: AL AR AU BE BH BU CR CT FR GB GE GG GR IR IT LU MA MC
MD ME NL PT RO SB SL SP ST SZ UK N: CI A: IN LE TM TR
exiguus Mulsant, 1842: 210 (Aphodius)
martialis Mulsant, 1842: 193 (Aphodius)
nitidus Stephens, 1830: 196 (Aphodius)
nomas Kolenati, 1846: 14 (Aphodius)
vernus Mulsant, 1842: 193 (Aphodius)

convexus Erichson, 1848a: 810 (Aphodius) E: AB AR AU BH BU CR CT CZ FR GE GG GR HU IT NL PL RO SB SK
SL SP ST SZ UK N: AG MO TU A: ES FE IN MG TR WS
affinis P.H. Lucas, 1846a: pl. 24 (Aphodius) [HN]
ascendens Reiche, 1863: 75 (Aphodius)
emerichi Reitter, 1892e: 187 (Aphodius)
lucasi Harold, 1859: 224 (Aphodius) [RN]

hasegawai Nomura & Nakane, 1951: 41 (Aphodius) A: JA
aka Nakane, 1960: 4 (Aphodius)
akahane Nakane, 1960: 5 (Aphodius)

ibericus basilicatus A. Fiori, 1907: 118 (Aphodius) E: GR IT (Basilicata, Sicilia) A: TR
scheuerni Ádám, 1979: 5 (Aphodius)

ibericus ibericus Harold, 1874a: 203 (Aphodius) E: PT SP N: AG MO TU
jugurtha Balthasar, 1931c: 217 (Aphodius)

ishidai Masumoto & Kiuchi, 1987: 45 (Aphodius) A: JA
lindbergi Petrovitz, 1959a: 107 (Aphodius) A: AF
lungaiensis Petrovitz, 1962a: 121 (Aphodius) A: SCH SHA
madara Nakane, 1960: 4 (Aphodius) A: JA
monicae Stebnicka, 1982b: 338 (Aphodius) [DA] A: NP
monikae Král, 2013: 482 A: GAN QIN SCH
montisamator Balthasar, 1965a: 112 (Aphodius) A: NP
obliviosus Reitter, 1892e: 199 (Aphodius) A: QIN MG NMO XIN
virginalis Reitter, 1900d: 157 (Aphodius)

pseudolungaiensis Mencl & Rakovič, 2012: 221 A: SHA
punctator Reitter, 1892e: 196 (Aphodius) A: XIN
ritsukoae Kawai, 2004: 1 (Aphodius) A: JA (Shikoku)
shukronajevi Nikolajev, 1998: 41 (Aphodius) A: KI TD
spinulosus A. Schmidt, 1910c: 358 (Aphodius) A: LIA YUN ZHE
striatus A. Schmidt, 1910c: 359 (Aphodius) A: LIA YUN ZHE
surdus Boucomont, 1929: 785 (Aphodius) A: SCH
tashigaonae Emberson & Stebnicka, 2001: 408 (Aphodius) A: NP
tenax Balthasar, 1932e: 118 (Aphodius) A: FE SCH

SCARABAEIDAE

105

uniformis C.O. Waterhouse, 1875a: 84 (Aphodius) A: FE JA NC SC
etorofuensis Kôno, 1937: 7 (Aphodius)
hayachinensis Nomura & Nakane, 1951: 42 (Aphodius)
maritimus Nomura & Nakane, 1951: 42 (Aphodius)

wassuensis Petrovitz, 1962a: 120 (Aphodius) [DA] A: SCH
genus Ahermodontus Báguena, 1930: 315 type species Ahermodontus marini Báguena, 1930
ambrosi Pardo Alcaide, 1936: 392 (Aphodius) E: SP N: MO
bischoffi Všetečka, 1939a: 31 (Aphodius) E: AL
marini Báguena, 1930: 315 E: SP N: MO
genus Alocoderus A. Schmidt, 1913b: 127 type species Aphodius semenowi Reitter, 1887
carinifrons Reitter, 1892e: 214 (Aphodius) N: MO
digitalis D. Koshantschikov, 1894b: 102 (Aphodius) [RN] E: ST A: KZ TM UZ
digitatus Reitter, 1892a: 225 [=1893a: 104] (Aphodius)

elongatulus Fabricius, 1801a: 68 (Aphodius) A: HKG NP SD TAI UP ORR NARi
chinensis Harold, 1861: 105 (Aphodius)
testaceus Germar, 1813: 118 (Aphodius)
tuberifrons Fairmaire, 1889a: 13 (Aphodius)

holdereri Reitter, 1900d: 155 (Aphodius) A: NP PA QIN XIN XIZ
hydrochaeris Fabricius, 1798: 23 (Scarabaeus) E: AB AR AU BE CR CT FR GE GG GR HU IT KZ MA MD NL PL
PT RO SB SL SP ST SZ UK N: AG CI EG LB MO MR TU A: CY IN IQ IS LE SI SY TR WS
coloratus Mulsant, 1842: 218 (Aphodius)
discicollis Mulsant, 1842: 218 (Aphodius)
germanus Mulsant, 1842: 218 (Aphodius)
meridionalis A. Villa & G.B. Villa, 1835b: 48 (Aphodius)
tataricus Harold, 1866: 114 (Aphodius)

machatschkei Balthasar, 1960a: 3 (Aphodius) A: IQ SY
mineti Clément, 1981: 3 (Aphodius) N: MO
semenowi Reitter, 1887b: 220 (Aphodius) A: XIN XIZ
strigimargo Reitter, 1892e: 238 (Aphodius) E: ST A: IN KI KZ TM UZ XIN
fronticornis D. Koshantschikov, 1894c: 114 (Aphodius)

teyrovskyi Balthasar, 1935e: 255 (Aphodius) A: AF NP PA SD UP ORR
milvus Balthasar, 1956b: 416 (Aphodius)
pedongensis R. Paulian, 1936a: 65 (Aphodius)

turbatus Baudi di Selve, 1870: 66 (Aphodius) E: AB TR N: EG LB A: CY IN IQ IS JO LE SY TR
antilibanonicus Balthasar, 1961c: 366 (Aphodius)
lineimargo Reitter, 1892a: 227 [=1893a: 106] (Aphodius)

volubilis Balthasar, 1961c: 364 (Aphodius) A: AF
wolfgangi Stebnicka, 1990: 7 (Aphodius) A: NP

106

SCARABAEIDAE

genus Amidorus Mulsant & Rey, 1870c: 488 [=1871a: 248] type species Aphodius sericatus W.L.E. Schmidt,
1840 (=Scarabaeus obscurus Fabricius, 1792)
alagoezi Olsoufieff, 1918: 63 (Aphodius) E: AR GG ST A: IN
monticustos Balthasar, 1946c: 58 (Aphodius)
rollandi Kalashian & Lumaret, 2000: 497 (Aphodius)

cribrarius Brullé, 1832: 171 (Aphodius) E: AB AL BH BU CR GG GR MC SB ST A: CY IS LE SY TR
dalmatinus W.L.E. Schmidt, 1840b: 130 (Aphodius)
obscurus Kiesenwetter, 1859c: 188 (Aphodius)
purpuripennis Reitter, 1892e: 217 (Aphodius)
tarsensis Petrovitz, 1967b: 329 (Aphodius)

cribricollis P.H. Lucas, 1846c: pl. 23 (Aphodius) N: AG MO TU
barbaroides A. Schmidt, 1910b: 52 (Aphodius) [RN]
oranicus Balthasar, 1961c: 360 (Aphodius)

immaturus Mulsant, 1842: 263 (Aphodius) E: AU FR GE IT SZ
germandi J.-L. Nicolas & Riboulet, 1967: 114 (Aphodius)

moraguesi Baraud, 1978: 53 (Aphodius) N: MO TU
obscurus latinus G. Dellacasa, 1983a: 163 (Aphodius) E: IT (Appennino Centrale)
obscurus obscurus Fabricius, 1792a: 25 (Scarabaeus) E: AB AN AR AU BE BH BU CR CT CZ FR GE GG GR IT KO
LA LS LT MC MD ME PL RO SB SL SP ST SZ UK A: CY IN IS TR
asiaticus Faldermann, 1835a: 252 (Aphodius)
bubulcus Faldermann, 1835a: 258 (Aphodius)
dichrous Reitter, 1892e: 217 (Aphodius) [HN]
flavipennis L. Miller, 1883: 265 (Aphodius)
hyleus Gistel, 1857b: 595 (Aphodius)
kluchoris Roubal, 1918: 7 (Aphodius)
purpureus A. Schmidt, 1916: 96 (Aphodius) [RN]
sericatus Laporte, 1840: 95 (Aphodius)
sericatus W.L.E. Schmidt, 1840b: 128 (Aphodius)

thermicola Sturm, 1800: 44 (Aphodius) E: AB AR AU BH BU CR CZ FR GE GG GR HU IT MC ME PL RO SB SK
SL SP ST TR UK A: CY SY TR
meridionalis Mulsant, 1842: 265 (Aphodius)
ovium Roubal, 1936a: 330 (Aphodius)

genus Ammoecius Mulsant, 1842: 302 type species Scarabaeus elevatus A.G. Olivier, 1789
Digonorhinus Seabra, 1909: 12 type species Scarabaeus elevatus A.G. Olivier, 1789

amplicollis Peyerimhoff, 1939c: 279 (Aphodius) N: MO
brevis Erichson, 1848a: 907 (Aphodius) E: AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG HU IT KZ LA
LT MC MD ME NL NR NT PL RO SB SK SL SP ST SV SZ UK A: ES KZ WS XIN
dentatus A. Schmidt, 1908d: 504 (Aphodius) E: PT SP N: MO
dogueti Baraud, 1980d: 279 (Aphodius) N: AG
elevatus A.G. Olivier, 1789: no 3: 89 (Scarabaeus) E: FR IT (Liguria) PT SP N: AG MO TU

SCARABAEIDAE
chobauti Báguena, 1927: 179 (Aphodius) [HN]
corellai Balthasar, 1941a: 93 (Aphodius) [RN]
edentulus Mulsant, 1842: 303 (Aphodius)
fusciventris Mulsant, 1842: 303 (Aphodius)
persimilis Schatzmayr, 1946: 62 (Aphodius)

eli Petrovitz, 1961f: 347 (Aphodius) A: IQ
felscheanus Reitter, 1904d: 255 (Aphodius) N: AG MO
franzi Petrovitz, 1964e: 185 (Aphodius) N: MO
frigidus Brisout de Barneville, 1866: 374 (Aphodius) E: FR PT SP N: MO
galaicus Báguena, 1955: 277 (Aphodius)

lusitanicus Erichson, 1848a: 908 (Aphodius) E: PT SP N: MO
meurguesae Clément, 1975: 10 (Aphodius) E: AB A: IN
hyrcanius Petrovitz, 1980: 607 (Aphodius)

muchei Petrovitz, 1962b: 110 (Aphodius) E: GR A: TR
naviauxi Baraud, 1971d: 66 (Aphodius) A: SY
numidicus Mulsant, 1851c: 192 N: AG MO TU
rugifrons Aubé, 1850: 335 (Aphodius) E: IT (Sardegna, Sicilia) N: AG TU
levaillanti Godart, 1852: 297

satanas Carpaneto, 1976: 243 (Aphodius) A: TR
genus Anomius Mulsant & Rey, 1870c: 506 [=1871a: 266] type species Aphodius castaneus Illiger, 1803
annamariae Baraud, 1982a: 85 (Aphodius) E: PT SP
antii Gridelli, 1930: 312 (Aphodius) N: LB
apollonius Petrovitz, 1971b: 168 (Aphodius) N: LB
baeticus Mulsant & Rey, 1870c: 510 [=1871a: 270] (Aphodius) [RN] E: SP N: AG MO TU
badius Mulsant & Rey, 1870b: 214 [=1872c: 190] (Aphodius) [HN]
icosius Petrovitz, 1958a: 148 (Aphodius)
lekefensis Baraud, 1981b: 135 (Aphodius) [RN]
petrovitzi Baraud, 1976c: 401 (Aphodius) [HN]

castaneus Illiger, 1803: 194 (Aphodius) E: FR IT (Sardegna, Sicilia) MA PT SP N: AG MO TU
carbonarius Báguena, 1927: 182 (Aphodius)
gineri Báguena, 1930: 318 (Aphodius)
solieri Mulsant & Rey, 1870b: 212 [=1872c: 188] (Aphodius)

crovettii G. Dellacasa, 1983a: 173 (Aphodius) E: IT (Sardegna)
hamricola Clément, 1928: 100 (Aphodius) N: MO
neidae Petrovitz, 1971b: 170 (Aphodius) N: LB
petrovitzi Clément, 1976: 7 (Aphodius) [RN] N: MO
naviauxi Petrovitz, 1971b: 167 (Aphodius) [HN]
ouirganensis Baraud, 1979b: 47 (Aphodius) [RN]

peyerimhoffi Théry, 1925c: 347 (Aphodius) N: MO
saturnalis Ziani, 2011b: 121 A: IS

107

108
segonzaci Bedel, 1904: 225 (Aphodius) N: AG MO
pardoalcaidei Petrovitz, 1971b: 172 (Aphodius)

theryi Clément, 1962: 41 N: MO
permixtus Clément & Petrovitz, 1971: 168 (Aphodius) [RN]

genus Aparammoecius Petrovitz, 1958c: 138 type species Aphodius isaburoi Nakane, 1956
Paremadus Nakane, 1967: 1 type species Aphodius isaburoi Nakane, 1956

annapurnae Stebnicka, 1986b: 350 (Aphodius) A: NP
bagmatiensis Stebnicka, 1983: 318 (Caelius) A: NP
langtangicus Stebnicka, 1983: 319 (Caelius)

balangensis Petrovitz, 1958c: 139 (Aphodius) A: SCH SHN
chinensis Balthasar, 1945b: 106 (Caelius) A: FUJ
yenpingensis Stebnicka, 1986b: 346 (Aphodius) [RN]

gansuensis Stebnicka, 1997: 76 (Aphodius) A: GAN
heinzi Pittino, 1997: 301 (Aphodius) A: PA
holzschuhi Pittino, 1997: 294 (Aphodius) A: NP
isaburoi Nakane, 1956b: 119 (Aphodius) A: JA
masumotoi Nakane, 1967: 3 (Aphodius)

ivani Stebnicka, 1989: 27 (Aphodius) A: PA
kanglae Stebnicka, 1990: 12 (Aphodius) A: NP
kaskiensis D. Ahrens & Stebnicka, 1997: 6 (Aphodius) A: NP
lamjungi D. Ahrens & Stebnicka, 1997: 5 (Aphodius) A: NP
leisteri S.I. Medvedev, 1968a: 176 (Aphodius) E: GG ST
mahriensis Stebnicka, 1983: 318 (Caelius) A: HP
mizo Nakane, 1967: 2 (Aphodius) A: JA
mudukensis Stebnicka, 1989: 25 (Aphodius) A: PA
muzaffarabadensis Pittino, 1997: 304 (Aphodius) A: PA
nepalensis Petrovitz, 1968c: 37 (Caelius) A: NP
yaralensis Stebnicka, 1986b: 348 (Aphodius) [RN]

pallidiligonis C.O. Waterhouse, 1875a: 87 (Aphodius) A: JA JIL
nipponensis Balthasar, 1956a: 70 (Caelius)

phulcokiensis Stebnicka, 1986b: 347 (Aphodius) A: NP
sabhae Stebnicka, 1990: 9 (Aphodius) A: NP
schawalleri Stebnicka, 1989: 24 (Aphodius) A: KA
shogranicus Pittino, 1997: 299 (Aphodius) A: PA
sulcatus Balthasar, 1953b: 232 (Caelius) A: FUJ
nomurai Stebnicka, 1986b: 346 (Aphodius) [RN]

tarokensis Stebnicka, 1994: 78 (Aphodius) A: TAI
nishikawai Masumoto, 1996: 68 (Aphodius)

topali Pittino, 1997: 297 (Aphodius) A: HP
waniaramensis Pittino, 1997: 305 (Aphodius) A: HP

SCARABAEIDAE

SCARABAEIDAE

109

yangricus Stebnicka, 1983: 321 (Caelius) A: NP
zojilae Stebnicka, 1981a: 326 (Aphodius) A: KA
genus Aphodaulacus W. Koshantschikov, 1911b: 209 type species Aphodius turkestanicus Heyden, 1881
assyricus Petrovitz, 1971d: 218 (Aphodius) A: IN IQ
balachanicola Balthasar, 1973: 51 (Aphodius) A: TM
grumi Frolov, 2002b: 165 (Aphodius) A: QIN
ignobilis Reitter, 1887b: 223 (Aphodius) A: KZ QIN TM UZ XIZ
forsteri Reitter, 1900d: 158 (Aphodius)

kabaki Frolov, 2001c: 140 (Aphodius) A: SCH
kizeritskyi Frolov, 2002b: 163 (Aphodius) E: ST A: TM
koltzei Reitter, 1892e: 229 (Aphodius) A: ES FE JIL
kurenzovi Nikritin, 1969b: 1663 (Aphodius)
medvedevi Nikritin, 1969b: 1662 (Aphodius)

nigrotessellatus Motschulsky, 1866: 170 (Melinopterus) A: BEI ES FE FUJ JA JIL LIA NC NMO SC XIN XIZ
fundator Reitter, 1892e: 230 (Aphodius)
ignobilis Heyden, 1893: 72 (Aphodius) [HN]
obsoleteguttatus C.O. Waterhouse, 1875a: 86 (Aphodius)

perblandus Balthasar, 1933a: 59 (Aphodius) A: SCH
saraevi Nikolajev, 2004d: 98 (Aphodius) A: KZ
turkestanicus Heyden, 1881: 323 (Aphodius) A: AF KI KZ TM UZ XIN
bolassogloi König, 1889: 304 (Aphodius)
stomachosus Balthasar, 1970: 248 (Aphodius)

variabilis C.O. Waterhouse, 1875a: 90 (Aphodius) [DA] A: ES FE JA LIA NC NMO SC
genus Aphodiellus A. Schmidt, 1913b: 140 type species Aphodius impunctatus C.O. Waterhouse, 1875
impunctatus C.O. Waterhouse, 1875a: 85 (Aphodius) A: BEI ES FE HEI JIL JA NC SC SHX WS
laevigatus Harold, 1886b: 290 (Aphodius)

genus Aphodius Hellwig, 1798: 101 type species Scarabaeus fimetarius Linnaeus, 1758
Cytoderhinus Seabra, 1909: 15 type species Scarabaeus fimetarius Linnaeus, 1758
Platycephalus Cuvier, 1797: 517 [HN] type species Scarabaeus fimetarius Linnaeus, 1758

calichromus Balthasar, 1932e: 115 A: GUA FUJ SCH YUN
coniugatus Panzer, 1795b: 364 (Scarabaeus) E: AB AR AU BH CR CT FR GE GG GR HU IT MD RO SB SK SP ST
SZ UK A: IN TM TR
angustifasciatus Dalla Torre, 1879: 107
interruptus Dalla Torre, 1879: 107 [HN]

corallifer W. Koshantschikov, 1913b: 262 A: ZHE
elegans Allibert, 1847: 18 A: CHQ FUJ GAN GUA GUI HUB JIA JIX NC NMO QIN SC SCH SHG SHN TAI XIN XIZ
YUN ZHE ORR
expletus A. Schmidt, 1909b: 20

110

SCARABAEIDAE

fasciger Harold, 1881: 89 A: BT GUI NP QIN SCH SD SHN YUN XIN XIZ ORR
infestus A. Schmidt, 1907b: 570
schenklingi A. Schmidt, 1907a: 202
sexsignatus A. Schmidt, 1907b: 570

fimetarius Linnaeus, 1758: 348 (Scarabaeus) E: AU BE BH BU CR CZ DE FR GB GE GG GR HU IR IT KO MA ME
NL PL PT RO SP SZ TR N: AG EG LB MO MR TU A: CY IQ IS LE SY TR AURi NARi NTRi
bicolorellus A. Schmidt, 1922: 273 [RN]
bicolor Geoffroy, 1785: 9 (Scarabaeus)
bicolor Mulsant, 1842: 187 [HN]
bicoloratus A. Schmidt, 1916: 96 [HN]
cardinalis Reitter, 1892e: 186
castaneus Bouskell, 1901: 16
fimetivorus Gistel, 1857b: 588
hypopygialis Mulsant, 1842: 187
maculipennis Mulsant, 1842: 187
monticola Heer, 1841: 513
orophilus Charpentrier, 1825: 210
punctulatus Mulsant, 1842: 187
subluteus Mulsant, 1842: 187

foetidus Herbst, 1783: 7 (Scarabaeus) E: AL AN AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR IT
LA LT MC ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO TU A: CY IS LE SY TR WS
argillicolor Mulsant, 1842: 180
conflagratus Herbst, 1783: 5 (Scarabaeus)
coprinus Marsham, 1802: 12 (Scarabaeus)
dichrous W.L.E. Schmidt, 1840b: 134
ferrugineus Baudetlafarge, 1809: 66 [HN]
fuscus Dalla Torre, 1879: 107
kerkiranus Balthasar, 1932e: 120
lineatomaculatus Balthasar, 1932e: 120
nigricans Mulsant, 1842: 179
pallipes Mulsant, 1842: 180
panzeri Gistel, 1857b: 591 [RN]
testaceus Dalla Torre, 1879: 107

guangdongensis Maté, 2008: 66 A: FUJ GUA ZHE
irregularis Westwood, 1839b: lv A: HP KA NP PA SD UP XIZ ORR
krupkai Tesař, 1969b: 62

pedellus De Geer, 1774: 266 (Scarabaeus) E: AB AL AN AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG
GR HU IR IT KO KZ LA LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG EG MO
A: AF ES FE FUJ HP IN KA KI KZ MG NP PA TD TM TR UZ WS XIN XIZ ORR NARi NTRi
autumnalis Naezen, 1792: 167 (Scarabaeus)

SCARABAEIDAE

111

imperfectus Mulsant, 1842: 187
nodifrons Randall, 1838: 20

plasoni Käufel, 1914: 142 A: FE JA LIA
reginae Král, 1997c: 212 A: YUN
swaneticus Reitter, 1892e: 186 E: GG ST
genus Aphodobius Péringuey, 1901: 420 type species Aphodius inornatus A. Schmidt, 1908
Esymoides Balthasar, 1945a: 40 type species Aphodius amplipennis Balthasar, 1945

amplipennis Balthasar, 1945a: 41 (Aphodius) A: SCH
genus Apsteiniella A. Schmidt, 1916: 99 [RN] type species Jacobsonia georgii W. Koshantschikov, 1912
Jacobsonia W. Koshantschikov, 1912: 511 [HN] type species Jacobsonia georgii W. Koshantschikov, 1912

aralica Nikolajev, 1979a: 39 (Aphodius) A: KZ UZ
georgii W. Koshantschikov, 1912: 511 (Jacobsonia) A: TM
naviauxi Baraud, 1977b: 53 A: AE IQ QA SA
genus Biralus Mulsant & Rey, 1870c: 467 [=1871a: 227] type species Scarabaeus satellitius Herbst, 1789
equinus Faldermann, 1835a: 257 (Aphodius) E: AR A: IN IQ IS JO KI TR SY
mahunkaorum Ádám, 1983: 25 E: FR IT PT SP N: AG MO TU
menetriesi Ménétriés, 1849: 43 (Aphodius) E: AB CT ST KZ A: AF IN KI KZ TD TM TR UZ
satellitius Herbst, 1789: 289 (Scarabaeus) E: AB AR AU AZ BE BH BU CR CT FR GE GG GR HU KZ LU MD ME
NL PL RO SB SK SL ST SZ TR UK A: CY IN IQ IS KZ LE SY TM TR
affinis Brahm, 1790: 66 (Scarabaeus)
decipiens Schrank, 1798: 382 (Scarabaeus)
invisibilis Dalla Torre, 1879: 109 (Aphodius)
pecari Fabricius, 1792a: 38 (Scarabaeus)
planus W.L.E. Schmidt, 1840b: 170 (Aphodius)

genus Bodiloides M. Dellacasa & G. Dellacasa, 2005: 61 type species Scarabaeus ictericus Laicharting, 1781
asiricus Pittino, 1984b: 297 (Aphodius) A: IN SA
ictericus ghardimaouensis Balthasar, 1929b: 105 (Aphodius) E: CR FR GR IT MA PT SP TR N: CI AG EG LB MO
MR TU A: CY IN IS JO LE SA SY TR
lychnobius Petrovitz, 1954: 268 (Aphodius)

ictericus ictericus Laicharting, 1781: 14 (Scarabaeus) E: AB AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE
GG GR HU IR IT KZ LA LT MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK A: KZ TM TR WS XIN
crimensis W. Koshantschikov, 1913a: 186 (Aphodius)
gilvus W.L.E. Schmidt, 1840b: 136 (Aphodius)
melanescens Bouskell, 1901: 20 (Aphodius)
nitidulus Fabricius, 1792a: 30 (Scarabaeus)
pallescens Bouskell, 1901: 20 (Aphodius)

112

SCARABAEIDAE

genus Bodilopsis Ádám, 1994a: 5 type species Scarabaeus sordidus Fabricius, 1775
aquila A. Schmidt, 1916: 109 (Aphodius) A: FUJ HEB NMO SCH SHN YUN ORR
bella A. Schmidt, 1916: 110 (Aphodius)
pfefferi Balthasar, 1932e: 113 (Aphodius)

ogloblini Semenov & S.I. Medvedev, 1928a: 103 (Aphodius) E: AB ST A: KZ TM UZ
rufa Moll, 1782: 372 (Scarabaeus) E: AB AN AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG GR HU IR
IT KZ LA LT LU MA MC ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK A: CH ES IS KZ NP TR WS
arcuata Moll, 1785: 169 (Scarabaeus)
aurantiaca Mulsant, 1842: 221 (Aphodius)
castana Marsham, 1802: 12 (Scarabaeus)
cohrsi W. Kolbe, 1911: 11 (Aphodius)
foetens A.G. Olivier, 1789: no 3: 85 (Scarabaeus)
hypocophina E. Strand, 1917: 95 (Aphodius) [RN]
hypocyphta W.L.E. Schmidt, 1840b: 139 (Aphodius)
hypocopha Gistel, 1857b: 599 (Aphodius) [HN]
hypocopha Heer, 1841: 523 (Aphodius)
illigeri Dalla Torre, 1879: 108 (Aphodius)
melancholica Bouskell, 1901: 21 (Aphodius)
melanota Mulsant, 1842: 222 (Aphodius)
moscovitica Semenov, 1900a: 91 (Aphodius)
ochracea Stephens, 1830: 191 (Aphodius)
pertinens Gistel, 1857b: 602 (Aphodius)
piceola Semenov, 1900a: 92 (Aphodius)
rufescens Fabricius, 1801a: 74 (Aphodius)
rufifrons Dufour, 1851: 334 (Aphodius)
scropharum Gistel, 1857b: 577 (Aphodius)
unicolor Marsham, 1802: 11 (Scarabaeus)
ultramontana J.R. Sahlberg, 1926: 18 (Aphodius)

sordida changajica Endrődi, 1965: 195 (Aphodius) A: MG
sordida sordida Fabricius, 1775: 16 (Scarabaeus) E: AL AR AU BE BU BY CR CT CZ DE EN FI FR GB GE GG GR
HU IR IT (Alpi) KZ LA LT LU MD NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK A: BEI ES FE HUB JA
KI KZ LIA NC NMO QIN SC TR WS YUN
bipunctatella Mulsant, 1842: 221 (Aphodius)
creutzeri Dalla Torre, 1879: 107 (Aphodius)
equinaria Gistel, 1857b: 577 (Aphodius)
hungarica E. Strand, 1917: 95 (Aphodius) [RN]
limbatella Mulsant, 1842: 221 (Aphodius)
nigromaculata A. Schmidt, 1909b: 20 (Aphodius)
porcella Gistel, 1857b: 579 (Aphodius) [HN]
quadripunctata Panzer, 1789: 4 (Scarabaeus)
signatipennis Mulsant & Wachanru, 1852a: 6 [=1852b: 166] (Aphodius)
simplex Bouskell, 1901: 20 (Aphodius)

SCARABAEIDAE

113

genus Bodilus Mulsant & Rey, 1870c: 518 [=1871a: 278] type species Aphodius lugens Creutzer, 1799
Melaphodius Reitter, 1892e: 238 type species Aphodius limbatus Germar, 1824
Neobodilus Hollande & Thérond, 1999: 149 type species Aphodius lugens Creutzer, 1799
Paramelinopterus Rakovič, 1984: 213 type species Aphodius longipennis Rakovič, 1984
Therilus Mulsant & Rey, 1870b: 207 [=1872c: 175] type species Aphodius orophilus Mulsant & Rey, 1870

apiceopacus Petrovitz, 1959b: 113 (Aphodius) N: AG MO
ardescens A. Schmidt, 1916: 106 (Aphodius) A: PA
arsissaensis Petrovitz, 1968g: 465 (Aphodius) A: IN TR
barbarus Fairmaire, 1860a: 171 (Aphodius) E: SP N: AG MO TU
irritans Reitter, 1892e: 239 (Aphodius)

beduinus Reitter, 1892e: 193 (Aphodius) E: IT (Basilicata, Calabria, Puglia, Sardegna, Sicilia) MA SP N: AG LB
MO TU
circumcinctus W.L.E. Schmidt, 1840b: 143 (Aphodius) E: AU BU CR CT HU KZ MC MD RO SB SK ST UK A: KI
KZ TD TM UZ WS
limbatus Germar, 1824: 112 (Aphodius) [HN]

crassus A. Schmidt, 1916: 107 (Aphodius) A: MG NMO WS
furvus A. Schmidt, 1916: 108 (Aphodius) A: NP PA ORR
rawalpindianus Balthasar, 1960a: 1 (Aphodius)

gregarius Harold, 1871b: 112 (Aphodius) E: AB CT KZ ST UK A: ES IN KI KZ MG PA TD TM UZ WS XIN ORR
kaeufeli Roubal, 1915: 64 (Aphodius)
lineatosetosus Benderitter, 1911: 88 (Aphodius)
maculicollis Ballion, 1871a: 333 (Aphodius) [HN]
rudii Endrődi, 1968b: 71 (Aphodius)

hastatus Reitter, 1892a: 227 [=1893a: 106] (Aphodius) A: GAN KZ MG QIN TM UZ
insperatus Petrovitz, 1967e: 480 (Aphodius) A: IN PA
kaznakovi Frolov, 2001e: 444 (Aphodius) A: QIN XIN
kermanschahensis Petrovitz, 1959b: 114 (Aphodius) A: IN TR
liesenfeldti Petrovitz, 1958c: 132 (Aphodius) A: PA SD UP ORR
pakistanus Balthasar, 1961c: 369 (Aphodius)

longeciliatus Reitter, 1887b: 212 (Aphodius) A: IN JIA KZ MG NMO WS XIN XIZ
longipennis Rakovič, 1984: 214 (Aphodius) A: AF KI KZ
inylchekensis Frolov, 2001b: 92 (Aphodius)

longispina Küster, 1854: 33 (Aphodius) E: SP N: AG EG LB MO TU A: SI
lugens Creutzer, 1799b: 59 (Aphodius) E: AB AL AR AU BE BH BU BY CR CT CZ EN FR GE GG GR HU IT LA LT
LU MA MC MD ME NL NT PL PT RO SB SK SL SP ST SZ TR UK N: AG EG MO TU A: AF CY IN IQ IS JO KI
KZ SI SY TD TM TR UZ XIN
emarginalis Mulsant, 1842: 225 (Aphodius)
faldermanni Sperk, 1835: 157 (Aphodius)
indecorus Mulsant, 1842: 225 (Aphodius)

marani Balthasar, 1929b: 106 (Aphodius) [DA] N: TU
nigriventris Reitter, 1892e: 194 (Aphodius) A: XIN
noxius A. Schmidt, 1916: 98 (Aphodius) [RN] E: ST

114

SCARABAEIDAE
orophilus Mulsant & Rey, 1870b: 206 [=1872c: 174] (Aphodius) [HN]

punctipennis Erichson, 1848a: 821 (Aphodius) E: AB AR CT GG GR HU RO SB ST UK N: EG A: AF CY IN IQ IS
KI KZ LE SI SY TM TR UZ WS XIN
incertus Ballion, 1878: 282 (Aphodius)
intermedius Ballion, 1871a: 332 (Aphodius)

sabaeus Peyerimhoff, 1907: 45 (Aphodius) A: IS JO SI
saoudi R. Paulian, 1980: 144 (Aphodius) A: SA
sordescens Harold, 1869f: 431 (Aphodius) A: ES FE KZ MG NIN NMO WS XIZ
delirus Balthasar, 1965c: 448 (Aphodius)
pallididorsis Reitter, 1887b: 224 (Aphodius)

terminassianae Kolov & Nikolajev, 2011: 127 (Aphodius) A: KZ
vartiani Petrovitz, 1965d: 677 (Aphodius) A: IQ SY
veselyi Tesař, 1969b: 61 (Aphodius) A: PA
zarudnyi Frolov, 2001b: 91 (Aphodius) A: IN
zunbajanicus Stebnicka, 1975: 185 (Aphodius) A: MG
genus Boucomontiellus Balthasar, 1932c: 36 type species Aphodius pelliculosus Balthasar, 1932
pelliculosus Balthasar, 1932c: 37 (Aphodius) A: SCH
umbrinus W. Koshantschikov, 1913b: 264 (Aphodius) A: JIL
genus Brachiaphodius W. Koshantschikov, 1913a: 200 type species Aphodius pilosus Harold, 1874
Balthasarianus R. Paulian, 1934a: 111 type species Aphodius pilifer R. Paulian, 1934

ahrensi Král, 2000: 67 (Aphodius) A: NP
babori Balthasar, 1938a: 8 (Aphodius) A: SCH
eccoptus Bates, 1889: 297 (Aphodius) A: JA
koichianus Matsumura, 1934: 65 (Aphodius)

jendeki Král, 2000: 72 (Aphodius) A: YUN ORR
nainiensis Petrovitz, 1963d: 631 (Aphodius) A: NP UP ORR
pilifer R. Paulian, 1934a: 111 (Aphodius) A: NP SD ORR
fruhstorferi Petrovitz, 1970b: 19 (Aphodius)

pilosus Harold, 1874a: 190 (Aphodius) A: BT NP SD UP
abditus Petrovitz, 1970b: 18 (Aphodius)
gregori Balthasar, 1941c: 176 (Aphodius)

ruubpai Masumoto, 1991a: 31 (Aphodius) A: YUN ORR
taiwanicus Petrovitz, 1976: 6 (Aphodius) A: TAI
weibao Král, 2000: 84 (Aphodius) A: YUN
yuan Král, 2000: 87 (Aphodius) A: YUN ORR
genus Calamosternus Motschulsky, 1860f: 156 type species Scarabaeus granarius Linnaeus, 1767
Megalisus Mulsant & Rey, 1870b: 209 [=1872c: 185] type species Aphodius stercorarius Mulsant & Rey, 1870

algiricus Mariani & Pittino, 1983: 153 (Aphodius) E: IT FR (Corse) MA PT SP N: AG EG LB MO TU AFR

SCARABAEIDAE

115

perezi Harold, 1870a: 113 (Aphodius)

boreosinicus Červenka, 1994a: 14 (Aphodius) A: GAN QIN SCH
granarius Linnaeus, 1767: 547 (Scarabaeus) E: AB AL AN AR AU AZ BE BH BU BY CR CT CZ DE EN FI FR GB
GE GG GR HU IR IT KZ LA LS LT LU MA MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N:
AG CI EG LB MO MR TU A: AF CY ES IN IQ IS KA KI KZ LE SA SI SY TD TM TR UZ WS XIN COS
adelaidae Hope, 1846a: 146 (Aphodius)
apicalis Ragusa, 1883: 290 (Aphodius) [HN]
ater Thunberg, 1818: 400 (Aphodius) [HN]
aterrimus Melsheimer, 1844: 136 (Aphodius)
basilaris Dalla Torre, 1879: 107 (Aphodius)
brunnescens Reitter, 1892e: 189 (Aphodius)
carbonarius Sturm, 1805: 128 (Aphodius)
concolor Mulsant, 1842: 199 (Aphodius)
cribratus Mulsant, 1842: 199 (Aphodius)
elongatus Ménétriés, 1832: 182 (Aphodius)
emarginatus Stephens, 1830: 198 (Aphodius)
fusculus Bouskell, 1901: 19 (Aphodius)
haemorrhous Stephens, 1830: 196 (Aphodius)
hypocrita Mulsant & Rey, 1870c: 449 [=1871a: 209] (Aphodius)
inutilis Horn, 1887: 50 (Aphodius)
lucens Stephens, 1830: 196 (Aphodius)
melanopus Stephens, 1830: 198 (Aphodius)
metallicus Haldeman, 1848: 105 (Aphodius)
moestus Mulsant, 1842: 199 (Aphodius) [HN]
nama H.J. Kolbe, 1908b: 126 (Aphodius)
niger Creutzer, 1799b: 20 (Aphodius)
parcepunctatus Mulsant, 1842: 199 (Aphodius)
pecirkai Balthasar, 1932c: 40 (Aphodius)
pirinensis Balthasar, 1946c: 62 (Aphodius)
quadrituberculatus Fabricius, 1798: 23 (Scarabaeus)
retusus Waltl, 1835: 67 (Aphodius)
rugosulus Mulsant, 1842: 199 (Aphodius)
signatus A. Schmidt, 1916: 99 (Aphodius) [RN]
spretus Haldeman, 1848: 105 (Aphodius)
suturalis Faldermann, 1835a: 259 (Aphodius)
thoracicus Dalla Torre, 1879: 107 (Aphodius) [HN]
tilgranicus Nakane, 1966: 232 (Aphodius)
tomskianus Balthasar, 1973: 55 (Aphodius)
vagus Marseul, 1878: 56 (Aphodius)

hyxos Petrovitz, 1962a: 123 (Aphodius) E: GR N: EG A: CY IN IQ IS JO LE SI SY TR
machulkai Balthasar, 1935c: 122 (Aphodius) A: AF IN IQ

116

SCARABAEIDAE

mayeri Pilleri, 1953: 110 (Aphodius) E: CR IT FR GR MA PT SP N: AG EG LB MO TU AFR
maghribinicus Petrovitz, 1969a: 82 (Aphodius)

prusai Tesař, 1945: 68 (Aphodius) A: SC
pseudolucidus Rakovič, 1977a: 67 (Aphodius) [DA] A: UZ
sauteri Petrovitz, 1961c: 113 (Aphodius) A: TAI
spinosus D. Koshantschikov, 1894c: 107 (Aphodius) A: AF IQ TM TR UZ
acriculus Balthasar, 1938c: 96 (Aphodius)

srinagarensis Pittino, 2001: 143 (Aphodius) A: KA
stercorarius Mulsant & Rey, 1870b: 208 [=1872c: 176] (Aphodius) [DA] A: IQ
tricornifrons Reitter, 1909b: 76 (Aphodius) E: KZ A: KZ TM
trucidatus Harold, 1863: 351 (Aphodius) E: AB AR GR ST A: IN IQ IS KI KZ LE MG SY TD TM TR UZ WS
humeridens Reitter, 1898c: 341 (Aphodius)
suturifer Reitter, 1892e: 189 (Aphodius)

unicolor A.G. Olivier, 1789: no 3: 183 (Scarabaeus) E: FR MA PT SP N: AG EG MO TU A: IN
ferrugineus Mulsant, 1842: 233 (Aphodius)

uniplagiatus C.O. Waterhouse, 1875a: 84 (Aphodius) A: FUJ JA LIA NC SC SHG TAI YUN ZHE ORR
desuetus Balthasar, 1933c: 145 (Aphodius)

vexator Balthasar, 1933c: 145 (Aphodius) N: EG A: IN IQ SA
zmoskev Červenka, 1994a: 13 (Aphodius) A: GAN
genus Carinaulus Tesař, 1945: 66 type species Aphodius vseteckai Tesař, 1945
armandidavidi Červenka, 2000: 41 (Aphodius) A: SCH
coreensis J.I. Kim, 1986: 8 (Aphodius) A: SC
dierli Balthasar, 1967b: 123 (Aphodius) A: NP
dundarum Červenka, 2000: 40 (Aphodius) A: SCH
gansuicus Červenka, 2000: 42 (Aphodius) A: GAN SCH
imitabilis Červenka, 2000: 35 (Aphodius) A: YUN
inexspectatus Balthasar, 1935c: 121 (Aphodius) A: ES FE JA KZ NC SC
nigrocarinatus Nikolajev, 1979c: 99 (Aphodius)

jakli Červenka, 2000: 38 (Aphodius) A: GAN
krali Červenka, 2000: 37 (Aphodius) A: YUN
meghalayensis Červenka, 2000: 30 (Aphodius) A: UP
milaninikodymi Červenka, 2000: 33 (Aphodius) A: QIN
pseudovseteckai Červenka, 2003: 23 (Aphodius) A: SCH
pseudozhanglaensis Červenka, 2003: 24 (Aphodius) A: SCH
pucholti Balthasar, 1961c: 362 (Aphodius) A: FUJ TAI
tryznai Červenka, 2000: 34 (Aphodius) A: XIZ
vseteckai Tesař, 1945: 66 (Aphodius) A: SCH YUN
wrzecionkoi Červenka, 2000: 39 (Aphodius) A: XIZ
zhanglaensis Červenka, 2000: 31 (Aphodius) A: SCH SHA

SCARABAEIDAE

117

genus Chilothorax Motschulsky, 1860f: 156 type species Scarabaeus conspurcatus Linnaeus, 1758
albosetosus Pittino, 1995: 255 (Aphodius) A: TR
alexis Frolov, 2001a: 42 (Aphodius) A: MG
aljibei Blanco Villero, 1986: 388 (Aphodius) [DA] E: SP
altaicus Nikolajev, 1984: 149 (Aphodius) [RN] A: MG QIN XIN
gobiensis Balthasar, 1967b: 128 (Aphodius) [HN]
puntsagdulamae Nikolajev, 1984: 151 (Aphodius)

auliensis Balthasar, 1938d: 215 (Aphodius) A: KI KZ TM
badenkoi Nikolajev, 1987: 115 (Aphodius) E: ST A: KZ TM UZ XIN
baghlanicus Frolov, 1996: 28 (Aphodius) A: AF
bistriga Reitter, 1900d: 157 (Aphodius) A: GAN NMO QIN XIN XIZ
brancoi Baraud, 1981d: 271 (Aphodius) [DA] E: PT SP
cervorum Fairmaire, 1871c: 420 (Aphodius) E: BE FR PT SP
vernicosus Hoffmann, 1928a: 90 (Aphodius)

chandmanicus Frolov, 2001a: 45 (Aphodius) A: MG
clathratus Reitter, 1892e: 223 (Aphodius) E: AB AR ST A: IN KZ TD TM TR UZ
clausula W. Koshantschikov, 1910: 19 (Aphodius) A: KZ TD TM UZ XIN
eremobius Semenov & S.I. Medvedev, 1929a: 86 (Aphodius)
gussakovskii Semenov & S.I. Medvedev, 1929a: 87 (Aphodius)

comma Reitter, 1892e: 227 (Aphodius) E: CT A: ES FE HEI KZ MG NMO UZ WS XIN
incomma Reitter, 1906e: 440 (Aphodius)

conspurcatus Linnaeus, 1758: 348 (Scarabaeus) E: AB AU BE BU BY CR CT CZ DE EN FI FR GB GE HU IR IT LA
LT ME NL NR NT PL PT RO SB SL SP ST SV SZ UK
discedens A. Schmidt, 1907b: 567 (Aphodius) A: IS JO
distinctus distinctus O.F. Müller, 1776: 53 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR
GB GE GG GR HU IT KZ LA LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: LB TU
A: ES IN IS KI KZ MG SY TD TM TR UZ WS NARi
attaminatus Marsham, 1802: 13 (Scarabaeus)
anxius Mulsant, 1842: 245 (Aphodius)
auctus Mulsant, 1842: 246 (Aphodius)
baseolus Mulsant, 1842: 245 (Aphodius)
centrolineatoides E. Strand & Gulbis, 1957: 253 (Aphodius)
centrolineatus Panzer, 1798: 1 (Scarabaeus)
confluens Schilsky, 1888b: 311 (Aphodius)
distinctoides E. Strand & Gulbis, 1957: 254 (Aphodius)
foedatus Marsham, 1802: 14 (Scarabaeus)
fumosus Mulsant, 1842: 245 (Aphodius)
hemicyclus Mulsant, 1842: 245 (Aphodius)
humeralis Báguena, 1927: 182 (Aphodius)
hypocoprus Gistel, 1857b: 587 (Aphodius)

118

SCARABAEIDAE
inquinatulus Reitter, 1898b: 119 (Aphodius)
inquinatus Herbst, 1783: 6 (Scarabaeus)
interruptus Mulsant, 1842: 246 (Aphodius)
libyanus Balthasar, 1961c: 379 (Aphodius)
lopezromeui Báguena, 1927: 183 (Aphodius)
lunatus Mulsant, 1842: 245 (Aphodius)
maculipennis Melsheimer, 1844: 137 (Aphodius)
nubiloides E. Strand & Gulbis, 1957: 254 (Aphodius)
nubilus Panzer, 1798: 3 (Scarabaeus)
ophthalmicus Mulsant, 1842: 245 (Aphodius)
pauper Mulsant, 1842: 245 (Aphodius)
pseudonubilus E. Strand & Gulbis, 1957: 254 (Aphodius)
pulcheroides E. Strand & Gulbis, 1957: 253 (Aphodius)
scutellaris Mulsant, 1842: 245 (Aphodius)
striatulus Eichler, 1922: 67 (Aphodius)
subcinctus Mulsant, 1842: 246 (Aphodius)
subconfluens E. Strand & Gulbis, 1957: 254 (Aphodius)
trifasciatus E. Strand & Gulbis, 1957: 253 (Aphodius)

distinctus sevanicus Rakovič, 1991: 14 (Aphodius) E: AR
dobrovljanskyi W. Koshantschikov, 1913a: 195 (Aphodius) E: AR
dzongensis Petrovitz, 1976: 5 (Aphodius) A: QIN XIZ
equitis W. Koshantschikov, 1912: 519 (Aphodius) A: IQ SY
exclamationis Motschulsky, 1849: 108 (Aphodius) E: PT SP N: AG LB MO TU
exilimanus Kabakov, 1994a: 35 (Aphodius) E: CT
figuratus A. Schmidt, 1906: 410 (Aphodius) A: KI UZ XIN
ambitiosus A. Schmidt, 1916: 116 (Aphodius)

flavimargo Reitter, 1901a: 75 (Aphodius) E: ST A: AF KI KZ TD TM UZ XIN
flammulatus Harold, 1876: 122 (Aphodius) E: AB GG A: IN TM
deplanatus Reitter, 1892e: 223 (Aphodius)

fritschi Balthasar, 1933c: 144 (Aphodius) A: IQ
discrepitus Balthasar, 1933c: 144 (Aphodius)

glebi Frolov, 2006: 284 (Aphodius) A: MG
grafi Reitter, 1901a: 74 (Aphodius) A: ES FE MG WS
boreomongolicus Nikolajev, 1984: 155 (Aphodius)
grafianus Reitter, 1901a: 74 (Aphodius)
heinrichi Reitter, 1901a: 74 (Aphodius)
kryzhanovskii Nikritin, 1969b: 1661 (Aphodius)
schutovae Nikritin, 1969b: 1664 (Aphodius)

hahni Reitter, 1907d: 410 (Aphodius) [DA] E: CT KZ ST A: KZ RU (border with the North-West of KZ)
hieroglyphicus Klug, 1845: [6], pl. xlii (Aphodius) N: AG CI EG LB MO TU A: IN IS SY
maculosus Wollaston, 1864: 189 (Aphodius)

SCARABAEIDAE

119

magicus Fairmaire, 1875a: 507 (Aphodius)

hozuensis Ochi, Tsukamoto & Kawahara, 2015: 117 A: XIN
huangyuanensis Červenka, 1994a: 15 (Aphodius) A: QIN
hucklesbyi R. Paulian, 1942b: 60 (Aphodius) A: IS
ivanovi Lebedev, 1912: 348 (Aphodius) [DA] E: CT ST UK A: WS
jacobsoni W. Koshantschikov, 1911a: 20 (Aphodius) A: AF ES KZ MG NMO SHG TM XIN XIZ
alexidis W. Koshantschikov, 1911a: 20 (Aphodius)
georgii W. Koshantschikov, 1911a: 21 (Aphodius)
praenubilus Balthasar, 1933c: 143 (Aphodius)
stichai Balthasar, 1933c: 140 (Aphodius)

kandaharicus Balthasar, 1961c: 375 (Aphodius) A: AF
ulcerosus Balthasar, 1961c: 377 (Aphodius)

kardonensis R. Huang & Du, 1999: 288 (Aphodius) A: XIN
kerzhneri Nikolajev, 1984: 159 (Aphodius) A: ES KZ MG WS
kukunorensis Semenov, 1898a: 506 (Aphodius) A: QIN XIZ
adumbratus W. Koshantschikov, 1912: 521 (Aphodius)
petiolatus W. Koshantschikov, 1912: 521 (Aphodius)

lieni Masumoto, Kiuchi & T.-C. Wang, 2014a: 185 (Aphodius) A: TAI
lineolatus Illiger, 1803: 191 (Aphodius) E: AL AR BU CR FR GR IT MA PT SB SP TR N: AG MO TU A: CY IN IQ TR
assimulans Mackauer, 1958: 51 (Aphodius)
conjunctus Mulsant, 1842: 238 (Aphodius)
deletus Mulsant, 1842: 238 (Aphodius)
fuscicollis Mulsant, 1842: 238 (Aphodius)
lateralis Brullé, 1832: 171 (Aphodius)
nigrolineatus Rosenhauer, 1847: 30 (Aphodius)
virgatus A. Schmidt, 1916: 99 (Aphodius) [RN]
vittatus Mulsant, 1842: 238 (Aphodius) [HN]

logunovi Zinchenko, 2003: 191 (Aphodius) A: ES (Altai) WS
longetarsalis Pittino, 1988: 118 (Aphodius) N: AG
melanostictus W.L.E. Schmidt, 1840b: 153 (Aphodius) E: AB AR AU BE BH BU BY CR CT CZ EN FR GE GG GR
HU KZ LA LT LU MC MD ME NL NT PL PT RO SB SK SP ST SV SZ UK N: AG CI EG LB MO TU A: AF ES IN
IS KI KZ LE MG SI SY TD TM TR UZ WS XIN
basalis Báguena, 1927: 182 (Aphodius)
bifasciatus Dalla Torre, 1879: 108 (Aphodius)
biguttatus Dalla Torre, 1879: 108 (Aphodius) [HN]
castilianus J. Daniel, 1903: 251 (Aphodius)
catenatus Mulsant, 1842: 241 (Aphodius)
circumclusus A. Schmidt, 1916: 98 (Aphodius) [RN]
egenus Mulsant, 1842: 241 (Aphodius)
elbursicola Balthasar, 1961c: 377 (Aphodius)
graphicus Kolenati, 1846: 18 (Aphodius)

120

SCARABAEIDAE

lineatus Dalla Torre, 1879: 108 (Aphodius) [HN]
lituratus Rey, 1890a: 164 (Aphodius)
melanostictoides Balthasar, 1961c: 374 (Aphodius)
pillularis Gistel, 1857b: 571 (Aphodius)
septemmaculatus Mulsant, 1842: 241 (Aphodius)
sexmaculatus Mulsant, 1842: 241 (Aphodius)
simulans W. Koshantschikov, 1913a: 190 (Aphodius)
subannulatus Mulsant, 1842: 241 (Aphodius)
tescorum Semenov, 1898a: 508 (Aphodius)
transformis Balthasar, 1965c: 447 (Aphodius)

mongolaltaicus Nikolajev, 1984: 158 (Aphodius) A: KZ MG WS
mossulensis Mulsant & Godart, 1879: 121 (Aphodius) A: IQ
naevuliger Reitter, 1894e: 187 (Aphodius) N: AG MO TU
nigrivittis kongurensis Ochi, Tsukamoto & Kawahara, 2015: 115 A: XIN
nigrivittis nigrivittis Solsky, 1876a: 327 (Aphodius) A: AF KI KZ MG TD TM UZ XIN
impubis W. Koshantschikov, 1913a: 192 (Aphodius)
obliquatus Reitter, 1892e: 226 (Aphodius)
reitterellus W. Koshantschikov, 1911a: 22 (Aphodius)

ohishii Masumoto, 1975: 85 (Aphodius) A: JA
okadai kiritachiensis Ochi, Kawahara & Ohshio, 2012: 4 (Aphodius) A: JA (Kyushu)
okadai okadai Nakane, 1951: 71 (Aphodius) A: JA (Honshū)
pamirensis S.I. Medvedev, 1930: 55 (Aphodius) A: AF KI TD XIN
paykulli Bedel, 1908: 56 [RN] (Aphodius) E: AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG GR HU IR
IT LA LT LU ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: MO TU A: LE SY TR
amplificatus Mulsant, 1842: 252 (Aphodius)
appendiculatus Mulsant, 1842: 252 (Aphodius)
basalis Schilsky, 1888b: 313 (Aphodius)
connexus Mulsant, 1842: 252 (Aphodius)
dilatatus Mulsant, 1842: 252 (Aphodius)
gutheili Schilsky, 1888b: 313 (Aphodius)
intricatus Mulsant, 1842: 253 (Aphodius)
irregularis Mulsant, 1842: 252 (Aphodius)
scutellatus Mulsant, 1842: 252 (Aphodius)
tessulatus Paykull, 1798: 20 (Scarabaeus) [HN]
umbrosus Mulsant, 1842: 253 (Aphodius)

pictus Sturm, 1805: 24 (Aphodius) E: AU BE CR CT CZ DE EN FR GE HU LA LT NL PL RO SB SK SL SP SV SZ UK
N: AG
brumalis Mulsant, 1842: 249 (Aphodius)
flavidulus Mulsant, 1842: 249 (Aphodius)
indigens Mulsant, 1842: 249 (Aphodius)
pollutus Brahm, 1809: 234 (Aphodius)

SCARABAEIDAE
planus D. Koshantschikov, 1894d: 125 (Aphodius) E: KZ ST UK A: KZ TM WS
transvolgensis Semenov, 1898b: 597 (Aphodius)

plutschewskii D. Koshantschikov, 1894d: 123 (Aphodius) E: ST A: KI KZ
potanini Frolov, 2001d: 398 (Aphodius) A: GAN XIZ
propola Balthasar, 1946c: 56 (Aphodius) A: KI TD TM UZ
punctatus C.O. Waterhouse, 1875a: 87 (Aphodius) A: JA FUJ SC
kuatunensis Balthasar, 1961c: 380 (Aphodius)

shansianus Nakane & Shirahata, 1957: 231 (Aphodius) A: SHA
sinicus Červenka, 1994a: 17 (Aphodius) A: QIN
subpolitus Motschulsky, 1860e: 132 (Aphodius) [DA] A: ES
tanhensis Frolov, 2001a: 44 (Aphodius) A: MG WS
tenuimanus Sharp, 1878b: 171 (Aphodius) [DA] A: KA
variicolor D. Koshantschikov, 1894d: 124 (Aphodius) E: AB CT ST A: KI KZ
aralensis Stebnicka, 1978: 257 (Aphodius)

variipennis A. Schmidt, 1916: 99 (Aphodius) [RN] A: ES
variegatus Motschulsky, 1860e: 132 (Aphodius) [HN]

xanthellus Frolov, 2002a: 60 (Aphodius) A: KZ
xanthopterus Balthasar, 1938a: 11 (Aphodius) A: LE SY
zaissanicus Nikolajev, 1987: 114 (Aphodius) A: KZ
genus Chittius Tarasov, 2008: 189 type species Aphodius anatolicus Petrovitz, 1963c
anatolicus Petrovitz, 1963c: 241 (Aphodius) A: TR
genus Cnemargulus Semenov, 1903f: 354 type species Cnemargulus krulikovskyi Semenov, 1903
alkhubraicus Pittino, 1984b: 306 A: SA
krulikovskyi Semenov, 1903f: 354 A: TM
pusio Semenov, 1903f: 355 A: TM UZ
genus Cnemisus Motschulsky, 1868: 63 [RN] type species Cnemargus rufescens Motschulsky, 1845
Ahermes Reitter, 1891g: 254 type species Mendidius rufescens Reitter, 1888
Cnemargus Motschulsky, 1845a: 56 [HN] type species Cnemargus rufescens Motschulsky, 1845

ahngeri Semenov, 1903a: 9 (Ahermes) A: TM UZ
kaznakovi Semenov, 1903b: 27 (Ahermes) A: ES (Buryaria) MG NMO
kaszabi Endrődi, 1964: 425 (Cnemisus)

kozlovi Semenov, 1903b: 27 (Ahermes) A: NMO
rufescens Motschulsky, 1845a: 56 (Cnemargus) E: CT KZ ST A: KI KZ
rufescens Reitter, 1888d: 425 (Mendidius)

genus Colobopterus Mulsant, 1842: 165 type species Scarabaeus erraticus Linnaeus, 1758
brignolii Carpaneto, 1973: 23 (Aphodius) E: AB AR GG ST UK A: TR

121

122

SCARABAEIDAE

erraticus Linnaeus, 1758: 348 (Scarabaeus) E: AB AL AN AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE
GG GR HU IR IT KO KZ LA LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO
TU A: AF CY ES FE GAN IN IQ IS JIL JO KI KZ LE MG NIN NMO PA QIN SC SCH SY TD TM TR UZ WS XIN
XIZ YUN NARi ORR
bistrigatus Dalla Torre, 1879: 106 (Aphodius)
capraram Gistel, 1857b: 582 (Aphodius)
fumigatus Mulsant, 1842: 166 (Aphodius)
gonagricus Fischer von Waldheim, 1844: 45 (Aphodius)
korbeli Roubal, 1935: 245 (Aphodius)
limitinus A. Schmidt, 1922: 21 (Aphodius) [RN]
lineatus Dalla Torre, 1879: 106 (Aphodius) [HN]
melancholicus Dalla Torre, 1879: 106 (Aphodius)
nebulosus Mulsant, 1842: 166 (Aphodius)
pensvallensis Melsheimer, 1844: 135 (Aphodius)
striolatus Gebler, 1832: 50 (Aphodius)
submaculatus Mulsant, 1842: 166 (Aphodius)
taeniatus Schilsky, 1888b: 318 (Aphodius)
unicolor Dalla Torre, 1879: 106 (Aphodius)
zighanicus Eichler, 1922: 66 (Aphodius)

indagator Mannerheim, 1849: 233 (Aphodius) A: ES FE GAN HEI MG NC SC SHX
notabilipennis Petrovitz, 1972b: 31 (Aphodius) A: ES FE
propraetor Balthasar, 1932a: 2 (Aphodius) A: ES FE GAN HEI LIA MG NC SC SCH SHX WS
quadratus Reiche, 1850: 343 (Aphodius) A: FE (Kunashir) HEI JA JIL LIA NC NMO SC SCH
apicalis Harold, 1861: 96 (Aphodius) [HN]
haroldianus Balthasar, 1932a: 1 (Aphodius) [RN]

genus Coprimorphus Mulsant, 1842: 168 type species Scarabaeus scrutator Herbst, 1789
scrutator Herbst, 1789: 161 (Scarabaeus) E: AB AL AN AR AU AZ BE BH BU CR CT CZ FR GB GE GG GR HU IT
LS LT MC MD ME PL PT RO SB SK SL SP ST SZ TR UK N: CI A: IN KI TR
aequinoctialis Harold, 1861: 98 (Aphodius)
angulatus Dalla Torre, 1879: 106 (Aphodius)
angustemarginatus Dalla Torre, 1879: 106 (Aphodius)
brevicornis Schrank, 1789: 62 (Scarabaeus)
brunnipes Mulsant, 1842: 169 (Aphodius)
capybara Gistel, 1857b: 588 (Aphodius)
latemarginatus Dalla Torre, 1879: 106 (Aphodius)
nigricollis Mulsant, 1842: 169 (Aphodius)
rubidus A.G. Olivier, 1789: no 3: 77 (Scarabaeus)
submaculatus Mulsant, 1842: 169 (Aphodius)
tristissimus Dalla Torre, 1879: 106 (Aphodius)

SCARABAEIDAE

123

genus Coptochiroides Balthasar, 1938d: 210 type species Aphodius guentherschmidti Balthasar, 1938
(=Aphodius subcostatus H.J. Kolbe, 1886)
subcostatus subcostatus H.J. Kolbe, 1886: 189 (Aphodius) A: FE JA NC SC TAI YUN ORR
guentherschmidti Balthasar, 1938d: 210 (Aphodius)

genus Emadiellus A. Schmidt, 1913b: 138 type species Aphodius rufopustulatus Wiedemann, 1823
Biraloides Balthasar, 1945a: 41 type species Aphodius pistor Balthasar, 1945

jhelumensis Rakovič, 1991: 9 (Aphodius) [DA] A: PA
rufopustulatus rufopustulatus Wiedemann, 1823: 27 (Aphodius) A: PA ORR
inconsultus Balthasar, 1945a: 42 (Aphodius)

genus Erytus Mulsant & Rey, 1870c: 506 [=1871a: 266] type species Aphodius brunneus Klug, 1845
aequalis A. Schmidt, 1907a: 201 (Aphodius) [RN] E: AB AR BU CT FR (Corse) GG GR IT (Calabria, Puglia,
Sardegna) RO ST UK N: AG LB TU A: AF CY IN IQ IS JO KI KZ MG SA SY TD TM TR UZ XIN XIZ
nitidus Ballion, 1871a: 333 (Aphodius) [HN]
stepicola Balthasar, 1941a: 88 (Aphodius)

bucharicus Petrovitz, 1961c: 108 (Aphodius) A: AF IN PA SA TM UZ
cognatus Fairmaire, 1860a: 172 (Aphodius) E: AB AR GG GR IT (Sicilia) SP ST N: AG CI EG LB MO TU A: AF CY
IN IQ IS JO KI KU KZ LE SY TD TM TR UZ AFR
brunneus Klug, 1845: [6], pl.xlii (Aphodius) [HN]
klugi A. Schmidt, 1910a: 93 (Aphodius) [RN]
unicolor P.H. Lucas, 1847a: 264 (Aphodius) [HN]

hormonzensis Petrovitz, 1980: 610 (Aphodius) A: AE IN IS KU KZ OM SA YE
lindemannae Balthasar, 1960a: 3 (Aphodius) A: IN IQ IS KI SY
mesopotamicus Petrovitz, 1961c: 109 (Aphodius)

opacior D. Koshantschikov, 1894b: 102 (Aphodius) [RN] N: AG EG LB MO TU A: IN IQ IS JO OM SA
irakensis Baraud, 1977b: 55 (Aphodius)
opacus Reitter, 1892e: 191 (Aphodius) [HN]

persicus Petrovitz, 1961c: 105 (Aphodius) E: ST A: AF IN JO KZ PA TR (Konya)
pruinosus Reitter, 1892e: 191 (Aphodius) E: AB ST N: AG EG LB MO TU A: AE AF IN IQ IS JO KI KU KZ LE OM
PA SA TD TM TR UZ YE AFR
buettikeri R. Paulian, 1980: 146 (Aphodius)

psammophilus Balthasar, 1941a: 87 (Aphodius) N: AG AFR
tekkensis Petrovitz, 1961c: 108 (Aphodius) E: ST A: AF IN IQ KI SA TM UZ
transcaspicus Petrovitz, 1961c: 105 (Aphodius) A: IN KZ PA TM UZ
genus Esymus Mulsant & Rey, 1870c: 519 [=1871a: 279] type species Scarabaeus merdarius Fabricius, 1775
Orodalus Mulsant & Rey, 1870c: 439 [=1871a: 199] type species Scarabaeus pusillus Herbst, 1789

alkani Petrovitz, 1963c: 242 (Aphodius) A: TR
filitarsis Reitter, 1898c: 341 (Aphodius) A: LE SY TR

124

SCARABAEIDAE

finitimus A. Schmidt, 1916: 99 (Aphodius) [RN] N: AG MO TU
terminatus Marseul, 1878: 57 (Aphodius) [HN]

helenaeliviae G. Dellacasa & Pittino, 1985: 68 (Aphodius) N: MO
himalayanus W. Koshantschikov, 1916: 201 (Aphodius) [DA] A: KA PA
hladili Chromý, 1993: 5 (Aphodius) A: TM
hyalinus Petrovitz, 1961a: 39 (Aphodius) A: AF
indulgens Balthasar, 1960a: 2 (Aphodius) A: AF PA TM
merdarius Fabricius, 1775: 19 (Scarabaeus) E: AB AL AN AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE
GG GR HU IR IT LA LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO A: IN
IQ IS JO KI TM TR WS
atricollis Mulsant, 1842: 232 (Aphodius)
collaris Dalla Torre, 1879: 109 (Aphodius)
exilis W.L.E. Schmidt, 1840b: 120 (Aphodius)
foriorum Panzer, 1798: 9 (Scarabaeus)
gelbinus Schrank, 1798: 391 (Scarabaeus)
melinopus Mulsant, 1842: 232 (Aphodius)
quisquilius Schrank, 1781: 18 (Scarabaeus)
smetaceki Roubal, 1911b: 134 (Aphodius)

ornatulus Harold, 1866: 121 (Aphodius) A: IN IS LE SY TR
vittipennis J.R. Sahlberg, 1908: 60 (Aphodius)

pokornyi Chromý, 1993: 2 (Aphodius) A: KI
pusillus pusillus Herbst, 1789: 155 (Scarabaeus) E: AB AL AN AR AU BE BH BU BY CR CT CZ DE EN FI FR GB
GE GG GR HU IR IT KO KZ LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK A:
IN IS JA KI KZ MG TD TM TR UZ WS
bicolor Dalla Torre, 1879: 109 (Aphodius)
coecus Mulsant, 1842: 213 (Aphodius)
coeni A. Schmidt, 1922: 259 (Aphodius) [RN]
coenosus A. Ahrens, 1812: 36 (Aphodius) [HN]
dechlamydatus Balthasar, 1932g: 167 (Aphodius)
foenellus Gistel, 1857b: 573 (Aphodius)
granum Gyllenhal, 1808: 19 (Aphodius)
macularis Mulsant, 1842: 213 (Aphodius)
ochripennis Reitter, 1892e: 212 (Aphodius)
parvulus Hochhuth, 1873b: 132 (Aphodius)
phaeopterus Stephens, 1830: 204 (Aphodius)
rufangulus C.O. Waterhouse, 1875a: 89 (Aphodius)
rufulus Mulsant, 1842: 213 (Aphodius)
suturalis Gerhardt, 1909: 422 (Aphodius)
tachypus Gistel, 1857b: 570 (Aphodius)

pusillus roubali Balthasar, 1932g: 167 (Aphodius) A: BEI ES EF GAN JIL LIA NC SC SCH SHG XIN
pyreti Penecke, 1911: 198 (Aphodius) [DA] E: UK

SCARABAEIDAE

125

sesquivittatus Fairmaire, 1883c: xliii (Aphodius) E: SP N: AG LB MO TU
fuentei Báguena, 1930: 320 (Aphodius)
oligopictus Báguena, 1930: 321 (Aphodius)

sicardi Reitter, 1892a: 226 [=1893a: 105] (Aphodius) N: AG MO TU
stribrnyi Tesař, 1945: 71 (Aphodius) A: HEI
suturinigra A. Schmidt, 1916: 99 (Aphodius) [RN] E: AR RO A: CY IN IQ IS LE SY TR
suturalis L. Redtenbacher, 1843: 986 (Aphodius) [HN]

genus Eudolus Mulsant & Rey, 1870c: 467 [=1871a: 227] type species Scarabaeus quadriguttatus Herbst,
1783
quadriguttatus Herbst, 1783: 10 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE FR GE GG GR HU IT
KZ LA LT LU MA MC MD ME NL PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO TU A: AF CH CY ES IN
IQ IS JO KI KZ LE SY TD TM TR UZ WS
angularis Mulsant, 1842: 261 (Aphodius)
astaurus Fuente, 1897: 130 (Aphodius)
cruciatus Mulsant, 1842: 261 (Aphodius)
jaczewskyi Balthasar, 1938a: 10 (Aphodius)
joliveti Baraud, 1980d: 280 (Aphodius)
nigrosuturalis J. Müller, 1902: 447 (Aphodius)
quadripustulatus Duftschmid, 1805: 125 (Aphodius)

quadrinaevulus Reitter, 1892e: 212 (Aphodius) E: AB AR GR TR A: TR
genus Euheptaulacus G. Dellacasa, 1983a: 419 type species Aphodius carinatus Germar, 1824
atlantis Peyerimhoff, 1925b: 106 (Heptaulacus) N: MO (Grand Atlas)
peyerimhoffi R. Paulian & Villiers, 1939: 49 (Heptaulacus)

carinatus brutius Luigioni, 1920: 203 (Heptaulacus) E: IT (Appennino Centrale)
carinatus carinatus Germar, 1824: 111 (Aphodius) E: AB AR AU BE BH BU CR CT CZ FR GE GG GR IT KZ MC
ME NT RO SB SK SL ST SZ UK A: ES FE KZ MG TR WS XIN
alpinus Drapiez, 1819: 49 (Aphodius) [HN]
nivalis Mulsant, 1842: 298 (Heptaulacus)
obenbergeri Balthasar, 1951: 146 (Heptaulacus)

carinatus esuriens G. Dellacasa, 1983a: 423 E: IT (Sicilia)
nemethi Théry, 1925c: 347 (Heptaulacus) N: MO (Moyen Atlas)
porcellus J. Frivaldszky, 1879: 5 (Aphodius) E: AU BH BU CZ HU IT (Liguria, Marche, Abruzzo) MC RO SK SL
sus Herbst, 1783: 9 (Scarabaeus) E: AB AR AU BE BH BU BY CT CR CZ DE EN FI FR GB GE GG GR HU IT KZ
LA LT LU MC MD ME NL NT PL RO SB SK SL SP ST SV SZ UK A: IN KZ SY TM TR WS
austerus Zirk, 1922: 126 (Heptaulacus)
interminis Zirk, 1922: 126 (Heptaulacus)
pubescens A.G. Olivier, 1789: no 3: 91 (Scarabaeus)
quisquilius Schrank, 1798: 391 (Scarabaeus)

126

SCARABAEIDAE

villosus Gyllenhal, 1806: 83 (Aphodius) E: AU BE BH CR CT CZ DE EN FI FR GB GE GR HU IT KO LA LU NL NR
PL RO SB SK SL ST SV SZ
genus Euorodalus G. Dellacasa, 1983a: 260 type species Scarabaeus coenosus Panzer, 1798
boiteli Théry, 1918: 144 (Aphodius) E: SP N: MO TU
baraudi Villarreal, 1975: 309 (Aphodius)

coenosus Panzer, 1798: 7 (Scarabaeus) E: AL AR AU BE BU BY CT CZ DE EN FI FR GB GE LA LT LU MC ME NL
NR NT PL PT RO SB SK SP ST SV SZ UK A: TR
fallax Mulsant, 1842: 209 (Aphodius)
mirandus Mulsant, 1842: 209 (Aphodius)
pellucidus Mulsant, 1842: 209 (Aphodius)
putridicola Gistel, 1857b: 598 (Aphodius)
scapularis Mulsant, 1842: 209 (Aphodius)
tristis Zenker, 1799: 1 (Scarabaeus)
vicinus Mulsant, 1842: 208 (Aphodius)

elephanthinus Petrovitz, 1967b: 326 (Aphodius) [DA] A: TR
longevittatus A. Schmidt, 1916: 98 (Aphodius) [RN] A: IS LE SY
luridipennis J.R. Sahlberg, 1908: 61 (Aphodius) [HN]

lucifer D. Koshantschikov, 1894c: 113 (Aphodius) [DA] A: KZ UZ
paracoenosus Balthasar & Hrubant, 1960: 254 (Aphodius) E: AB AL AR AU BH BU BY CR CT CZ FR GE GG GR
HU IT MC MD ME PL RO SB SK SL SP ST SZ TR UK A: IN TR
signifer Mulsant & Rey, 1870b: 216 [=1872c: 192] (Aphodius) N: EG A: IS LE SY
subtilitarsis Petrovitz, 1963c: 243 (Aphodius) A: TR
tersus Erichson, 1848a: 859 (Aphodius) E: IT MA PT SP N: AG EG LB MO TU
suturalis P.H. Lucas, 1847a: 263 (Aphodius) [HN]
tersus Rosenhauer, 1856: 130 (Aphodius) [HN]

genus Eupleurus Mulsant, 1842: 170 type species Scarabaeus subterraneus Linnaeus, 1758
antiquus Faldermann, 1835b: 367 (Aphodius) A: ES FE JIL MG NMO WS
subterraneus krasnojarskicus G. Dellacasa, 1986: 264 (Aphodius) A: WS
subterraneus subterraneus Linnaeus, 1758: 348 (Scarabaeus) E: AB AR AU BE BH BU BY CR CT CZ DE EN FI
FR GB GE GG GR HU IR IT KZ LA LT LU MD NL NR NT PL RO SB SK SL SP ST SV SZ TR UK N: AG MO A:
AF BEI ES FE IN IQ IS KI KZ LIA MG NC NMO SC TD TM TR UZ WS XIN XIZ NARi
coerulescens Harold, 1861: 103 (Aphodius)
fuscipennis Mulsant, 1842: 171 (Aphodius)
fuscus Dalla Torre, 1879: 106 (Aphodius)
ruber Dalla Torre, 1879: 106 (Aphodius)
rufus Dalla Torre, 1879: 106 (Aphodius)
subcarpathicus Roubal, 1936a: 324 (Aphodius)
sulcatus Fabricius, 1792a: 24 (Scarabaeus) [HN]

SCARABAEIDAE

127

genus Gilletianus Balthasar, 1933c: 139 type species Aphodius proclivis Balthasar, 1933
assamensis Petrovitz, 1976: 8 (Aphodius) A: AP ORR
atsushii Ochi, 1986: 55 (Aphodius) A: JA (Ryukyus)
comatus A. Schmidt, 1920: 140 (Aphodius) A: FE FUJ JA JIL LIA NC SC
chokaiensis Nomura & Nakane, 1951: 40 (Aphodius)

commatoides Balthasar, 1961c: 371 (Aphodius) A: FUJ
costatulus A. Schmidt, 1908b: 47 (Aphodius) A: BT HAI HKG HP NP SD UP TAI ORR
dindigalensis Balthasar, 1941c: 179 (Aphodius)
subpilosus Petrovitz, 1965a: 163 (Aphodius)

fukiensis Balthasar, 1953b: 234 (Aphodius) A: FUJ TAI
kratochvili Balthasar, 1941a: 86 (Aphodius) A: SCH
nigrovirgatus A. Schmidt, 1911a: 49 (Aphodius) [RN] A: NP YUN ORR
nigrosulcatus A. Schmidt, 1909e: 189 (Aphodius) [HN]

proclivis Balthasar, 1933c: 139 (Aphodius) A: NC YUN
rangoonensis Petrovitz, 1970b: 20 (Aphodius) A: BT NP ORR
pokharensis Stebnicka, 1980b: 535 (Aphodius)

reichei Harold, 1859: 210 (Aphodius) A: FUJ GUI SCH TAI AUR ORR
segmentaroides A. Schmidt, 1909a: 11 (Aphodius) A: UP TAI ORR
yunnanensis Král, Rakovič & Mencl, 2015: 312 A: YUN
genus Gonaphodius Reitter, 1892e: 240 type species Aphodius postangulus Reitter, 1887
scoparius Harold, 1877c: 112 (Aphodius) A: ES MG WS XIN XIZ
postangulus Reitter, 1887b: 207 (Aphodius)

genus Grandinaphodius Ziani, 2002: 101 type species Grandinaphodius inferorum Ziani, 2002
inferorum Ziani, 2002: 102 A: LE
smoliki Käufel, 1914: 143 (Aphodius) A: SY
genus Guanyinaphodius Masumoto & Kiuchi, 2001: 119 type species Aphodius kishimotoi Masumoto &
Kiuchi, 2001
kishimotoi Masumoto & Kiuchi, 2001: 120 (Aphodius) A: SCH
genus Heptaulacus Mulsant, 1842: 296 type species Scarabaeus testudinarius Fabricius, 1775
Heptaulaculus G. Dellacasa & Baraud, 1978: 62 type species Scarabaeus testudinarius Fabricius, 1775

algarbiensis Branco & Baraud, 1984: 35 (Heptaulaculus) E: PT SP
brancoi Baraud, 1976b: 133 E: PT SP N: MO
gadetinus Baraud, 1973c: 165 E: SP
koshantschikoffi A. Schmidt, 1911a: 48 A: KI KZ TD
pirazzolii Fairmaire, 1881d: cvl (Aphodius) N: LB TU
rasettii Carpaneto, 1978: 135 E: IT (Toscana)

128

SCARABAEIDAE

syrticola Fairmaire, 1882c: clxxvii (Aphodius) N: AG LB TU
testudinarius Fabricius, 1775: 19 (Scarabaeus) E: AR AU BE BY CR CT CZ DE EN FR GB GE HU IT KZ LA LT LU
MD NL PL PT RO SB SK SL SP ST SV SZ UK A: KZ UZ
genus Koreoxyomus J.I. Kim, 1996: 102 type species Koreoxyomus koreanus J.I. Kim, 1996
koreanus J.I. Kim, 1996: 103 A: SC
genus Koshantschikovius A. Schmidt, 1913b: 149 type species Aphodius substriatus A. Schmidt, 1911
andrewesi A. Schmidt, 1908b: 46 (Aphodius) A: NP UP
chovdgolensis Endrődi, 1983: 114 (Aphodius) A: MG
peculator Balthasar, 1971b: 56 (Aphodius) A: BT NP
bhutanensis Petrovitz, 1975c: 217 (Aphodius)

schusteri Balthasar, 1935a: 73 (Aphodius) A: SA AFR
zinderi R. Paulian, 1942a: 80

genus Labarrus Mulsant & Rey, 1870c: 516 [=1871a: 276] type species Scarabaeus lividus A.G. Olivier, 1789
Pseudocalamosternus Balthasar, 1936a: 1 type species Aphodius rigidus Balthasar, 1936

digitatus Harold, 1871d: 278 (Aphodius) N: EG A: AF IN IQ IS TR UZ
matthiesseni Reitter, 1907d: 407 (Aphodius)

hoabinhensis Balthasar, 1946c: 67 (Aphodius) A: BT NP SHN TAI ORR
insularis Petrovitz, 1961c: 106 (Aphodius)

kasyi Petrovitz, 1965d: 678 (Aphodius) A: AF
lividus A.G. Olivier, 1789: no 3: 86 (Scarabaeus) E: AB AL AR AU AZ BE BH BU BY CR CT CZ EN FI FR GB GE
GG GR HU IT KZ LA LT LU MA MC MD ME NL NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG CI EG LB
MO MR TU A: AE AF BEI CY ES IN IQ IS JA KI KU KZ LE MG NP OM PA QA SA SI SY TR TD TM UP UZ YE
YUN AFR NAR NTR ORR
anachoreta Fabricius, 1801a: 74 (Aphodius) [HN]
bilituratus Marsham, 1802: 15 (Scarabaeus)
discus Wiedemann, 1823: 28 (Aphodius)
fimetifex Gistel, 1857b: 590 (Aphodius)
hircinus Gistel, 1857b: 584 (Aphodius)
innumerabilis A. Schmidt, 1911a: 31 (Aphodius)
limbatus Wiedemann, 1821: 129 (Aphodius)
limicola Panzer, 1798: 6 (Scarabaeus)
maculicollis Montrouzier, 1860: 268 (Aphodius)
matusitai Nakane, 1961: 151 (Aphodius)
obsoletus Fabricius, 1801a: 70 (Aphodius)
scutellaris Roth, 1851: 132 (Aphodius)
sequens Walker, 1858: 207 (Aphodius)
spilopterus Germar, 1848: 189 (Aphodius)

SCARABAEIDAE

129

suturalis Fabricius, 1792a: 28 (Scarabaeus)
vespertinus Creutzer, 1799a: no. 67: 3 (Scarabaeus)

nabeleki Balthasar, 1936a: 2 (Aphodius) A: SHN
nefandus Reitter, 1907d: 408 (Aphodius) A: JIL SCH
praestes Balthasar, 1946c: 66 (Aphodius)

paludani Petrovitz, 1955: 279 (Aphodius) A: AF IN
rigidus Balthasar, 1936a: 1 (Aphodius) A: SHN
rugosocapita Mittal, 1993: 242 (Aphodius) A: UP
splendidus Petrovitz, 1955: 278 (Aphodius) N: EG A: AF IN IS JO KZ OM QA PA SA TD TM YE
sublimbatus Motschulsky, 1860e: 132 (Aphodius) [DA] A: ES FE HEI JA JIL NC NMO SC TAI WS ORR
pallidicinctus C.O. Waterhouse, 1875a: 85 (Aphodius)
ussuriensis Petrovitz, 1961c: 112 (Aphodius)

translucidus Petrovitz, 1961c: 103 (Aphodius) E: ST N: EG A: AE AF IN IQ OM PA SA TM UP
genus Limarus Mulsant & Rey, 1870c: 504 [=1871a: 264] type species Aphodius zenkeri Germar, 1813
hirtipennis P.H. Lucas, 1846c: pl. 20 (Aphodius) N: AG TU
maculatus Sturm, 1800: 42 (Aphodius) E: AB AR AU BE BH BU BY CR CT CZ FR GE GG GR HU LU MC NL PL
RO SB SK SL ST SZ UK A: IN TR
brevifascia A. Schmidt, 1916: 97 (Aphodius) [RN]
fasciatus Dalla Torre, 1879: 108 (Aphodius) [HN]
immaculatus Schilsky, 1888b: 320 (Aphodius)
involutus A. Schmidt, 1916: 98 (Aphodius) [RN]
obsoletus Dalla Torre, 1879: 108 (Aphodius) [HN]
styriacus Grimmer, 1841: 40 (Aphodius)

zenkeri Germar, 1813: 118 (Aphodius) E: AU BE BH BY CR CZ DE FR GB GE IT LU NL PL SK SL ST SV SZ UK
ceresai Schatzmayr, 1941: 86 (Aphodius)
marchicus Scholz, 1935: 172 (Aphodius)
rupestris Gistel, 1857b: 592 (Aphodius)

genus Lindbergianus Petrovitz, 1961a: 40 type species Aphodius solitarius Petrovitz, 1961
solitarius Petrovitz, 1961a: 41 (Aphodius) A: AF
pseudatricolor Nikritin & Kabakov, 1979: 760 (Aphodius)

genus Liothorax Motschulsky, 1860f: 156 type species Scarabaeus plagiatus Linnaeus, 1767
Ataeniomorphus Balthasar, 1953a: 22 type species Aphodius isikdagensis Balthasar, 1953

isikdagensis Balthasar, 1953a: 22 (Aphodius) E: GR A: CY IN IS TR
bytinskisalzi Petrovitz, 1971d: 219 (Aphodius)
cypricola Balthasar, 1971b: 57 (Aphodius)
ressli Petrovitz, 1962a: 126 (Aphodius)

kraatzi Harold, 1868a: 84 (Aphodius) E: AB AL AR AU BH BU CR CT GG GR HU KZ MD ME RO SB SK ST UK
A: AF IN KA KZ TD TM TR UZ WS

130

SCARABAEIDAE

niger Illiger, 1798: 24 (Aphodius) E: AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG HU IT LA LT NL NR
NT PL PT RO SB SK SL SP ST SV SZ TR UK A: IN KI KZ TM TR WS XIN XIZ
muscorum Ádám, 1994a: 6 (Aphodius)
wilsonae Maté & Angus, 2005: 229 (Aphodius)

plagiatus Linnaeus, 1767: 559 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR
HU IR KZ LA LT LU MA MC NL NR NT PL PT RO SB SK SP ST SV SZ TR UK N: TU A: AF BEI ES FE GUI IQ
IS JO KI KZ LE MG SCH SY TD TM TR UZ WS YUN
concolor Schilsky, 1888b: 311 (Aphodius)
discoides A. Schmidt, 1916: 96 (Aphodius) [RN]
discus Reitter, 1892e: 204 (Aphodius) [HN]
elongatus Stephens, 1830: 209 (Aphodius)
hungaricus Endrődi, 1955a: 48 (Aphodius)
immaculatus Dalla Torre, 1879: 108 (Aphodius)
jakutorum Balthasar, 1938d: 213 (Aphodius)
longulus Ménétriés, 1849: 44 (Aphodius)
parvomaculatus G. Schmidt, 1938: 43 (Aphodius)

rusakovi Gusakov, 2004: 6 (Aphodius) E: CT ST UK A: KZ WS
genus Loboparius A. Schmidt, 1913b: 123 type species Aphodius globulus Harold, 1859
drumonti Rakovič & Mencl, 2010: 234 (Aphodius) A: HP
dunchensis Stebnicka, 1982b: 333 (Aphodius) A: NP
globulus Harold, 1859: 207 (Aphodius) A: FUJ GAN GUA GUI GUX HEB HKG JA QIN SCH SHN TAI YUN ORR
bisectus C.O. Waterhouse, 1875a: 80 (Aphodius)
perpunctatus R. Paulian, 1934a: 110 (Aphodius)

hazarae Stebnicka, 1985: 361 (Aphodius) A: PA
immarginatus cordieri Boucomont, 1921a: 62 (Aphodius) A: PA ORR
immarginatus immarginatus A. Schmidt, 1907a: 201 (Aphodius) [RN] A: NP SD ORR
crassulus Fairmaire, 1893e: 305 (Aphodius) [HN]
vitalisi Boucomont, 1921a: 62 (Aphodius)

immarginatus kumaoensis Petrovitz, 1962a: 106 (Aphodius) A: UP
kotoensis Miwa, 1930: 174 (Aphodius) A: TAI
mirificus Balthasar, 1933a: 57 (Aphodius) A: NP SD YUN ORR
scheibei Balthasar, 1956b: 419 (Aphodius) A: AF NP PA SD
schereri Petrovitz, 1963d: 630 (Aphodius) A: NP PA UP
genus Loraphodius Reitter, 1892e: 187 type species Aphodius suarius Faldermann, 1835
corrugatus Petrovitz, 1954: 267 (Aphodius) A: IN
kashmirensis Sharp, 1878b: 171 (Aphodius) A: AF KA PA QIN XIZ
tshingchaicus Balthasar, 1971b: 58 (Aphodius)

latisulcus Reitter, 1892e: 188 (Aphodius) E: GG ST UK (Krym) A: IN
secundus Roubal, 1911a: 103 (Aphodius)

SCARABAEIDAE

131

suarius Faldermann, 1835a: 254 (Aphodius) E: AB AL AU BH BU CR FR GG GR HU IT MC MD RO SB SK SL ST
TR UK A: CY IN IS SY TR
ukrainicus Balthasar, 1946c: 59 (Aphodius)

genus Loraspis Mulsant & Rey, 1870c: 418 [=1871a: 178] type species Scarabaeus sulcatus auct. (=Aphodius
frater Mulsant & Rey, 1870)
frater Mulsant & Rey, 1870b: 203 [=1872c: 179] (Aphodius) E: AB AU BU CR CT CZ DE FR GE GG HU KZ NL PL
RO SK SP ST SV SZ UK A: KI KZ SCH TR UZ WS XIN XIZ
erythropterus d’Orbigny, 1896b: 205 (Aphodius)
pannonicus Petrovitz, 1961f: 346 (Aphodius)

genus Lunaphodius Balthasar, 1964a: 62 type species Aphodius lunifer Solsky, 1876
edgardi Solsky, 1876a: 340 (Aphodius) A: AF KI KZ QIN SCH TD TM UZ XIZ
medvedevi Iablokoff-Khnzorian, 1970: 54 (Aphodius) [HN]
sartus Iablokoff-Khnzorian, 1971: 184 (Aphodius) [RN]

lunifer Solsky, 1876a: 336 (Aphodius) A: AF KI KZ TD TM UZ
genus Macroretrus Péringuey, 1908: 632 type species Macroretrus singularis Péringuey, 1908
confusus Harold, 1862a: 154 (Aphodius) N: EG AFR
genus Mecynodes Mulsant & Rey, 1870c: 465 [=1871a: 225] type species Aphodius parallelus Mulsant, 1843
(=Aphodius striatulus Waltl, 1845)
Esimaphodius Reitter, 1892e: 208 type species Aphodius leucopterus Klug, 1845
Meraphodius D. Koshantschikov, 1894c: 112 type species Aphodius kisilkumi Solsky, 1876

alaiensis Reitter, 1892a: 226 [=1893a: 105] (Aphodius) A: AF KI KZ TD TM UZ
nasalis D. Koshantschikov, 1894c: 112 (Aphodius)

anemurensis Petrovitz, 1968g: 467 (Aphodius) A: TR
angulosus Harold, 1869f: 432 (Aphodius) N: AG EG LB TU A: IS LE SY TR
asmaricus Balthasar, 1956b: 421 (Aphodius) A: AF
becvari Červenka, 1995: 91 (Aphodius) A: JO
demoflysi Baraud, 1977b: 56 (Aphodius) N: AG MO TU
gilvipennis Balthasar, 1946c: 60 (Aphodius) A: AF TR UZ
kisilkumi Solsky, 1876a: 332 (Aphodius) E: AB AR ST A: IN IQ KI KZ TD TM TR UZ WS
mendidioides Reitter, 1892e: 209 (Aphodius)
selenocarus D. Koshantschikov, 1894c: 111 (Aphodius)

leucopterus Klug, 1845: [5], pl. xlii (Aphodius) E: SP N: AG EG LB MO TU A: CY IS LE
forcipatus Harold, 1868a: 84 (Aphodius)
luridipennis Mulsant, 1851a: 138 (Aphodius)

stereotypus D. Koshantschikov, 1894c: 109 (Aphodius) A: AF IN KZ TM UZ
augias Obenberger, 1915b: 83 (Aphodius)
kudukensis Obenberger, 1914e: 111 (Aphodius)

132

SCARABAEIDAE

striatulus Waltl, 1835: 67 (Aphodius) E: AL BH CR FR GR IT ME PT SL SP N: AG LB MO TU A: IN TR
fenestratus Reitter, 1892e: 211 (Aphodius)
parallelus Mulsant, 1843: 277 (Aphodius)
parisii Schatzmayr, 1937: 395 (Aphodius)
tormes Graëlls, 1858: 60 (Aphodius)

trochilus Reitter, 1892e: 208 (Aphodius) A: IS JO SY
genus Megatelus Reitter, 1892e: 179 type species Aphodius scolytiformis Reitter, 1892 (=Aphodius contractus
Klug, 1845)
contractus contractus Klug, 1845: [5], pl. xlii (Aphodius) A: AF CY IN IQ OM PA SI SY TR AFR ORR
scolytiformis Reitter, 1892e: 179 (Aphodius)

contractus scolytoides P.H. Lucas, 1846a: pl. 24 (Otophorus) E: IT (Lampedusa) N: AG EG LB MO TU
genus Melinopterus Mulsant, 1842: 282 type species Scarabaeus prodromus Brahm, 1790
abeillei Sietti, 1903: 66 (Aphodius) E: PT SP N: AG MO TU
dissimilis Petrovitz, 1968e: 227 (Aphodius)

consputus Creutzer, 1799b: 41 (Aphodius) E: AL AR AU BE BH BU CR CT CZ EN FR GB GE GG GR HU IT LA LU
MA MC ME NL PL PT RO SB SK SL SP ST SZ TR UK N: AG MO TU A: IN IS SY TR NARi
coriarius Gistel, 1857b: 595 (Aphodius)
griseus W.L.E. Schmidt, 1840b: 135 (Aphodius)
impunctatus Mulsant, 1842: 259 (Aphodius)
mancus Dalla Torre, 1879: 109 (Aphodius)
mendicus Mulsant, 1842: 258 (Aphodius)
metallescens Mulsant, 1842: 258 (Aphodius)
obsoletus Dalla Torre, 1879: 109 (Aphodius)
prodromus Duftschmid, 1805: 109 (Aphodius)

dellacasai Ávila, 1986: 101 (Aphodius) E: SP
edithae Reitter, 1906e: 437 (Aphodius) E: AR A: IN IQ
gissaricus Akmetova & Frolov, 2012: 65 (Aphodius) [DA] A: TD
guillebeaui Reitter, 1891a: 180 (Aphodius) [RN] E: FR SL
pectoralis Guillebeau, 1888: 190 (Aphodius) [HN]

imamae Stebnicka, 1988: 962 (Aphodius) A: IN
meuseli Reitter, 1906e: 435 (Aphodius) A: ES WS
prodromus Brahm, 1790: 3 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG GR
HU IR IT KZ LA LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO A: AF CY
ES FE IN IQ IS KI KZ LE MG SY TD TM TR UZ WS XIN NARi
abscurus Eichler, 1922: 67 (Aphodius)
angustatus Mulsant, 1842: 284 (Aphodius)
discoidalis Gistel, 1857b: 589 (Aphodius) [HN]
flavogriseus Mulsant, 1842: 284 (Aphodius)

SCARABAEIDAE

133

gisteli E. Strand, 1917: 95 (Aphodius) [RN]
griseolus Mulsant, 1842: 284 (Aphodius)
griseus Dalla Torre, 1879: 109 (Aphodius)
lunulatus Eichler, 1922: 67 (Aphodius)
mulsanti d’Orbigny, 1896b: 233 (Aphodius) [RN]
obliquus Mulsant, 1842: 285 (Aphodius)
rapax Faldermann, 1835a: 256 (Aphodius)
restrictus Mulsant, 1842: 284 (Aphodius)
semilunus Mulsant, 1842: 285 (Aphodius)
semipellitus Solsky, 1876a: 235 (Aphodius)
syriacus Mulsant & Rey, 1870b: 220 [=1872c: 196] (Aphodius) [HN]
weberi Reitter, 1906e: 436 (Aphodius)

pubescens Sturm, 1800: 40 (Aphodius) E: AB AL AR AU BE BH BU CR CT CZ GE GG GR HU LT MC ME PL RO
SK SL ST SV SZ UK N: TU A: FE IN IQ IS LE SY TR UZ WS
nebulosus Schilsky, 1888b: 316 (Aphodius)
tabidus Erichson, 1848a: 876 (Aphodius)

punctatosulcatus hirtipes Fischer von Waldheim, 1844: 45 (Aphodius) E: AB AR BH BY CT EN FI GG GR KZ LA
NR NT RO ST SV TR UK A: ES FE IN IQ KI KZ MG TD TM TR UZ WS XIN
arenicola Möller, 1890: 115 (Aphodius)
ponticus Petrovitz, 1967b: 328 (Aphodius)
pseudoinclusus Balthasar, 1963d: 283 (Aphodius)
sabulicola C.G. Thomson, 1868: 16 (Aphodius)
similis D. Koshantschikov, 1894c: 116 (Aphodius)

punctatosulcatus punctatosulcatus Sturm, 1805: 15 (Aphodius) E: AU CZ FR GB GE HU IR IT (Veneto, TrentinoAlto Adige) NL PL PT RO SK SP N: MO
convexifrons Rey, 1890a: 164 (Aphodius)
laeviceps Rey, 1890a: 164 (Aphodius)
funebris Reitter, 1892e: 233 (Aphodius)

reyi Reitter, 1892e: 233 (Aphodius) E: AU BH BU CZ DE FR GE GR HU IT LU NL PL RO SP SK ST SZ TR UK A: TR
scuticollis Semenov, 1898b: 598 (Aphodius) A: AF KI KZ QIN TD TM WS UZ XIN XIZ
ingenticollis W. Koshantschikov, 1912: 520 (Aphodius)
triens W. Koshantschikov, 1911a: 23 (Aphodius)

sertavulensis Pittino, 1988: 115 (Aphodius) A: TR
sphacelatus Panzer, 1798: 5 (Scarabaeus) E: AB AL AN AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG
GR HU IR IT LA LT LU MC MD NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: AG MO TU A: ES IQ IS KI
KZ TM TR WS
extensus Mulsant, 1842: 285 (Aphodius)
fimicola Gebler, 1833: 284 (Aphodius)
hirtellus Laporte, 1840: 95 (Aphodius)
marginalis Stephens, 1830: 203 (Aphodius)

134

SCARABAEIDAE

obscurellus Schilsky, 1888b: 315 (Aphodius)

stolzi Reitter, 1906e: 439 (Aphodius) E: AL CR FR (Corse) GR IT ME TR N: LB A: TR
tingens Reitter, 1892e: 233 (Aphodius) E: FR IT (Toscana, Sardegna, Sicilia) PT SP N: AG MO TU
litigiosus Báguena, 1930: 321 (Aphodius)

villarreali Baraud, 1975a: 47 (Aphodius) E: SP N: AG TU
wittmeri Petrovitz, 1975c: 219 (Aphodius) A: BT
genus Mendidaphodius Reitter, 1901a: 73 type species Aphodius spinifrontis Reitter, 1901 (=Mendidius brancsiki Reitter, 1899)
adustus Klug, 1855: 656 (Aphodius) A: SA AFR
rubroplagiatus Balthasar, 1935a: 72 (Aphodius)

angustatus Klug, 1845: [7], pl. xlii (Aphodius) N: EG A: IS AFR
armiger Harold, 1871d: 259 (Aphodius) E: GR A: IS SY TR
brancsiki Reitter, 1899c: 200 (Mendidius) E: AB ST A: AF KZ PA TM UZ
adolfischmidti Reitter, 1909b: 75 (Aphodius)
spinifrontis Reitter, 1901a: 73 (Aphodius)

fischeri Petrovitz, 1962a: 109 (Aphodius) A: TM UZ
lepidulus Harold, 1866: 119 (Aphodius) E: AR A: AF IN IQ IS TM TR
bagdadensis Petrovitz, 1971d: 216 (Aphodius)
fuscolimbatus Harold, 1880a: 155 (Aphodius)

linearis Reiche & Saulcy, 1856: 394 (Aphodius) E: BY CT GR ST UK N: EG A: IN IS SY TR
cylindricus Reiche, 1856b: 396 (Aphodius)
elegantulus Hochhuth, 1873b: 131 (Aphodius)
lgockii Roubal, 1913b: 70 (Aphodius)
malinovskyi W. Koshantschikov, 1913a: 183 (Aphodius)
roseus W. Koshantschikov, 1913a: 184 (Aphodius)
rubronotatus Balthasar, 1935c: 124 (Aphodius)
sareptanus Balthasar, 1935c: 123 (Aphodius)

nodulifer D. Koshantschikov, 1894c: 110 (Aphodius) A: UZ
palaestinensis Petrovitz, 1963d: 632 (Aphodius) A: IS JO
paganettii Petrovitz, 1963d: 641 (Aphodius) E: PT SP N: MO
lajonquierei Baraud, 1977a: 124 (Aphodius)

porrectus A. Schmidt, 1913b: 177 (Aphodius) A: SA AFR
pseudoadustus Petrovitz, 1958c: 135 (Aphodius) A: PA ORR
rutilus Klug, 1845: [7], pl. xlii (Aphodius) N: EG A: SA SI YE AFR
lesnei A. Schmidt, 1911b: 101 (Aphodius)

steinbergi Nikritin, 1973: 619 (Aphodius) A: TM
timurkirgizi G. Dellacasa & M. Dellacasa, 2009: 87 A: TR
genus Mendidius Harold, 1868a: 86 [RN] type species Aphodius fimbriolatus Mannerheim, 1849
Aphodosugrames Endrődi, 1964: 424 type species Aphodosugrames gobiensis Endrődi, 1964
Cnemargus Harold, 1866: 123 [HN] type species Aphodius fimbriolatus Mannerheim, 1849

SCARABAEIDAE
assectator Balthasar, 1961c: 368 (Aphodius) N: EG LB A: IQ IS KU SA
pseudofimbriolatus Petrovitz, 1961h: 445 (Aphodius)

atricolor Reitter, 1892e: 183 (Aphodius) A: AF TD
baigakumi W. Koshantschikov, 1911a: 18 (Aphodius) E: ST A: KZ UZ
beluchistanicus Petrovitz, 1962a: 118 (Aphodius) A: IN SA
berbericus Balthasar, 1941a: 84 (Aphodius) N: AG LB MO
bidens Solsky, 1876a: 346 (Aphodius) E: AR KZ A: AF ES IN KI MG TM TR UZ
bispinifrons Reitter, 1889b: 103 (Aphodius) E: AR ST A: TM TR XIN
deserticola S.I. Medvedev & Nikritin, 1976: 1913 (Aphodius)

bivittatus A. Schmidt, 1922: 100 (Aphodius) A: HEB HEI JIL LIA TIA
vlacili Tesař, 1945: 69 (Aphodius)

calliger J.R. Sahlberg, 1908: 57 (Aphodius) [DA] E: EG A: IS LE SY
curtulus Harold, 1866: 125 (Aphodius) E: ST
diffidens Reitter, 1892e: 182 (Aphodius) E: AB AR A: IN TR
endroedii Balthasar, 1967b: 137 (Aphodius) A: ES
fimbriolatus Mannerheim, 1849: 235 (Aphodius) A: ES JIL MG TM
bidentellus A. Schmidt, 1910c: 353 (Aphodius) [RN]

gobiensis Endrődi, 1964: 424 (Aphodius) A: MG
granulifrons Fairmaire, 1883c: xlii (Aphodius) N: AG EG MO TU A: SA
heikertingeri Balthasar, 1938a: 12 (Aphodius) E: AB (Naxçıvan)
kabakovi Nikritin, 1979b: 762 (Aphodius) A: AF
laevicollis Harold, 1866: 123 (Aphodius) N: AG EG LB AFR
mamajevi S.I. Medvedev, 1968b: 1249 (Aphodius) A: UZ
multiplex Reitter, 1897l: 241 (Aphodius) E: AR KZ ST A: AF IN KI KZ MG NMO TM UZ XIN
breviciliatus Poppius, 1904b: 366 (Aphodius)
granulifrons Reitter, 1887b: 221 (Aphodius) [RN]
reitteri Fairmaire, 1894b: 313 (Aphodius) [HN]
reitterianus Heyden, 1896: 44 (Aphodius) [RN]

nelsinae S.I. Medvedev, 1968b: 1247 (Aphodius) E: ST A: KZ XIN
olexai Král, 1996b: 195 (Aphodius) A: KZ
osiris W. Koshantschikov, 1916: 201 (Aphodius) [DA] N: EG
palmetincolus Karsch, 1881: 45 (Aphodius) N: AG CI EG LB MO TU A: IS SI
pseudobidens Balthasar, 1941a: 85 (Aphodius) A: HEI JIL
repetekensis Nikritin, 1971: 142 (Aphodius) A: TM
kuznetzovi Nikritin, 1971: 144 (Aphodius)

rutilinus Reitter, 1892e: 182 (Aphodius) N: AG EG
samarcandicus Nikritin, 1979a: 66 (Aphodius) A: UZ
sijazovi Lebedev, 1932: 59 (Aphodius) A: IN KZ UZ XIN
willbergi Reitter, 1891g: 255 A: UZ
xerses Petrovitz, 1963c: 243 (Aphodius) A: IN
zhantievi Gusakov, 1997: 15 (Aphodius) A: TM

135

136

SCARABAEIDAE

genus Mesontoplatys Motschulsky, 1864: 462 type species Mesontoplatys rufolaterus Motschulsky, 1864
arabicus Harold, 1875a: 89 (Aphodius) N: EG A: AE IN IS KU OM SA SI YE
dorsalis Klug, 1855: 656 (Aphodius) A: PA SA YE AFR
probes Péringuey, 1901: 401 (Aphodius)
tenuitarsis Fairmaire, 1897b: 98 (Aphodius)

parvulus Harold, 1871a: 17 (Aphodius) A: HP PA AFR ORR
rufolaterus Motschulsky, 1864: 462 (Aphodius) A: PA UP ORR
roissi Petrovitz, 1961c: 111 (Aphodius)
tranquebaricus Harold, 1874a: 208 (Aphodius)

genus Mothon Semenov & S.I. Medvedev, 1927: 182 type species Aphodius sarmaticus Semenov & S.I.
Medvedev, 1927
sarmaticus Semenov & S.I. Medvedev, 1927: 182 (Aphodius) E: RO ST UK
genus Mozartius Nomura & Nakane, 1951: 46 type species Mozartius testaceus Nomura & Nakane, 1951
jugosus horii Ochi, Kawahara & Inagaki, 2006a: 145 A: JA (Fukue-jima)
jugosus jugosus Lewis, 1895c: 383 (Oxyomus) A: JA
jugosus shingoi Ochi, Kawahara & Inagaki, 2006b: 35 A: JA (Tsushima-shi)
kyushuensis Ochi, Kawahara & Kawai, 2002: 46 A: JA
testaceus testaceus Nomura & Nakane, 1951: 46 A: JA
testaceus shikokuensis Masumoto, 1984b: 173 A: JA
uenoi hadai Kawai, 2003: 47 A: JA (Oita Pref.)
uenoi uenoi Masumoto, 1984a: 25 A: JA
genus Neagolius W. Koshantschikov, 1912: 517 type species Aphodius falcispinis W. Koshantschikov, 1912
abchasicus Reitter, 1892e: 241 (Aphodius) E: AR GG ST A: TR
amblyodon K. Daniel, 1900: 139 (Aphodius) E: FR (Southeastern Alps) IT (Alpi Occidentali) SZ
aragatsi Shokhin & Kalashian, 2015: 315 E: AR
bilimeckii Seidlitz, 1891: 149 (Aphodius) E: AL AU BH CR CZ (Jeseníky) GE GR IT (Lombardia, Veneto, FriuliVenezia Giulia) MC ME SB SL SZ
bernhaueri Reitter, 1896g: 269 (Aphodius)
danielorum Semenov, 1902b: 294 (Aphodius) [RN]
limbolarius Reitter, 1892e: 242 (Aphodius)
picturatus J. Daniel, 1902: 80 (Aphodius)

dzhungaricus Nikolajev & Frolov, 1997: 281 (Aphodius) A: KZ
falcispinis W. Koshantschikov, 1912: 517 (Aphodius) A: ES GAN MG NMO QIN WS XIN
haroldi D. Koshantschikov, 1894a: 98 (Aphodius) A: KI KZ XIN
hengduanensis Král, 1997a: 54 (Aphodius) A: YUN ORR
heydeni Harold, 1871b: 112 (Aphodius) E: PT SP
interstitialis D. Koshantschikov, 1894c: 115 (Aphodius) A: TD UZ
roniticus Stebnicka, 1975: 186 (Aphodius)

SCARABAEIDAE

137

liguricus J. Daniel, 1902: 90 (Aphodius) E: FR (Southeastern Alps) IT (Alpi Marittime, Appennino
Tosco-Emiliano)
montanus Erichson, 1848a: 887 (Aphodius) E: AL AN AU BH BU CR CT FR GE GR IT MC ME PL RO SB SL SP
ST SZ
deubeli Reitter, 1897c: 76 (Aphodius)
sterbai Rambousek, 1915: 88 (Aphodius)

montisjuldi Pittino, 1988: 113 (Aphodius) A: XIN
montivagus Erichson, 1848a: 889 (Aphodius) E: AU GE IT (Alpi) SL
brunneus Schilsky, 1888b: 316 (Aphodius) [HN]
cenisius J. Daniel, 1902: 85 (Aphodius)
cingulatus A. Schmidt, 1916: 96 (Aphodius) [RN]

moronensis Frolov, 2009: 625 (Aphodius) A: MG
orinus W. Koshantschikov, 1912: 516 (Aphodius) A: KA
ovitensis laticeps Pittino & Ballerio, 1996: 143 (Aphodius) A: TR (Western Çanik Mts.)
ovitensis ovitensis Pittino & Ballerio, 1994: 237 (Aphodius) A: TR (Eastern Çanik Mts.)
penninus J. Daniel, 1902: 89 (Aphodius) E: IT (Alpi Pennine)
petri Frolov, 2014: 449 (Aphodius) [RN] A: MG XIN XIZ
kozlovi Frolov, 2009: 627 (Aphodius) [HN]

pollicatus Erichson, 1848a: 888 (Aphodius) E: AU CR GE IT (Alpi Orientali, Prealpi) SL
praecox Erichson, 1848a: 889 (Aphodius) E: AU (Steyerische Alpen)
hibernus Gistel, 1857b: 593 (Aphodius)
picimanus Erichson, 1848a: 890 (Aphodius)
schmidti Rosenhauer, 1847: 115 (Aphodius)

savitskiyi Gusakov, 2004: 3 (Aphodius) A: KZ
schlumbergeri consobrinus K. Daniel, 1900: 139 (Aphodius) E: AU IT (Alpi e Prealpi Bergamasche, Prealpi
Venete, Alpi Apuane)
schlumbergeri samniticus J. Daniel, 1902: 83 (Aphodius) E: IT (Appennino Meridionale e Centrale)
schlumbergeri schlumbergeri Seidlitz, 1888b: 150 [Arten] (Aphodius) E: AN FR (Pyrénées Centrales) SP
(Pirineos)
schlumbergeri temperei Hoffmann, 1938: 3 (Aphodius) E: FR (Pyrénées Orientales, Massif Central) SP
(Cantabria)
suschkini W. Koshantschikov, 1913a: 197 (Aphodius) A: WS
takin Král, 1995b: 105 (Aphodius) A: SCH
zangbo Král, 1997a: 55 (Aphodius) A: XIZ
genus Neocalaphodius Bordat, 1990: 47 type species Aphodius moestus Fabricius, 1801
moestus Fabricius, 1801a: 78 (Aphodius) A: AF BT HP IN IQ JA NP OM PA SA TAI TD TR UP AFR ORR
madagascariensis Harold, 1859: 208 (Aphodius)
mutans Walker, 1858: 207 (Aphodius)
subvittatus Fairmaire, 1896d: 452 (Aphodius)

mussooriensis Mittal, 1993: 243 (Aphodius) A: UP

138

SCARABAEIDAE

genus Nialaphodius H.J. Kolbe, 1908b: 128 type species Aphodius nigrita Fabricius, 1801
nigrita Fabricius, 1801a: 73 (Aphodius) N: EG AFR NAR NTR ORR
binominatus A. Schmidt, 1907a: 202 (Aphodius) [RN]
cuniculus Chevrolat, 1864: 411 (Aphodius)
expertus Harold, 1871a: 19 (Aphodius)
guadeloupensis Fleutiaux & Sallé, 1889: 396 (Aphodius)
nigritulus Boheman, 1857: 338 (Aphodius)
paivanus Wollaston, 1867: 90 (Aphodius)
pervicax Balthasar, 1942a: 119 (Aphodius)
tuberifrons Fairmaire, 1903: 187 (Aphodius) [HN]
vestiarius Horn, 1870: 121 (Aphodius)

genus Nialosternus Hollande & Thérond, 1999: 96 type species Aphodius sitiphoides d’Orbigny, 1896
sitiphoides d’Orbigny, 1896a: 149 (Aphodius) N: AG LB MO TU A: IQ IS QA SA SY AFR
irakanus Petrovitz, 1965d: 675 (Aphodius)
mothomimus R. Paulian, 1980: 148 (Aphodius)
tschadensis Petrovitz, 1963d: 636 (Aphodius)

genus Nialus Mulsant & Rey, 1870c: 456 [=1871a: 216] type species Aphodius varians Duftschmid, 1805
jhavanicus Balthasar, 1971a: 18 (Aphodius) A: AP NP ORR
cobharensis Stebnicka, 1985: 363 (Aphodius)

politus Mulsant & Rey, 1870b: 204 [=1872c: 172] (Aphodius) [DA] A: SY
varians Duftschmid, 1805: 93 (Aphodius) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FR GB GE GG GR HU
IT KO KZ LA LS LT LU MC MD ME NL NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO A: IN IQ IS JO
KI KZ SY TD TM TR UZ WS XIN
ambiguus Mulsant, 1842: 202 (Aphodius)
bimaculatus Fabricius, 1787a: 8 (Scarabaeus) [HN]
fabricii d’Orbigny, 1896b: 215 (Aphodius) [RN]
immaculatus Hochhuth, 1873b: 130 (Aphodius)
kocae Reitter, 1897g: 204 (Aphodius)
matutinalis Gistel, 1857b: 601 (Aphodius)
niger Panzer, 1797a: 1 (Scarabaeus) [HN]
punctatellus Mulsant, 1842: 202 (Aphodius)
punctulatus Marsham, 1802: 30 (Scarabaeus)
punctulatus Mulsant & Rey, 1870c: 461 [=1871a: 221] (Aphodius)
purpurascens Eichler, 1922: 67 (Aphodius)
rugosopunctatus Petrovitz, 1971e: 567 (Aphodius)
venyigei Endrődi, 1969a: 275 (Aphodius)

genus Nimbus Mulsant & Rey, 1870c: 578 [=1871a: 338] type species Aphodius obliteratus Panzer, 1823
affinis Panzer, 1823: 1 (Aphodius) E: AU CT CZ UK

SCARABAEIDAE

139

ciliaris W.L.E. Schmidt, 1840b: 164 (Aphodius)
hoberlandti Tesař, 1945: 72 (Aphodius)

anyerae Ruiz, 1998b: 308 (Aphodius) N: MO
cartalinius Olsoufieff, 1918: 66 (Aphodius) E: AB GG A: TR
contaminatus Herbst, 1783: 9 (Scarabaeus) E: AU BE BH BU CR CT CZ DE FI FR GB GE GG GR HU IR IT LA LT
LU NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: TU A: IS TR
casarum Gistel, 1857a: 5 [=1857b: 517] (Aphodius)
ciliaris Marsham, 1802: 14 (Scarabaeus)
incoloratus Mulsant, 1842: 292 (Aphodius)
indistinctus Mulsant, 1842: 292 (Aphodius)
miser Mulsant, 1842: 292 (Aphodius)

franzinii Pittino, 1978: 302 (Aphodius) E: FR (Corse) IT (Sardegna)
harpagonis Reitter, 1890f: 147 (Aphodius) A: IS JO LE SY
johnsoni Baraud, 1976a: 80 (Aphodius) E: AL BH BU CR GR IT ME SL A: TR
libanonensis Petrovitz, 1958c: 137 (Aphodius) [DA] A: IS LE SY TR
makowskyi D. Koshantschikov, 1891: 438 (Aphodius) [DA] A: KI KZ TM UZ
alatavicus Nikolajev, 1974: 82 (Aphodius)
bartosi Balthasar, 1938d: 213 (Aphodius)
madari Tesař, 1945: 67 (Aphodius)

marianii Pittino, 1978: 306 (Aphodius) E: FR
obliteratus Panzer, 1823: 3 (Aphodius) E: AB AU BE BH BU CR CT CZ DE FR GB GE GG GR HU IR IT LA LU MA
MC ME NL PL RO SB SK SL SP ST SV SZ A: IS TR
fulveolus Mulsant, 1842: 289 (Aphodius)
insubidus Germar, 1824: 110 (Aphodius)

orbignyi Clouët des Pesruches, 1896b: 371 (Aphodius) N: AG CI MO TU
proximus Ádám, 1994a: 7 E: BE FR PT SP
richardi Veiga, 1984: 181 (Aphodius) E: SP N: MO
genus Nipponaphodius Nakane, 1959: 32 type species Aphodius gotoi Nomura & Nakane, 1951
gotoi Nomura & Nakane, 1951: 41 (Aphodius) A: JA
genus Nipponoagoliinus Ochi & Kawahara, 2001: 59 type species Aphodius yasutakai Ochi & Kawahara, 2001
yasutakai Ochi & Kawahara, 2001: 60 (Aphodius) A: JA
genus Nobiellus M. Dellacasa & G. Dellacasa, 2005: 53 type species Aphodius bonnairei Reitter, 1892
bonnairei Reitter, 1892e: 234 (Aphodius) E: FR SP N: AG MO
cuniculorum Mayet, 1904: 131 (Aphodius)

circumductus Solsky, 1876a: 338 (Aphodius) E: AB ST A: KZ TM UZ
bachofeni Reitter, 1906e: 441 (Aphodius)

inclusus Reitter, 1892e: 235 (Aphodius) E: AB AR GG ST A: AF IN TD TM TR
glasunovi D. Koshantschikov, 1894c: 108 (Aphodius)

140

SCARABAEIDAE

kulikulanus Balthasar, 1970: 251 (Aphodius)

pustulifer Reitter, 1892e: 228 (Aphodius) E: ST A: AF KZ TM UZ
malyi Obenberger, 1914e: 111 (Aphodius)

genus Nobius Mulsant & Rey, 1870c: 563 [=1871a: 323] type species Scarabaeus serotinus Creutzer, 1799
bosaki Tesař, 1945: 70 (Aphodius) A: SCH SHN
chaldaeus Petrovitz, 1971d: 217 (Aphodius) A: IN IQ TM
moreleti Baraud, 1980c: 140 (Aphodius)

costatellus A. Schmidt, 1916: 96 (Aphodius) [RN] A: ES FE
costulatus Reitter, 1896b: 33 (Aphodius) [HN]

dosangi Akhmetova & Frolov, 2008: 408 (Aphodius) E: ST A: KZ
gresseri Semenov, 1899b: 611 (Aphodius) E: AR ST UK A: KZ
korgaldzhensis Nikolajev, 1987: 118 (Aphodius)

rhodiensis Baraud, 1976c: 402 (Aphodius) A: GR (Ródos)
serotinus Panzer, 1799b: 2 [NP] (Scarabaeus) E: AB AR AU BH BU BY CR CT EN FI FR GG GR HU IT KZ LA MD
NT RO SK SL ST SZ UK A: CY ES FE HEI IQ IS KZ MG TD TM TR WS
minutus Herbst, 1783: 10 [NO] (Scarabaeus)
serotinus Creutzer, 1799a: no. 67: 2 [HN] (Scarabaeus)
x-signum Reitter, 1892e: 235 (Aphodius)

genus Odontacrossus G. Dellacasa, Král, M. Dellacasa & Bordat, 2014: type species Aphodius obenbergeri
Balthasar, 1932
obenbergeri Balthasar, 1932h: 209 (Aphodius) A: GAN SCH
pseudoobenbergeri Červenka, 1995: 89 (Aphodius) A: XIZ YUN ORR
trisuliensis Stebnicka, 1986a: 28 (Aphodius) A: AP NP ORR
genus Orodaliscus Reitter, 1900c: 84 type species Aphodius rotundangulus Reitter, 1900
angulatulus A. Schmidt, 1916: 95 (Aphodius) [RN] A: UZ
angulatus A. Schmidt, 1907a: 201 (Aphodius) [RN] [HN]
discolor Solsky, 1876a: 345 (Aphodius) [HN]
ghilarovi S.I. Medvedev, 1968c: 1565 (Aphodius)

rotundangulus Reitter, 1900c: 84 (Aphodius) E: BU CT PL RO ST UK A: KI KZ XIN
batesoni Semenov & S.I. Medvedev, 1928a: 101 (Aphodius)
fortimargo Reitter, 1907d: 409 (Aphodius)
makolskii Roubal, 1936b: 99 (Aphodius)

transaralicus Nikolajev, 1983: 538 (Aphodius) A: KZ TM UZ
zangi A. Schmidt, 1906: 411 (Aphodius) [DA] E: CT KZ NT ST UK A: ES KI KZ WS XIN
jureceki Balthasar, 1932e: 113 (Aphodius)

genus Oromus Mulsant & Rey, 1870c: 435 [=1871a: 195] type species Scarabaeus alpinus Scopoli, 1793
alpinus Scopoli, 1763: 9 (Scarabaeus) E: AN AU BE BH CR CZ FR GE GR IT (Alpi) PL RO SB SK SL SP SZ

SCARABAEIDAE

141

carthusianus Mulsant, 1842: 189 (Aphodius)
dilatatus W. L. E. Schmidt, 1840b: 105 (Aphodius)
festinans Gistel, 1857b: 581 (Aphodius)
mulsanti Dalla Torre, 1879: 108 (Aphodius) [RN]
nigerrimus Dalla Torre, 1879: 108 (Aphodius)
onthobius Gistel, 1857b: 570 (Aphodius)
picescens Fairmaire, 1903: 187 (Aphodius)
rubens Comolli, 1837: 23(Aphodius)
rubens Mulsant, 1842: 189 (Aphodius) [HN]
rupicola Mulsant, 1842: 189 (Aphodius)
schmidti Heer, 1841: 513 (Aphodius)
turficola Gistel, 1857b: 576 (Aphodius)

genus Osmanius Branco & Baraud, 1988: 33 type species Paracoptochirus balthasari Petrovitz, 1963
balthasari Petrovitz, 1963c: 244 (Paracoptochirus) E: GR A: IN TR
dellacasai Petrovitz, 1970a: 1 (Paracoptochirus) N: LB
persicus Branco & Baraud, 1988: 36 A: IN
genus Otophorus Mulsant, 1842: 178 type species Scarabaeus haemorrhoidalis Linnaeus, 1758
haemorrhoidalis Linnaeus, 1758: 348 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB
GE GG GR HU IT KO KZ LA LT LU MD NL NR NT ME PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO A:
BT CY ES FE GAN HP IN JA JIL KI KZ LIA MG NC NMO NP SC SCH SY TD TM TR UZ WS XIN XIZ NARi
NTRi
crudus A. Schmidt, 1916: 97 (Aphodius) [RN]
humeralis Mulsant, 1842: 174 (Aphodius)
rubidus Mulsant, 1842: 174 (Aphodius) [HN]
sanguinolentus Herbst, 1783: 6 (Scarabaeus)

genus Oxyaphodius Balthasar, 1965b: 314 type species Aphodius sikkimensis Balthasar, 1965
sikkimensis Balthasar, 1965b: 314 (Aphodius) A: BT NP SD
genus Oxyomus Dejean, 1833: 147 type species Scarabaeus porcatus Fabricius, 1775 (=Scarabaeus sylvestris
Scopoli, 1763)
Amidorinus W. Koshantschikov, 1912: 519 type species Amidorinus rosti W. Koshantschikov, 1912

arunae Stebnicka, 1985: 364 A: NP
cameratus A. Schmidt, 1908b: 47 A: UP
ishidai Nakane, 1977b: 167 A: JA
masumotoi Nomura, 1973b: 42 A: TAI
miliaris A. Schmidt, 1908c: 53 A: NP SD ORR
rosti W. Koshantschikov, 1912: 519 (Amidorinus)

nubigenus Petrovitz, 1968c: 36 A: NP SD UP

142

SCARABAEIDAE

simillimus A. Schmidt, 1908b: 48 A: UP
sylvestris Scopoli, 1763: 5 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG GR
HU IT KO LA LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: TU A: CY IN IS TM TR
NARi
fenestralis Schrank, 1781: 17 (Scarabaeus)
foveolatus Moll, 1785: 170 (Scarabaeus)
foveolatus Mulsant, 1842: 309 [HN]
opacifrons Horn, 1871a: 284
platycephalus Marsham, 1802: 56 (Ips)
porcatus Fabricius, 1775: 20 (Scarabaeus)

taipingensis Masumoto, Kiuchi & T.-C. Wang, 2014c: 267 A: TAI
genus Parabodilus Hollande & Thérond, 1999: 159 type species Aphodius wollastoni Harold, 1862
wollastoni iranicus Balthasar, 1946c: 65 (Aphodius) A: AE IN IQ IS JO KU OM PA SA SI SY TM YE ORR
radjastanicus Petrovitz, 1962a: 122 (Aphodius)

wollastoni wollastoni Harold, 1862b: 397 (Aphodius) N: AG CI EG LB MO TU AFR
genus Paracoptochirus Balthasar, 1963d: 277 type species Coptochirus singularis Harold, 1868
kozanensis Pittino, 2001: 86 E: GR (Makedonia)
petrovitzi Branco & Baraud, 1988: 30 A: TR
singularis Harold, 1868d: 396 (Coptochirus) E: GR (Thessalia) RO (Constanţa) TR A: IS
haemonius Reitter, 1903b: 176 (Coptochirus)

vignai Carpaneto & Piattella, 1990: 88 E: GR
genus Paracrossidius Balthasar, 1932c: 30 type species Aphodius instigator Balthasar, 1932
impressiusculus Fairmaire, 1888e: cci (Aphodius) A: BEI GUA NMO NP SCH XIN XIZ
impressicollis Boucomont, 1929: 776 (Aphodius)
viturati Reitter, 1907d: 411 (Aphodius)

instigator Balthasar, 1932c: 33 (Aphodius) A: QIN SCH
kozlovi W. Koshantschikov, 1916: 197 (Aphodius) [DA] A: MG
laticapitus Y.-W. Zhang, 1992b: 492 (Aphodius) [DA] A: SCH
melanodiscus Y.-W. Zhang, 1992b: 491 (Aphodius) [DA] A: SCH
planipennis Petrovitz, 1961f: 345 (Aphodius) A: SCH
sichuanensis Y.-W. Zhang, 1992b: 491 (Aphodius) [DA] A: SCH
suvorovi Kabakov, 1996: 881 (Aphodius) [DA] A: ES FE
truculentus Balthasar, 1963d: 280 (Aphodius) A: QIN SCH XIZ
transmontanus Balthasar, 1932h: 211 (Aphodius) A: SCH YUN
genus Paradidactylia Balthasar, 1937: 199 type species Paradidactylia atra Balthasar, 1937
biseriata A. Schmidt, 1908b: 46 (Aphodius) A: NP UP ORR
dehradunensis Petrovitz, 1961e: 147 (Aphodius)

SCARABAEIDAE

143

brivioi Petrovitz, 1961e: 146 (Aphodius) A: UP ORR
carinulata Motschulsky, 1864: 461 (Aphodius) A: GUA TAI ORR
fastigata A. Schmidt, 1908a: 41 (Aphodius) A: OM AFR
haafi Petrovitz, 1961c: 101 (Aphodius) A: NP PA ORR
kathmandui Stebnicka, 1985: 360 (Aphodius) A: NP PA
shantungensis Balthasar, 1941a: 88 (Aphodius) A: SHN ORR
wichei Petrovitz, 1961c: 110 (Aphodius) A: PA ORR
coimbatorensis Balthasar, 1973: 52 (Aphodius)

genus Parammoecius Seidlitz, 1888a: 143 [Arten] type species Aphodius gibbus Germar, 1816
Amalus Mulsant & Rey, 1870: 597 [=1871a: 357] [HN] type species Ammoecius pyrenaeus Jacquelin du Val, 1860

amanicus Stebnicka, 1978: 258 (Aphodius) A: TR
asphaltinus Kolenati, 1846: 15 (Aphodius) E: AB AR GG ST A: TR
falsarius Reitter, 1892e: 197 (Aphodius)

brevithorax Sumakov, 1903: 47 (Aphodius) E: GG ST A: TR
osseticus Iablokoff-Khnzorian, 1972: 281 (Aphodius)
interfinius Ziani, 1999: 1 (Aphodius)

corvinus Erichson, 1848a: 831 (Aphodius) E: AU BE BH BU BY CR CZ FR GE HU IT NL PL RO SK SL SP ST SZ UK
rufipennis G. Schmidt, 1938: 43 (Aphodius)
vertumnus Gistel, 1857b: 589 (Aphodius)
totorufus G. Schmidt, 1938: 43 (Aphodius)

gibbus Germar, 1816: 2 (Aphodius) E: AU BH CZ FR GE IT (Alto-Adige) LA PL RO SK SL ST SZ UK
anthracinus W.L.E. Schmidt, 1840b: 111 (Aphodius)
transsylvanicus Küster, 1849b: 54 (Ammoecius)

pyrenaeus Jacquelin du Val, 1860: 130 [Catalogue] (Ammoecius) E: AN FR IT (Alpi Occidentali) SP SZ
davidi Hoffmann, 1954b: 130 (Aphodius)
intactus A. Schmidt, 1916: 98 (Aphodius) [RN]
jugicola Harold, 1863: 379 (Aphodius)
monticola Mulsant, 1842: 215 (Aphodius) [HN]
reitterianus Schwarz, 1897: 32 (Aphodius)
rugiceps Mulsant & Rey, 1870c: 605 [=1871a: 365] (Aphodius)

yamato Nakane, 1960: 3 (Aphodius) A: JA
genus Paulianellus Balthasar, 1938a: 6 type species Aphodius maderi Balthasar, 1938
asahinai Nakane, 1951: 70 (Aphodius) A: JA NC SC
decoctor Balthasar, 1965a: 109 (Aphodius) A: NP
jacksoni Petrovitz, 1975b: 113 (Aphodius) A: NP SD
khandui Stebnicka, 1980b: 533 (Aphodius) A: BT NP
maderi Balthasar, 1938a: 7 (Aphodius) A: ANH SCH SHA XIZ YUN
yunlingensis Petrovitz, 1961h: 443 (Aphodius)

mureensis Stebnicka, 1980b: 531 (Aphodius) A: NP SD

144

SCARABAEIDAE

neomaderi Mencl, Král & Rakovič, 2015: 67 A: GAN SCH SHA YUN
nepalensis Balthasar, 1965a: 110 (Aphodius) A: NP
taiwanensis Mencl, Král & Rakovič, 2015: 69 A: TAI
genus Phaeaphodius Reitter, 1892e: 205 type species Aphodius solskyi Harold, 1871 (=Calamosternus rectus
Motschulsky, 1866)
Limaroides Balthasar, 1932e: 109 type species Aphodius kiulungensis Balthasar, 1932

costalis Gebler, 1848: 83 (Aphodius) E: KZ ST A: ES KZ TM WS
jakovlevi W. Koshantschikov, 1912: 514 (Aphodius)
sibiricus Harold, 1863: 341 (Aphodius)

dauricus Harold, 1863: 337 (Aphodius) E: CT ST A: AF AR ES FE KI KZ MG NMO SD TD TM UZ WS XIN XIZ
acutangulus Reitter, 1892e: 206 (Aphodius)
aeger Sharp, 1878b: 170 (Aphodius)
albociliatus Reitter, 1892e: 206 (Aphodius)
alleoni d’Orbigny, 1896a: 149 (Aphodius)
margianus Semenov & S.I. Medvedev, 1928a: 102 (Aphodius)
semiluteus Reitter, 1887b: 225 (Aphodius)
truncatangulus Reitter, 1894e: 186 (Aphodius)

fusculus Reitter, 1892e: 206 (Aphodius) [DA] A: TR
himalorectus D. Ahrens & Stebnicka, 1997: 12 (Aphodius) A: NP
jouravliowi Reitter, 1907d: 408 (Aphodius) A: ES KZ MG
latonius Reitter, 1907d: 409 (Aphodius)

kiulungensis Balthasar, 1932e: 110 (Aphodius) A: FUJ SCH XIZ
novikovi Kabakov, 1998: 6 (Aphodius) E: ST UK
plutenkoi Král, 2002a: 47 (Aphodius) A: SHA
davidkrali M. Dellacasa, G. Dellacasa & Bordat, 2002a: 7 (Aphodius)

pupillus Balthasar, 1961c: 361 (Aphodius) A: AF
rectus Motschulsky, 1866: 169 (Calamosternus) E: CT NT UK A: BEI ES FE FUJ GAN HEI HEN IN JA JIA JIL KI
KZ LIA MG NC NMO QIN SC SCH SHA SHN TAI WS XIN XIZ NAR
agraphus Mader, 1953: 78 (Aphodius)
biformis Reitter, 1892e: 205 (Aphodius)
castaneipennis C.O. Waterhouse, 1875a: 83 (Aphodius)
controversus Balthasar, 1932e: 112 (Aphodius)
diabolicus Balthasar, 1932e: 112 (Aphodius)
diversus C.O. Waterhouse, 1875a: 82 (Aphodius)
edai Balthasar, 1956a: 68 (Aphodius)
lividipennis C.O. Waterhouse, 1875a: 81 (Aphodius)
semicolor Reitter, 1892e: 205 (Aphodius)
solskyi Harold, 1871d: 251 (Aphodius)
tshuensis Protsenko, 1979: 15 (Aphodius)
vitta Motschulsky, 1866: 170 (Chilothorax)
zichyi Csiki, 1901: 107 (Aphodius)

SCARABAEIDAE

145

roddi W. Koshantschikov, 1911a: 19 (Aphodius) [DA] A: KZ WS
tauricola Hrubant, 1961: 97 (Aphodius) E: AR A: IN TR
genus Phalacronothus Motschulsky, 1860f: 156 type species Scarabaeus quadrimaculatus Linnaeus, 1760
Caenaphodius Balthasar, 1936a: 5 type species Aphodius asperolineatus Balthasar, 1936
Emadus Mulsant & Rey, 1870c: 449 [=1871a: 209] type species Scarabaeus quadrimaculatus Linnaeus, 1760
Paragolius Balthasar, 1945a: 42 type species Aphodius botulus Balthasar, 1945

ambulans Petrovitz, 1971e: 566 (Aphodius) A: TR
biguttatus Germar, 1824: 111 (Aphodius) E: AL AR AU BE BH BU CR CT CZ FR GE GG GR HU IT LA LU MC MD
NL PL RO SB SK SL SP ST SZ TR UK N: AG MO TU A: KZ TR WS
apicalis Schilsky, 1888b: 315 (Aphodius) [HN]
conjunctulus Reitter, 1892e: 213 (Aphodius)
discoloratus A. Schmidt, 1916: 95 (Aphodius) [RN]
dubius Schilsky, 1889: 337 (Aphodius)
fallax Schilsky, 1888b: 315 (Aphodius) [HN]
lutigradus Gistel, 1857b: 599 (Aphodius)
petulans A. Schmidt, 1916: 97 (Aphodius) [RN]
rusticus Gistel, 1857b: 566 (Aphodius)
sanguinolentus Panzer, 1797b: 4 (Scarabaeus)
similis Schilsky, 1888b: 315 (Aphodius)
tyrolensis Rosenhauer, 1847: 29 (Aphodius)

botulus Balthasar, 1945a: 43 (Aphodius) A: HEI JIL JA NC SC
manschuriensis Petrovitz, 1958c: 138 (Aphodius)
naraensis Nakane, 1956b: 120 (Aphodius)

citellorum Semenov & S.I. Medvedev, 1929a: 88 (Aphodius) E: AU BU CT CZ HU RO SK ST UK A: KZ TR WS
supermaculatus Mader, 1953: 92 (Aphodius)

flavolimbatus Petrovitz, 1963c: 240 (Aphodius) [DA] A: IN
fumigatulus Reitter, 1892e: 208 (Aphodius) E: AB AR BU GG GR MC A: IN IS LE SY TM TR
asperolineatus Balthasar, 1936a: 5 (Aphodius)
sculpturatus Reitter, 1892e: 208 (Aphodius)

hilaris Harold, 1869f: 433 (Aphodius) [DA] E: AR A: IN IS SY TR
pseudoavunculus Masumoto, 1977: 29 (Aphodius) A: TAI
putoni Reitter, 1894e: 187 (Aphodius) E: PT SP
quadrimaculatus Linnaeus, 1760: 138 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ EN FI FR GB GE
GG GR HU IT LA ME NL NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO TU A: ES IS KZ LE SY TM TR
UZ WS
caudatus Mulsant, 1842: 206 (Aphodius)
diecki Harold, 1870a: 114 (Aphodius)
lunulatus d’Orbigny, 1896b: 214 (Aphodius)
macrii O.G. Costa, 1839b: 39 (Aphodius)
prolongatus Mulsant, 1842: 207 (Aphodius)

146

SCARABAEIDAE

quadripustulatus Fabricius, 1792a: 36 (Scarabaeus)
quadrisignatus Brullé, 1832: 172 (Aphodius)
sanguinolens Mulsant, 1842: 206 (Aphodius)
sexpustulatus Gebler, 1841b: 588 (Aphodius)

semiangulus Balthasar, 1952: 225 (Aphodius) A: FUJ
genus Pharaphodius Reitter, 1892e: 172 type species Scarabaeus marginellus Fabricius, 1781
aegyptiacus Petrovitz, 1964e: 177 (Aphodius) [DA] N: EG
attritus Balthasar, 1933a: 55 (Aphodius) A: YUN ORR
calo Balthasar, 1971a: 17 (Aphodius) [DA] A: NP ORR
campestris Mittal, 1993: 240 (Aphodius) [DA] A: UP
corax Balthasar, 1937: 194 (Aphodius) [DA] A: SA YE AFR
cornutus Wiedemann, 1823: 26 (Aphodius) [DA] A: AF NP PA ORR
crenatus Harold, 1862a: 142 (Aphodius) [DA] A: HP NP TAI ORR
desertus Klug, 1845: [6], pl. xlii (Aphodius) [DA] N: EG A: SA SI AFR
perplexus Harold, 1862a: 151 (Aphodius)

jodhpurensis Petrovitz, 1958c: 134 (Aphodius) [DA] A: BT PA
marginellus Fabricius, 1781a: 21 (Scarabaeus) A: BT GUI HAI HKG HP JA NP PA TAI UP ORR
orientalis Harold, 1862a: 147 (Aphodius) [DA] A: GUA JA NP TAI ORR
pallescens Walker, 1871: 11 (Aphodius) [DA] N: EG A: SA YE AFR
gibbifrons Fairmaire, 1892a: 92 (Aphodius)

parvatia Mittal, 1993: 242 (Aphodius) [DA] A: UP
pereirai Balthasar, 1956b: 415 (Aphodius) [DA] A: AF NP PA
priscus Motschulsky, 1858d: 56 (Aphodius) A: NC NP SC TAI ORR
putearius Reitter, 1895f: 208 (Aphodius) [DA] A: BEI FUJ SHG TAI ORR
redargutus Balthasar, 1971c: 153 (Aphodius) [DA] A: NP ORR
robustus Walker, 1858: 207 (Aphodius) [DA] A: AF NP PA ORR
rugosostriatus C.O. Waterhouse, 1875a: 92 (Aphodius) [DA] A: ANH BEI ES FE FUJ GAN GUA GUI GUX HAI
HEB HEI HEN HUB HUN JA JIA JIL JIX LIA NC NIN NMO SC SCH SHA SHG SHN SHX TAI TIA YUN ZHE
raddei Berlov, 1989b: 393 (Aphodius)

tenerus Petrovitz, 1959b: 112 (Aphodius) [DA] A: SA AFR
vernilis A. Schmidt, 1911a: 6 (Aphodius) [DA] A: OM SA YE ORR
genus Plagiogonus Mulsant, 1842: 306 type species Scarabaeus arenarius A.G. Olivier, 1789
Eccoptaphodius Iablokoff-Khnzorian, 1964: 63 type species Aphodius avetissiani Iablokoff-Khnzorian, 1964
Oloperus Mulsant & Rey, 1870c: 610 [=1871a: 370] type species Aphodius nanus Fairmaire, 1860

arenarius A.G. Olivier, 1789: no 3: 96 (Scarabaeus) E: AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG
GR HU IT LA LT LU MD ME NL PL RO SB SK SL SP ST SV SZ UK A: CY ES IN KZ TM TR WS
avetissiani Iablokoff-Khnzorian, 1964: 61
nitidus Küster, 1849b: 55 (Ammoecius)

SCARABAEIDAE

147

pusillus Preyssler, 1790: 104 (Scarabaeus)
rhododactylus Marsham, 1802: 29 (Scarabaeus)
sabulicola Mulsant, 1842: 306 (Aphodius)

bortkevitschi W. Koshantschikov, 1916: 195 (Aphodius) A: ES JIL MG WS
gurjevae Nikolajev, 1984: 147 (Aphodius)

burgaltaicus Csiki, 1901: 107 (Aphodius) A: FE MG NMO
chan Endrődi, 1967a: 138 (Aphodius) A: MG
peregii Endrődi, 1983: 114 (Aphodius)

culminarius Reitter, 1900d: 156 (Aphodius) A: BEI FE GAN GUA KI LIA MG SC XIN XIZ ZHE
esimoides Reitter, 1892a: 226 [=1893a: 105] (Aphodius) N: AG LB MO TU
farai Balthasar, 1956b: 420 (Aphodius) A: AF IN PA
inflatus Balthasar, 1938d: 214 (Aphodius) A: SCH XIN
tesarianus R. Paulian, 1942b: 59 (Aphodius)

martensi Stebnicka, 1986a: 36 (Aphodius) A: NP
nanoides Balthasar, 1961c: 363 (Aphodius) [DA] A: IS LE TR SY
nanus Fairmaire, 1860a: 172 (Aphodius) E: SP N: AG CI EG LB MO TU
algiricus Harold, 1863: 388 (Aphodius)
theryi Clouët des Pesruches, 1896a: 54 (Aphodius)

oberthueri R. Paulian, 1936a: 63 (Aphodius) A: SD UP
nigrescens R. Paulian, 1936a: 63 (Aphodius)
rubricatus R. Paulian, 1936a: 63 (Aphodius)

palea Balthasar, 1967b: 128 (Aphodius) A: NP SD
praeustus Ballion, 1871a: 333 (Aphodius) E: AB AR ST A: AF CY IS KI KZ NMO SY TD TM TR WS UZ
quadriclypealis Červenka, 1995: 90 (Aphodius) A: YUN ORR
ramamiensis Stebnicka, 1982b: 336 (Aphodius) A: NP SD
reitteri D. Koshantschikov, 1894a: 97 (Aphodius) A: KI KZ UZ
kricheldorffi W. Koshantschikov, 1910: 18 (Aphodius)

shirahatai Nakane, 1951: 70 (Aphodius) A: SHA SHX
syriacus Harold, 1863: 389 (Aphodius) E: AB AR GG ST A: IN IQ IS JO IS LE SY TR
zobeideae Petrovitz, 1980: 609 (Aphodius) A: IN
genus Planolinellus M. Dellacasa & G. Dellacasa, 2005: 75 type species Aphodius vittatus Say, 1825
vittatus Say, 1825: 191 (Aphodius) E: AB AR BU CT GR KZ RO ST TR UK A: CY ES FE IN IS KI KZ MG SCH SY TD
TM TR UZ WS XIN NAR NTR
mundus Reitter, 1892e: 199 (Aphodius)
rufoplagiatus Reitter, 1892e: 198 (Aphodius)
sellatus Mannerheim, 1852a: 299 (Aphodius)
semiruber Motschulsky, 1860e: 131 (Aphodius)
tjanshanicus Balthasar, 1956a: 70 (Aphodius)
vitiosus Reitter, 1892e: 199 (Aphodius)

148

SCARABAEIDAE

genus Planolinoides M. Dellacasa & G. Dellacasa, 2005: 77 type species Aphodius borealis Gyllenhal, 1827
borealis Gyllenhal, 1827: 248 (Aphodius) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG GR HU IR IT
KZ LA LT MC MD ME NL NR NT PL PT RO SK SL SP ST SV SZ UK A: ES FE KZ MG TR WS NAR
errans W.J. Brown, 1930b: 243 (Aphodius)
evanescens Bouskell, 1901: 23 (Aphodius)
gyllenhalii Seidlitz, 1888b: 143 [Arten] (Aphodius)
hespericus Gistel, 1857b: 589 (Aphodius)
sedulus Harold, 1871c: 119 (Aphodius)

genus Planolinus Mulsant & Rey, 1870c: 426 [=1871a: 186] type species Scarabaeus foetidus Fabricius, 1792
(=Scarabaeus fasciatus A.G. Olivier, 1789)
fasciatus A.G. Olivier, 1789: no 3: 90 (Scarabaeus) E: AR AU BE BH BU BY CR CT CZ DE EN FA FI FR GB GE GG
HU IC IR IT LA LT LU MC NL NR NT PL PT RO SB SK SL SP ST SV SZ UK A: CY ES FE IN KI KZ MG NC NMO
SC SHA TR WS
foetidus Fabricius, 1792a: 40 (Scarabaeus) [HN]
hirundinarius Gistel, 1857b: 569 (Aphodius)
putridus Herbst, 1789: 160 (Scarabaeus)
satunini Olsoufieff, 1918: 58 (Aphodius)
sus Kugelann, 1792a: 267 (Scarabaeus) [HN]
transitus Reitter, 1892e: 200 (Aphodius)
uliginosus Hardy, 1847: 382 (Aphodius)
vitulinus Gistel, 1857b: 571 (Aphodius)

nikolajevi Berlov, 1989b: 401 (Aphodius) [DA] A: ES FE
pratensis Nomura & Nakane, 1951: 43 (Aphodius) [DA] A: ES FE (Sakhalin) JA
rufoanalis Petrovitz, 1961c: 112 (Aphodius) [DA] A: KA PA
genus Platyderides A. Schmidt, 1916: 99 [RN] type species Aphodius arrowi A. Schmidt, 1910
Platyderus A. Schmidt, 1913b: 122 [HN] type species Aphodius arrowi A. Schmidt, 1910
Thaiaphodius Masumoto, 1991a: 27 type species Aphodius doiangkhangensis Masumoto, 1991

arrowi A. Schmidt, 1910c: 353 (Aphodius) A: LIA SCH ZHE
jengi Masumoto, Kiuchi & T.-C. Wang, 2014b: 337 (Aphodius) A: TAI
klapperichi Balthasar, 1942a: 122 (Aphodius) [DA] A: FUJ
genus Pleuraphodius A. Schmidt, 1913b: 121 type species Aphodius mutilus A. Schmidt, 1911
ilamensis Stebnicka, 1990: 8 (Aphodius) [DA] A: NP
lewisi C.O. Waterhouse, 1875a: 92 (Aphodius) [DA] A: BEI FUJ GUX JA NC NMO NP PA SC TAI UP AUR ORR
genus Pseudacrossus Reitter, 1892e: 218 type species Aphodius grombczewskyi D. Koshantschikov, 1891
absconditus Balthasar, 1932c: 38 (Aphodius) A: SCH
caspius Ménétriés, 1832: 181 (Aphodius) E: AB AR CT GG KZ MD RO ST UK A: ES IN KI KZ MG TM UZ WS

SCARABAEIDAE

149

grebenscikovi Balthasar, 1961c: 372 (Aphodius) A: ES MG WS
grombczewskyi D. Koshantschikov, 1891: 439 (Aphodius) A: KI KZ TD UZ XIN
juxtus Petrovitz, 1972c: 163 (Aphodius) A: HUB SC
kalabi Král, 1997b: 132 (Aphodius) A: QIN
koshantschikovi Jakobson, 1911: 145 (Aphodius) [RN] [DA] E: AB AR GG GR ST A: CY IS KI SY TR
atratellus A. Schmidt, 1916: 97 (Aphodius) [RN]
fimicola Reiche & Saulcy, 1856: 402 (Aphodius) [HN]
rubens Olsoufieff, 1918: 64 (Aphodius)

kuskai Stebnicka, 1982b: 335 (Aphodius) A: NP
nasutus Reitter, 1887b: 221 (Aphodius) A: BEI ES MG NMO QIN SHG WS XIN XIZ
consors Reitter, 1892a: 225 [=1893a: 104] (Aphodius)
kelleri W. Koshantschikov, 1911a: 21 (Aphodius)
mehelyi Csiki, 1901: 108 (Aphodius)
necopinus Balthasar, 1932g: 165 (Aphodius)
permodicus Reitter, 1895f: 208 (Aphodius)
roschlapili Csiki, 1901: 108 (Aphodius)

przewalskyi Reitter, 1887b: 226 (Aphodius) A: QIN SCH XIN XIZ
friedrichi Petrovitz, 1958c: 131 (Aphodius)

qinghaiensis Král, 1997b: 138 (Aphodius) A: QIN
serrimargo W. Koshantschikov, 1913a: 188 (Aphodius) [DA] A: GUA SHA
sharpi Harold, 1874a: 205 (Aphodius) E: PT SP N: AG MO TU
smetanai Král, 2011: 250 (Aphodius) A: HUB
spalacophilus Novikov, 1996: 75 (Aphodius) E: ST UK
subsericeus Ballion, 1878: 283 (Aphodius) A: KI KZ XIN XIZ
impolitus A. Schmidt, 1909d: 104 (Aphodius)

suffertus A. Schmidt, 1916: 96 (Aphodius) [RN] E: GR IT (Basilicata, Puglia, Calabria, Sicilia) TR A: CY IS TR SY
ampliatus Reitter, 1892e: 220 (Aphodius)
defiorei A. Fiori, 1907: 124 (Aphodius)
dilatatus Reiche & Saulcy, 1856: 399 (Aphodius) [HN]

tenebricosus A. Schmidt, 1916: 96 (Aphodius) [RN] A: ES KZ WS
caminarius Reitter, 1892e: 219 (Aphodius) [HN]

wewalkai Petrovitz, 1971d: 217 (Aphodius) A: TR
zuercheri Reitter, 1908e: 247 (Aphodius) A: TR
genus Pseuderytus Hollande & Thérond, 1999: 76 type species Aphodius chobauti Clouët des Pesruches, 1896
chobauti Clouët des Pesruches, 1896b: 371 (Aphodius) N: AG EG MO LB TU A: IN IQ IS JO SA SY AFR
moltonii Schatzmayr, 1938b: 367 (Aphodius)
oasis Baraud, 1976c: 402 (Aphodius)
pseudounicolor Balthasar, 1960a: 4 (Aphodius)
saharicus Balthasar, 1941a: 84 (Aphodius)

150

SCARABAEIDAE

genus Pseudesymus d’Orbigny, 1896b: 218 type species Aphodius lucidus Klug, 1845
lucidus Klug, 1845: [5], pl. xlii (Aphodius) E: ST N: AG CI EG LB MO TU A: AF IN IQ IS JO KU KZ SA SI TM
TR UZ
taeniatus Wollaston, 1864: 189 (Aphodius)

genus Pseudoahermodontus G. Dellacasa & M. Dellacasa, 1997: 407 type species Pseudoahermodontus
benesi G. Dellacasa & M. Dellacasa, 1997
benesi G. Dellacasa & M. Dellacasa, 1997: 409 A: GAN
genus Pseudoheptaulacus M. Dellacasa & G. Dellacasa, 2000: 28 type species Heptaulacus iniquus
A. Schmidt, 1909
iniquus A. Schmidt, 1909b: 20 (Heptaulacus) A: YUN ORR
nepalensis Pittino, 2004a: 137 A: NP
genus Pseudoteuchestes G. Dellacasa, 1986: 193 type species Aphodius schirparakensis Petrovitz, 1955
schirparakensis Petrovitz, 1955: 277 (Aphodius) A: AF KZ
parabrachysomus Balthasar, 1956b: 417 (Aphodius)

genus Pubinus Mulsant & Rey, 1870c: 489 [=1871a: 249] type species Scarabaeus tomentosus O.F. Müller, 1776
tomentosus O.F. Müller, 1776: 55 (Scarabaeus) E: AU BE BY CT DE EN FI GE IT (Trentino Alto-Adige) KZ LA LT
NL NT PL RO ST SV SZ UK A: ES KZ WS
immundus Fabricius, 1801a: 77 (Aphodius)
lutarius Fabricius, 1792a: 35 (Scarabaeus)
modicus Semenov, 1899a: 89 (Aphodius)
tunicatus Reitter, 1894e: 188 (Aphodius)

genus Qingaphodius Král, 1997b: 143 type species Aphodius nikodymi Král, 1997
nikodymi Král, 1997b: 144 (Aphodius) A: QIN
ochotonarum Král, 1997b: 146 (Aphodius) A: QIN
genus Renaudius Balthasar, 1951: 153 type species Heptaulacus tesari R. Paulian, 1942
tesari R. Paulian, 1942b: 60 (Heptaulacus) A: CHQ SCH YUN
genus Rhodaphodius Ádám, 1994a: 5 type species Scarabaeus foetens Fabricius, 1787
foetens Fabricius, 1787a: 8 (Scarabaeus) E: AB AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG HU
IR IT KZ LA LS LT LU MC NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: MO A: ES KI KZ MG TR UZ
WS XIN
aestivalis Stephens, 1839: 160 (Aphodius)
analis Gistel, 1857b: 590 (Aphodius) [HN]
conjugatulus E. Strand & Gulbis, 1957: 256 (Aphodius)
foetens Stephens, 1830: 190 (Aphodius) [HN]

SCARABAEIDAE

151

fuscipes Mulsant, 1842: 184 (Aphodius)
hypocastaneus Bouskell, 1901: 16 (Aphodius)
limbatus Dalla Torre, 1879: 107 (Aphodius)
nigricollis Mulsant, 1842: 184 (Aphodius) [HN]
obscuricollis A. Schmidt, 1922: 274 (Aphodius) [RN]
obscuriellus A. Schmidt, 1916: 98 (Aphodius) [RN]
pseudanalis E. Strand, 1917: 95 (Aphodius) [RN]
rufiventris Preller, 1862: 75 (Aphodius)
sanguinipennis Mulsant, 1842: 184 (Aphodius)
vaccinarius Herbst, 1789: 138 (Scarabaeus)

genus Sahlbergianus M. Dellacasa & G. Dellacasa, 2002: 109 type species Aphodius longissimus J.R. Sahlberg,
1908
longissimus J.R. Sahlberg, 1908: 58 (Aphodius) N: EG A: IS SY
genus Serraphodius Kabakov, 1996: 878 type species Aphodius circassicus Reitter, 1892
circassicus Reitter, 1892e: 215 (Aphodius) E: GG ST
lederi Harold, 1876: 121 (Aphodius) E: GG ST A: TR
genus Sigorus Mulsant & Rey, 1870c: 489 [=1871a: 249] type species Scarabaeus porcus Fabricius, 1792
porcus Fabricius, 1792a: 26 (Scarabaeus) E: AB AU BE BH BU CR CZ DE EN FI FR GB GE GG GR HU IR IT
LT LU MD NL NR NT PL PT RO SB SK SL SP ST SV SZ UK A: TR
anachoreta Creutzer, 1796: 1 (Scarabaeus)
fulvicrus Marsham, 1802: 16 (Scarabaeus)
haemorrhoideus Mulsant, 1842: 267 (Aphodius)
ruficrus Marsham, 1802: 16 (Scarabaeus)
turpis Marsham, 1802: 15 (Scarabaeus)

genus Sinaphodius Červenka, 1994b: 111 type species Aphodius yunnanus Červenka, 1994
chitwanensis Schoolmeesters & van den Heuvel, 1999: 141 (Aphodius) A: NP
sinuatus Harold, 1860: 614 (Aphodius) A: TAI ORR
yunnanus Červenka, 1994b: 112 (Aphodius) A: YUN
genus Sinodiapterna G. Dellacasa, 1986: 265 type species Aphodius troitzkyi Jakobson, 1897
gorodinskiyi Gusakov, 2006: 33 (Aphodius) A: FE
hammondi G. Dellacasa, 1986: 266 (Aphodius) A: FE HEI
songrini Stebnicka & Galante, 1992: 11 (Aphodius) A: NC SC
troitzkyi Jakobson, 1897: 87 (Aphodius) A: ES FE JA NC SC
donghariensis Stebnicka, 1973: 119 (Aphodius)

zeni Ochi, 1991: 53 (Aphodius) A: TAI

152

SCARABAEIDAE

genus Sitiphus Fairmaire, 1894b: 313 type species Sitiphus brisouti Fairmaire, 1894 (=Aphodius brevitarsis
Reitter, 1894)
ammodites Peyerimhoff, 1939a: 45 (Aphodius) [DA] N: AG
brevitarsis Reitter, 1894e: 186 (Aphodius) N: AG TU
brisouti Fairmaire, 1894b: 313

sefrensis Petrovitz, 1958d: 153 (Aphodius) [DA] N: EG AG MO AFR
genus Stenotothorax A. Schmidt, 1913b: 122 type species Aphodius oblongus Say, 1823
Stenobronchus A. Schmidt, 1916: 99 [RN] type species Aphodius oblongus Say, 1823
Stenothorax A. Schmidt, 1916: 99 [RN] type species Aphodius oblongus Say, 1823

hibernalis hibernalis Nakane & Tsukamoto, 1956: 176 (Aphodius) A: JA
hibernalis saghalinensis Nakane & Tsukamoto, 1956: 176 (Aphodius) A: FE (Sakhalin)
genus Subrinus Mulsant & Rey, 1870c: 511 [=1871a: 271] type species Aphodius illigeri Mulsant & Rey, 1870
(=Aphodius sturmi Harold, 1870)
clermonti Reitter, 1907d: 407 (Aphodius) E: AB AR A: IN IQ TM TR
feculentus Fairmaire, 1892a: 93 (Aphodius) [DA] A: IS AFR
motschulskyi Nikritin, 1969a: 171 (Aphodius) A: ES FE
scintillius Nikritin & Kabakov, 1979: 762 (Aphodius) [DA] A: AF
sturmi Harold, 1870a: 106 (Aphodius) [RN] E: AB AR AU BH BU CR CT CZ FR GB GE GG GR HU IT MA MC
MD ME PL PT RO SB SK SL SP ST SZ TR UK N: AG MO MR TU A: ES FE IN IS JA KZ LIA MG NC NMO SC
TM TR
illigeri Mulsant & Rey, 1870b: 511 (Aphodius) [HN]
inouei Nomura, 1942: 116 (Aphodius)
rufus Sturm, 1805: 144 (Aphodius) [HN]

subtilis D. Koshantschikov, 1894d: 126 (Aphodius) [DA] E: ST
vitellinus Klug, 1845: [6], pl. xlii (Aphodius) E: FR IT (Sicilia, Lampedusa, Linosa, Pantelleria) SP N: AG EG
LB MO TU A: IN IQ IS SA SI SY AFR
ephippiger Mulsant & Rey, 1870b: 210 [=1872c: 186] (Aphodius)
nitens Mulsant & Rey, 1870b: 211 [=1872c: 187] (Aphodius)

yemeniticus Petrovitz, 1970d: 387 (Aphodius) A: OM SA YE
genus Sugrames Reitter, 1894e: 184 type species Sugrames hauseri Reitter, 1894
Pseudosugrames Kiseritzky, 1928b: 113 type species Sugrames reichardti Kiseritzky, 1928

auriculatus Reitter, 1892e: 181 (Aphodius) A: KZ TM UZ
hauseri Reitter, 1894e: 185 E: ST A: AF KZ TM UZ
uvarovi Semenov & S.I. Medvedev, 1927: 181

johni Kiseritzky, 1928b: 113 A: KZ
reichardti Kiseritzky, 1928b: 114 A: KZ TM

SCARABAEIDAE

153

genus Teuchestes Mulsant, 1842: 176 type species Scarabaeus fossor Linnaeus, 1758
analis Fabricius, 1787a: 8 (Scarabaeus) A: BT CHQ FUJ GUA GUI HKG NP SD XIZ AUR ORR
beckeri Mader, 1938: 56 (Aphodius)
sorex Fabricius, 1792a: 23 (Scarabaeus)

brachysomus Solsky, 1874b: 13 (Aphodius) A: ES FE FUJ GUA HEI JIL NC SC SCH JA WS XIN YUN ZHE
major C.O. Waterhouse, 1875a: 80 (Aphodius)
quinquemaculatus A. Schmidt, 1913b: 177 (Aphodius)

fossor Linnaeus, 1758: 348 (Scarabaeus) E: AB AN AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG GR
HU IR IT KO LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: CI AG A: ES FE KI
KZ MG SC TD TM UZ WS XIN NARi
brunneus Mulsant, 1842: 177 (Aphodius)
femoralis J.R. Sahlberg, 1926: 14 (Aphodius)
fossorius Gistel, 1857b: 599 (Aphodius)
scutellearis Depoli, 1938: 251 (Aphodius)
sylvaticus A. Ahrens, 1812: 33 (Aphodius)
triangulum Piller & Mitterpacher, 1783: 97 (Scarabaeus)

guangdong Rakovič & Mencl, 2012: 296 A: GUA
sinofraternus G. Dellacasa & Johnson, 1983: 528 (Aphodius) A: FUJ GAN GUA GUI GUX JIX SCH XIN ZHE
uenoi Ochi, Kawahara & Kon, 2006: 37 A: FUJ GUA
genus Trichaphodius A. Schmidt, 1913b: 135 type species Aphodius humilis Roth, 1851
andreinii Balthasar, 1939a: 14 (Aphodius) A: SA AFR
foveiventris Raffray, 1877: 326 (Aphodius) A: SA YE AFR
sericeus A. Schmidt, 1909d: 107 (Aphodius)

hindustanicus Balthasar, 1935e: 256 (Aphodius) A: HP NP PA SD UP ORR
jirianus Balthasar, 1965a: 110 (Aphodius)

niloticus Petrovitz, 1967c: 388 (Aphodius) N: EG AFR
genus Trichonotulus Bedel, 1911b: 378 [RN] type species Scarabaeus scrofa Fabricius, 1787
Trichonotus Mulsant, 1842: 294 [HN] type species Scarabaeus scrofa Fabricius, 1787

amdoensis Král, Rakovič & Mencl, 2015: 316 A: QIN
castetsi R. Paulian, 1936b: 363 (Aphodius) A: NP ORR
echinatus Petrovitz, 1961e: 146 (Aphodius)

dzamosanicus Stebnicka, 1973: 119 (Aphodius) A: NC SC
mongolicus Mannerheim, 1852a: 300 (Aphodius) [DA] A: ES FE MG NC NMO
osetinus S.I. Medvedev & Dzhambazishvili, 1977: 669 (Aphodius) E: GG ST
scrofa Fabricius, 1787a: 11 (Scarabaeus) E: AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG GR HU IT LA
LT LU MC MD NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: MO A: AF CY ES FE IN KI KZ MG NC
NMO PA TD TM TR UZ WS NARi
cinereus Mulsant & Rey, 1870b: 218 [=1872c: 194] (Aphodius)

154

SCARABAEIDAE

fuscus P. Rossi, 1792: 8 (Scarabaeus)
merdaroides Reitter, 1896h: 288 (Aphodius)
minutus Herbst, 1789: 269 (Scarabaeus)
setiger Mulsant, 1842: 294 (Trichonotus)
stercoris Gistel, 1857b: 574 (Aphodius)
tomentosus Kugelann, 1792a: 269 (Scarabaeus) [HN]

vultuosus Balthasar, 1971b: 58 (Aphodius) A: KA NP PA
genus Tristaphodius Balthasar, 1932c: 34 type species Aphodius mortuorum Balthasar, 1932
asperulus Červenka, 1994a: 18 (Aphodius) A: GAN
mortuorum Balthasar, 1932c: 35 (Aphodius) A: SCH SD
genus Turanella Semenov, 1905b: 140 [RN] type species Aphodius latevittis Reitter, 1887
Isochirus Reitter, 1892e: 171 [HN] type species Aphodius latevittis Reitter, 1887

latevittis Reitter, 1887d: 509 (Aphodius) E: AB AR A: IN KI KZ TD TM UZ XIN
genus Vladimirellus M. Dellacasa, G. Dellacasa & Bordat, 2002b: 273 type species Aphodius socors
Balthasar, 1967
socors Balthasar, 1967b: 127 (Aphodius) A: WS
genus Volinus Mulsant & Rey, 1870c: 537 [=1871a: 297] type species Scarabaeus sticticus Panzer, 1798
sticticus Panzer, 1798: 4 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB
GE GG GR HU IR IT KO LA LT LU MC MD ME NL NR PL RO SB SK SL SP ST SV SZ TR UK N: CI MO A:
IN TR
clypeolatus Mulsant, 1842: 256 (Aphodius)
confusus Mulsant, 1842: 256 (Aphodius)
equestris Panzer, 1798: 2 (Scarabaeus)
nemoralis Creutzer, 1799b: 1 (Scarabaeus)
occellatus Mulsant, 1842: 256 (Aphodius)
pallescens Mulsant, 1842: 256 (Aphodius)
prolongatus Mulsant, 1842: 256 (Aphodius)
striolatus Mulsant, 1842: 256 (Aphodius)

subtribe Aphodiina, species incertae sedis
beloni Mulsant & Godart, 1879: 123 (Aphodius) A: IQ
clypeatus Fischer von Waldheim, 1821: 11 (Aphodius) A: UZ
compressus Fischer von Waldheim, 1842b: 23 (Aphodius) A: ES
haagi Becker, 1867: 108 (Aphodius) A: ST
marginatus Fischer von Waldheim, 1842a: 11 (Aphodius) A: WS
pedrosi Wollaston, 1854: 226 (Aphodius) N: MR
thoracicus Fischer von Waldheim, 1842b: 23 (Aphodius) A: ES WS

155

SCARABAEIDAE
subtribe Didactyliina Pittino, 1985

genus Didactylia d’Orbigny, 1896b: 247 type species Aphodius pallicolor Fairmaire, 1886
arabica Pittino, 1984b: 310 A: AE OM SA YE
buxtoni R. Paulian, 1939b: 314 A: OM YE AFR
capitulata Clouët des Pesruches, 1898: 186 (Aphodius) N: AG
clementi Peyerimhoff, 1929a: 122 N: AG MO AFR
volkonskyi R. Paulian, 1942c: 76 N: AG AFR
genus Oxycorythus Solsky, 1876a: 395 type species Oxycorythus morawitzi Solsky, 1876
morawitzi Solsky, 1876a: 397 A: KI KZ TD TM UZ
solskyi Wilkins, 1886: 51
solskyi Dokhtouroff, 1887: 345 [HN]

genus Pseudomothon Pittino, 1984b: 311 type species Pseudomothon arabicus Pittino, 1984
arabicus Pittino, 1984b: 312 A: SA
genus Trigonoscelus Petrovitz, 1963d: 633 type species Aphodius coelebs Petrovitz, 1963
afghanus Petrovitz, 1963d: 635 (Aphodius) A: AF IN
coelebs Petrovitz, 1963d: 634 (Aphodius) A: PA
elbursensis Stebnicka & Galante, 1991: 728 A: IN
narayaniensis Ilčíková & Král, 2004: 86 A: NP

tribe Corythoderini A. Schmidt, 1910

genus Chaetopisthes Westwood, 1847b: 242 type species Chaetopistes fulvus Westwood, 1847
subgenus Chaetopisthes Westwood, 1847b: 242 type species Chaetopistes fulvus Westwood, 1847
latipes Tangelder & Krikken, 1982: 52 A: UP
subgenus Neochaetopisthes Wasmann, 1918: 16 type species Chaetopisthes assmuthi Wasmann, 1911
brevipes Wasmann, 1923: 580 A: UP
longulus Wasmann, 1923: 581 A: UP
saetiger Tangelder & Krikken, 1982: 59 A: UP ORR
semisulcatus Tangelder & Krikken, 1982: 65 A: UP
septentrionalis Tangelder & Krikken, 1982: 74 A: KA
genus Termitopisthes Wasman, 1918: 13 type species Chaetopisthes wasmanni A. Schmidt, 1911
forficulus Tangelder & Krikken, 1982: 81 A: UP
kistneri Tangelder & Krikken, 1982: 88 A: UP
parallelus Wasmann, 1923: 578 A: UP
schmidti G.C. Champion, 1923c: 571 A: UP
submissus Tangelder & Krikken, 1982: 83 A: UP
wasmanni A. Schmidt, 1911a: 33 (Chaetopisthes) A: UP ORR

156

SCARABAEIDAE
tribe Eupariini A. Schmidt, 1910

genus Ataenius Harold, 1867d: 100 type species Ataenius scutellaris Harold, 1867
Hexalus Mulsant & Rey, 1870a: 200 [=1872b: 169] type species Hexalus simplicipes Mulsant & Rey, 1870
Pseudammoecius A. Schmidt, 1913b: 160 type species Aphodius australasiae Boheman, 1858

australasiae Boheman, 1858: 50 (Aphodius) A: FUJ HAI JA AUR ORR
cancelliventris Motschulsky, 1858d: 55 (Aphodius)
castanicolor Motschulsky, 1858d: 54 (Aphodius)
fukiensis Balthasar, 1942a: 124
gestroi Harold, 1877a: 87 (Aphodius)
malayanus Petrovitz, 1961f: 354
nigricans R. Paulian, 1933b: 136
okinawensis Nakane, 1960: 5
tweedensis Blackburn, 1904a: 160

brevicollis Wollaston, 1854: 229 (Oxyomus) Ei: AZ PT Ni: CI MR NTR
chopardi R. Paulian, 1937a: 75 (Pleurophorus)
frankenbergeri Balthasar, 1938b: 56
lherminieri R. Paulian, 1947: 43
sulcatula Chevrolat, 1864: 413 (Auperia)

garamas Peyerimhoff, 1929a: 122 N: AG A: OM SA YE AFR
alticola Villiers, 1950: 226 (Pleurophorus)
basilevskyi Endrődi, 1955b: 103 (Saprosites)
kratochvili Balthasar, 1942b: 118
morogorensis Balthasar, 1947: 53

gracilis Melsheimer, 1844: 137 (Oxyomus) Ei: AZ Ai: JA NAR NTR
chilensis Solier, 1851: 72
nocturnus Nomura, 1943: 77

heinekeni Wollaston, 1854: 228 (Oxyomus) Ni: MR NAR NTR
floridanus W.J. Brown, 1930a: 3 (Ataenius)
rhyticephalus Chevrolat, 1864: 413 (Auperia)
vexator Harold, 1869b: 103 (Ataenius)

horticola Harold, 1869f: 429 E: AB AR BU CR GE GR MC ST TR UK (Krym) N: EG A: AF CY IN IQ IS JO LE SY
TD TR
alleoni Fairmaire, 1875b: cxciii (Psammodius)
cypria Baudi di Selve, 1870: 68 (Euparia)
multipunctatus Marseul, 1878: 58 (Psammodius)

maghribinicus Baraud, 1985a: 210 N: AG
picinus Harold, 1867e: 281 Ei: FR IT Ai: JA (Takara-jima) TAI AUR NAR NTR
alegrus Balthasar, 1947: 50
boucomonti R. Paulian, 1937b: 41
darlingtoni Hinton, 1937b: 179
duplopunctatus Lea, 1923: 6

SCARABAEIDAE

157

paracognatus Balthasar, 1961: 123
queroisii R. Paulian, 1934b: 219
rugosus Richards, 1959: 41 (Saprosites)
salutator Fall, 1930: 99

simplicipes Mulsant & Rey, 1870a: 200 (Hexalus) [=1872b: 169] Ei: FR NTR
spretulus Haldeman, 1848: 106 (Aphodius) Ei: PT NAR
consors Fall, 1930: 104 [HN]
falli Hinton, 1934: 119 [RN]

genus Cnematoplatys A. Schmidt, 1913b: 173 type species Cnematoplatys benardi A. Schmidt, 1913
Neosaprosites Endrődi, 1951a: 162 type species Neosaprosites cylindricus Endrődi, 1951

numensis Stebnicka, 1986a: 44 A: NP ORR
genus Parataenius Balthasar, 1961a: 121 type species Parataenius mirabilis Balthasar, 1961 (=Aphodius ­derbesis
Solier, 1851)
Brancoataenius R. Paulian, 1979: 66 type species Ataenius lusitanicus R. Paulian, 1979

simulator Harold, 1868a: 85 (Ataenius) Ei: PT AUR NAR NTR
granuliceps Petrovitz, 1971a: 102
lusitanicus R. Paulian, 1979: 66 (Ataenius)
schwarzi Linell, 1896: 721 (Psammodius)
simulatrix Burmeister, 1877: 412 (Euparia)

genus Saprosites L. Redtenbacher, 1858: 436 type species Saprosites peregrinus L. Redtenbacher, 1858
Cylindropterus Motschulsky, 1864: 464 type species Cylindropterus pleurophoroides Motschulsky, 1864
Passalaphodius Martinez, 1953: 80 type species Passalaphodius cartwrighti Martinez, 1953

arabicus Pittino, 1984b: 316 A: SA
holzschuhi Pittino, 2008: 117 A: NP
japonicus C.O. Waterhouse, 1875a: 93 A: GUA JA NP SC SCH SD TAI YUN ORR
coomani R. Paulian, 1933b: 135
imperfuscus A. Schmidt, 1922: 99
formosensis Nomura, 1939: 36
sumatranus Petrovitz, 1961b: 116

mendax Blackburn, 1892a: 36 (Ataenius) Ei: GB AUR
punctatus Richards, 1959: 45

narae Lewis, 1895c: 382 A: JA ORR
tonkinensis Balthasar, 1933e: 46

natalensis Péringuey, 1901: 449 (Pleurophorus) Ei: GB AFR AUR
nepalensis Pittino, 2008: 118 A: NP
peregrinus L. Redtenbacher, 1858: 437 Ei: AU NTR
yanoi Nomura, 1939: 36 A: TAI

158

SCARABAEIDAE

genus Setylaides Stebnicka, 1994: 75 type species Aulonocnemis monstrosa Harold, 1877
foveatus A. Schmidt, 1909c: 61 (Dialytes) A: JA

tribe Odochilini Rakovič, 1987

genus Odochilus Harold, 1877a: 97 type species Odochilus syntheticus Harold, 1877
subgenus Parodochilus Rakovič, 1997: 163 type species Odochilus balthasari Rakovič, 1987
convexus Nomura, 1971: 64 A: JA (Iriomote)

tribe Psammodiini Mulsant, 1842
subtribe Psammodiina Mulsant, 1842

genus Brindalus Landin, 1960: 52 type species Phycochus azoricus Landin, 1960 (=Aphodius porcicollis
Illiger, 1803)
granulicollis Pittino, 1980b: 352 (Psammodius) N: LB
maderae Pittino, 1983b: 98 (Psammodius) N: MR
porcicollis Illiger, 1803: 195 (Aphodius) E: AL AZ FR GB GR IT MA ME PT SP N: AG EG LB MO TU A: IS CY LE
SY TR
azoricus Landin, 1960: 52 (Phycochus)
rugulosus Mulsant, 1842: 323 (Psammodius)

rotundipennis Reitter, 1892e: 159 (Psammodius) E: IT (Sardegna) SP (Andalucía) N: AG MO TU
rotundicollis d’Orbigny, 1896b: 254 (Psammobius)

schatzmayri Pittino, 1980b: 345 (Psammodius) N: CI MO
genus Diastictus Mulsant, 1842: 318 type species Aphodius vulneratus Sturm, 1805
vulneratus Sturm, 1805: 175 (Aphodius) E: AU BE BH BU CR CT CZ DE EN FR GB GE GR HU IT LA LT NL PL RO
SB SK SL ST SV SZ UK A: WS
latitans Mulsant, 1842: 319
semipunctatus Bonelli, 1812: 155 (Aphodius)

genus Granulopsammodius Rakovič, 1981a: 63 type species Psammodius plicatulus Fairmaire, 1892
centralasiae Rakovič, 1978: 123 (Psammodius) A: KZ TM UZ
loebli Pittino, 1980a: 77 (Psammodius) N: AG MO TU
mesopotamicus Petrovitz, 1971d: 222 (Psammobius) A: IN IQ
mongol Endrődi, 1969b: 295 (Rhyssemus) A: MG
gobiensis S.I. Medvedev, 1976: 160

persicus Rakovič, 1998: 43 A: IN
plicatuloides Pittino, 1984a: 18 (Psammodius) A: XIN
plicatulus Fairmaire, 1892a: 95 (Psammobius) N: AG EG LB MO A: OM SA YE AFR
sinicus Rakovič, 1986: 11 (Psammodius) A: CH
tienshanicus Pittino, 1984a: 20 (Psammodius) A: XIN
transcaspicus Petrovitz, 1961b: 134 (Psammobius) E: ST A: TM UZ

SCARABAEIDAE

159

genus Leiopsammodius Rakovič, 1981a: 16 type species Psammodius laevicollis Klug, 1845
aegialius Ádám, 1986: 13 (Psammodius) A: TR
belloi Pierotti, 1981: 411 (Psammodius) E: GR SB A: TR
desertorum Fairmaire, 1868a: 482 (Aegialia) N: AG LB
gestroi Clouët des Pesruches, 1900: 13 (Psammobius) A: AF AP NP PA SC UP ORR
haruspex Ádám, 1980: 17 (Psammodius) E: GR HU
indicus Harold, 1877a: 100 (Psammodius) A: PA TAI YE AFR ORR
japonicus Harold, 1878a: 69 (Psammobius) A: JA SC
nomurai Tesař, 1944a: 60 (Diastictus)

jelineki Rakovič, 1977b: 318 (Psammodius) A: IN
laevicollis Klug, 1845: [7], pl. xlii (Psammodius) N: AG EG LB MO TU A: IS OM SA SY YE AFR
chobauti d’Orbigny, 1898b: 148 (Psammodius)
pallidus Reitter, 1892e: 161 (Psammodius)

laevis R. Paulian, 1942a: 130 (Psammobius) A: YE AFR
nomurai Masumoto, 2012: 126 A: TAI
pelluscens Petrovitz, 1961b: 133 (Psammobius) A: PA SA ORR
scabrifrons Walker, 1871: 12 (Psammobius) N: EG
strumae Chromý, 1983: 210 (Psammodius) E: BU
genus Psammodius Fallén, 1807: 37 type species Aphodius sulcicollis Illiger, 1802 (=Scarabaeus asper
Fabricius, 1775)
Psammobius Heer, 1841: 531 type species Aphodius sulcicollis Illiger, 1802 (=Scarabaeus asper Fabricius, 1775)

asper Fabricius, 1775: 83 (Scarabaeus) E: AB AR AU BE BU BY CR CT CZ DE EN FI FR GB GE HU IT KZ LA LT
LU MD ME NL NR NT PL RO SK SL ST SV SZ UK A: ES (Minusinsk) IN KZ MG WS
canaliculatus Mulsant, 1842: 321
costatus Stierlin, 1864b: 489
sulcicollis Illiger, 1802b: 20 (Aphodius)

asperoides Pittino, 2007: 99 E: BU GR
basalis Mulsant & Rey, 1870c: 639 [=1871a: 396] (Psammobius) E: AZ BU FR GR IT RO SP ST TR UK A: CY NARi
besucheti Petrovitz, 1975d: 616 (Psammobius) N: AG LB TU
bilyi Rakovič, 1994: 47 A: NP
bulgaricus Mencl, 1982: 310 E: BU
caucasicus Pittino & Shokhin, 2006: 23 E: GG ST A: TR
convexus C.O. Waterhouse, 1875a: 94 A: JA
danubialis Ádám, 1989: 5 E: AU HU SB SK
flavolittoralis J.I. Kim, 1980a: 8 A: SC
generosus Reitter, 1892e: 161 E: AB AR ST A: KZ TM UZ
hangangnensis J.I. Kim, 1980b: 10 A: SC
kobayashii Nomura, 1973b: 43 A: JA (Ryukyus) TAI
kondoi Masumoto, 1984a: 81

160

SCARABAEIDAE

laevipennis A. Costa, 1844: 18 E: AB AL AZi BH BU CR FR GR HU IT MC PT RO SB SK SL SP ST TR UK N:
AG MO TU A: CY IS JO LE SY TR NARi
ciliatus Küster, 1849b: 51
rugicollis Erichson, 1848a: 916
scutellaris Mulsant & Wachanru, 1859a: 521 [=1859b: 187]

maceki Rakovič, 1994: 49 A: NP
macnamarae Pittino, 1984a: 15 A: NP UP
maruyamai Ochi, Kawahara & Inagaki, 2011: 69 A: JA
nepalensis Balthasar, 1971a: 18 A: NP
nocturnus Reitter, 1892e: 160 E: IT N: AG TU A: CY IS LE TR
brevior Normand, 1936: 374
lacoi Roubal, 1929a: 32

pierottii Pittino, 1979: 598 E: AL AU FR GE GR HU IT MA SK SP SZ TR N: AG TU NARi
plicicollis Erichson, 1848a: 916 E: FR GR IT PT N: AG
accentifer Mulsant & Rey, 1859a: 236 [=1859b: 172]
insculptus Mulsant & Rey, 1870c: 638 [=1871a: 398]
planipennis Reitter, 1892a: 224 [=1893a: 103]

sefrensis Petrovitz, 1961b: 130 (Psammobius) N: AG TU
seiferti Rakovič, 1994: 50 A: NP
sungshinarum J.I. Kim, 1980a: 7 A: SC
tesari Rakovič, 1977b: 317 A: NP PA ORR
thailandicus Balthasar, 1965c: 445 (Psammobius) A: JA (Ryukyus) ORR
genus Rakovicius Pittino, 2006a: 13 type species Psammobius ainu Lewis, 1895
ainu Lewis, 1895c: 384 (Psammodius) A: JA
coreanus J.I. Kim, 1980b: 9 (Psammodius) A: JA (Tsushima) SC
lanae Masumoto, 2012: 126 A: TAI
subopacus Nomura, 1973b: 44 (Psammodius) A: TAI
kawaii Pittino, 2006a: 17

genus Sicardia Reitter, 1896i: 318 type species Sicardia psammodiformis Reitter, 1896
Bipammosus Normand, 1938b: 340 type species Bipammosus boiteli Normand, 1938

boiteli Normand, 1938b: 340 (Bipammosus) N: AG
pierrei Hollande & Thérond, 1999: 235 (Psammobius)

psammodiformis Reitter, 1896i: 318 N: LB TU
genus Tesarius Rakovič, 1981b: 27 type species Phycochus sulcipennis Lea, 1904
caelatus LeConte, 1857b: 42 (Aegialia) Ei: GB NAR
mcclayi Cartwright, 1955: 448 (Psammodius) Ei: GB NAR

161

SCARABAEIDAE
subtribe Rhyssemina Pittino & Mariani, 1986

genus Bordatius Pittino & Mariani, 1986: 148 type species Bordatius capensis Pittino & Mariani, 1986
tingitanus Pittino & Mariani, 1986: 153 N: MO
genus Messyrhus Rakovič & Král, 1997: 235 type species Messyrhus brachypterus Rakovič & Král, 1997
brachypterus Rakovič & Král, 1997: 235 A: QIN
genus Myrhessus Balthasar, 1956b: 425 type species Myrhessus mirabilis Balthasar, 1956
afghanus Balthasar, 1939b: 42 (Rhyssemus) A: AF TD
kubotai Ochi, Kawahara & Inagaki, 2012: vi A: JA (Yaku-shima)
mirabilis Balthasar, 1956b: 426 A: AF TD
nishinoi Ochi, Kawahara & Inagaki, 2012: v A: JA (Honshū)
samurai Balthasar, 1941a: 92 (Rhyssemus) A: JA
tuberculatus Nomura & Nakane, 1951: 45 (Rhyssemus)

yorikoae Ochi, Kawahara & Inagaki, 2006a: 143 A: JA
genus Neotrichiorhyssemus Rakovič & Král, 1997: 239 type species Trichiorhyssemus hirsutus Clouët des
Pesruches, 1901
esakii Nomura, 1943: 80 (Trichiorhyssemus) A: JA
boninensis Nakane, 1960: 5 (Trichiorhyssemus)

hauseri Balthasar, 1933f: 116 (Rhyssemus) A: TAI ORR
hirsutus Clouët des Pesruches, 1901: 35 A: TAI AUR ORR
kentingensis Masumoto, Ochi & Lan, 2013: 243 A: TAI
genus Pararhyssemus Balthasar, 1956b: 423 type species Rhyssemus coluber Mayet, 1887
coluber Mayet, 1887a: lxxxix (Rhyssemus) N: AG EG MO TU A: AE AF IN SA SI TM YE AFR ORR
caesoides Reitter, 1894e: 184 (Rhyssemus)
laevifrons Fairmaire, 1892a: 94 (Rhyssemus)
paradoxus Balthasar, 1956b: 424
parisii Balthasar, 1939a: 25 (Rhyssemus)

genus Pittinius Rakovič & Král, 1997: 242 type species Pittinius omnisetosus Rakovič & Král, 1997
farkaci Rakovič, Král & Mencl, 2014: 578 A: NP
omnisetosus Rakovič & Král, 1997: 242 A: NP UP
genus Platytomus Mulsant, 1842: 310 type species Platytomus sabulosus Mulsant, 1842 (=Aphodius tibialis
Fabricius, 1798)
jailensis Apostolov & Maltzev, 1986: 88 (Pleurophorus) E: UK (Krym)
laevistriatus Perris, 1870: 13 (Psammodius) E: FR IT (Sardegna, Sicilia) MA N: AG TU
poricollis Fairmaire, 1871a: 374 (Psammodius)

162

SCARABAEIDAE

mongolicus S.I. Medvedev, 1974: 113 (Pleurophorus) A: BEI FE (“Siberia, Amur reg.”) KZ MG XIN
tibialis Fabricius, 1798: 24 (Scarabaeus) E: AL AZi BH CR FR GR IT MA RO N: AG CI EG LB MO MR TU A: CY
IS LE SY TR AUR NAR
insculptus Küster, 1849b: 49 (Psammodius)
ovipennis Desbrochers des Loges, 1884: 98
sabulosus Mulsant, 1842: 310

variolosus Kolenati, 1846: 20 (Oxyomus) E: AB AR KZ ST UK (Krym) A: AF FE (“Siberia, Amur reg.”) KZ TD TM
TR UZ XIN
afghanus Balthasar, 1956b: 423 (Pleurophorus)
foveicollis Ballion, 1871a: 333 (Psammobius)

yadai Ochi, Kawahara & Inagaki, 2006b: 31 (Leiopsammodius) A: JA (Okinawa-shotō) QA
qatarensis Bordat, 2007: 80

genus Pleurophorus Mulsant, 1842: 312 type species Scarabaeus caesus Panzer, 1796
akamasicus Pittino & Miessen, 2007: 75 A: CY
anatolicus Petrovitz, 1961g: 394 E: AB AR GR (Ródos) TR A: CY IN IQ IS JO LE SY TM TR
apicipennis Reitter, 1892e: 167 A: AF IN KI KZ PA TD TM TR UZ XIN
arabicus Pittino & Mariani, 1986: 76 E: AB ST A: IN IQ JO OM PA SA TM
caesus Panzer, 1796: no. 35: 2 (Scarabaeus) E: AB AU AZ BE BH BU CR CZ FR GB GE GG GR HU IT KZ LA LU
MA MC MD ME PL PT RO SB SK SL SP ST SV SZ TR UK N: AG CI EG MO MR TU A: CY IS JA KZ TR UZ AFRi
NARi NTRi
elongatulus Mulsant, 1842: 312

formosanus Pittino & Mariani, 1986: 120 A: TAI
ledouxi Pittino & Mariani, 1986: 128 A: AF
maghrebinicus Pittino & Mariani, 1986: 89 E: GR IT (Sicilia) N: AG MO TU
mediterranicus Pittino & Mariani, 1986: 82 E: BE BH FR GE GR IT PT SP N: AG MO TU
nepalensis Pittino & Mariani, 1986: 131 A: NP
opacus Reitter, 1892a: 224 [=1893a: 103] E: IT N: AG LB MO TU
pannonicus Petrovitz, 1961g: 394 E: AB AU BE BH BU CR CZ FR GR HU IT MC RO SL SK ST SZ UK A: CY IN
TR UZ
tonkinensis Balthasar, 1933e: 47 A: TAI ORR
torretassoi Schatzmayr, 1930: 111 N: AG LB TU
genus Rhyssemodes Reitter, 1892e: 161 type species Rhyssemus tenuisculptus Reitter, 1892
alutaceus Reitter, 1892e: 162 A: KI KZ TM UZ
bouvieri Clouët des Pesruches, 1901: 22 A: BEI MG NMO TIA
malyi Rakovič, 1982: 10 A: MG
nox Iablokoff-Khnzorian, 1971: 185 A: TM
orientalis Mulsant & Godart, 1875a: 75 [=1875b: 411] (Rhyssemus) E: AB AR SP N: AG CI EG LB MO TU A: AF IN
IS KZ LE MG SA SI TD TM TR UZ YE AFR
aspericeps Chevrolat, 1861b: 266 (Rhyssemus)

SCARABAEIDAE

163

gemmifer Marseul, 1878: 58 (Rhyssemus)
hybridus Reitter, 1892e: 167 (Rhyssemus)
obsoletus Reitter, 1887b: 227 (Rhyssemus)
parvus Balthasar, 1963d: 289
reitteri d’Orbigny, 1896b: 254
transversus Reitter, 1892e: 162

sindicus Pittino, 1984a: 22 A: BEI HEN IN NP PA TAI UP YUN ORR
tenuisculptus Reitter, 1892e: 162 A: AF KZ TM UZ
opacus Reitter, 1892a: 223 [=1893a: 102]

transcaspicus Rakovič, 1982: 15 E: ST A: UZ
turcicus Pittino & Rakovič, 1984: 167 A: TR
wittmeri Pittino, 1984b: 325 A: SA
genus Rhyssemus Mulsant, 1842: 314 type species Scarabaeus asper Fabricius, 1775 sensu Mulsant, 1842
(=Ptinus germanus Linnaeus, 1767)
ahrensi Rakovič & Král, 2001: 348 A: NP
algiricus P.H. Lucas, 1846a: pl. 24 E: IT SP N: AG MO TU
annaedicatus Pierotti, 1980: 22 E: AL BU GR IT MC ME A: TR SY
asperocostatus Fairmaire, 1892a: 94 A: SA YE AFR
bedeli Clouët des Pesruches, 1901: 94 N: AG TU AFR
berytensis Marseul, 1878: 57 E: BU GR (Makedonia) A: CY IN IQ IS LE TR
mediterranicus Petrovitz, 1971e: 568

brevitarsis Pittino, 1984b: 332 A: OM SA YE
buettikeri Pittino, 1984b: 334 A: SA YE
cantabricus Balthasar, 1961b: 125 E: SP
chinensis Pittino, 1996: 63 A: XIN
convexus Bénard, 1911b: 296 E: SP (Andalucía) A: MO
seminulum Balthasar, 1945b: 110

exaratus Marseul, 1878: 57 A: SA YE N: EG AFR
falcatus Petrovitz, 1961c: 113 A: KA
feae Clouët des Pesruches, 1901: 91 A: TAI ORR
lunatus Petrovitz, 1963a: 44

freudei Balthasar, 1960a: 5 A: NP PA UP ORR
germanus Linnaeus, 1767: 566 (Ptinus) E: AB AL AR AU BE BH BU BY CR CT CZ DE FR GE GG GR HU IT KZ
LA LT LU MC MD ME NL NT PL PT RO SB SK SL SP ST SV SZ TR UK N: MO TU A: CH FE IN KZ MG SY TD
TM TR UZ NARi
funebris Roubal, 1912b: 155
geminatus Reitter, 1890s: 390
obsoletus Rey, 1890a: 171
pertinax Balthasar, 1961b: 127

granosus Klug, 1842: 7 (Oxyomus) N: EG A: AE OM QA SA YE AFR

164
grossepunctatus Balthasar, 1961b: 126 N: AG
hamatus Petrovitz, 1963a: 43 A: NP PA ORR
helenae Pittino, 1983a: 114 A: PA
histrio Balthasar, 1961b: 128 E: AB A: AF IN TM
histrioides Petrovitz, 1963a: 36 A: AF
indicus Clouët des Pesruches, 1901: 82 A: NP UP ORR
inscitus Walker, 1858: 207 (Psammodius) A: IN NP PA UP TAI AFR AUR ORR
insularis Pittino, 1984a: 28 E: GR A: CY
interruptus Reitter, 1892a: 165 E: AB AR ST A: KI KZ TD UZ
reitteri D. Koshantschikov, 1894c: 117

koreanus Stebnicka, 1980a: 248 A: NC
limbolarius Petrovitz, 1963a: 33 E: AU FR IT SL
arvernensis Baraud, 1976c: 403
celejensis Petrovitz, 1967c: 401

macedonicus Bénard, 1923: 243 E: GR (Makedonia) A: IN TR
marqueti Reiche, 1863: 76 E: FR SP
pyrenaeus Balthasar, 1961b: 124

mayeti Clouët des Pesruches, 1901: 110 N: EG AFR
meridionalis Reitter, 1890s: 391 E: GR “Caucasus” A: TR
caucasicus Clouët des Pesruches, 1901: 91

mesopotamicus Petrovitz, 1963a: 34 A: IQ
morgani Bénard, 1911a: 214 E: AB A: IN
murghabensis Balthasar, 1967a: 110 (Myrhessus) A: AF
nanshanchicus Masumoto, 1977: 30 A: TAI
obliviosus Petrovitz, 1963a: 41 A: TM
osmanlis W. Koshantschikov, 1916: 204 A: TR
parallelus Reitter, 1892e: 166 E: IT (Sardegna, Sicilia) N: AG MO TU
perissinottoi Pittino, 1983a: 106 A: IN
plicatus Germar, 1817: 184 (Aphodius) E: AL BH CR FR GR IT MA SP N: AG MO TU
arenarius A. Costa, 1844: 17 (Oxyomus)
kocheri Petrovitz, 1967a: 232
sulcigaster Mulsant & Rey, 1959a: 238 [=1859b: 174]

ponticus Petrovitz, 1962b: 112 A: TR
psammobiiformis Petrovitz, 1963a: 39 E: PT SP
leonensis Petrovitz, 1963a: 41

punctatissimus Pittino, 1984b: 49 A: SA YE AFR
puncticollis Brown, 1929: 91 E: AU CZ GE LA PL NARi
purkynei Balthasar, 1961b: 128 E: SP (Valencia)
ressli Petrovitz, 1965a: 167 A: TR
rubeolus Harold, 1871a: 25 A: SA YE (Suqutra) AFR
arabicus Rakovič, 1982: 11 (Rhyssemodes)

SCARABAEIDAE

165

SCARABAEIDAE
saoudi Pittino, 1984b: 336 A: SA YE
sardous Pierotti, 1980: 23 E: FR (Corse) IT (Sardegna)
sinuaticollis Pittino, 1984a: 54 A: HP ORR
sulcatus A.G. Olivier, 1789: no 4: 11 (Trox) E: FR IT SP N: AG MO TU
corrugatulus Reitter, 1892e: 164
costipennis Reitter, 1894e: 183
godarti Mulsant, 1846: suppl. 7

syriacus Petrovitz, 1971d: 220 A: JO SY
thailandicus Pittino, 1996: 66 A: GUX ORR
vaulogeri Clouët des Pesruches, 1901: 84 A: AG TU
vinodolensis Petrovitz, 1963a: 39 E: CR GR
verrucosus Mulsant, 1842: 316 E: FR IT SP
xerxes Pittino, 1983a: 112 A: IN
genus Trichiorhyssemus Clouët des Pesruches, 1901: 15 type species Rhyssemus riparius Horn, 1871
adhabharicus Pittino, 1983a: 116 (Rhyssemus) A: NP
asperulus C.O. Waterhouse, 1875a: 94 (Rhyssemus) A: JA SC
dalmatinus Petrovitz, 1967c: 398 E: CR IT
elegans Petrovitz, 1963a: 35 (Rhyssemus) A: IN
harmoziensis Petrovitz, 1980: 612 (Rhyssemus)

fokiensis Petrovitz, 1968a: 6 A: FUJ
kitayamai Ochi, Kawahara & Kawai, 2001: 51 A: JA (Ryukyus) TAI
lasionotus Clouët des Pesruches, 1901: 26 A: TAI TIA
microtrichius Ochi., Kawahara & Inagaki, 2011: 72 A: JA
nepalensis Pittino, 1983a: 119 (Rhyssemus) A: NP
setulosus Reitter, 1892e: 163 (Rhyssemus) E: IT (Sicilia)
taiwanus Ochi, Masumoto & C.-F. Lee, 2015: 331 A: TAI
yumikoae Pittino & Kawai, 2007: 67 A: FUJ (Namoa Island) JA (Okinawa-shotō) TAI

tribe Rhyparini A. Schmidt, 1910

genus Lioglyptoxenus Pittino, 2006b: 93 type species Lioglyptoxenus mirabilis Pittino, 2006
mirabilis Pittino, 2006b: 96 A: NP
genus Rhyparus Agassiz, 1846: 328 type species Rhyparus helephoroides Fairmaire, 1893
Ryparus Westwood, 1845b: 93 [HN] type species Ryparus desjardinsi Westwood, 1843

azumai azumai Nakane, 1956b: 122 A: JA
azumai loebli R. Paulian, 1983: 618 A: TAI
chinensis Balthasar, 1953b: 230 A: FUJ
helophoroides Fairmaire, 1893b: 145 A: JA TAI ORR
amamianus Nakane, 1956b: 123
australiae Lea, 1923: 19

166

SCARABAEIDAE

orousseti R. Paulian, 1981: 111
risbeci R. Paulian, 1934b: 220

kitanoi kitanoi Miyake, 1982a: 65 A: JA
kitanoi taiwanus Ochi, 2001: 3 A: TAI
klapperichorum R. Paulian, 1983: 620 A: TAI
nepalensis Balthasar, 1971a: 19 A: NP
peninsularis Arrow, 1905: 537 A: TAI ORR
schachti Balthasar, 1971a: 21 A: NP
genus Sybacodes Fairmaire, 1896a: 84 type species Sybacodes lutulentus Fairmaire, 1896
simplicicollis Fairmaire, 1896a: 84 A: BT YUN ORR

subfamily Aulonocneminae A. Janssens, 1946

genus Palnia Stebnicka, 1985: 650 type species Palnia loebli Stebnicka, 1985
cambeforti Král, 1999a: 468 A: YUN

subfamily Chironinae C.É. Blanchard, 1845

genus Chiron W.S. Macleay, 1819: 107 type species Scarites cylindrus Fabricius, 1798
cylindrus Fabricius, 1798: 44 (Scarites) Ei: FR IT (Sicilia) A: HP NP PA UP ORR
cylindricus Fabricius, 1799: 45 (Scarites)
digitatus Fabricius, 1801b: 377 (Sinodendron)

genus Theotimius Huchet, 2000: 21 type species Chiron grandis Gory, 1830
subgenus Theotimius Huchet, 2000: 21 type species Chiron grandis Gory, 1830
grandis Gory, 1830: pl. 22 (Chiron) N: EG AFR

subfamily Scarabaeinae Latreille, 1802
tribe Ateuchini Perty, 1830

genus Delopleurus Erichson, 1847a: 761 type species Delopleurus pullus Boheman, 1857
mencli Král, 2014: 134 A: YE (Suqutra)
parvus Sharp, 1875a: 47 (Coptorhina) A: NP “India bor.” ORR
cardoni R. Paulian, 1934c: 162

striatus Arrow, 1931: 411 A: UP
genus Onychothecus Boucomont, 1912b: 275 type species Onychothecus ateuchoides Boucomont, 1912
ateuchoides Boucomont, 1912b: 276 A: YUN ORR
corniclypeus Chang, 1980a: 303 A: YUN ORR
genus Parachorius Harold, 1873: 103 type species Parachorius thomsoni Harold, 1873
fungorum Kryzhanovskij & S.I. Medvedev, 1966: 394 A: YUN
globosus Arrow, 1931: 360 A: SD
hookeri Arrow, 1931: 359 A: SD

167

SCARABAEIDAE
krali Utsunomiya & Masumoto, 2001: 125 A: YUN
thomsoni Harold, 1873: 103 A: FUJ NP PA SD YUN ORR
genus Paraphytus Harold, 1877a: 42 type species Paraphytus doriae Harold, 1877
Maraxes Lewis, 1895c: 376 type species Maraxes dentifrons Lewis, 1895
Xynophron Harold, 1880c: 199 type species Xynophron ritsemae Harold, 1880

dentifrons Lewis, 1895c: 376 (Maraxes) A: JA (Ryukyus) ORR
hindu Arrow, 1913b: 193 A: AP NP ORR
ritsemae Harold, 1880c: 201 A: AP AFR ORR

tribe Canthonini van Lansberge, 1875

genus Cassolus Sharp, 1875a: 40 type species Cassolus nudus Sharp, 1875
fukiensis Balthasar, 1960c: 90 A: FUJ ORR
gotoi Masumoto, 1986a: 85 A: TAI
humeralis Arrow, 1907b: 416 A: SD YUN ORR
nudus Sharp, 1875a: 40 A: FUJ YUN ORR
quadridentatus Balthasar, 1952: 223 (Panelus)

genus Ochicanthon Vaz de Mello, 2003: 25 type species Phacosoma dytiscoides Boucomont, 1914
Phacosoma Boucomont, 1914b: 249 [HN] type species Phacosoma dytiscoides Boucomont, 1914

obscurum Boucomont, 1920b: 307 (Phacosoma) A: AP ORR
thailandicus Masumoto, 1989: 32 (Phacosoma) A: YUN ORR
tristoides R. Paulian, 1983: 616 (Phacosoma) A: SD
genus Panelus Lewis, 1895c: 378 type species Temnoplectron parvulum C.O. Waterhouse, 1874
assamensis Arrow, 1907b: 417 A: SD ORR
crenatus Nomura, 1973b: 40 A: TAI
deuvei R. Paulian, 1992: 122 A: NP
kubotai Kawahara, Inagaki & Ochi, 2007: 61 A: JA (Yaku-shima)
maedai Nomura, 1973b: 40 A: TAI
manmiaoae Masumoto, Ochi & Tsai, 2011: 82 A: TAI
ovatus Nomura, 1973a: 13 A: JA (Tokara-rettō)
parvulus C.O. Waterhouse, 1874b: 175 (Temnoplectron) A: JA SC
rufulus Nomura, 1973a: 13 A: JA (Ishigaki-jima)
tonkinensis R. Paulian, 1939a: 69 A: YUN ORR
wangi Masumoto, Ochi & Tsai, 2011: 80 A: TAI
watanabei Masumoto & Yin, 1993: 85 A: YUN

tribe Coprini Leach, 1815

genus Catharsius Hope, 1837: 21 type species Scarabaeus molossus Linnaeus, 1758
birmanensis van Lansberge, 1874a: 11 A: BT NP SD UP ORR
capucinus Fabricius, 1781a: 25 (Scarabaeus) A: NP PA ORR

168

SCARABAEIDAE

granulatus Sharp, 1875a: 41 A: NP PA SD ORR
javanus van Lansberge, 1886a: 7 A: FUJ SCH TAI ORR
molossus Linnaeus, 1758: 347 (Scarabaeus) A: AF AP BEI CHQ FUJ GUI HEB HEN HKG HP NP PA SD SCH SHG
TAI UP YUN XIZ ZHE ORR
abbreviatus Herbst, 1789: 53 (Scarabaeus)
berbicaeus Herbst, 1789: 228 (Scarabaeus)
kangeanus R. Paulian, 1936c: 395
timorensis van Lansberge, 1879: cxlviii
ursus Fabricius, 1801a: 43 (Copris)

pithecius Fabricius, 1775: 21 (Scarabaeus) A: AP CE HP NP PA UP ORR
cribricollis Walker, 1858: 208 (Copris)
nanus Fabricius, 1792a: 42 (Scarabaeus)
sabaeus Fabricius, 1781a: 23 (Scarabaeus)
sinensis Hope, 1842a: 60 [=1843a: 62, 1845: 6] (Copris)

platypus Sharp, 1875a: 42 A: PA ORR
nathani Frey, 1956: 655

sagax Quensel, 1806: 43 (Copris) A: BT HP NP UP ORR
sesostris C.O. Waterhouse, 1888a: 86 N: EG A: IS JO SA SI SY AFR
pylades Péringuey, 1901: 325

genus Copris Geoffroy, 1762: 87 type species Scarabaeus lunaris Linnaeus, 1758
subgenus Copris Geoffroy, 1762: 87 type species Scarabaeus lunaris Linnaeus, 1758
Insulicopris Tsukamoto, 1974: 1 type species Copris brachypterus Nomura, 1964
Litocopris C.O. Waterhouse, 1891a: 53 type species Litocopris punctiventris C.O. Waterhouse, 1891
Pilularius Schrank, 1798 type species Pilularius lunus Schrank, 1798 (=Scarabaeus lunaris Linnaeus, 1758

acutidens Motschulsky, 1861a: 13 A: FUJ HUB JA SCH
frankebergeri Balthasar, 1934c: 149
glabriceps Felsche, 1910: 346
muelleri Balthasar, 1939b: 42

angusticornis Arrow, 1933: 427 A: CHQ FUJ HUB SCH YUN ORR
armeniacus Faldermann, 1835a: 238 E: AR A: IN TR
felschei Reitter, 1892a: 243 [=1893a: 95]

arrowi Felsche, 1910: 350 A: “South China” ORR
bengalensis Gillet, 1911a: 289 A: GUI XIZ YUN ORR
brachypterus brachypterus Nomura, 1964a: 50 A: JA (Amami-Ôshima)
brachypterus toguchii Ochi & Kawahara, 2012: i A: JA (Tokunoshima)
cariniceps Felsche, 1910: 348 A: TAI (Lanyu) “South China”
carinicus Gillet, 1910: 8 A: YUN ORR
confucius Harold, 1877a: 48 A: FUJ GUA GUI HKG SCH YUN ORR
corpulentus Gillet, 1910: 13 A: AP FUJ SCH YUN ORR
davidis Deyrolle, 1878: 95 (Catharsius) A: HUB SCH YUN

SCARABAEIDAE

169

delicatus Arrow, 1931: 117 A: AP SD YUN ORR
frankenbergerianus Balthasar, 1958: 477 A: SCH
fricator Fabricius, 1787a: 15 (Scarabaeus) A: PA ORR
indicus Gillet, 1910: 5

fukiensis Balthasar, 1953b: 225 A: FUJ TAI ZHE
hispanus cavolinii V. Petagna, 1792: 140 (Scarabaeus) E: AB AL AR AU BH BU CR GR IT MC ME ST UK A: AF CY
IN IS JO KI KZ PA SY TD TM TR UZ
persianus Mikšić, 1958a: 1
sinuatus Mulsant, 1842: 68
tridens Mulsant, 1842: 69

hispanus hispanus Linnaeus, 1764: 12 E: FR IT (Sardegna, Pantelleria) PT SP N: AG EG LB MO TU AFR
hispanicus Poiret, 1789: 297 (Scarabaeus)
laevicollis Mulsant, 1842: 69
marianus Petiver, 1764: 27 (Scarabaeus)
paniscus Fabricius, 1775: 24 (Scarabaeus)
pseudolunaris Ferreira, 1969: [pl. 1, 1]
retusus Mulsant, 1842: 68

illotus Balthasar, 1942c: 189 A: FUJ
inaequabilis Y.-W. Zhang, 1997: 758 A: HUB
klapperichi Balthasar, 1942a: 116 A: FUJ
lunaris Linnaeus, 1758: 346 (Scarabaeus) E: AB AL AR AU BH BU BY CR CT DE EN FR GB GE GG GR HU IT KO
KZ LA LT LU MC MD ME NL NR PL PT RO SB SK SL SP ST SV SZ TR UK A: IN KI KZ TD TR UZ XIN AURi
belisama Schrank, 1798: 394 (Pilularius)
bifidus Poda von Neuhaus, 1761: 18 (Scarabaeus)
castaneus Mulsant, 1842: 73
corniculatus Mulsant, 1842: 73
deletus Mulsant, 1842: 73
emarginatus A.G. Olivier, 1789: no 3: 115 (Scarabaeus)
jenisonianus Gistel, 1857b: 586
gistelianus Gistel, 1857b: 587
lunus Schrank, 1798: 394 (Pilularius)
obliteratus Mulsant, 1842: 73

magicus Harold, 1881: 88 A: AP CHQ GUI HUB HUN NP SCH SD XIN YUN ORR
manni Ochi, Kon & Bai, 2009: 211 A: GUX SCH SHX
medogensis Y.-W. Zhang, 1988: 228 A: XIZ
numa van Lansberge, 1886a: 19 A: AP (Mishmi Hills) ORR
potanini Semenov, 1891: 310 A: GAN
pseudosinicus Balthasar, 1958: 475 A: CH
pueli Mollandin de Boissy, 1905: 110 N: AG TU
punctatus formosanus Gillet, 1921: 126 A: TAI
punctatus punctatus Gillet, 1910: 4 A: FUJ SCH ORR

170

SCARABAEIDAE

repertus Walker, 1858: 208 A: AP HP SD NP UP YUN ORR
claudius Harold, 1877a: 48

ruricola Balthasar, 1933d: 266 A: SCH
sarpedon Harold, 1868c: 104 [RN] A: KA HP NP PA UP ORR
pompilius C.O. Waterhouse, 1875a: 75
sexdentatus Kollar & L. Redtenbacher, 1844: 520 [HN]

schoolmeestersi Ochi & Kon, 2010: 127 A: NP
siangensis Biswas, 1979b: 339 A: AP
sinicus Hope, 1842a: 60 [=1843a: 62, 1845: 6] A: FUJ HUB JIX NP SCH SHG YUN ORR
sulcicollis van Lansberge, 1886a: 10

sorex Balthasar, 1942a: 117 A: FUJ ORR
subdolus Balthasar, 1958: 476 A: FUJ
szechouanicus Balthasar, 1958: 479 A: CHQ FUJ GUI HUN SCH ZHE
tripartitus C.O. Waterhouse, 1875a: 74 A: CHQ FUJ HEI JA NC SC SCH YUN ORR
umbilicatus Abeille de Perrin, 1901b: 68 E: AL BU CR FR GR HU IT RO SK
zhangi Ochi, Kon & Bai, 2009: 211 A: GUI QIN SCH
subgenus Sinocopris Ochi, Kon & Bai, 2009: 207 type species Copris uenoi Hanboonsong, Ochi & Masumoto,
2007
barclay Ochi, Kon & Bai, 2009: 209 A: YUN ORR
basipunctatus Balthasar, 1942a: 115 A: FUJ
cheni Ochi, Kon & Bai, 2007: 137 A: XIZ
laevigatus Gillet, 1927: 252 A: SCH YUN ORR
obenbergeri Balthasar, 1933d: 263 A: HEN SCH YUN
ochus Motschulsky, 1861a: 13 (Catharsius) A: CE FE FUJ GAN GUA HEB HEI HEN JA NE NMO NC NS SC ZHE
“Manchuria”
pecuarius continentalis Ochi & Kon, 2010: 130 A: NC
pecuarius pecuarius Lewis, 1884: 17 A: FE JA
yamamotoi Tesař, 1937: 101

pseudochus Ochi & Kon, 2004: 343 A: GUI
quasilaevigatus Ochi, Kon & Bai, 2007: 142 A: SCH XIZ
sabinus Gillet, 1910: 17 A: HP NP UP ORR
sacontala Kollar & L. Redtenbacher, 1844: 519 A: HP KA NP SD UP XIZ ORR
yangi Ochi, Kon & Bai, 2007: 140 A: SCH XIZ
genus Heliocopris Hope, 1837: 23 type species Scarabaeus gigas Linnaeus, 1758
bucephalus Fabricius, 1775: 24 (Scarabaeus) A: AF HP HAI NP SCH YUN ORR
dominus Bates, 1868: 88 A: AP PA ORR
mouhotus Sharp, 1878a: 155

gigas Linnaeus, 1758: 348 (Scarabaeus) N: EG A: OM QA SA YE AFR
antenor Péringuey, 1901: 312
isidis Laporte, 1840: 76 (Copris)

midas Fabricius, 1775: 21 (Scarabaeus) A: HP IN PA UP ORR

171

SCARABAEIDAE
genus Metacatharsius Montreuil, 1998: 137 type species Scarabaeus ferrugineus A.G. Olivier, 1789
inermis Laporte, 1840: 78 (Copris) N: EG A: AE IN IQ KU OM PA SA YE AFR ORR
genus Microcopris Balthasar, 1958: 474 type species Scarabaeus reflexus Fabricius, 1787
apicepunctatus Balthasar, 1942c: 190 (Copris) A: FUJ
propinquus Felsche, 1910: 347 (Copris) A: FUJ SCH TAI YUN ORR
minutissimus Kryzhanovskij & S.I. Medvedev, 1966: 392 (Copris)

reflexus Fabricius, 1787a: 16 (Scarabaeus) A: CHQ FUJ HAI HUB SCH YUN ORR
oryctes Herbst, 1789: 215 (Scarabaeus)

vitalisi Gillet, 1921: 124 (Copris) A: SCH YUN ORR
genus Paracopris Balthasar, 1939a: 2 type species Copris punctulatus Wiedemann, 1823
andrewesi C.O. Waterhouse, 1891b: 521 (Copris) A: NP ORR
cariniceps Felsche, 1910: 348 (Copris) A: HKG JIA JIX SCH SHG TAI YUN ORR
compressipennis Gillet, 1910: 14 (Copris) A: SD ORR
excisus C.O. Waterhouse, 1891b: 521 (Copris) A: HP PA UP ORR
furciceps Felsche, 1910: 348 (Copris) A: AP SD ORR
imitans Felsche, 1910: 347 (Copris) A: NP ORR
punctulatus Wiedemann, 1823: 11 (Copris) A: AP YUN ORR
punjabensis Gillet, 1921: 126 (Copris) A: HP NP PA UP ORR
surdus Arrow, 1931: 132 (Copris) A: UP ORR
genus Synapsis Bates, 1868: 89 type species Copris brahminus Hope, 1831
Homalocopris Solsky, 1871b: 136 type species Ateuchus tmolus Fischer von Waldheim, 1821

birmanica Gillet, 1907: 600 A: AP YUN ORR
brahmina Hope, 1831: 22 (Copris) A: BU NP SD ORR
batesi Sharp, 1875a: 43

davidis Fairmaire, 1878a: 96 A: CHQ FUJ GAN GUI HUB SHA SCH TAI
gilleti Arrow, 1931: 83 A: NP SD
masumotoi Ochi, 1992: 9 A: TAI
naxiorum Král & Rejsek, 2000: 268 A: YUN
simplex Sharp, 1875a: 45 A: YUN ORR
strnadi Král, 2002b: 283 A: YUN ORR
tmolus Fischer von Waldheim, 1821: 11 (Ateuchus) A: IN KI KZ TD TM UZ XIN
tridens Sharp, 1881: xcii A: GUI SD SCH YUN ORR
yunnana Arrow, 1933: 428

tribe Gymnopleurini Lacordaire, 1856

genus Allogymnopleurus A. Janssens, 1940b: 9 type species Gymnopleurus chloris Klug, 1855 (=Gymnopleurus
splendidus Bertoloni, 1849)
anthracinus Klug, 1845: [2], pl. xli (Gymnopleurus) A: SA AFR
hildebrandti Harold, 1875d: 218 (Gymnopleurus)

172

SCARABAEIDAE

maculosus W.S. Macleay, 1821: 517 (Scarabaeus) A: NP UP ORR
exanthema Wiedemann, 1823: 22 (Gymnopleurus)

genus Garreta A. Janssens, 1940b: 9 type species Ateuchus azureus Fabricius, 1801
dejeanii Laporte, 1840: 70 (Gymnoleurus) A: HP PA UP ORR
inconspicuus C.O. Waterhouse, 1890b: 371 (Gymnoleurus)

mombelgi Boucomont, 1929: 760 A: SCH YUN (Gymnopleurus)
morosus Fairmaire, 1886: 319 (Gymnopleurus) A: SCH YUN
mundus Wiedemann, 1819: 162 (Gymnopleurus) A: CH PA ORR
capicola Laporte, 1840: 70 (Gymnopleurus)

opacus Kollar & L. Redtenbacher, 1844: 516 (Gymnopleurus) A: HP KA NP UP ORR
ruficornis Motschulsky, 1854d: 63 (Gymnopleurus) A: AF HP SHG UP YUN ORR
subtilis C.O. Waterhouse, 1890b: 372 (Gymnopleurus)

genus Gymnopleurus Illiger, 1803: 199 type species Scarabaeus geoffroyi Fuessly, 1775
subgenus Gymnopleurus Illiger, 1803: 199 type species Scarabaeus geoffroyi Fuessly, 1775
Spinigymnopleurus Shipp, 1897: 168 type species Gymnopleurus tristis Laporte, 1840

aciculatus Gebler, 1841a: 372 E: AB ST A: AF IN KI KZ TD TM UZ XIN
violaceus Ballion, 1871a: 331

aeruginosus Harold, 1867d: 94 N: EG AFR
atratus Klug, 1845: [2], pl. xli A: “Arabia deserta”
bombayensis Arrow, 1931: 65 A: PA ORR
cyaneus Fabricius, 1798: 34 (Copris) A: HP NP PA UP ORR
impressus Laporte, 1840: 73
indicus Laporte, 1840: 73

elegans Klug, 1845: [3], pl. xli A: AE OM SA YE
flagellatus asperatus Motschulsky, 1849: 103 E: GG A: IN SY TR
flagellatus calignosus Balthasar, 1934c: 148 A: IS LE SY
flagellatus flagellatus Fabricius, 1787a: 17 (Scarabaeus) E: AB AR FR GG GR IT PT SP ST TR UK N: AG EG LB
MO TU AURi
clypeolatus Mulsant, 1842: 58
confusus Mulsant, 1842: 58
coriarius Herbst, 1789: 309 (Scarabaeus)
minor Seabra, 1907: 75
rugulosus Mulsant, 1842: 58
suturalis Mulsant, 1842: 58
rufipes Seabra, 1907: 75
variolosus Motschulsky, 1849: 102

flagellatus hornei C.O. Waterhouse, 1890c: 410 A: AF PA “N.W. India”
flagellatus serratus Fischer von Waldheim, 1821a: 11 E: AB KZ ST A: IN KI KZ MG TD TM UZ XIN

SCARABAEIDAE

173

barovskyi Kiseritzky, 1928a: 45
conflagratus Motschulsky, 1849: 102
cribellatus Motschulsky, 1849: 102

geoffroyi Fuessly, 1775: 2 (Scarabaeus) E: AL AR AU BH BU CR CZ FR GE GR HU IT KO MC MD ME PL SK RO
SB SL ST SZ TR UK N: AG EG A: CY IN IS KU SI TR
atratulus Motschulsky, 1849: 100
bidentatus Mulsant, 1842: 55
cantharus Illiger, 1803: 201
cyanescens Motschulsky, 1849: 101
sinuatus Geoffroy, 1785: 15 (Copris) [HN]
tuberculatus Mulsant, 1842: 55

lacunosus Klug, 1845: [2], pl. xli A: SA
mimus Balthasar, 1934c: 148 A: IN
mopsus mopsus Pallas, 1781: 3 (Scarabaeus) E: AB AL AR AU BH BU CR CT FR GE GG GR HU IT KO KZ MC MD
ME PT RO SB SK SL SP ST SZ TR UK A: AF CY IN IS KI KU KZ MG NC NMO SC SI SY TD TM TR UZ
castanonota Seabra, 1907: 70
dorsalis Mulsant, 1842: 55
hortobagyensis Papp, 1943: 453
indistinctus Mulsant, 1842: 55
laevifrons Mulsant, 1842: 55

mopsus sinensis Balthasar, 1934c: 148 A: BEI FUJ HEB JIX SHD “Manchuria”
naviauxi Montreuil, 2009: 420 A: IQ
persianus Reitter, 1909b: 79 A: AE IN IQ SA SY
arabs Garreta, 1914b: 357

plicatulus Fairmaire, 1890: 547 A: YE AFR
qurosh Montreuil, 2011: 414 E: AR GG A: IN IS LE SY TR
sericeifrons Fairmaire, 1887a: 108 A: EG AFR
aeneipes Fairmaire, 1893c: 147
cuprarius H.J. Kolbe, 1914: 320
cyaneus H.J. Kolbe, 1914: 320
kruegeri H.J. Kolbe, 1895a: 336
subauratus H.J. Kolbe, 1914: 320

sindensis Arrow, 1931: 66 A: PA
sturmii W.S. Macleay, 1821: 512 (Scarabaeus) E: AL BH BU CR FR GR IT KO MC ME PT RO SB SP TR N: AG EG
LB MO TU A: IS JO SY TR
atronitidus W.S. Macleay, 1821: 513 (Scarabaeus)
convexiusculus Motschulsky, 1849: 101
glabriusculus Mulsant, 1842: 55
laeviusculus Mulsant, 1842: 55
minor Ladeiro, 1950: 4
transpyrenaeus Gistel, 1857b: 595

174

SCARABAEIDAE

virescens Seabra, 1907: 75 [HN]

tuxeni Petrovitz, 1955: 276 A: AF
subgenus Metagymnopleurus Kabakov, 2006a: 78 type species Scarabaeus miliaris Fabricius, 1775
bicallosus Felsche, 1909: 751 A: AF IN PA
gemmatus Harold, 1871c: 117 A: AP NP UP ORR
koenigii Fabricius, 1775: 29 (Scarabaeus) A: NMO XIZ ORR
guttatus Gmelin, 1790: 1558 (Scarabaeus)
scriptus Pallas, 1781: 7 (Scarabaeus)

miliaris Fabricius, 1775: 817 (Scarabaeus) A: AF BT HP KA PA UP ORR
parvus W.S. Macleay, 1821: 517 (Scarabaeus) A: UP ORR
genus Paragymnopleurus Shipp, 1897: 166 type species Scarabaeus sinuatus A.G. Olivier, 1789
Progymnopleurus Garreta, 1914a: 52 type species Scarabaeus sinuatus A.G. Olivier, 1789

ambiguus A. Janssens, 1943: 1 A: TAI
japonicus Balthasar, 1955a: 395 (Gymnopleurus)

brahminus C.O. Waterhouse, 1890c: 411 (Gymnopleurus) A: CHQ FUJ GUI HUB HUN JIA JIX SCH SHG YUN
ZHE XIZ ORR
martinezi Balthasar, 1955a: 393 (Gymnopleurus) A: SCH
melanarius Harold, 1867c: 76 (Gymnopleurus) A: FUJ HKG SCH SHG TAI YUN ORR
aethiops Sharp, 1875a: 34 (Gymnopleurus)

sinuatus assamensis C.O. Waterhouse, 1890c: 411 (Gymnopleurus) A: AP HP NP PA SD UP ORR
sinuatus sinuatus A.G. Olivier, 1789: no 3: 160 (Scarabaeus) A: ANH BEI FUJ GUA GUI HUB HAI HUN JIA NC
SHG SHN TAI YUN ORR
sinuatus szechouanicus Balthasar, 1934c: 149 (Gymnopleurus) A: CHQ SCH XIZ ORR

tribe Oniticellini H.J. Kolbe, 1905
subtribe Drepanocerina van Lansberge, 1875

genus Eodrepanus Barbero, Palestrini & Roggero, 2009: 1846 type species Drepanocerus parallelus
Raffray, 1877
integriceps A. Janssens, 1953: 23 (Drepanocerus) A: GUI HEN SCH YUN ORR
liuchungloi Kryzhanovskij & S.I. Medvedev, 1966: 396 (Drepanocerus) A: GUI YUN
striatulus R. Paulian, 1945: 140 (Drepanocerus) A: AP FUJ GUI NP SCH YUN ORR
genus Paraixodina Roggero, Barbero & Palestrini, 2015: 164 type species Drepanocerus runicus Arrow, 1909
runica Arrow, 1909a: 93 (Drepanocerus) A: YUN ORR
genus Sinodrepanus Simonis, 1985: 95 type species Oniticellus falsus Sharp, 1875
besucheti Simonis, 1985: 100 A: FUJ SCH
falsus Sharp, 1785a: 52 (Oniticellus) A: SCH ORR
rex Boucomont, 1912b: 277 (Drepanocerus) A: FUJ HUN SCH XIZ YUN
arrowi Balthasar, 1932d: 64 (Drepanocerus)

175

SCARABAEIDAE
rosannae Simonis, 1985: 100 A: FUJ
similis Simonis, 1985: 102 A: FUJ GUI ORR
tsaii Masumoto, M.-M. Yang & Ochi, 2004: 360 A: TAI
uenoi Ochi, Kon & Masumoto 2004: 348 A: GUI
genus Tibiodrepanus Krikken, 2009b: 16 type species Copris setosus Wiedemann, 1823
Sulcodrepanus Krikken, 2009b: 17 type species Drepanocerus sulcicollis Laporte, 1840

hircus Wiedemann, 1823: 19 (Copris) A: HKG ORR
setosus Wiedemann, 1823: 19 (Copris) A: BT HKG HP NP PA SD UP ORR
setosa Motschulsky, 1864: 459 (Ixodina)

simplex Kabakov, 2006a: 142 (Drepanocerus) A: AF
sinicus Harold, 1868c: 104 [RN] (Drepanocerus) A: BT FUJ GUA GUI GUX HAI HKG HP HUB NP PA UP YUN
ORR
setosus Boheman, 1858: 50 [HN] (Drepanocerus)

subtribe Oniticellina H.J. Kolbe, 1905

genus Euoniticellus A. Janssens, 1953: 41 type species Scarabaeus fulvus Goeze, 1777
fulvus Goeze, 1777: 74 (Scarabaeus) [NP] E: AB AL AR AU BE BH BU BY CR CT CZ FR GE GG GR HU IT KZ MC
MD ME PL PT RO SB SK SL SP ST SZ TR UK N: AG EG LB MO TU A: AF CY IN IQ KI KZ LE MG SY TD TM
TR UZ WS XIN AURi
flavipes Fabricius, 1792a: 70 (Scarabaeus)
fulvicollis Mulsant, 1842: 100 (Oniticellus)
fulvipterus Mulsant, 1842: 100 (Oniticellus)
maculatus Mulsant, 1842: 100 (Oniticellus)
minuta Seabra, 1907: 150 (Oniticellus)
subcornutus Mulsant, 1842: 99 (Oniticellus)
thoracocircularis Laicharting, 1781: 24 (Scarabaeus)

intermedius Reiche, 1850: 337 (Oniticellus) A: SA AFR AURi NTRi
clavatus Roth, 1851: 127 (Oniticellus)
speciosus A. Costa, 1853b: 28 (Oniticellus)

nasicornis Reiche, 1850: 339 (Oniticellus) A: YE AFR
pallens A.G. Olivier, 1789: no 3: 170 (Scarabaeus) E: FR (Corse) IT (Sardegna, Sicilia) SP N: AG EG LB MO TU A:
AE AF HP IQ PA TR UP YE AFR
concinnus Gené, 1836: 185 (Oniticellus)
revelierei Mulsant & Rey, 1871c: 469 [=1871a: 725, 1872d: 430] (Oniticellus)
speculifer Ménétriés, 1849: 58 (Oniticellus)

pallipes Fabricius, 1781a: 33 (Scarabaeus) E: AB AL AR BH BU CR FR GG GR HU KZ MC MD ME
IT PT RO SB SP ST TR UK N: MO A: AF CY HP IN IS KA KI KZ NMO PA SY TD TM TR UP UZ
XIN AURi
nitidicollis Arrow, 1908a: 179 (Oniticellus)
subdeletus Mulsant, 1842: 96 (Oniticellus)

176

SCARABAEIDAE

genus Liatongus Reitter, 1892a: 167 [=1893a: 45] type species Onthophagus phanaeoides Westwood, 1839
Onthosphaenus Motschulsky, 1860f: 154 type species Scarabaeus vertagus Fabricius, 1798

affinis Arrow, 1908a: 181 (Oniticellus) A: YUN ORR
capricornis Arrow, 1931: 368

ancorifer Král & Rejsek, 1999: 2 A: YUN
appositicornis Král & Rejsek, 1999: 5 A: GAN SCH
bucerus Fairmaire, 1891f: cxciv (Oniticellus) A: GUI SCH YUN ZHE
clypeocornis Scheuern, 1988: 305 A: BT
davidi Boucomont, 1919b: 601 (Oniticellus) A: SCH XIZ
denticornis Fairmaire, 1887b: 101 (Oniticellus) A: CHQ SCH YUN
boucomonti Balthasar, 1932d: 68 (Oniticellus)
concavicollis Fairmaire, 1891f: cxciv (Oniticellus)
semenovi Balthasar, 1934a: 20 (Oniticellus)

endroedii Balthasar, 1956a: 66 A: SCH YUN
fairmairei Boucomont, 1921a: 27 (Oniticellus) A: CE SHG
gagatinus Hope, 1831: 22 (Onthophagus) A: AP CHQ GUI HP KA NP SCH SD UP XIZ YUN ORR
brama Kollar & L. Redtenbacher, 1844: 521 (Onthophagus)

hastatus Král & Rejsek, 1999: 11 A: NP
imitator Balthasar, 1938c: 98 A: GAN SCH
incurvicornis Fairmaire, 1887b: 100 (Oniticellus) A: SCH XIZ YUN
medius Fairmaire, 1889c: 655 (Oniticellus) A: GAN SCH XIZ
mergacerus Hope, 1831: 22 (Onthophagus) A: AP BT NP SD UP ORR
minutus Motschulsky, 1861a: 13 (Phanaeus) A: GAN FE JA NC SC SCH TAI YUN
fumiroi Yohena, 1937: 7
yohenai Matsumura, 1934: 66 (Oniticellus)

phanaeoides Westwood, 1839b: lv (Onthophagus) A: AP CHQ FUJ GUI HEB HEN HP JA NC PA SC SCH SHX UP
YUN ORR
excavatus Kollar & L. Redtenbacher, 1844: 523 (Onthophagus)

pugionatus Boheman, 1858: 44 (Onthophagus) A: HKG
rhinoceros Arrow, 1931: 363 A: HP NP SD ORR
triacanthus Boucomont, 1920a: 264 (Oniticellus) A: SD XIZ ORR
tridentatus Boucomont, 1919b: 602 (Oniticellus) A: YUN ORR
vertagus Fabricius, 1798: 30 (Copris) A: FUJ GUA GUI HKG HP SHG TAI UP XIZ YUN ORR
bifurcalis White, 1844: 423 (Onthophagus)
bifurcatus Erichson, 1833: 235 (Oniticellus)
dichrous Boheman, 1858: 43 (Onthophagus)
pictisternum Fairmaire, 1891f: cxcvii (Oniticellus)
scutellaris Reitter, 1892a: 167 [=1893a: 46]
tonkineus Fairmaire, 1893e: 305 (Oniticellus)

vseteckai Balthasar, 1933a: 63 (Oniticellus) A: FUJ SCH

177

SCARABAEIDAE
genus Oniticellus Dejean, 1821: 53 type species Scarabaeus cinctus Fabricius, 1775

Oniticellus Le Peletier de St. Fardeau & Audinet-Serville, 1828: 356 [HN] type species Scarabaeus cinctus Fabricius, 1775
Pseudoniticellus Kraatz, 1895d: 143 type species Oniticellus planatus Laporte, 1840

cinctus Fabricius, 1775: 30 (Scarabaeus) A: AP FUJ HP TAI UP YUN ORR
diadema Wiedemann, 1819: 159 (Onitis)
serratipes Drury, 1773: 82 (Scarabaeus)

gayeni Biswas & Chatterjee, 1985: 156 A: AP
namdaphaensis Biswas & Chatterjee, 1985: 152 [DA] A: AP
puberulus Y.-W. Zhang, 1988: 228 [DA] A: XIZ
subhendui Biswas & Chatterjee, 1985: 154 [DA] A: AP
genus Paroniticellus Balthasar, 1963a: 76 type species Onitis festivus Steven, 1809
festivus Steven, 1809: 31 (Onitis) E: AB AR GG ST A: IN TM TR
specularis Fischer von Waldheim, 1844: 44 (Onthophagus)
steveni Billberg, 1815: 273 (Onitis)

genus Tiniocellus Péringuey, 1902: 116 type species Oniticellus spinipes Roth, 1851
imbellis Bates, 1891a: 13 (Oniticellus) A: HP NP PA UP ORR
modestus Arrow, 1908a: 182 (Oniticellus)

tribe Onitini Laporte, 1840

genus Bubas Dejean, 1833: 143 type species Scarabaeus bison Linnaeus, 1767
bison Linnaeus, 1767: 547 (Scarabaeus) E: AL BH BU CR FR GR IT ME PT SP TR N: AG MO TU AURi
brevicornis Mulsant, 1842: 78
castaneus Mulsant, 1842: 78
dentifrons Mulsant, 1842: 78
lineifrons Mulsant, 1842: 78
trispinosus L. Petagna, 1819: 26 (Scarabaeus)

bubaloides A. Janssens, 1938a: 8 N: AG EG LB MO TU A: CY IN IS JO LE TR SY
bubalus A.G. Olivier, 1811: 492 (Onitis) E: FR PT SP
brunnipterus Mulsant, 1842: 81
inermifrons Mulsant, 1842: 81
integricornis Mulsant, 1842: 81
simplicifrons Mulsant, 1842: 81

genus Cheironitis van Lansberge, 1875b: 19 [NP] type species Scarabaeus furcifer P. Rossi, 1792
Chironitis A. Janssens, 1937: 152 [unjustified emendation]
Uposlotus A. Costa, 1853b: 22 [NO] type species Scarabaeus furcifer P. Rossi, 1792

arrowi A. Janssens, 1937: 159 (Chironitis) A: AF NP PA SA ORR
candezei van Lansberge, 1875b: 21 A: IQ
eumenes Gebler, 1860: 465 (Onitis) E: ST UK “Caucasus” A: KI KZ MG NW UZ

178

SCARABAEIDAE

kerzhneri S.I. Medvedev, 1974: 114
phoebus Reitter, 1892a: 222 [=1893a: 101]

furcifer P. Rossi, 1792: 7 (Scarabaeus) E: AL BH BU CR GR IT MC SP TR N: AG EG LB MO TU A: IS SY TR
metasternalis Reitter, 1894g: 302
pugil A. Costa, 1853b: 22 (Onitis)
syphax Fairmaire, 1875a: 506
theryi Tondu, 1907: 275

granulipennis Reitter, 1909b: 80 A: IN
haroldi Ballion, 1871a: 331 (Onitis) E: AB AR BU GG GR ST TR A: AF CY IN KI KZ TD TM TR UZ
luctuosus van Lansberge, 1875b: 21
rotundicoxis Reitter, 1892a: 219 [=1893a: 98]

irroratus P. Rossi, 1790: 7 (Scarabaeus) E: FR (Corse) IT PT SP N: AG EG MO LB TU A: IS SI SY TR
calcaratus A.G. Olivier, 1811: 490 (Onitis)
inversus A. Costa, 1853b: 2 (Onitis)
lophus Fabricius, 1798: 26 (Onitis)

klapperichi Balthasar, 1956b: 411 (Chironitis) A: AF IN
moeris Pallas, 1781: 3 (Scarabaeus) E: GG ST A: AF IN KI KZ LE SY TD TM UZ
sophax Fischer von Waldheim, 1830: 185 (Onitis)

osiridis Reiche, 1856a: 118 (Onitis) N: AG EG LB A: AE OM PA YE AFR
pamphilus Ménétriés, 1849: 57 (Onitis) E: AB AR GG GR ST TR A: AF CY IN IS LE SI SY TD TM TR UZ
ponticus van Lansberge, 1875b: 36

socotranus Gahan, 1900: 10 A: YE (Suqutra)
sterculius Ballion, 1871a: 331 (Onitis) A: AF IN KI KZ TD TM UZ
hauseri Reitter, 1892a: 221 [=1893a: 100]
pygidialis Koshantchikov, 1894: 104

ungaricus Herbst, 1789: 230 (Scarabaeus) E: AU BH BU CR FR GR HU IT MC MD ME RO SB SP ST TR UK
alexis Mulsant, 1842: 88 (Onitis) [HN]
amyntas Steven, 1806: 165 [=1812: 128] (Onitis)
clinias Fabricius, 1792a: 19 (Scarabaeus)
meliboeus Mulsant, 1842: 88 (Onitis)
tityrus Mulsant, 1842: 88 (Onitis)

genus Onitis Fabricius, 1798: 2 type species Scarabaeus sphinx A.G. Olivier, 1789 [HN] (=Onitis belial Fabricius,
1798)
alexis septentrionalis Balthasar, 1942c: 195 E: FR (Corse) GR IT (Sicilia) MC TR N: AG EG LB MO TU A: CY IS
LE SA SY TR
belial Fabricius, 1798: 27 E: FR IT PT SP N: AG MO TU AURi
fuscus Mulsant, 1842: 85
inermicrus Mulsant, 1842: 85
olivierii Illiger, 1800a: 245 (Copris)
planifrons Mulsant, 1842: 85

SCARABAEIDAE

179

subcostalis Mulsant, 1842: 85
subtuberculatus Mulsant, 1842: 85

brevidens Bai, X.-K. Yang & Y.-W. Zhang, 2006: 391 A: YUN
castaneus Kollar, 1844: 517 A: AP HP KA NP PA UP
crassus Sharp, 1875b: 48 A: PA UP
vischnu Preudhomme de Borre, 1881a: xxxix

damoetas Steven, 1806: 163 [=1812: 126] E: AB AR BU GG GR MC MD RO ST TR UK A: CY IN IS JO LE SY TR
laevigatus A.G. Olivier, 1811: 492
steveni Brullé, 1832: 170

excavatus Arrow, 1931: 391 A: AP FUJ HKG HP NP PA SCH UP YUN ORR
ezechias Reiche & Saulcy, 1856: 390 N: EG A: IN IS JO LE SY TR
falcatus Wulfen, 1786: 14 (Scarabaeus) A: AP BEI CHQ FUJ GUA GUI HAI HEB HEN HP HUB JIA JIX NP PA
SCH SHX TAI UP YUN ZHE ORR
himaleyicus Kollar, 1844: 518

feae Felsche, 1907: 293 A: AP YUN ORR
humerosus Pallas, 1771: 462 (Scarabaeus) E: AB AR GG KZ ST TR A: AF CY IN IS JO KI KZ LE PA SY TD TM TR
UZ XIN
chevrolatii P.H. Lucas, 1846c: pl. 23
menalcas Pallas, 1781: 4 (Scarabaeus)
violaceus J.R. Sahlberg, 1913: 119

ion A.G. Olivier, 1789: no 3: 186 (Scarabaeus) E: FR IT SP N: AG MO TU A: IS
granulatus Mulsant, 1842: 92
infuscata Seabra, 1907: 94
trispinus Mulsant, 1842: 92
vandelli Fabricius, 1801a: 28

intermedius J. Frivaldszky, 1892: 122 A: SHG ORR
lama van Lansberge, 1875b: 123 A: PA UP ORR
naviauxi Cambefort, 1988: 189 A: NP
miesseni Gusakov & Klimenko, 2009: 3 N: MO
numida marginicollis Balthasar, 1929c: 113 A: “Turkestan”
numida numida Laporte, 1840: 90 N: AG LB MO TU
strigatus Erichson, 1841a: 170

pers Gusakov & Klimenko, 2009: 5 A: IN
philemon Fabricius, 1801a: 30 A: AP HP NP PA SE SW UP ORR
distinctus van Lansberge, 1875b: 138
minor van Lansberge, 1875b: 139

punctatostriatus A. Janssens, 1937: 48 A: PA SD
ringenbachi Keith, 2007e: 15 N: LB
spinipes Drury, 1773: 79 (Scarabaeus) A: “South China” ORR
subopacus Arrow, 1931: 395 A: AF AP HP KA NP PA UP YUN ORR
tingaudi Keith & Moretto, 2008: 177 A: SA

180

SCARABAEIDAE

virens van Lansberge, 1875b: 135 A: CH PA UP ORR
amplectens van Lansberge, 1875b: 136

viridulus Boheman, 1857: 253 A: YE AFR AURi
consanguineus Felsche, 1911: 140
insulanus H.J. Kolbe, 1914: 306

tribe Onthophagini Burmeister, 1846

genus Anoctus Sharp, 1875a: 49 type species Anoctus laevis Sharp, 1875
siamensis Balthasar, 1942c: 199 A: YUN ORR
foveolatus R. Paulian, 1945: 61

genus Caccobius C.G. Thomson, 1859: 80 type species Scarabaeus schreberi Linnaeus, 1767
subgenus Caccobius C.G. Thomson, 1859: 80 type species Scarabaeus schreberi Linnaeus, 1767
Histeridium Motschulsky, 1860f: 150 type species Scarabaeus schreberi Linnaeus, 1767

brevis C.O. Waterhouse, 1875a: 73 A: BEI FE HEB JA NC NE SC SCH SHX “Manchuria”
atomarius Balthasar, 1935d: 185
matsuoi Matsumura & Yohena, 1937a: 120
vacerrosus Lewis, 1895c: 378 (Onthophagus)

chokakurainus Matsumura, 1938: 56 (Onthophagus) A: TAI
chujoi R. Paulian, 1942b: 58
flavolimbatus Balthasar, 1942c: 199

demangei Boucomont, 1920b: 320 A: SCH YUN ORR
denticollis Harold, 1867a: 5 A: AF HP KA NP PA SD UP XIZ
excavatus Bai, Y.-W. Zhang & X.-K. Yang, 2007: 131 A: XIZ
gonoderus Fairmaire, 1888a: 17 (Onthophagus) A: CHQ FUJ GUI HUN SCH TAI YUN ORR
histeroides Ménétriés, 1832: 180 (Onthophagus) E: AB AL AR BU CR GG GR HU KZ MC ME RO SB ST TR UK
(Krym) A: CY IN IQ IS KZ SY TM TR
anthracinus Faldermann, 1835a: 247 (Onthophagus)

imitans Balthasar, 1933a: 67 A: SCH XIZ YUN
jessoensis Harold, 1867d: 100 A: FE (Kunashir, Sakhalin) HEN JA
amagisanus Matsumura, 1937a: 121
hirayamai Matsumura, 1936: 62
koichii Matsumura, 1934: 67 (Onthophagus)
microcephalus Harold, 1877e: 349
sapporensis Matsumura, 1936: 64
yubariensis Matsumura, 1936: 65

mundus Ménétriés, 1839: 23 (Onthophagus) E: AB AR BU GR TR A: IN IQ IS JO LE SY TM TR
pantherinus Arrow, 1931: 154 A: HP PA ORR
scheuerni Cambefort, 1990: 303 A: NP
schreberi Linnaeus, 1767: 551 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FR GE GG GR HU
IT KO KZ LA LT MC MD ME PL PT RO SB SK SL SP ST SV SZ TR UK N: AG EG LB MO TU A: AF IN IQ IS KZ
SY TM TR

SCARABAEIDAE
bidentatus Mulsant, 1842: 143 (Onthophagus)
bimaculatus Mulsant, 1842: 144 (Onthophagus)
corsicus O. Schneider, 1902: 52
haemorrhoidalis Geoffroy, 1785: 15 (Copris)
indistinctus Mulsant, 1842: 144 (Onthophagus)
juvenilis Mulsant, 1842: 144 (Onthophagus)
mixtus Mulsant, 1842: 144 (Onthophagus)
obscurus Mulsant, 1842: 144 (Onthophagus)

subgenus Caccophilus Jekel, 1872: 410 type species Caccobius himalayanus Jekel, 1872
aterrimus Fabricius, 1798: 35 (Copris) A: HP NP UP ORR
haroldi Jekel, 1872: 416

balthasari Tesař, 1969b: 63 A: PA
boucomonti Balthasar, 1935d: 186 A: GAN SCH SHA YUN
christophi Harold, 1879b: 229 A: FE HEB LIA MG NC NO SC SCH SHX YUN “Manchuria”
asininus Heyden, 1887b: 301
crenatostriatus Balthasar, 1935d: 188

diminutivus Walker, 1858: 208 (Copris) A: SD UP ORR
setulosus Motschulsky, 1864: 459

fukiensis Balthasar, 1942a: 119 A: FUJ SHG
himalayanus Jekel, 1872: 411 A: HP KA UP XIZ
indicus Harold, 1867a: 12 A: KA HP NP UP ORR
inermis Arrow, 1931: 147 A: AP HP SD UP ORR
kelleri Olsoufieff, 1907b: 192 (Onthophagus) A: BEI ES FE HEI HEN LIA NC NO
sibiricus Balthasar, 1935d: 188

masumotoi Cambefort, 1990: 305 A: TAI YUN ORR
yunnanicus Kabakov, 1994b: 302

meridionalis Boucomont, 1914a: 239 A: HP UP ORR
nikkoensis Lewis, 1895c: 378 (Onthophagus) A: JA
kasuganus Matsumura & Yohena, 1936: 63

pulicarius Harold, 1875c: 214 A: SY “Palestina”
pullus Jekel, 1872: 416 A: CH
sordidus Harold, 1886a: 141 A: BEI ES FE GAN HEB HEN NC NO SC SCH SHX “S. Manchuria”
koltzei Reitter, 1892a: 213 [=1893a: 92]

suzukii Matsumura, 1936: 65 A: JA
torticornis Arrow, 1931: 144 A: UP
tortus matsudai Ochi & Araya, 1996: 4 A: TAI (Layau)
tortus tortus Sharp, 1875a: 50 A: GUA ORR
ultor Sharp, 1875a: 50 A: HP UP ORR
unicornis Fabricius, 1798: 33 (Copris) A: BEI FUJ GAN HUB HUN JA NC SC SCH SHG SHX TAI YUN ORR
nitidiceps Fairmaire, 1893e: 304 (Onthophagus)
yamauchii Matsumura, 1936: 66

181

182

SCARABAEIDAE

viridicollis Fåhraeus, 1857: 325 (Onthophagus) A: YE AFR
minimus d’Orbigny, 1898c: 160
semicoeruleus d’Orbigny, 1905: 386

vulcanus Fabricius, 1801a: 41 (Copris) A: HP NP UP ORR
bicuspis Wiedemann, 1823: 11 (Onthophagus)
mutans Sharp, 1875a: 51

genus Cleptocaccobius Cambefort, 1984: 3 type species Caccobius signaticollis d’Orbigny, 1902
scrofa Balthasar, 1942a: 118 (Caccobius) A: FUJ SHX TAI
shansicus Nakane, 1951: 69

simplex Boucomont, 1923a: 9 (Caccobius) A: YUN ORR
genus Digitonthophagus Balthasar, 1959b: 464 type species Scarabaeus bonasus Fabricius, 1775
bonasus Fabricius, 1775: 23 (Scarabaeus) A: AF HP IN NP PA NP UP ORR
gazella Fabricius, 1787b: 377 (Scarabaeus) Ni: EG Ai: CH HP IN JA NP PA UP YE AFR AURi NARi NTRi ORRi
antilope Fabricius, 1798: 32 (Copris)
dorcas A.G. Olivier, 1789: no 3: 121 (Scarabaeus)
intermedius Reiche, 1840: 243 (Onthophagus)
metallicus Fabricius, 1798: 28 (Copris)

genus Euonthophagus Balthasar, 1959b: 467 type species Scarabaeus amyntas A.G. Olivier, 1789
aeneobrunneus Kabakov, 1977: 390 (Onthophagus) A: IN
markaziensis Petrovitz, 1980: 601 (Onthophagus)

amyntas alces Fabricius, 1792a: 56 (Scarabaeus) E: AL AU BH BU CR GG GR HU KO KZ MC MD ME PL RO SB
SK ST TR UK A: AF CY IS JO SY TR
hybneri Fabricius, 1792a: 61 (Scarabaeus)

amyntas amyntas A.G. Olivier, 1789: no 3: 127 (Scarabaeus) E: AU FR IT PT SL SP SZ
tages A.G. Olivier, 1789: no 3: 143 (Scarabaeus)

amyntas auchenia L. Redtenbacher, 1850: 48 (Onthophagus) A: IN
aspadanaensis Petrovitz, 1965d: 671 (Onthophagus)
rechingeriorum Mandl, 1976: 372 (Onthophagus)

amyntas subviolaceus Ménétriés, 1832: 177 (Onthophagus) E: AB AR ST A: IN KI KZ TD TM UZ XIN
atramentarius Ménétriés, 1832: 179 (Onthophagus) E: AB AL AR BU GG GR MC ST SY TR N: EG A: CY IN IQ IS
SY TR UZ
felschei Reitter, 1891: 243 (Onthophagus)
orcas Ménétriés, 1838: 24 (Onthophagus)
peropacus Kirschenblatt, 1935: 201 (Onthophagus)

bedeli bedeli Reitter, 1891f: 241 (Onthophagus) N: AG EG LB MO TU A: AF IS SI SY
bedeli conterminus Petrovitz, 1971d: 215 N: TU A: IN IS JO SI SY
rapillyi Baraud, 1984b: 105

carbonarius Klug, 1855: 653 (Onthophagus) A: OM SA YE

SCARABAEIDAE

183

crocatus clementi Baraud, 1960: 232 (Onthophagus) N: MO
crocatus crocatus Mulsant & Godart, 1872: 315 [=1873: 13] (Onthophagus) E: SP N: AG LB MO TU
imitator Reitter, 1891f: 243 (Onthophagus)

dorbignyi Olsoufieff, 1900: 268 (Onthophagus) A: AF IN
gibbosus Scriba, 1790: 56 (Copris) E: AB AL AR BH BU FR GG GR IT MC ME PT RO SB SP ST SZ TR UK A: AF
CY ES IN IS KI KZ MG SY TD TM TR UZ XIN XIZ
difformis Mulsant, 1842: 106 (Onthophagus)
dubius Mulsant, 1842: 106 (Onthophagus)
ganglbaueri Reitter, 1891f: 244 (Onthophagus)
sycophanta Mulsant, 1842: 106 (Onthophagus)
umbrinus Mulsant, 1842: 106 (Onthophagus)
unituberculatus Mulsant, 1842: 106 (Onthophagus)
weisei Reitter, 1891f: 244 (Onthophagus)

hazariensis Kabakov, 1977: 387 (Onthophagus) A: AF
koshantschikoffi Reitter, 1891f: 245 (Onthophagus) A: AF IN KI KZ TD TM UZ XIN XIZ
ochripennis d’Orbigny, 1898a: 138 (Onthophagus)
semirufus d’Orbigny, 1898a: 138 (Onthophagus)

laghmanicus Kabakov, 1977: 386 (Onthophagus) A: AF PA
loeffleri Petrovitz, 1965d: 672 (Onthophagus) A: IN IQ
maindroni d’Orbigny, 1898e: 232 (Onthophagus) N: EG AFR
mostafatsairi Palestrini, Varola & Zunino, 1979: 431 A: IN IQ
mostafatsairii Petrovitz, 1980: 599 (Onthophagus) [HN]

pentaceros Olsoufieff, 1900: 266 (Onthophagus) A: AF IN
pertinax Balthasar, 1959b: 467 [=1960b: 187] (Onthophagus) A: AF IN PA
schnabeli Splichal, 1910: 567 (Onthophagus) A: AF KA PA XIZ
sulcicollis Reitter, 1892d: 135 (Onthophagus) A: AF IN KI KZ PA TD TM UZ
tissoni Reitter, 1906b: 33 (Onthophagus) A: IS JO SY
hannas J.R. Sahlberg, 1908: 62 (Onthophagus)

verberatus Ziani, 2006: 103 A: IN
yama Ziani, 2006: 99 A: IN
yemenicus Balthasar, 1946b: 16 (Onthophagus) A: YE
genus Haroldius Boucomont, 1914b: 253 type species Haroldius rugatulus Boucomont, 1914
Afroharoldius A. Janssens, 1949b: 183 type species Afroharoldius ennearthrus A. Janssens, 1949
Cyclotrogus Wasmann, 1918: 2 type species Cyclotrogus heimi Wasmann, 1918
Formicdubius Philips & Scholtz, 2000: 227 type species Formicdubius convexus Philips & Scholtz, 2000: 230
Ponerotrogus Silvestri, 1924: 583 type species Ponerotrogus annandalei Silvestri, 1924

heimi Wasmann, 1918: 4 (Cyclotrogus) A: NP PA ORR
nigrita Wassmann, 1918: 4 (Cyclotrogus)

jechaer Král & Hájek, 2012: 208 A: YUN
lassallei Cambefort, 1986: 69 A: NP

184

SCARABAEIDAE

perroti R. Paulian, 1939a: 70 A: TAI ORR
stevensi Arrow, 1931: 416 A: NP SD ORR
turnai Král, 2003: 194 A: HUB
uenoi Masumoto & Yin, 1993: 84 A: YUN
genus Onthophagus Latreille, 1802a: 141 type species Scarabaeus taurus Schreber, 1759
subgenus Altonthophagus Kabakov, 1990: 28 type species Onthophagus sibiricus Harold, 1877
cervenkai Kabakov, 2008: 417 A: GAN QIN SCH SHA SHX
concolor Sharp, 1878b: 170 A: AF HP KA PA UP ORR
cupreiceps Arrow, 1907b: 427 A: BT NP SD XIZ ORR
inelegans Balthasar, 1935f: 315 A: GAN SCH YUN
kozlovi Kabakov, 1990: 36 A: GAN QIN SCH SHA XIZ
kukunorensis Kabakov, 1990: 36 A: GAN QIN
marmotae Kabakov, 1990: 33 A: AF TD
sibiricus Harold, 1877d: 335 A: AF ES KI KZ MG TD UZ WS XIN
deletus Reitter, 1892a: 187 [=1893a: 66]
nocturnus Reitter, 1892a: 204 [=1893a: 65]
nakatomii Matsumura, 1937c: 4
shoyozanus Matsumura, 1937c: 5

tibetanus Arrow, 1907b: 428 A: KA UP XIZ
turpidoides Kabakov, 2008: 415 A: QIN SCH SHA YUN
turpidus Reitter, 1887b: 219 A: GAN MG NP QIN SCH XIN XIZ
sublineatus d’Orbigny, 1898a: 176

uniformis Heyden, 1886b: 275 A: BEI FE GAN HEB HEI LIA NC NO SC SHX “Manchuria”
subgenus Amphionthophagus Martín Piera & Zunino, 1983: 60 type species Onthophagus numidicus
d’Orbigny, 1908
falzonii Goidanich, 1926a: 37 A: IN IS JO LE SY TR
melitaeus Fabricius, 1798: 30 (Copris) E: PT SP N: AG MO
numidicus d’Orbigny, 1908: 165 N: AG TU
subgenus Colobonthophagus Balthasar, 1963a: 164 type species Scarabaeus tragus Fabricius, 1792
aenescens Wiedemann, 1823: 13 (Copris) A: HP NP PA UP ZIX ORR
armatus C.É. Blanchard, 1853: 98 A: AP FUJ GUA HKG ORR
luzonicus van Lansberge, 1883b: 51
pseudoarmatus Balthasar, 1944: 93

arunensis Scheuern, 1995: 436 A: NP SD
bengalensis Harold, 1886a: 146 A: UP ORR
dama Fabricius, 1798: 32 (Copris) A: AP BT HP NP SD UP ORR
aeneus A.G. Olivier, 1789: no 3: 131 (Scarabaeus) [HN]
cervicornis Kirby, 1825: 565
zubaci Balthasar, 1932f: 151

hindu Arrow, 1931: 289 A: AF KA PA ORR

SCARABAEIDAE

185

lahorensis Kabakov, 2008: 412 A: PA
lunatus Harold, 1868b: 84 A: FUJ GUA HAI HKG TAI AUR ORR
lamellatus Boucomont, 1924: 115 [RN]
laminatus Taschenberg, 1879: 107 [HN]
taschenbergi Gillet, 1925a: 10 [RN]

nagasawai Matsumura, 1938: 60 A: TAI
paliceps Arrow, 1931: 287 A: PA UP
piffli Petrovitz, 1961c: 103 A: PA
ramosus Wiedemann, 1823: 13 (Copris) A: HP KA PA UP ORR
tragoides Boucomont, 1914a: 230 A: YUN ORR
tragus Fabricius, 1792a: 56 (Scarabaeus) A: CHQ FUJ HEB SC SCH SHX TAI YUN ORR
fulvicornis Motschulsky, 1854c: 45
hypertropis Olsoufieff, 1900: 269
ibex Motschulsky, 1854d: 63 [HN]
mandarinus Harold, 1859: 224 [RN]
neptunulus Van Lansberge, 1883b: 50
tsunoellus Matsumura, 1938: 55 (Caccobius)

subgenus Eremonthophagus Zunino, 1979: 7 type species Onthophagus sticticus Harold, 1867
heydeni Harold, 1875c: 216 A: IN IQ SY
infuscatus Klug, 1845: [4], pl. xli A: OM SA YE AFR
oberthueri d’Orbigny, 1898c: 161 A: IN IQ SA
trihastatus Reitter, 1909b: 78

semicinctus d’Orbigny, 1898e: 242 A: AF PA ORR
bufo Arrow, 1907b: 420

sticticus Harold, 1867b: 50 N: AG EG MO TU A: AE IN SA SI AFR
biarcuatus van Lansberge, 1882: xxiii

transcaspicus König, 1889: 302 N: AG EG LB MO TU A: AE AF IN IQ IS OM PA SA SI TD TM TR UZ AFR ORR
lineatus Reitter, 1889b: 102
obsoletus Fairmaire, 1875a: 487

subgenus Exonthophagus Kabakov, 2006a: 293 type species Onthophagus haroldi Ballion, 1871
haroldi Ballion, 1871a: 332 A: AF IN KI KZ TD TM UZ WS XIZ
viridilimbatus Olsoufieff, 1907b: 195

viriditinctus Reitter, 1892a: 183 [=1893a: 62] A: IN
subgenus Furconthophagus Zunino, 1979: 10 type species Scarabaeus furcatus Fabricius, 1781
amicus Gillet, 1925b: 230 (Caccobius) A: NP SD UP ORR
centricornis Fabricius, 1798: 33 (Copris) A: AF AP HP PA UP ORR
luteipennis Wiedemann, 1823: 20 (Copris)
minutus Motschulsky, 1858d: 54 [HN]
promissus Harold, 1868c: 104 [RN]

dapcauensis Boucomont, 1921a: 30 A: GUA HKG TAI ORR

186

SCARABAEIDAE

furcatus Fabricius, 1781a: 30 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ FR GBi GE GG GR HU IT
KO KZ MC MD ME PL PT RO SB SK SL SP ST SZ TR UK N: MO A: IN IQ IS KZ SA SY TM TR
bicornutus Mulsant, 1842: 149
bidentarus Mulsant, 1842: 149
degener Mulsant, 1842: 149
fanatus Laporte, 1840: 87
fossatus Boucomont & Gillet, 1927: 122 [unjustified emendation]
laminiger Mulsant, 1842: 149
mniszekii Hochhuth, 1873b: 127
rubellus Mulsant, 1842: 150
rutilipennis Reitter, 1892a: 194
terminatus Faldermann, 1835a: 249
vitulus Laicharting, 1781: 26 (Scarabaeus) [HN]

parvulus Fabricius, 1798: 36 (Copris) A: PA ORR
schoolmeestersi Ochi & Kon, 2007: 81 A: NP
sellatus Klug, 1845: [4], pl. xli [DA] N: EG LB A: CY IS JO LE SA SY AFR
analis P.H. Lucas, 1846c: pl. 23
euthymi Mulsant & Godart, 1875a: 73 [=1875b: 409]
pygialis J. Thomson, 1858: 55

troglodyta Wiedemann, 1823: 20 (Copris) A: AF PA UP ORR
spitiensis Gillet, 1925b: 235

variegatus Fabricius, 1798: 36 (Copris) N: EG A: AE AF IN IQ OM PA QA SA YE AFR ORR
venustulus Erichson, 1843: 233

subgenus Gibbonthophagus Balthasar, 1963a: 163 type species Onthophagus atripennis C.O. Waterhouse, 1875
amamiensis Nomura, 1965: 1 A: JA (Amami-Ôshima)
apicetinctus d’Orbigny, 1898a: 148 A: JA
yaeyamanus Nomura, 1964: 47

atripennis C.O. Waterhouse, 1875a: 77 A: BEI FE FUJ JA NC NO SC SCH SHA
akirai Matsumura, 1937b: 153
cicatricosus Balthasar, 1935f: 317
ibonus Matsumura, 1937b: 155
kogatanus Matsumura, 1937b: 158
rubrotinctus d’Orbigny, 1898a: 148
shigeoi Matsumura, 1937b: 163

balthasari Všetečka, 1939b: 43 A: SHG YUN ZHE ORR
dissentaneus Balthasar, 1964b: 289 A: GAN YUN
dubernardi Boucomont, 1914a: 226 A: CHQ GUI HP HUB SCH SD TAI XIZ YUN ORR
duporti Boucomont, 1914a: 228 A: AP HP UP YUN ORR
gracilipes Boucomont, 1914a: 219 A: YUN ORR
incollaris Kabakov, 2008: 414 A: YUN
kentingensis Nomura, 1973b: 41 A: TAI

SCARABAEIDAE
kleinei Balthasar, 1935f: 314 A: FUJ SCH
limbatus Herbst, 1789: 207 (Scarabaeus) A: YUN ORR
analis Germar, 1813: 115
nuchidens Fabricius, 1798: 31 (Copris)

luridipennis Boheman, 1858: 45 A: AP HKG TAI UP YUN ORR
fuscopunctulatus Boucomont, 1914b: 293 [RN]
fuscopunctatus van Lansberge, 1883b: 48 [HN]

proletarius Harold, 1875a: 89 A: FUJ HAI HKG TAI YUN ORR
mendicus Gillet, 1924: 67

rectecornutus van Lansberge, 1883b: 49 A: AP BT FUJ GUA GUI NP TAI YUN ORR
luridus R. Paulian, 1933a: 98

solivagus Harold, 1886b: 290 A: BEI CHQ FE FUJ HEN HUB NC NO SC SCH SHX
confucius Goidanich, 1926e: 135

susterai Balthasar, 1952: 225 A: FUJ
taurinus White, 1844: 424 A: FUJ HKG HUB SHG TAI ORR
egurianus Matsumura, 1938: 57
mushensis Matsumura, 1938: 59

viduus Harold, 1874b: 291 A: BEI FUJ GAN HEI JA JIX LIA NC NO SC SCH SHG TAI ZHE
aequiperus Matsumura & Yohena, 1937b: 152
amenthus Matsumura, 1937c: 2
chibanus Matsumura & Yohena, 1937b: 153
chosensis Matsumura, 1937c: 2
flavonotatus d’Orbigny, 1898a: 147
futsukaichianus Matsumura & Yohena, 1937b: 152
hyuganus Matsumura, 1937b: 155
jedensis Matsumura & Yohena, 1937b: 156
jononis Matsumura, 1936: 63 (Caccobius)
komabellus Matsumura, 1937b: 159
micellus Matsumura, 1937c: 3
minoi Matsumura & Yohena, 1937b: 160
misujianus Matsumura, 1937b: 161
miyazakianus Matsumura, 1937b: 161
narashinensis Matsumura & Yohena, 1936: 64 (Caccobius)
oishii Matsumura, 1937b: 162
okushirianus Matsumura, 1937b: 162
putealis Matsumura, 1938: 61
rubromaculatus H.J. Kolbe, 1886: 187
shurianus Matsumura & Yohena, 1937b: 164
spurius Matsumura, 1937b: 165
takabayashii Matsumura, 1937b: 165
yumigatanus Matsumura, 1937b: 169

187

188

SCARABAEIDAE

subgenus Indachorius Balthasar, 1941b: 161 type species Indachorius hirtus Balthasar, 1941 (=Onthophagus
nefarius Balthasar, 1963)
arai Masumoto, 1989: 37 A: SCH ORR
chinensis Balthasar, 1953b: 224 (Indachorius) A: FUJ
hsui Masumoto, K.-M. Chen & Ochi, 2004: 126 A: TAI
koshunensis Balthasar, 1941d: 183 A: TAI
magnini R. Paulian, 1933c: 207 A: YUN ORR
nefarius Balthasar, 1963a: 448 [RN] A: NP ORR
hirtus Balthasar, 1941b: 162 (Indachorius) [HN]

platypus Y.-W. Zhang, 1997: 763 A: HUB
pumilus Kabakov, 2014: 52 A: YUN
suginoi Ochi, 1984: 65 A: JA
yaoi Masumoto, Ochi & C.-F. Lee, 2014: 261 A: TAI
subgenus Indonthophagus Kabakov, 2006a: 154 type species Scarabaeus mopsus Fabricius, 1792
ensifer Boucomont, 1914a: 220 A: PA ORR
hastifer van Lansberge, 1885b: 380 A: FUJ GUA HAI TAI ORR
agilis Matsumura, 1938: 56
puetalis Matsumura, 1938: 61
turmalis Gillet, 1924: 66

mopsus Fabricius, 1792a: 58 (Scarabaeus) A: AF KA PA SD UP ORR
gracilicornis Germar, 1813: 114

nitidulus Klug, 1845: [4], pl. xli N: EG A: AE AF IN IQ PA SA AFR
mediofasciatus d’Orbigny, 1898a: 203
tetraspilus Fairmaire, 1887a: 114

subgenus Macronthophagus Ochi, 2003a: 263 type species Onthophagus uenoi Ochi, 1995
cludtsi Ochi, 2003b: 294 A: YUN ORR
diabolicus Harold, 1877a: 78 A: GUX YUN ORR
manipurensis Arrow, 1907b: 426 A: AP XIZ YUN ORR
rubricollis Hope, 1831: 22 A: AP BT HP NP SD UP XIZ YUN ORR
subgenus Matashia Matsumura, 1938: 63 type species Matashia mushana Matsumura, 1938 (=Onthophagus
yubarinus Matsumura, 1937)
fossor Arrow, 1931: 300 A: SD
kuluensis Bates, 1891a: 12 A: HP KA NP UP XIZ ORR
lutosopictus Fairmaire, 1897a: 69 A: JA (Ishigaki-jima)
ohbayashii Nomura, 1939: 35 A: JA NC SC
hvangeus Stebnicka, 1973: 117

yubarinus Matsumura, 1937b: 168 A: TAI
ivae Balthasar, 1963b: 92
mushana Matsumura, 1938: 63 (Matashia)
taiheizanus Matsumura, 1937b: 168

SCARABAEIDAE

189

subgenus Micronthophagus Balthasar, 1963: 165 type species Onthophagus hystrix Boucomont, 1914
hystrix Boucomont, 1914: 238 A: SD UP ORR
incertus d’Orbigny, 1898e: 241 A: PA
konoi Matsumura, 1938: 58 A: FUJ TAI
liui Masumoto, Ochi & C.-F. Lee, 2014: 263 A: TAI
melanocephalus Klug, 1845: [3] pl. xli N: AG EG LB TU A: IS SI AFR
ochreatus d’Orbigny, 1898e: 241 A: AE PA SA AFR ORR
vigilans Boucomont, 1921a: 38 A: YUN ORR
wangi Masumoto, K.-M. Chen & Ochi, 2004: 127 A: TAI
subgenus Onthophagiellus Balthasar, 1935f: 308 type species Onthophagus crassicollis Boucomont, 1913
andreji Prokofiev, 2014: 12 A: NP
crassicollis Boucomont, 1913: 418 [RN] A: AP YUN ORR
nanus van Lansberge, 1883b: 67 [HN]

popovi Kabakov, 1994b: 306 A: YUN ORR
taiyaruensis Masumoto, 1977: 28 A: TAI
unguiculatus Kabakov, 1994b: 306 A: YUN
subgenus Onthophagus Latreille, 1802a: 141 type species Scarabaeus taurus Schreber, 1759
bivertex Heyden, 1887b: 302 A: BEI FE FUJ GAN HEB HEI JA MG NC NO SC SCH SHX YUN
brevicornis Yohena, 1936: 435
kozunonis Matsumura, 1937b: 159
minokuchianus Matsumura, 1934: 67
oniellus Matsumura, 1937b: 163
shinanensis Matsumura, 1934: 68
yohenai Matsumura, 1937b: 167 [RN]

expansicornis Bates, 1891a: 11 A: AF HP PA UP ORR
hajimei Masumoto, 1985: 78 A: TAI
hayashii Masumoto, 1991b: 153 A: TAI
illyricus Scopoli, 1763: 11 (Scarabaeus) E: AB AL AR AU AZ BE BH BY CR CT CZ LU FR GE GG GR HU IT KZ MC
MD ME PL PT RO SB SK SL SP ST SV SZ UK A: AF CY IN IQ IS LE SY TR
brisouti d’Orbigny, 1898e: 233
menetriesii Faldermann, 1835a: 239
piliger Mulsant, 1842: 140
rufipes Mulsant, 1842: 140
urus Ménétriés, 1832: 175

masumotoi Ochi, 1985: 49 A: TAI
neofurcatus Goidanich, 1926b: 39 [RN] A: FUJ HEI ORR
furcatoides Reitter, 1892a: 176 [=1893a: 55] [HN]

quaestus Sharp, 1875b: 55 A: HP UP ORR
roubali Balthasar, 1935f: 316 A: SCH TAI
sakainoi Masumoto, 1991b: 155 A: TAI

190

SCARABAEIDAE

sauteri Gillet, 1924: 68 A: TAI
matsuokai Matsumura & Yohena, 1937b: 160

setchan Masumoto, 1984a: 80 A: TAI
tatsienluensis Balthasar, 1942a: 121 A: FUJ SCH TAI ORR
taurus Schreber, 1759: 7 (Scarabaeus) E: AB AL AR AU AZ BE BH BU BY CR CT CZ FR GB GE GG GR HU IT KZ
LT LU MC MD ME NL PL PT RO SB SK SL SP ST SV SZ TR UK N: AG EG LB MO TU A: AF CY IN IQ IS KI KZ
SY TD TM TR UZ XIN AURi NARi
alternans d’Orbigny, 1898a: 149
alternatus d’Orbigny, 1898a: 149
athenae Goidanich, 1925a: 23
bos A. Villa & G.B. Villa, 1833: 34
bovillus Mulsant, 1842: 139
capra Fabricius, 1787a: 15 (Scarabaeus)
capreolus Mulsant, 1842: 139
castanonotus Seabra, 1907: 104
circumcinctus Escalera, 1914a: 150
corniger Geoffroy, 1785: 16 (Copris)
cruoreus Schrank, 1798: 397 (Scarabaeus)
femineus Mulsant, 1842: 140
fuscipennis Mulsant, 1842: 140
glareanus Gistel, 1857b: 577
mendax Mulsant, 1842: 140
morio Brullé, 1832: 169
nigrovirescens Mulsant, 1842: 140
quadrum Kugelann, 1792a: 276 (Scarabaeus)
recticornis Leske, 1785: 45 (Scarabaeus)
reichardti S.I. Medvedev, 1930: 57
rubripennis Goidanich, 1924: 176
rugosus Poda von Neuhaus, 1761: 20 (Scarabaeus)
saharae Bedel, 1911a: 29

terminatus Eschscholtz, 1822: 32 (Copris) A: TAI (Lanyu, Lutao) ORR
yangi Masumoto, Tsai & Ochi, 2006: 140 A: TAI
subgenus Palaeonthophagus Zunino, 1979: 8 type species Scarabaeus vacca Linnaeus, 1767
Bubalonthophagus Ádám, 1994a: 8 type species Onthophagus furciceps Marseul, 1869
Citellonthophagus Ádám, 1994a: 8 type species Scarabaeus vitulus Fabricius, 1777
Haplonthophagus Ádám, 1994a: 9 type species Scarabaeus lemur Fabricius, 1781
Kisonthophagus Ádám, 1994a: 8 type species Scarabaeus ovatus Linnaeus, 1767
Lucidonthophagus Ádám, 1994a: 8 type species Copris lucidus Illiger, 1800
Troglonthophagus Ádám, 1994a: 8 type species Scarabaeus semicornis Panzer, 1798

aerarius Reitter, 1892a: 205 [=1893a: 84] N: AG EG LB MO TU E: AB AR A: AF IN IS JO SA SY
diversicornis Kirschenblatt, 1935: 201

SCARABAEIDAE

191

afghanus Petrovitz, 1961a: 36 A: AF TD
akinini König, 1889: 303 A: TD TM UZ
albarracinus Baraud, 1979a: 24 E: SP (Teruel)
aleppensis aleppensis L. Redtenbacher, 1843: 985 A: AF CY IN IQ JO LE SY TR
aleppensis hyrcanicus Kabakov, 1982: 15 A: IN
amirus Kabakov, 1982: 12 A: AF IN
anatolicus Petrovitz, 1962b: 108 A: TR
andalusicus andalusicus Waltl, 1835: 66 E: IT MA PT SP N: AG MO TU
italicus Goidanich, 1925d: 104
marginatus Seabra, 1907: 122 [HN]
maurus P.H. Lucas, 1846c: pl. 23

andalusicus persianus Olsoufieff, 1900: 270 E: AB AR ST A: AF IN
angorensis Petrovitz, 1963c: 235 E: AB AR BU GG GR HU MC RO TR A: IN IS LE SY TM TR
arnoldii Kabakov, 1982: 8 A: TD TM
baraudi Nicolas, 1964: 8 E: AU FR GE IT SZ
basipustulatus Heyden, 1889a: 327 E: AB AR A: AF KI SY TD TR UZ
bonsae Zunino, 1976: 97 A: LE
bytinskii Balthasar, 1960d: 201 A: IS LE
carpanetoi Pittino, 1982a: 189 A: IN JO TR
clitellifer Reitter, 1894e: 189 A: BEI ES FE GUI HEI LIA MG NIN NMO NC NO QIN
transbaicalicus Csiki, 1901: 109

coenobita Herbst, 1783: 11 (Scarabaeus) E: AL AR AU BE BH BU BY CR CT CZ DE FR GB GE GG GR HU IT KO
MC MD ME LA LT LU NL PL PT RO SB SK SL SP ST SV SZ TR UK A: IS TM TR
cuspidiusculus Mulsant, 1842: 128
fulgens Brahm, 1790: 38 (Scarabaeus)
fulvipes Faldermann, 1835a: 243
rufipes Ménétriés, 1832: 178
subprominulus Mulsant, 1842: 129
tenuicornis Preyssler, 1790: 44 (Scarabaeus)
tricuspis Mulsant, 1842: 128

conspersus Reitter, 1892a: 195 [=1893a: 74] E: AB A: AF IN KA TD TM
kabulicus Balthasar, 1971b: 56

cruciatus Ménétriés, 1832: 178 E: AB AR GG GR (Ródos) A: CY IN IQ IS JO LE SY TR
capilossus d’Orbigny, 1905: 482
centromaculatus L. Redtenbacher, 1843: 985

dacatrai Pittino, 2004b: 180 A: JO TR
dellacasai Pittino & Mariani, 1981: 98 E: AL BU GR MC TR
dorsosignatus d’Orbigny, 1898e: 237 E: AB (Naxçivan) AR A: AF IN IQ TR
flavofasciatus d’Orbigny, 1898a: 195

eulaminicornis Pittino, 2006c: 109 [RN] E: BU GR
laminicornis Pittino, 2004b: 175 [HN]

192

SCARABAEIDAE

excubitor Ziani & Gudenzi, 2006: 227 A: IN
finschi Harold, 1877d: 333 A: KI KZ MG TD UZ XIN
capreolus Ballion, 1878: 279
paraprotzenkoi Tesař, 1967: 137
protzenkoi Tesař, 1967: 135

fissicornis Steven, 1809: 34 (Copris) E: AB AL AR BU CR GG GR MC MD ME RO SB ST TR TR UK (Krym) A: IN
IQ IS SY TM TR
quadridentatus Ménétriés, 1832: 176 [HN]

fissinasus Fairmaire, 1895c: cix E: GR (Sámos) A: IN IQ IS LE SY TR
flagrans Reitter, 1892a: 199 [=1893a: 78] A: IN KI KZ TD TM UZ
formaneki Reitter, 1897g: 204 E: AB AR A: IN SY TR
albinae Reitter, 1897g: 204
bifenestratus Reitter, 1898c: 342

fortigibber Reitter, 1909b: 79 E: AB GG ST A: TR
tricuspis Semenov, 1900a: 93 [HN]
unxovi Bogatchev, 1930: 87

fracticornis Preyssler, 1790: 99 (Scarabaeus) E: AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG GR
HU IT KO LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK A: CY IN KZ SY TR
WS “Palestina”
anonymus G. Delabie, 1956: 176
assimilis Hoppe, 1795: 28 (Scarabaeus)
flavescens Seabra, 1907: 129
herbsti Brahm, 1790: 39 (Scarabaeus)
irroratus Faldermann, 1835a: 242
marginatus Mulsant, 1842: 121
nasutus Mulsant, 1842: 120
pauperatus Mulsant, 1842: 120
semiflavus Reitter, 1892a: 192 [=1893a: 71]
sublaminatus Mulsant, 1842: 120
subrecticornis Mulsant, 1842: 119
tricuspidus Mulsant, 1842: 120
virescens Seabra, 1907: 129

furciceps Marseul, 1869b: 379 A: IN LE TR
gibbulus gibbulus Pallas, 1781: 7 (Scarabaeus) E: AU BY CR CT CZ EN FI GE HU IT KZ LA LT MD NT PL RO SB
SK SL ST SV SZ UK A: BEI ES FE GAN HEB HEI IN IQ JA KI KZ MG NC NIN NMO SC SCH SHX SY TD TM
TR UZ WS XIN
austriacus Panzer, 1793d: 6 (Scarabaeus)

gibbulus rostrifer Reitter, 1892a: 192 [=1893a: 71] E: AB AR GG A: IN TR “Kurdistan”
glasunowi D. Koshantschikov, 1894c: 105 A: KI TD UZ
gorodinskii Kabakov, 2008: 418 A: SCH
grossepunctatus Reitter, 1905c: 245 E: AL AR AU BH BU CR CT CZ FR GG GR HU IT KZ MC MD ME PL PT RO
SB SK SL SP ST TR UK A: TR

SCARABAEIDAE

193

hermonensis Baraud, 1982b: 237 A: IS LE SY
hissariensis Kabakov, 2006a: 261 A: TD
isikdaghensis Pittino, 2004b: 194 A: TR
joannae Goljan, 1953: 66 E: AU BE BU CR CZ DE FR GB GE GR HU IT LU NL NR PL PT RO SK SP SV SZ UK
kindermanni Harold, 1877b: 99 E: BU GR HU MC MD RO SB TR UK A: TR
kolenatii Reitter, 1892a: 200 [=1893a: 79] E: AB
koryoensis J.I. Kim, 1985: 8 A: SC
laticornis Gebler, 1823: 128 A: ES FE MG NE NIN NMO
latigena d’Orbigny, 1898e: 234 E: PT SP N: MO
distinctus Fuente, 1910: 448

lemur Fabricius, 1781: 495 (Scarabaeus) E: AB AL AR AU BE BH BU CR CZ FR GE GR HU IT LS LU MC MD ME
PL PT RO SB SK SL SP ST SZ TR UK A: IN SY TM TR
cinctus Dalla Torre, 1879: 105
cruciatus Depoli, 1924: 162
curvicinctus Mulsant, 1842: 109
decempunctatus Schaller, 1783: 237 (Scarabaeus)
depolii Goidanich, 1925c: 95
egenus Mulsant, 1842: 109
germari Depoli, 1917: 191
glandicolor Mulsant, 1842: 109
ibrydus O.G. Costa, 1828: 92 (Scarabaeus)
inquinatus Goidanich, 1925c: 95
lineolatus Mulsant, 1842: 109
mutabilis Mulsant, 1842: 109
plurimacula Depoli, 1924: 162
quadrituberculatus Laicharting, 1781: 23 (Scarabaeus)

lemuroides d’Orbigny, 1898d: 177 A: IN IQ TR
leucostigma Steven, 1806: 158 [=1812: 121] (Copris) E: CT KZ ST UK A: AF KI KZ TD TM TR UZ
leucomelas Solsky, 1876a: 309

lucidus Illiger, 1800b: 106 (Copris) E: AB AL AR AU BU CR GG GR HU MC MD ME RO SB SK ST TR UK A: CY
IN IQ IS JO SY TR
lucidus Sturm, 1800: 95 (Copris) [HN]
nitidicollis Brullé, 1832: 168
punctatominiatus Goidanich, 1925b: 78

macedonicus Mikšić, 1959b: 39 E: MC (Petrina planina)
marginalis marginalis Gebler, 1817: 315 (Copris) E: AB AR BU GR KZ MC RO ST TR A: AF BEI CY ES FE HEB HEI
IN IS KI KZ LIA MG NC NMO SC SCH SHX SY TD TM TR WS
circumscriptus Faldermann, 1835a: 241
marmoratus Ménétriés, 1832: 176

marginalis nigrimargo Goidanich, 1926c: 54 A: AF HP KA NP PA UP XIZ ORR
marginalis przewalskii Kabakov, 2006a: 225 A: KI KZ XIN
obtusicornis Motschulsky, 1859b: 305 [HN]

194

SCARABAEIDAE

massai Baraud, 1975d: 292 E: IT (Sicilia)
medius Kugelann, 1792a: 271 (Copris) E: AB AU AZ BE BH BU CR CZ DE FR GB GE GG GR HU IT LU MC ME
NL PL PT RO SB SK SL SP ST SZ TR A: IN KZ TR
affinis Sturm, 1800: 102 (Copris)
confluens Gistel, 1857b: 573
intermedius Mulsant, 1842: 133 [HN]
medius Panzer, 1796a: 4 (Scarabaeus) [HN]
propinquus Mulsant, 1842: 134
sublineolatus Mulsant, 1842: 134
vicinus Mulsant, 1842: 133

medvedevi Kabakov, 1982: 9 A: KZ TD UZ
merdarius Chevrolat, 1865: 350 E: PT SP
mediofasciatus d’Orbigny, 1898a: 203

muelleri P. Novak, 1921: 100 A: IN IS JO LE SY E: TR
novaki Gillet & Boucomont, 1927: 125 [RN]

nebulosus Reiche, 1864: 239 N: AG EG LB MO TU A: IN IQ IS JO SA SY YE
nikolajevi Kabakov, 2006a: 257 A: GAN SCH
nuchicornis Linnaeus, 1758: 347 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG
GR HU IR IT KO KZ LA LS LT LU MD NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK A: BEI ES FE KZ
MG TM TR WS XIN NARi
acornis Geoffroy, 1785: 14 (Copris)
alpinus Kolenati, 1846: 5
dillwynii Stephens, 1830: 174 (Scarabaeus)
immaculatus Mulsant, 1842: 116
indistinctus Mulsant, 1842: 116
planicornis Herbst, 1789: 210 (Scarabaeus)
rhinoceros Melsheimer, 1844: 134
rubripes Mulsant, 1842: 116
submarginalis J.R. Sahlberg, 1926: 11
trituberculatus Schrank, 1798: 397 (Scarabaeus)
vulneratus Mulsant, 1842: 116
xiphias Fabricius, 1792a: 59 (Scarabaeus)

nurestanicus Kabakov, 1982: 12 A: AF
ocellatopunctatus C.O. Waterhouse, 1875a: 79 A: JA
olsoufieffi Boucomont, 1924: 114 [RN] A: BEI ES FE JA LIA NC NO SC
kandai Matsumura, 1937b: 156
penicillatus Olsoufieff, 1907b: 191 [HN]
uedanus Matsumura, 1937b: 166

opacicollis Reitter, 1892a: 192 [=1893a: 71] E: AB AL BH BU CR FR GR HU IT MC ME PT SK SL SP ST SZ TR N:
AG MO TU A: CY IN IS SY TR
opacicollis d’Orbigny, 1898e: 236 [HN]
schatzmayri Pierotti, 1959: 115

SCARABAEIDAE

195

osellai Pittino, 1982a: 197 A: IN SY TR
ovatus Linnaeus, 1767: 551 (Scarabaeus) E: AB AL AU BE BH BU BY CR CT CZ DE EN FR GE GG HU IT KZ LA
LS LU MC MD NL PL PT RO SB SK SL SP ST SV SZ UK A: IN KZ TR
fucatrus Mulsant, 1842: 152
subaenescens Goidanich, 1926d: 76
subaeneus Ménétriés, 1832: 179

panici Petrovitz, 1964a: 25 E: AL BH GR MC ME
parmatus Reitter, 1892a: 181 [=1893a: 60] E: AB AR HU MD RO ST A: IN SY TR
petrovitzianus Pittino, 2006c: 109 [RN] A: TR
citellorum Petrovitz, 1967b: 325 [HN]
petrovitzi Pittino, 2004b: 168 [HN]

pljushtchi Ivanova, 2012b: 45 A: AF
ponticus Harold, 1883: 434 E: AB AR BU GG GR MD RO ST UK A: IN IQ IS LE SY TR
citellorum S.I. Medvedev, 1965a: 186
furcicornis Reitter, 1892a: 202 [=1893a: 81]
granulosus D. Koshantschikov, 1894b: 102 [RN]
gorhami Shipp, 1895a: 178 [RN]
granulifer Reitter, 1892a: 204 [=1893a: 204] [HN]
krali Balthasar, 1963a: 603

pseudocaccobius Reitter, 1889b: 102 A: TD TM UZ
psychopompus Ziani & Gharakhloo, 2010: 34 A: IN
pygargus Motschulsky, 1845a: 56 E: AB (Naxçivan) A: AF IN KI KZ TD TM UZ XIN
saiga Ballion, 1871a: 332

quadrinodus Reitter, 1896h: 287 A: KZ TD UZ
rachelis Martín Piera, 1985: 442 A: XIN
rakovici Pittino, 2004b: 192 E: AR A: IN TR
rodentium Pittino, 2004b: 155 E: GR
ruficapillus Brullé, 1832: 169 E: AB AL AR AU BH BU CR CT CZ FR GG GR HU IT KO KZ MC MD ME PL PT RO
SB SK SL SP ST SZ TR UK A: CY IN IQ IS JO KZ SY TM TR
cutai Roubal, 1933b: 139
guilanensis Pittino, 1982b: 517

rufimanus Kabakov, 1982: 11 A: TM
sacharovskii Olsoufieff, 1918: 83 E: AR GG A: IN TR
samai Ziani, 2011a: 47 A: IN
scabriusculus Harold, 1873: 104 A: ES FE KZ MG NC NE NMO NO
semicornis Panzer, 1798: 10 (Scarabaeus) E: AU BU CR CT CZ FR GE GR HU IT KZ MC MD PL PT RO SB SK SL
SP ST SZ TR UK A: KZ MG TM UZ WS
angulicornis Mulsant, 1842: 146
decipiens Mulsant, 1842: 146

sericatus Reitter, 1892a: 192 [=1893a: 71] E: AB AL BH BU CR GR MC ME ST TR “Caucasus” A: IN IS LE SY TR
shapovalovi Gusakov, 2012: 3 A: KZ
shirakii Nakane, 1960: 1 A: JA (Amami-Ôshima)

196

SCARABAEIDAE

silus Balthasar, 1960c: 93 A: KI KZ TM UZ
kirgisicus Protsenko, 1963: 16

similis Scriba, 1790: 56 (Copris) E: AU BE BH BU BY CT CZ DE EN FI FR GB GE GR HU IR IT LT MC ME NL NT
PL PT SK SL SP SV SZ TR UK N: AG LB MO TU A: IS SY TR
speculifer Solsky, 1876a: 308 E: AR GG A: AF IN IS KI KZ TD TM TR
strabo Reitter, 1892a: 181 [=1893a: 60] A: TR
stylocerus Graëlls, 1851: 128 E: PT SP
rubrescens Seabra, 1907: 110

surmelii Petrovitz, 1963c: 236 E: AR A: IN TR
sutleinensis Splichal, 1910: 567 A: AF HP KA PA XIZ ORR
oncursator Balthasar, 1960c: 91
marxi Splichal, 1912: 467

suturellus Brullé, 1832: 168 E: AB AR GR ST TR UK A: IN SY TM TR “Palestina”
trochiscobius Kolenati, 1846: 7

tesquorum Semenov & S.I. Medvedev, 1927: 183 E: HU RO TR UK
trigibber Reitter, 1892a: 206 [=1893a: 85] E: SP N: AG MO TU
trispinus Reitter, 1892b: 62 A: TD TM UZ
umbilicopunctatus S.I. Medvedev, 1959: 869

truchmenus iranicus Kabakov, 2006a: 235 A: IN
truchmenus truchmenus Kolenati, 1846: 8 E: AB AR GG ST A: IN LE SY TR
araxicola Reitter, 1892a: 190 [=1893a: 69]
latiplaga Reitter, 1897h: 219

truchmenus turkmenicus Kabakov, 2006a: 235 A: TM
truchmenus xerxes Petrovitz, 1965d: 674 A: AF IN
vacca Linnaeus, 1767: 547 (Scarabaeus) E: AB AL AR AU AZ BE BH BU BY CR CT CZ FR GE GG GR HU IT KO
LA LT LU MC MD ME PT RO SB SK SL SP ST TR UK N: AG MO A: IN SY KZ TM TR “Palestina” AURi
aeruginosus Schrank, 1798: 339 (Scarabaeus)
antilope Motschulsky, 1845a: 57 [HN]
basalis Mulsant, 1842: 134
conspurcatus Geoffroy, 1785: 14 (Copris)
difficilis Mulsant, 1842: 133
lusitanica Seabra, 1907: 133
similis Mulsant, 1842: 133
tricornis Fischer von Waldheim, 1844: 43

verticicornis Laicharting, 1781: 22 (Scarabaeus) [NP] E: AL AR AU BE BH BU CR CZ FR GBi GE GG GR IT KO
LU MC MD ME PL PT RO SB SK SL SP ST SZ UK A: IN SY TR “Turkestan”
bituberculatus Scriba, 1790: 57 (Copris)
distinguendus Mulsant, 1842: 125
infucatus Mulsant, 1842: 125
nutans Fabricius, 1787a: 15 (Scarabaeus)
subconvexus Seabra, 1907: 108

SCARABAEIDAE

197

viridis Ménétriés, 1832: 177 E: AB A: IN
vitulus Fabricius, 1777: 209 (Scarabaeus) E: AB AR AU BU CR CT CZ FR GE GG GR HU IT KZ MD PL RO SB SK
ST SZ UK A: KI KZ MG TR WS
camelus Fabricius, 1787a: 13 (Scarabaeus)
humeralis Mikšić, 1950: 167
kaszabi Balthasar, 1963b: 91
thracum Pittino, 2004b: 167

vlasovi S.I. Medvedev, 1958: 717 A: AF IN KZ TD TM UZ
zuninoi Martín Piera, 1985: 436 A: KZ
zuvandi Qarjagdy, 1939: 86 E: AB A: AF IN TM TR
kryzhanovskii Kabakov, 1982: 16
rechingeri Petrovitz, 1980: 605

subgenus Paraphanaeomorphus Balthasar, 1959b: 465 type species Onthophagus phanaeicollis van Lansberge,
1883
acuticornis Endrődi, 1973b: 124 A: FUJ ORR
argyropygus Gillet, 1927: 256 A: HKG JA (Ryukyus) TAI ORR
bifasciatus Fabricius, 1781: 25 (Scarabaeus) A: NP SD UP ORR
birmanicus Harold, 1879a: 226

carinensis Boucomont, 1914a: 223 A: SCH YUN ORR
missor Balthasar, 1938c: 97

cyaneus Kabakov, 2006b: 127 A: PA
frugivorus Arrow, 1931: 336 A: SD ORR
jeannelianus shokhini Kabakov, 2014: 53 A: SCH YUN
phanaeiformis Boucomont, 1914a: 224 A: AP ORR
sobrius Balthasar, 1960b: 188 A: FUJ
trituber jacobsoni Kabakov, 2006a: 330 A: LIA ZHE
trituber trituber Wiedemann, 1823: 17 (Copris) A: JA GUA HKG SC SHG TAI YUN ORR
suturalis White, 1844: 424
umeonoi Matsumura, 1938: 61

vaulogeri Boucomont, 1923a: 10 A: AP UP YUN ORR
subgenus Parascatonomus R. Paulian, 1932b: 212 [RN] type species Coomanius politus R. Paulian, 1932
(=Onthophagus renaudpauliani Ochi & Araya, 1996)
Coomanius R. Paulian, 1932a: 205 [HN] type species Coomanius politus R. Paulian, 1932 (=Onthophagus renaudpauliani
Ochi & Araya, 1996)
Pseudonthophagus Balthasar, 1959b: 466 type species Onthophagus penicillatus Harold, 1789

acuticollis acuticollis Gillet, 1927: 255 A: HUN TAI ZHE
marani Všetečka, 1942: 255

acuticollis sakishimanus Nomura, 1976a: 29 A: JA (Ishigaki-jima, Iriomote-jima)
anguicorius Boucomont, 1914a: 211 A: FUJ GUA HAI HKG TAI ORR
midorianus Matsumura, 1938: 59

aokii Nomura, 1976a: 31 A: JA

198

SCARABAEIDAE

discedens Sharp, 1875b: 49 A: FUJ SCH SD YUN ORR
formosanus Gillet, 1924: 67 A: TAI
yanoi Matsumura, 1938: 62

funebris funebris Boucomont, 1920b: 309 A: YUN ORR
horni Balthasar, 1935f: 314 A: FUJ SCH “Korea”
impar Kabakov, 2006b: 130 (Pseudonthophagus) A: PA
itoi Nomura, 1976a: 28 A: JA (Okinawa-shotō)
kiuchianus Masumoto, M.-M. Yang & Ochi, 2004: 358 A: TAI
klapperichi Balthasar, 1953b: 228 A: FUJ GUA TAI
kuatunensis Balthasar, 1942a: 120 A: FUJ SCH TAI YUN
lakyim Masumoto, 1990: 50 A: YUN ORR
ohmomoi Miyake, 2000: 106
panfilovi Kabakov, 1992: 205 (Parascatonomus)

laotianus Boucomont, 1919a: 314 A: YUN ORR
miyakei Ochi & Araya, 1992: 89 A: GUX JA (Ryukyus) TAI ORR
mulleri van Lansberge, 1883b: 56 A: YUN ORR
murasakianus carnarius Nomura, 1976a: 32 A: JA (Amami-Ôshima)
okinawanus Nomura, 1976a: 33

murasakianus miyakoinsularis Ochi, Y. Miyake & Kusui, 1999: 35 A: JA (Ryukyus)
murasakianus murasakianus Nomura, 1976a: 30 A: JA (Ishigaki-jima, Iriomote-jima)
naviculifer Boucomont, 1920b: 315 A: YUN ORR
nitidus nitidus C.O. Waterhouse, 1875a: 78 A: HKG JA LIA SC SHG TAI
yakushimanus Nomura, 1976a: 31

penicillatus Harold, 1879a: 225 A: FUJ GUI SCH SD YUN ORR
rudis Sharp, 1875b: 58 A: AP SCH YUN ORR
asper Sharp, 1875b: 59
foveolatus Harold, 1877a: 68

shibatai Nakane, 1960: 2 A: JA (Amami-Ôshima)
tricornis Wiedemann, 1823: 10 (Copris) A: FUJ HUB JA SHG TAI YUN ORR
crassus Sharp, 1875b: 51
rectinicornis Fairmaire, 1878a: 96

subgenus Parentius Zunino, 1979: 5 type species Copris punctatus Illiger, 1803
Relictonthophagus Kabakov, 1979: 74 type species Copris punctatus Illiger, 1803

atricapillus d’Orbigny, 1908: 74 N: MO
emarginatus Mulsant, 1942: 154 E: FR IT SP
cupreolus Reitter, 1894g: 301

mauritii Boucomont, 1920b: 311 A: AP ORR
nigellus Illiger, 1803: 207 (Copris) E: PT SP N: AG MO TU
punctator Reitter, 1892a: 179 [=1893a: 58] A: BEI ES FE GAN HEB HUB LIA MG NC SC SCH SHX
miyabei Matsumura, 1937c: 4
sunanicus Stebnicka, 1973: 118

SCARABAEIDAE
punctatus Illiger, 1803: 208 (Copris) E: PT SP N: MO
hispanicus Baraud, 1963: 262

zagrosicus Kabakov, 2006a: 160 A: IN
subgenus Phanaeomorphus Balthasar, 1963a: 161 type species Onthophagus sycophanta Fairmaire, 1887
ater C.O. Waterhouse, 1875a: 76 A: JA
chuzenjianus Matsumura, 1937b: 154
granulipennis J. Frivaldszky, 1892: 123
kawarinus Matsumura, 1937b: 157

cernyi Balthasar, 1935f: 312 A: FUJ GUI HEN NC SCH TAI YUN ZHE
dorsofasciatus Fairmaire, 1893e: 304 A: GUI SCH SHA ORR
fodiens C.O. Waterhouse, 1875a: 75 A: FE FUJ HEB HEI JA JIX NC NO QIN SC SCH SHG SHX
hikosanus Matsumura, 1937b: 154
sobosanus Matsumura, 1937b: 164
ushiodai Matsumura, 1937b: 166
yugianus Matsumura, 1937b: 168

gagates Hope, 1831: 22 A: AP HP KA FUJ NP SCH TAI UP XIZ YUN ORR
angulatus Kollar & L. Redtenbacher, 1844: 522

gagatoides Kabakov, 2006b: 124 A: HP UP
ginyunensis Všetečka, 1942: 255 A: ZHE
hingstoni Arrow, 1931: 278 A: SD
obliquus A.G. Olivier, 1789: no 3: 165 (Scarabaeus) N: EG AFR
potanini hiurai Ochi, 1984: 63 A: TAI
potanini potanini Kabakov, 1979: 80 A: LIA NE SCH SHA
procurvus Balthasar, 1935f: 309 A: SCH YUN
pseudojaponicus fukiensis Balthasar, 1942a: 120 A: FUJ
pseudojaponicus pseudojaponicus Balthasar, 1941a: 90 A: SCH
schaefernai Balthasar, 1935f: 312 A: GUX SCH YUN
strandi Balthasar, 1935f: 311 A: FUJ HUN SCH YUN
sycophanta Fairmaire, 1887b: 100 A: BEI FUJ GAN GUI HUN SCH SHA SHX YUN ORR
taiwanus Nomura, 1973b: 41 A: TAI
zavreli Balthasar, 1935f: 310 A: CHQ HUB SCH YUN
subgenus Proagoderus van Lansberge, 1883a: 14 type species Onthophagus ritsemae van Lansberge, 1883
Onthotrogus Motschulsky, 1860f: 152 type species Copris auratus Fabricius, 1801

amplexus Sharp, 1875b: 53 A: CH HP PA UP ORR
smeei Harold, 1875b: 137

pactolus Fabricius, 1787a: 12 (Scarabaeus) A: HP NP UP ORR
quadriarmatus Fairmaire, 1892a: 91 A: SA YE AFR
tirapensis Biswas & Chatterjee, 1985: 160 A: AP
yunnanus Boucomont, 1912b: 278 A: FUJ GUI HUB SCH YUN
subgenus Serrophorus Balthasar, 1963a: 160 type species Scarabaeus seniculus Fabricius, 1781
atropolitus d’Orbigny, 1902: 148 A: HP SD UP ORR
impuncticollis Frey, 1950: 301

199

200

SCARABAEIDAE

laevis asiaticus Boucomont, 1919b: 604 A: AP SD UP YUN ORR
laevis laevis Harold, 1880b: 194 A: CH ORR
laevis lampromelas Fairmaire, 1891f: cxciii A: HUB SCH YUN ORR
laevis stevensi Arrow, 1931: 172 A: SD
oblongus Y.-W. Zhang, 1997: 762 A: HUB
sagittarius Fabricius, 1775: 24 (Scarabaeus) A: FUJ NP TAI UP ORR
erectus Wiedemann, 1819: 157 (Copris)
javanus Fabricius, 1801a: 33 (Copris)
obtusus Wiedemann, 1819: 157 (Copris)
oryx Fabricius, 1792a: 56 (Scarabaeus)

senex Boucomont, 1914a: 212 A: GUA HKG ORR
seniculus Fabricius, 1781: 23 (Scarabaeus) A: GUA HKG TAI YUN ORR
brevipes Herbst, 1783: 10 (Scarabaeus)
cornicollis d’Orbigny, 1902: 146
novemmaculatus Gmelin, 1790: 1544 (Scarabaeus)

subgenus Sinonthophagus Kabakov, 2006a: 307 type species Onthophagus rugulosus Harold, 1886
poeophagus Kabakov, 1979: 82 A: NE
productus Arrow, 1907b: 423 A: CHQ HP SCH SD UP YUN ORR
rugulosus Harold, 1886b: 289 A: AP BEI FE FUJ HUB LIA NC NE NO SC SCH TAI YUN ORR
expansicollis Fairmaire, 1891f: cxciii
sonani Miwa, 1930: 171

subgenus Strandius Balthasar, 1963a: 160 type species Onthophagus lenzii Harold, 1874
changshouensis Y.-W. Zhang, 1997: 763 A: CHQ SCH
japonicus Harold, 1874b: 290 A: FE GUI JA NC SC SHX
kuraruanus Matsumura, 1938: 58 A: TAI (Lanyu)
lenzii lenzii Harold, 1874b: 290 A: FUJ HEI JA NC NE NO SC SCH
basicruentatus Goidanich, 1926f: 136
marginithorax Všetečka, 1939b: 44

oshimanus Nakane, 1960: 2 A: JA (Amami-Ôshima)
subansiriensis Biswas, 1979a: 912 A: AP
yakuinsulanus Nakane, 1984: 596 [RN] A: JA (Yaku-shima)
yakushimanus Nakane, 1983b: 139 [HN]

subgenus Sunenaga Ochi, 2003a: 264 type species Onthophagus anguliceps Boucomont, 1914
anguliceps Boucomont, 1914a: 210 A: BT SD TAI XIZ YUN ORR
actaeon Balthasar, 1946a: 2
aviculus Arrow, 1931: 179

avocetta Arrow, 1933: 421 [RN] A: SD ORR
digitatus Arrow, 1931: 177 A: SD
subgenus Trichonthophagus Zunino, 1979: 6 type species Copris hirtus Illiger, 1803
hirtus Illiger, 1803: 203 (Copris) E: PT SP N: AG MO
conjugata Seabra, 1907: 140

SCARABAEIDAE
hirtulus Reitter, 1892a: 197 [=1893a: 76]
infuscata Seabra, 1907: 140

maki Illiger, 1803: 204 (Copris) E: FR IT PT SP N: AG LB MO TU
glabra Seabra, 1907: suppl. 1
intercepta Seabra, 1907: 141
lineatus Mulsant, 1842: 112
obscuratus Heyden, 1890: 67
strigatus Mulsant, 1842: 112
variabilis Mulsant, 1842: 112

genus Onthophagus, species incertae sedis
abacus Boucomont, 1921b: 44 A: NP UP ORR
abreui Arrow, 1931: 239 A: UP ORR
acrisius Balthasar, 1946a: 1 A: FUJ ORR
agaricophilus Arrow, 1931: 333 A: UP
amycoides Kabakov, 2014: 55 A: SCH
arcifer d’Orbigny, 1904: 206 N: EG AFR
arboreus Arrow, 1931: 225 A: UP ORR
arunachalensis Biswas & Chatterjee, 1985: 162 A: AP
balawaicus Scheuern, 1995: 445 A: PN SD
beesoni Arrow, 1931: 319 A: UP
bengali Gordon & Oppenheimer, 1975: 497 A: AP ORR
bicolor Raffray, 1877: 324 A: PA SA YE ORR
bicristatus d’Orbigny, 1905: 462 N: EG
bituberculatus A.G. Olivier, 1789: no 3: 131 (Scarabaeus) N: EG A: SA AFR NARi
discoideus A.G. Olivier, 1789: no 3: 171 (Scarabaeus)

blanchardi Harold, 1869a: 67 N: EG AFR
boucomontianus Balthasar, 1935f: 318 A: FUJ SCH YUN ORR
brittoni R. Paulian, 1948: 149 A: YE
brutus Arrow, 1931: 215 A: SD YUN ORR
cervus Fabricius, 1798: 31 (Copris) A: HN NP UP ORR
corvus Fabricius, 1798: 31 (Copris) [incorrect original spelling]

chrysurus Arrow, 1931: 211 A: SD ORR
circulifer Arrow, 1931: 274 A: KA SD UP ORR
compactus Arrow, 1933: 421 [RN] A: UP
solidus Arrow, 1931: 265 [HN]

convexicollis Boheman, 1858: 46 A: HKG
coracinoides Kabakov, 1994b: 313 A: YUN ORR
coracinus Boucomont, 1914a: 214 A: FUJ HKG YUN ORR
curvispina Reitter, 1892a: 173 [=1893a: 52] A: BEI
deflexicollis van Lansberge, 1883b: 72 A: AP GUI SCH SD UP YUN ORR
mutabilis van Lansberge, 1883c: 148

201

202
dynastoides Arrow, 1931: 182 A: “North India”
exquisitus Arrow, 1931: 225 A: SD UP
falcifer Harold, 1880d: 351 A: SD
cultrifer Harold, 1886a: 145

falsus Gillet, 1925b: 236 A: AF HP PA UP ORR
fasciatus Boucomont, 1914a: 231 A: HP UP ORR
furcicollis Arrow, 1931: 276 A: AP BT HP NP SD UP ORR
furcillifer Bates, 1891a: 11 A: HP KA NP UP ORR
germanus Gillet, 1927: 254 A: SD UP ORR
gratus Arrow, 1931: 212 A: HP UP ORR
griseosetosus Arrow, 1931: 192 A: HP UP ORR
hamaticeps Arrow, 1931: 308 A: UP
heyrovskyi Všetečka, 1944: 176 A: SCH
jucundus Arrow, 1931: 188 A: AP ORR
jugicola d’Orbigny, 1902: 82 A: YE AFR
kashmirensis Balthasar, 1966a: 47 A: KA
koma Matsumura, 1937c: 3 A: SC
komareki Balthasar, 1935f: 316 A: SCH
kulti Balthasar, 1953b: 227 A: FUJ ORR
kumaonensis Arrow, 1931: 272 A: HP KA NP UP ORR
lapillus Arrow, 1931: 267 A: HP SD UP ORR
latipennis d’Orbigny, 1898e: 238 A: SA AFR
lilliputanus van Lansberge, 1883b: 69 A: HP KA UP ORR
carbunculus Boucomont, 1921b: 46

margaritifer d’Orbigny, 1898a: 162 N: EG A: SA YE AFR
inermifrons d’Orbigny, 1902: 219

millingeni d’Orbigny, 1898a: 178: A: IQ SA YE
mirandus Arrow, 1931: 290 A: HP UP ORR
delkeskampi Frey, 1954: 742

moreleti Baraud, 1980c: 140 A: IQ
nasalis Arrow, 1931: 288 A: HP NP UP ORR
necrophagus Arrow, 1931: 196 A: UP
noctivagus Harold, 1877b: 99 N: EG
obliviosus Balthasar, 1935f: 317 A: SHN
orientalis Harold, 1868b: 83 A: AF AP FUJ GUA GUI HKG NP SCH SD UP TAI YUN ORR
perroti Paulian, 1978: 40

pacificus van Lansberge, 1885a: 17 A: AP UP YUN ORR
peguanus Boucomont, 1914: 215

porcus Arrow, 1931: 325 A: AP ORR
pseudobrutus Kabakov, 1994b: 312 A: YUN ORR

SCARABAEIDAE

SCARABAEIDAE
pseudoconvexicollis Bai & X.-K. Yang, nom. nov. [see new Acts] A: BEI SHX
sinicus Y.-W. Zhang & Wang, 1997a: 209 [HN]

pupillatus H.J. Kolbe, 1886: 187 A: SC
purpurascens Boucomont, 1914: 215 A: YUN ORR
quadridentatus Fabricius, 1798: 34 (Copris) A: HP PA UP ORR
quadricornis Fabricius, 1801a: 54 (Copris)
moerens Walker, 1858: 209

ramosellus Bates, 1891a: 11 A: AF AP HP NP PA SD UP ORR
capella Hope, 1831: 22 [HN]
hrubanti Balthasar, 1956b: 414

remotus Harold, 1879a: 226 A: AP ORR
royi Biswas & Chatterjee, 1985: 172 A: AP ORR
rubripennis Arrow, 1907b: 419 A: AP BT SD ORR
subcribratus Arrow, 1907b: 420
vespillo Boucomont, 1920b: 314

rugulipennis Fairmaire, 1887a: 114 A: YE AFR
saudiensis Frey, 1962d: 618 A: SA
scaber Roth, 1851: 125 N: EG AFR
schawalleri Scheuern, 1996: 3 A: NP
sikkimensis Gillet, 1925b: 234 A: SD ORR
simplicifrons Reitter, 1892a: 169 [=1893a: 48] A: FE NE
sinicus Hope, 1842a: 60 [=1843a: 62, 1845: 6] A: ZHE
smetanai Balthasar, 1953b: 226 A: FUJ
songsokensis Biswas & Chatterjee, 1985: 166 A: AP
sparsulus Reitter, 1892a: 177 [=1893a: 56] N: EG
spinifex Fabricius, 1781: 29 (Scarabaeus) A: HP NP PA UP ORR
aeneus Fabricius, 1781: 34 (Scarabaeus)
bifossus d’Orbigny, 1902: 145
reflexicornis L. Redtenbacher, 1868: 57
spinifer A.G. Olivier, 1789: no 3: 148 (Scarabaeus) [HN]
truncaticornis Herbst, 1789: 209 (Scarabaeus)

sternalis Arrow, 1931: 222 A: UP ORR
sternax Balthasar, 1960b: 189 A: CH
sulci Balthasar, 1935f: 315 A: FUJ SCH
tabidus Balthasar, 1935f: 313 A: SCH
tarandus Fabricius, 1792a: 48 (Scarabaeus) A: HP PA UP ORR
biexcavatus d’Orbigny, 1898e: 239
suturatus Germar, 1824: 105

tessulatus Harold, 1871a: 13 A: IQ SA YE AFR
triceratops Arrow, 1913b: 195 A: AP ORR
tricolor Boucomont, 1914b: 280 A: FUJ GUA HKG ORR

203

204

SCARABAEIDAE

tripolitanus Heyden, 1890: 72 N: LB TU
tritinctus Boucomont, 1914a: 217 A: HKG ORR
turfanicus Balthasar, 1971b: 55 A: XIN
umenoi Matsumura, 1938: 61 A: TAI
unifasciatus Schaller, 1783: 240 (Scarabaeus) A: UP ORR
prolixus Walker, 1858: 208

vinctus Erichson, 1843: 234 N: EG AFR
obesus Fåhraeus, 1857: 298

zimmermanni Balthasar, 1960b: 192 A: SCH YUN ORR
zolotarevi Olsoufieff, 1907b: 193 A: FE
genus Onthophagus, nomina dubia
circulator Reitter, 1891f: 245 A: IS LE SY
excisus Reiche & Saulcy, 1856: 388 E: GR (Peloponnísos)
ibex Fabricius, 1792a: 55 (Scarabaeus) A: “India orientalis”
necessarius Reitter, 1892a: 179 [=1893a: 58] A: SC
pictus Reitter, 1892a: 196 [=1893a: 75] N: EG A: SY AFR
simius Reitter, 1892a: 176 [=1893a: 55] A: “Syria”
zinovskyi Qarjagdy, 1939: 86 E: AB
genus Phalops Erichson, 1848a: 764 type species Copris ciconia Fabricius, 1801 (=Scarabaeus iphis A.G. Olivier,
1789)
Copriodes Gistel, 1857b: 603 type species Onthophagus fimbriatus Klug, 1835
Ephillopus Reitter, 1892a: 168 [=1893a: 47] type species Scarabaeus iphis A.G. Olivier, 1789

candezei van Lansberge, 1883d: 165 A: NP ORR
chalybeus Klug, 1845: [3] pl. xli (Onthophagus) A: “Arabia deserta”
cyanescens d’Orbigny, 1898e: 243 (Onthophagus) A: PA ORR
divisus Wiedemann, 1823: 12 (Copris) A: HP NP PA SD UP ORR
olivaceus van Lansberge, 1883d: 164 A: AF PA YE ORR

tribe Scarabaeini Latreille, 1802

genus Ateuchetus Bedel, 1892: 282 type species Scarabaeus laticollis Linnaeus, 1767
armeniacus Ménétriés, 1832: 173 (Ateuchus) E: AB AR BU GG GR MC ST A: CY IN IQ SI SY TR
nigrociliatus Motschulsky, 1849: 107 (Scarabaeus)

cicatricosus P.H. Lucas, 1846c: pl. 23 (Ateuchus) E: PT SP N: AG MO
indistinctus Seabra, 1907: 59 (Scarabaeus)
minor Seabra, 1907: 60 (Scarabaeus)
sanguinolentus Seabra, 1907: 59 (Scarabaeus)
semilunatoides Báguena Corella, 1955: 275 (Scarabaeus)

laticollis Linnaeus, 1767: 549 (Scarabaeus) E: FR IT PT SP SZ N: AG MO

SCARABAEIDAE

205

alluaudi Théry, 1932c: 143 (Scarabaeus)
laevicollis Mulsant, 1842: 52 (Scarabaeus)
minutus Seabra, 1907: 63 (Scarabaeus)
semilunatus Xambeu, 1893: 227 (Ateuchus)
serratus Geoffroy, 1785: 13 (Copris)
striatopunctatus Leoni, 1911a: 242 (Scarabaeus)
striolatus Reitter, 1893a: 42 (Scarabaeus)

puncticollis Latreille, 1819: 255 (Ateuchus) E: PT SP N: AG EG LB MO TU A: IQ IS JO LE TR
hypocrita Laporte, 1840: 64 (Ateuchus)
nudifrons Fischer von Waldheim, 1824: 211 (Ateuchus)
parumpunctatus Klug, 1845: [1], pl. xli (Ateuchus)
sericeus Motschulsky, 1849: 107(Scarabaeus)

semipunctatus Fabricius, 1792a: 63 (Scarabaeus) E: AL CR FR HU IT MA ME SP N: AG EG LB MO TU A:
“Palestina”
castaneipennis A. Costa, 1853b: 6 (Scarabaeus)
subinermis Mulsant, 1842: 50 (Scarabaeus)
substriatus Mulsant, 1842: 50 (Scarabaeus)

variolosus Fabricius, 1787a: 16 (Scarabaeus) E: AL BH BU CR GR IT MA ME N: AG MO TU A: TR
genus Escarabaeus Zídek & Pokorný, 2011: 89 [RN] type species Scarabaeus cristatus Fabricius, 1775
Mesoscarabaeus Zídek & Pokorný, 2008: 5 [HN] type species Scarabaeus cristatus Fabricius, 1775

andrewesi Felsche, 1907: 275 (Sebasteos) A: AF IN PA ORR
bannuensis A. Janssens, 1940a: 31 (Scarabaeus) N: AG LB MO TU A: AE IN IQ JO KU OM PA QA SA AFR
janssensianus Auber, 1952: 127 (Scarabaeus)

brahminus Laporte, 1840: 64 (Ateuchus) A: BT KA IN PA ORR
cristatus cristatus Fabricius, 1775: 27 (Scarabaeus) N: EG LB A: AE AF IN IS PA SA “Palestina” AFR
compressicornis Klug, 1845: [1], pl. xli (Ateuchus)
cornifrons Laporte, 1840: 64 (Ateuchus)

cristatus villiersi Baraud, 1973a: 11 (Scarabaeus) A: IN
senaariensis Laporte, 1840: 66 (Ateuchus) N: EG AFR
genus Kheper A. Janssens, 1940a: 59 type species Ateuchus aegyptiorum Latreille, 1827
aegyptiorum Latreille, 1827: 279 (Ateuchus) N: EG AFR
corinthius Fairmaire, 1887a: 102 (Scarabaeus)
cuprescens Gillet, 1907: 592 (Scarabaeus)
fulgidus Gillet, 1907: 592 (Scarabaeus)
nigricans Gillet, 1907: 592 (Scarabaeus)
stigmaticus Fairmaire, 1887a: 102 (Scarabaeus)

devotus Kollar & L. Redtenbacher, 1844: 515 (Ateuchus) A: AF NP “Punjab” ORR
erichsoni Harold, 1867d: 94 (Ateuchus) A: “India” ORR

206

SCARABAEIDAE

sanctus Fabricius, 1798: 34 (Copris) A: PA ORR
genus Mnematidium Ritsema, 1889: 207 type species Ateuchus multidentatus Klug, 1845
Neoctodon Bedel, 1892: 283 [RN] type species Ateuchus multidentatus Klug, 1845
Octodon van Lansberge, 1875a: 183 [HN] type species Ateuchus multidentatus Klug, 1845

multidentatum Klug, 1845: [1], pl. xli (Ateuchus) N: EG LB A: IS JO SI SY
genus Mnematium W.S. Macleay, 1821: 504 type species Mnematium ritchiei W.S. Macleay, 1821
ritchiei W.S. Macleay, 1821: 506 N: AG LB TU
bottoi Gridelli, 1930: 324 (Scarabaeus)

silenus G. R. Gray, 1832a: pl. 40: 2 A: IQ SA SI SY
rotundipenne Holdhaus, 1920: 54 (Scarabaeus)

genus Scarabaeus Linnaeus, 1758: 345 type species Scarabaeus sacer Linnaeus, 1758
Actinophorus Creutzer, 1799b: 79 type species Scarabaeus sacer Linnaeus, 1758
Ateuchus Fabricius, 1801a: 54 [HN] type species Scarabaeus sacer Linnaeus, 1758
Heliocantharus W.S. Macleay, 1821: 497 type species Scarabaeus sacer Linnaeus, 1758
Drepanopodus A. Janssens, 1940a: 73 type species Ateuchus costatus Wiedemann, 1823
Madateuchus R. Paulian, 1953: 24 type species Madateuchus viettei R. Paulian, 1953
Neateuchus Gillet, 1911b: 309 type species Ateuchus proboscideus Guérin-Méneville, 1844
Neomnematium A. Janssens, 1938b: 71 type species Scarabaeus sevoistra Alluaud, 1902
Parateuchus Shipp, 1895b: 221 type species Scarabaeus morbillosus Fabricius, 1792
Sebasteos Westwood, 1847a: 225 type species Sebasteos galenus Westwood, 1847

acuticollis Motschulsky, 1849: 104 (Ateuchus) E: AB A: AF IN IQ KI KU KZ SY TD TM UZ
aegyptiacus Stolfa, 1938: 143 N: AG EG LB MO TU A: SA
babori Balthasar, 1934c: 147 A: AF IN JO KI KZ PA TD TR TM UZ
carinatus Gebler, 1841a: 371 (Ateuchus) A: AF IN KI KZ TD TM UZ
dioscoridis Petrovitz, 1962b: 107 A: YE (Suqutra)
gangeticus Laporte, 1840: 64 (Ateuchus) N: EG A: AF PA ORR
indicus Motschulsky, 1864: 457 (Ateuchus)

irakensis Stolfa, 1938: 143 A: IN IQ IS JO QA SA SY TR
isidis Laporte, 1840: 64 (Ateuchus) A: OM SA YE AFR
krutai Malý & Montreuil, 2011: 5 A: SA
pius Illiger, 1803: 202 (Ateuchus) E: AB AL AR BH BU CR FR GG GR HU IT KZ MC ME RO SB SP ST A: AF IN IS
JO KZ LE SY TD TM TR UZ
digitatus Motschulsky, 1849: 105 (Ateuchus)
infirmus Fischer von Waldheim, 1823: pl. XXVII [=1824: 211] (Ateuchus)
monachus Faldermann, 1835a: 237 (Ateuchus)
punctulatus Mulsant, 1842: 46
subsulcatus Mulsant, 1842: 46

207

SCARABAEIDAE

sacer Linnaeus, 1758: 347 E: AB AL AR BU FR GG GR IT ME PT RO SB SP ST “Caucasus” UK N: AG EG LB MO
TU A: CY IQ IS JO SA SI SY TR
confluidens Fleischer, 1925: 20
crenatus De Geer, 1778: 638
degeeri W.S. Macleay, 1821: 502
dufresnii W.S. Macleay, 1821: 502
edentulus Mulsant, 1842: 46
europaeus Motschulsky, 1849: 103 (Ateuchus)
inermis Mulsant, 1842: 46
platychilus Fischer von Waldheim, 1824: 211 (Ateuchus)
retusus Brullé, 1832: 166 (Ateuchus)
rufipes Seabra, 1907: 54

transcaspicus Stolfa, 1938: 142 A: AF IN KZ TD TM UZ
typhon Fischer von Waldheim, 1823: pl. XXVII [=1824: 210] (Ateuchus) E: AB AL AR AU BH BU CR FR GG GR
HU IT KO KZ MC MD ME PT RO SB SK SP ST TR UK A: AF CY IN IQ IS JO KZ LE MG NC NMO SC SY TM
TR UZ XIN XIZ
affinis Brullé, 1832: 165 (Ateuchus)
peregrinus H.J. Kolbe, 1886: 189 (Ateuchus)
spencii W.S. Macleay, 1821: 502

wilsoni C.O. Waterhouse, 1890b: 366 A: IN “Syria”
frankenbergeri Balthasar, 1934c: 146
ganglbaueri Reitter, 1892a: 162 [=1893a: 41]

winkleri Stolfa, 1938: 144 A: AF IN KZ PA TM UZ XIN
zagrosensis Montreuil, 2006a: 446 A: IN IQ SY
tribe Scarabaeini, nomina dubia,
Ateuchus clypeatus Motschulsky, 1849: 106 E: ST
Scarabaeus impius Herbst, 1789: 302 [distribution unknown]

tribe Sisyphini Mulsant, 1842

genus Sisyphus Latreille, 1806: 79 type species Scarabaeus schaefferi Linnaeus, 1758
subgenus Neosisyphus J. Müller, 1942: 86 type species Sisyphus atratus Klug, 1855
bowringi White, 1844: 423 A: HKG SHG
subgenus Sisyphus Latreille, 1806: 79 type species Scarabaeus schaefferi Linnaeus, 1758
crispatus hirtus Wiedemann, 1823: 24 A: PA ORR
prominens Walker, 1859b: 219
setosulus Walker, 1858: 208
subsidens Walker, 1858: 208

denticrus Fairmaire, 1886: 320 A: SD SCH SW YUN ORR
indicus Hope, 1831: 22 A: AP GUA HP KA NP SCH SD XIZ ORR
caschmirensis Kollar & L. Redtenbacher, 1844: 516

208

SCARABAEIDAE

longipes A.G. Olivier, 1789: no 3: 164 (Scarabaeus) A: PA ORR
minutus Fabricius, 1792a: 70 (Scarabaeus)
helwigi Fabricius, 1798: 35 (Scarabaeus)

moltaensis Vazirani, 1967: 90 A: UP
neglectus Gory, 1833a: 14 A: NP PA SCH SD UP YUN ORR
popovi Kryzhanovskij & S.I. Medvedev, 1966: 389 A: YUN
schaefferi morio Arrow, 1909a: 93 A: BEI FE GAN HEB HEI NC NMO SC
schaefferi schaefferi Linnaeus, 1758: 349 (Scarabaeus) E: AB AL AR AU BE BH BU CR CZ FR GE GG GR
HU IT KO KZ LU MC MD ME NL PL PT RO SB SK SL SP ST SZ TR UK N: AG MO TU A: CY IN IS JO KZ SY
TM TR
arachnoides Geoffroy, 1785: 15 (Copris)
boschnakii Fischer von Waldheim, 1824: 210
capensis Gory, 1833a: 12
longipes Scopoli, 1763: 11 (Scarabaeus)
minutus Seabra, 1907: 66
subemarginatus Mulsant, 1842: 62
subinermis Mulsant, 1842: 62
tauscheri Fischer von Waldheim, 1823: pl. XXVII [=1824: 209]

subfamily Dynamopodinae Arrow, 1911
tribe Dynamopodini Arrow, 1911

genus Orubesa Reitter, 1895d: 150 type species Orubesa perforata Reitter, 1895
Dynamopus Semenov, 1896d: 336 type species Dynamopus athleta Semenov, 1896
Xanthelaeus Fairmaire, 1897c: 109 type species Xanthelaeus plicifrons Fairmaire, 1897

ata Semenov & S.I. Medvedev, 1929d: 173 (Dynamopus) A: IN
athleta Semenov, 1896d: 340 (Dynamopus) E: ST A: TD TM UZ
gladiator Balthasar, 1968: 147 (Dynamopus) A: AF
luctator Semenov & S.I. Medvedev, 1929d: 176 (Dynamopus) A: IN IQ PA ORR
perforata Reitter, 1895d: 150 A: AF IN
plicifrons Fairmaire, 1897b: 109 (Xanthelaeus) N: MO AFR
semenowi Arrow, 1911c: 612 (Dynamopus) N: LB A: AE SA AFR
sudanica Balthasar, 1971d: 1 (Dynamopus) A: AE OM AFR

tribe Thinorycterini Semenov & Reichardt, 1925

genus Thinorycter Semenov & Reichardt, 1925: 83 type species Thinorycter chlamydatus Semenov &
Reichardt, 1925
balthasari S.I. Medvedev, 1969: 1486 A: KZ
chlamydatus Semenov & Reichardt, 1925: 85 A: TM
diamesus Semenov & S.I. Medvedev, 1928b: 107

chorasmius Semenov & S.I. Medvedev, 1929c: 169 A: TM
mamajevi S.I. Medvedev, 1969: 1488 A: KZ TM UZ

209

SCARABAEIDAE
medvedevi Nikolajev, 1975b: 280

redikortzevi Kiseritzky & Reichardt, 1927: 255 A: KZ TM

subfamily Orphninae Erichson, 1847
tribe Orphnini Erichson, 1847

genus Chaetonyx Schaum, 1862b: 265 type species Chaetonyx robustus Schaum, 1862
binaghii Mariani, 1946: 81 E: GR MC
robustus italicus Mariani, 1946: 79 E: IT
robustus liguricus Mariani, 1946: 78 E: IT (Liguria)
robustus robustus Schaum, 1862b: 266 E: AL BU GR RO SB TR
schatzmayri Mariani, 1946: 80 E: BU MC
genus Hybalus Dejean, 1833: 149 type species Geobius cornifrons Brullé, 1832
Geobius Brullé, 1832: 172 [HN] type species Geobius cornifrons Brullé, 1832

algiricus Petrovitz, 1968d: 183 N: AG
alonsoi López-Colón & de la Puebla, 2014: 193 E: SP (Cádiz)
ameliae López-Colón, 1986b: 209 E: SP (Andalucía)
angustatus P.H. Lucas, 1855b: 558 N: AG
arenicola Baraud, 1991a: 12 N: AG
atlanticus López-Colón, 1992d: 361 N: MO (West)
baguenae López-Colón, 1986b: 210 E: SP (Andalucía)
barbarus Laporte, 1840: 108 (Geobius) N: AG TU
baudoni Petrovitz, 1964b: 51 N: MO (West)
benoitii Tournier, 1864: 266 E: IT (Sicilia, Calabria)
bigibber Reitter, 1892e: 260 E: IT (Sicilia) N: AG TU
bletoni Baraud, 1991a: 14 N: MO (Moyen Atlas)
constantini Baraud, 1979d: 227 N: AG
cornifrons Brullé, 1832: 173 (Geobius) E: GR
digitatus Petrovitz, 1963b: 119 N: AG
dorcas Fabricius, 1798: 31 (Copris) N: MO (North-West)
quedenfeldti Petrovitz, 1964b: 56
reclinans Fairmaire, 1879c: 172
tingitanus Fairmaire, 1852: 84 (Geobius)

doursii P.H. Lucas, 1853: xxii N: AG
gazella Raffray, 1873: 370 [HN]
raffrayi Arrow, 1911b: 394 [RN]

glabratus Fabricius, 1792a: 68 (Scarabaeus) N: AG MO
capreolus Mulsant, 1842: 340 (Geobius)
gazella Mulsant, 1842: 340 (Geobius)

granicornis Fairmaire, 1877a: 141 N: AG

210

SCARABAEIDAE

biretusus Marseul, 1878: 59

kocheri Petrovitz, 1964b: 52 N: MO (West)
maroccanus Petrovitz, 1964b: 51 N: MO (Moyen Atlas)
normandi Baraud, 1980a: 60 N: TU
numidicus Petrovitz, 1964b: 55 N: AG
parvicornis P.H. Lucas, 1855b: 556 N: AG TU
petrovitzi Baraud, 1991a: 21 N: MO
punicus Baraud, 1991a: 22 N: TU
pygmaeus Quensel, 1806: 28 (Scarabaeus) N: AG
ramicornis Reitter, 1892e: 261 N: AG
reflexus Petrovitz, 1964b: 54 N: MO (West)
rotroui peyerimhoffi López-Colón, 1992d: 365 N: MO (Moyen Atlas)
rotroui rotroui Petrovitz, 1964b: 54 N: MO (East Rif)
saezi López-Colón, 1992b: 185 E: SP (Andalucía)
servulus Normand, 1949: 86 N: TU
demoflysi Baraud, 1980a: 61

subcornutus Fairmaire, 1871a: 375 N: AG MO
sulcatus Baraud, 1991a: 24 N: AG
tricornis P.H. Lucas, 1846a: pl. 24 (Geobius) N: AG (North-East)
tuberculicornis Reitter, 1892e: 259 N: AG
varians Petrovitz, 1964b: 53 N: MO (North-West Rif)
genus Orphnus W.S. Macleay, 1819: 119 type species Scarabaeus bicolor Fabricius, 1801
subgenus Orphnus W.S. Macleay, 1819: 119 type species Scarabaeus bicolor Fabricius, 1801
Drepanognathus van Lansberge, 1886b: 92 [HN] type species Drepanognathus mandibularis van Lansberge, 1886
Sissantobius Ritsema, 1889: 217 [RN] type species Drepanognathus mandibularis van Lansberge, 1886

impressus Westwood, 1846: 177 A: UP ORR

subfamily Melolonthinae Leach, 1819
tribe Ablaberini C.É. Blanchard, 1850

genus Cyrtocamenta Brenske, 1897b: 104 type species Cyrtocamenta pygmaea Brenske, 1897
Microcamenta Brenske, 1898a: 334 type species Microcamenta transvaalensis Brenske, 1898

pygidialis Frey, 1968: 309 (Microcamenta) A: SA YE AFR
sundanica Frey, 1970a: 5 (Camenta)

pallida D. Ahrens, 2000e: 186 A: SA
genus Empecamenta Brenske, 1895: 218 type species Empecamenta pilifera Brenske, 1895
Isocamenta H.J. Kolbe, 1914: 348 type species Isocamenta montana H.J. Kolbe, 1914

arabica D. Ahrens, 2000e: 181 A: SA
buettikeri D. Ahrens, 2000e: 181 A: SA

211

SCARABAEIDAE
tribe Chasmatopterini Lacordaire, 1856

genus Chasmatopterus Dejean, 1821: 60 type species Melolontha hirtula Illiger, 1803
almeriensis Baraud, 1965c: 277 E: SP
annamariae Branco, 2004: 27 E: PT
canosus Báguena, 1955b: 282 E: SP
cobosi barrancoi Ruiz & Ávila, 1994: 100 E: SP
cobosi cobosi Baraud, 1965c: 276 E: SP
esuris Baraud, 1965c: 280 E: PT SP
hirtulus baraudi Pardo Alcaide, 1976: 88 N: MO
hirtulus hirtulus Illiger, 1803: 222 (Melolontha) E: PT SP
hispidulus Graëlls, 1846: 218

hirtus C.É. Blanchard, 1850: 57 E: PT SP
illigeri Perris, 1855: 280 E: PT SP
jaenensis Baraud, 1965c: 283
pardoi Baraud, 1965c: 282

pilosulus Illiger, 1803: 224 (Melolontha) E: PT SP
uxorum Baraud & Branco, 1985: 627 E: SP
villosulus Illiger, 1803: 223 (Melolontha) E: PT SP
ardoini Baraud, 1965c: 278
nigrocinctus Wollaston, 1854: 236

zonatus Escalera, 1925: 435 E: SP N: MO
genus Chasmatopterus, nomen dubium
parvulus Graëlls, 1851: 137 E: SP

tribe Diphycerini S.I. Medvedev, 1952

genus Dichelomorpha Burmeister, 1855: 480 type species Dichelomorpha ochracea Burmeister, 1855
Dejeania C.É. Blanchard, 1850: 96 [HN] type species Dejeania alsiosia C.É. Blanchard, 1850
Sinochelus Fairmaire, 1900b: 618 type species Sinochelus limbatus Fairmaire, 1900

delauneyi Fleutiaux, 1887: 62 (Dejeania) A: SE TAI ORR
limbata Fairmaire, 1900b: 618 (Sinochelus) A: FUJ
ochracea Burmeister, 1855: 480 A: HKG
chinensis Boheman, 1858: 52 (Dichelus)

genus Diphycerus Fairmaire, 1878a: 100 type species Diphycerus davidis Fairmaire, 1878
alcedo Arrow, 1920: 452 A: SD
cupripennis Arrow, 1925: 219 A: YUN
davidis Fairmaire, 1878a: 100 A: SCH GAN
jucundus Arrow, 1920: 452 A: SCH
reitteri Semenov, 1891: 328 A: GAN

212

SCARABAEIDAE

genus Metaceraspis Frey, 1962a: 63 type species Metaceraspis fukiensis Frey, 1962
fukiensis Frey, 1962a: 63 A: FUJ
genus Xenoceraspis Arrow, 1920: 453 type species Xenoceraspis dispar Arrow, 1920 (=Dichelomorpha
k­ urseongana Moser, 1917)
calcaratus Y.-W. Zhang, 1988: 246 A: XIZ
kurseonganus Moser, 1917: 95 (Dichelomorpha) A: SD
dispar Arrow, 1920: 454

longimacularius Y.-W. Zhang, 1988: 245 A: AP XIZ
tribe Diphycerini, genera incertae sedis
genus Hyperius Fairmaire, 1878a: 104 type species Hyperius davidis Fairmaire, 1878
castanescens Fairmaire, 1891f: cxcviii A: SCH
davidis Fairmaire, 1878a: 104 A: SCH
densiventris Fairmaire, 1878a: 105 A: CH (“Chine centrale”)
sparsutus Fairmaire, 1878a: 104 A: SHA
genus Photyna Brenske, 1897b: 97 type species Photyna rugicollis Brenske, 1897
rugicollis Brenske, 1897b: 97 A: CH (“Ho-chan”)
tomentosa Brenske, 1897b: 98 A: CH (“Ho-chan”)

tribe Diplotaxini Kirby, 1837

genus Apogonia Kirby, 1819a: 401 type species Apogonia gemellata Kirby, 1819 (=Melolontha rauca
Fabricius, 1781)
aenescens Hope, 1831: 23 A: AP NP UP ORR
aeneomicans Moser, 1913e: 424
brunnea Hope, 1831: 23

aerata Moser, 1916a: 179 A: BT ORR
convexicollis Frey, 1962c: 610

amida Lewis, 1896: 332 A: JA TAI
apicalis Moser, 1908c: 331 A: GUX YUN ORR
basalis Moser, 1915a: 124 A: FUJ
belutschistanica Petrovitz, 1958a: 4 A: IN PA
bicarinata Lewis, 1896: 333 A: JA TAI
okinawana Nomura, 1964b: 63
sauteri Moser, 1918: 232
yaeyamana Nomura, 1965: 3

cribricollis Burmeister, 1855: 256 A: CHQ FUJ GAN GUA GUI GUX HAI HKG NC NE SCH ORR
miurai Kobayashi, 2009a: 33
reticulata Frey, 1969: 184

SCARABAEIDAE

213

cupreoviridis H.J. Kolbe, 1886: 193 A: ANH BEI FE FUI GAN GUA GUX HEI HUB HUN JA (Miyako-jima) JIA JIX
LIA NC NE SCH SHA SHG SHN TIA YUN ZHE
fusana H.J. Kolbe, 1886: 193
miyakona Nomura, 1965: 3
nigroolivacea Heyden, 1886d: 292

diversicollis Moser, 1914b: 151 A: SD
formosana Moser, 1913e: 430 A: TAI
hongkongica Miyake, 1989: 38 A: HKG
ishiharai ishiharai Sawada, 1940: 272 A: JA
iriei Miyake, 1986a: 1
yakushimana Nomura, 1959b: 48

kamiyai Sawada, 1940: 276 A: JA TAI
kusuii Kobayashi & Bezděk, 2011: 55 A: YUN ORR
loi Kobayashi, 2007a: 8 A: TAI
major bicavata Arrow, 1913c: 399 A: JA (Ryukyus)
shibuyai Sawada, 1950b: 264

major major C.O. Waterhouse, 1875a: 103 A: JA
nigricans Hope, 1831: 23 A: HP NP
niponica Lewis, 1895c: 395 A: BEI GAN GUI HUB NC SC SCH SHA
chinensis Moser, 1918: 231

pilifera Moser, 1913e: 415 A: FUJ GUI HAI HUN JIX
pingtungensis Kobayashi, 2015b: 40 A: TAI
rufobrunnea Moser, 1915a: 121 A: FUJ TAI
rufofusca Moser, 1916a: 180 A: HKG
sequens Frey, 1975b: 233 A: UP
setosa Arrow, 1921a: 7 A: HP
splendida Boheman, 1858: 55 A: HKG
taiwana Kobayashi, 2007a: 7 A: TAI
takasagoensis Sawada, 1940: 270 A: TAI
tanigawaensis Sawada, 1940: 271 A: JA
ikedai Kobayashi, 1983b: 21

uniformis C.É. Blanchard, 1851: 229 A: SD ORR
ventralis Moser, 1913e: 422 A: GUA GUI GUX HKG YUN ORR
vicina Burmeister, 1855: 256 A: SD
wittmeri Frey, 1975b: 232 A: BT
yunnana Moser, 1915a: 122 A: YUN
genus Apogonia, nomina dubia
squamipennis Karsch, 1882: 123 A: “Persien”
obscura C.É. Blanchard, 1851: 229 A: “Indes septentr.”

214

SCARABAEIDAE

genus Liogenys Guérin-Méneville, 1831b: pl. 3 type species Melolontha palpalis Eschscholz, 1822
Peritryssus Reitter, 1918b: 77 type species Peritryssus excisus Reitter, 1918

excisa Reitter, 1918b: 78 (Peritryssus) Ei: IT (Sicilia)
tribe Diplotaxini, nomina dubia
genus Adoretops Kraatz, 1883c: 151 type species Melolontha pexa Zoubkov, 1833
pexus Zoubkov, 1833: 323 (Melolontha) A: KZ
genus Schismatocera Gautier des Cottes, 1872: 520 type species Adoretus nitidulus Gautier des Cottes, 1870
nitidula Gautier des Cottes, 1870: 104 (Adoretus) A: ES

tribe Euchirini Hope, 1840

genus Propomacrus Newman, 1837: 255 type species Propomacrus arbaces Newman, 1837 (=Scarabaeus
bimucronatus Pallas, 1781)
Macropropus Agassiz, 1846: 309 type species Propomacrus arbaces Newman, 1837 (=Scarabaeus bimucronatus
Pallas, 1781)
Porropus Laporte, 1840: 113 type species Scarabaeus bimucronatus Pallas, 1781

bimucronatus Pallas, 1781: 13 (Scarabaeus) E: BU GR MC TR A: IN IS LE SY TR
arbaces Newman, 1837: 256

cypriacus Alexis & Makris, 2002: 105 A: CY
davidi Deyrolle, 1874a: 110 [=1874b: 448] A: FUJ JIX ZHE
fujianensis Wu & Wu, 2008: 827

muramotoae Fujioka, 2007: 99 A: BT NP XIZ ORR
genus Cheirotonus Hope, 1840a: 78 [=1840b: 300, 1841: 594] type species Cheirotonus macleayii Hope, 1840
battareli Pouillaude, 1913f: 470 A: GUX YUN ORR
formosanus Ohaus, 1913: 142 A: TAI
fujiokai Muramoto, 1994: 8 A: GAN SHA
gestroi gestroi Pouillaude, 1913f: 471 A: YUN ORR
corompti Pouillaude, 1913f: 478
henrici Pouillaude, 1913f: 472

jambar Kurosawa, 1984: 74 A: JA (Okinawa-shotō)
jansoni Jordan, 1898: 419 (Propomacrus) A: CHQ FUJ GUA GUI GUX HAI HUB HUN JIA JIX SCH SHA YUN
ZHE ORR
nankinensis S.T. Yu, 1936: 3 (Propomacrus)
szetshuanus S.I. Medvedev, 1960: 14

macleayii Hope, 1840a: 78 [=1840b: 300, 1841c: 595] A: BT NP SD XIZ ORR
macleayanus Burmeister, 1842: 702 [RN]

parrii J.E. Gray, 1848: 59 A: HP SD UP ORR
chiangdaoensis Minet, 1987: 53

terunumai Muramoto, 2008: 85 A: XIZ YUN ORR

215

SCARABAEIDAE
tribe Heptophyllini S.I. Medvedev, 1951

genus Heptophylla Motschulsky, 1858a: 32 type species Heptophylla picea Motschulsky, 1858
Hypochrus Fairmaire, 1891f: cci type species Hypochrus brevicollis Fairmaire, 1891

brevicollis Fairmaire, 1891f: cci (Hypochrus) A: HUB SCH
calcarata Y.-W. Zhang, 1981: 349 A: XIZ
dimidiata Arrow, 1934: 9 A: JIA
laticollis Y.-W. Zhang, 1995: 243 A: ZHE
longilamella Y.-W. Zhang, 1981: 350 A: XIZ
picea iriei Kusui, 1971: 117 A: JA (Shimo-koshiki-shima)
picea maenamii Nomura, 1969: 73 A: JA (Izu--shotō)
picea picea Motschulsky, 1858a: 33 A: FE (Kuril’skie ostrova) JA SC
tosana Miyatake, 1963: 23
transversa Motschulsky, 1861a: 15 (Holotrichia)

genus Hexataenius Fairmaire, 1891d: ix type species Hexataenius protensus Fairmaire, 1891
ezakii Niijima & Kinoshita, 1923: 70 (Gymnogaster) A: TAI
protensus Fairmaire, 1891d: ix A: CHQ HUB JA SCH
genus Hilyotrogus Fairmaire, 1886: 325 type species Hilyotrogus unguicularis Fairmaire, 1886
Melichrus Brenske, 1892c: 156 type species Melichrus kolbei Brenske, 1892

bicoloreus Heyden, 1887a: 265 (Lachnosterna) A: CHQ FE HEB NC NE SCH
calcaratus Frey, 1971a: 132 (Melichrus) A: NP
changbaishanensis J. K. Li, 1992: 77 A: JIL
flavescens Y.-W. Zhang, 1997: 770 (Melichrus) A: CHQ
flavosericeus Brenske, 1897d: 348 (Holotrichia) A: SD
formosanus Niijima & Kinoshita, 1927: 28 (Heptophylla) A: TAI
holosericeus Kollar & L. Redtenbacher, 1844: 524 (Ancylonycha) A: HP NP
brevicollis C.É. Blanchard, 1851: 132 (Ancylonycha)
rufoflavus Moser, 1921: 65
stridulans Sharp, 1878b: 172 (Lachnosterna)

iridipennis Fairmaire, 1886: 326 A: YUN
kolbei Brenske, 1892c: 156 (Melichrus) A: BT SD
longiclavis Bates, 1891b: 74 A: GAN SCH XIN
luteosericea Brenske, 1897d: 347 (Holotrichia) A: SD
mangkamensis Y.-W. Zhang, 1981: 351 A: XIZ
nitens Frey, 1975b: 235 A: BT
piceosericeus Moser, 1913c: 281 A: YUN
pilicollis Moser, 1913c: 280 A: YUN
pilifer Moser, 1921: 66 A: SHN
sericeus Moser, 1915a: 144 A: SCH
setiger Moser, 1915a: 145 A: SHN

216

SCARABAEIDAE

setipennis Moser, 1913c: 281 A: CH
sikkimensis Moser, 1913c: 277 A: SD
stolidus Fairmaire, 1886: 326 A: YUN
unguicularis Fairmaire, 1886: 325 A: YUN
wittmeri Frey, 1975b: 233 A: BT
yasuii Nomura, 1970: 68 (Brahmina) A: JA (Iriomote-shima)
genus Taiwanotrichia Kobayashi, 1990d: 76 type species Taiwanotrichia longicornis Kobayashi, 1990
dorsopilosa C.-L. Li & P.-S. Yang, 1991: 320 A: TAI
hainanensis Keith & J. Li, 2009: 164 A: HAI
longicornis Kobayashi, 1990d: 76 A: TAI
similis C.-L. Li & P.-S. Yang, 1991: 323 A: TAI
sinocontinentalis fujianensis Keith, 2009b: 234 A: FUJ
sinocontinentalis sinocontinentalis Keith, 2009b: 233 A: GUA
genus Toxospathius Fairmaire, 1878a: 102 type species Toxospathius inconstans Fairmaire, 1878
auriventris Bates, 1891b: 73 A: SCH XIZ
brevicollis Arrow, 1934: 8 A: GAN SCH
inconstans Fairmaire, 1878a: 103 A: NO NW

tribe Hopliini Latreille, 1829

genus Ectinohoplia L. Redtenbacher, 1868: 63 type species Ectinohoplia sulphuriventris L. Redtenbacher, 1868
Spinohoplia Sabatinelli, 1997a: 67 type species Spinoplia ahrensis Sabatinelli, 1997

affinis Arrow, 1921b: 273 A: YUN ORR
ahrensis Sabatinelli, 1997a: 68 (Spinohoplia) A: BT NP SD
auriventris Moser, 1915a: 150 A: FUJ
balthasari Tesař, 1963: 94 A: SCH
davidis Fairmaire, 1889a: 15 A: HUB JIX SCH
hispidula Reitter, 1903a: 110

formosana arrowi Miwa, 1931a: 287 [RN] A: TAI
formosana Arrow, 1921b: 269 [HN]

formosana formosana Moser, 1919: 362 A: TAI
gracilipes Lewis, 1895c: 389 (Hoplia) A: JA
guttaticollis Fairmaire, 1900b: 620 A: SCH ORR
tonkinensis Moser, 1912a: 306

huettenbacheri Nonfried, 1892d: 258 A: “Himalaya borealis”
indica Moser, 1912a: 305 A: “Himalaya”
inscripta Arrow, 1921b: 269 A: FUJ
kuatunensis Tesař, 1963: 97 A: FUJ GAN
nantouensis Kobayashi, 1995c: 147 A: TAI
obducta Motschulsky, 1858a: 33 (Hoplia) A: JA

SCARABAEIDAE

217

caminaria Reitter, 1903a: 109
sabulicola Motschulsky, 1858a: 34 (Hoplia)
variolosa C.O. Waterhouse, 1875a: 99

paivai Wollaston, 1859: 430 (Hoplia) A: HUB SCH SHG
ichangensis Reitter, 1903a: 110
nigrotincta Fairmaire, 1897f: 243
variabilis Reitter, 1903a: 110
variegata Preudhomme de Borre, 1886b: 84

puella Endrődi, 1952b: 47 A: FUJ
quadrituberculata Preudhomme de Borre, 1886b: 86 A: SHG
rufipes Motschulsky, 1860e: 133 (Decamera) A: FE GAN HEI JA NC SC
simianshana Kobayashi & Fujioka, 2015: 80 A: CHQ
simillima Kobayashi & Fujioka, 2015: 74 A: CHQ GUX ORR
soror Arrow, 1921b: 270 A: FUJ GAN
sulphuriventris L. Redtenbacher, 1868: 63 A: FUJ HKG SHG
suturalis Preudhomme de Borre, 1886b: 85 A: CE ORR
chrysura Reitter, 1903a: 109
diabolica Reitter, 1903a: 109
luteostriata Brenske, 1894a: 34

trichota Jakobson, 1914: 1 A: BEI
triplagiata Fairmaire, 1897e: 211 A: SCH
viridilineata Kobayashi & Fujioka, 2015: 75 A: YUN ORR
yoi Sawada, 1939a: 42 A: TAI
yunnana Moser, 1919: 361 A: YUN
genus Himalhoplia Sabatinelli, 1983a: 205 type species Himalhoplia furcata Sabatinelli, 1983
furcata Sabatinelli, 1983a: 205 A: NP
genus Hoplia Illiger, 1803: 226 type species Scarabaeus farinosus Linnaeus, 1760 (=Scarabaeus argenteus Poda
von Neuhaus, 1761)
Diphydactylus J. Thomson, 1858: 58 type species Diphydactylus singularis J. Thomson, 1858

subgenus Decamera Mulsant, 1842: 503 type species Melolontha pulverulenta Fabricius, 1775 sensu Mulsant,
1842 (=Scarabaeus philanthus Fuessly, 1775)
angulata Reitter, 1902b: 138 A: KI KZ UZ
medvedevi Wainstein, 1956: 84

bezdeki Keith, 2002c: 145 A: SY
bucharica Reitter, 1898a: 10 A: TD UZ
cincticollis Faldermann, 1833: 49 (Anisoplia) A: ES FE HEB MG NW
corallipes Reitter, 1884a: 8 E: AB
corniculata Reitter, 1890r: 376 E: BU GR A: CY LE TR
semirufa Pic, 1908c: 65

218

SCARABAEIDAE

davidis Fairmaire, 1887d: 314 A: BEI GAN HEB QIN SCH
divina Endrődi, 1953a: 115 A: “Central China”
djukini Jakobson, 1914: 2 A: FE NC
endrodii Balthasar, 1955: 432 A: AF
euphratica Zaitzev, 1924: 103 A: TR
fiorii D’Amore Fracassi, 1906: 57 (Decamera) E: IT FR
meridionalis D’Amore Fracassi, 1906: 57 (Decamera)
micans D’Amore Fracassi, 1906: 57 (Decamera)

hauseri Reitter, 1904c: 157 A: KI KZ UZ
jakobsoni Reitter, 1903a: 113 A: IN TR
leonhardi Reitter, 1909a: 56

philanthus gagates Bedel, 1911a: 145 E: SP N: MO
philanthus philanthus Fuessly, 1775: 3 (Scarabaeus) E: AU BE BU BY CR CZ DE FR GB GE HU IT LS LU NL PL
SB SK SL SP SV SZ UK
fuesslini Moller, 1782: 376 (Scarabaeus)
misella Schauffus, 1882b: 553
pulverulenta Fabricius, 1775: 39
ramburi Heyden, 1870a: 115
squamosa Fabricius, 1792b: 173 (Melolontha)
tumulorum du Buysson, 1906: 18
varians Mulsant, 1842: 506

potanini Heyden, 1889c: 668 A: NMO
praticola Duftschmid, 1805: 180 E: AL AU BE BH BU CR CZ FR GE GR HU IT KO LA LS MD ME PL RO SB SK
SL SZ UK
bosnica Reitter, 1903a: 113
palustris Heer, 1842: 544
ripicola Mulsant, 1841: 509 (Decamera)
splendida Pic, 1907c: 169

pseudophilanthus Petrovitz, 1967b: 338 E: BU A: TR
semicastanea Fairmaire, 1887d: 315 A: BEI GAN NIN NMO TAI XIN ORR
subgenus Euchromoplia S.I. Medvedev, 1952: 235 type species Scarabaeus aureolus Pallas, 1781
Victoria Miyahara, 1941: 2 [HN] type species Victoria maritomnium Miyahara, 1941 (=Hoplia communis C.O. Waterhouse,
1875)

aureola Pallas, 1781: 18 (Scarabaeus) A: ES FE HEB JA MG NC QIN SHX WS XIN
duodecimpunctata A.G. Olivier, 1789: no 5: 121 (Melolontha)
flavicollis Reitter, 1903a: 119
impunctata Kraatz, 1879a: 232
macularis Reitter, 1903a: 118
maculata Reitter, 1890r: 378 [HN]
maculicollis Reitter, 1903a: 119
maculosa Reitter, 1897b: 48 [RN]

SCARABAEIDAE

219

octopunctata Kraatz, 1879a: 232
sibirica A.G. Olivier, 1789: no 5: 121 (Melolontha)

communis C.O. Waterhouse, 1875a: 100 A: JA
maculata Bates, 1889: 298
maritomnium Miyahara, 1941: 2 (Victoria)

hirticollis Heyden, 1889c: 666 A: SCH
reinii Heyden, 1879a: 87 [=1879b: 339] A: JA
reitteri Dalla Torre, 1912: 380 [RN] A: GAN QIN
fuliginosa Reitter, 1899c: 201 [HN]
imbella Reitter, 1903a: 119
nebulosa Reitter, 1903a: 119

rufocuprea S.I. Medvedev, 1952: 237 A: SCH
semenowi Heyden, 1889c: 667 A: GAN SCH
spectabilis S.I. Medvedev, 1952: 236 A: SCH
thibetana Dalla Torre, 1912: 382 [RN] A: QIN
oculata Reitter, 1899c: 200 [HN]

subgenus Hoplia Illiger, 1803: 226 type species Scarabaeus farinosus Linnaeus, 1760 (=Scarabaeus argenteus
Poda von Neuhaus, 1761)
Diphydactylus J. Thomson, 1858: 58 type species Diphydactylus singularis J. Thomson, 1858

africana africana Escalera, 1914a: 182 N: AG MO TU
cadmica Escalera, 1914a: 184

africana kocheri Baraud, 1981a: 133 N: AG MO TU
africana multiclava Normand, 1949: 89 N: TU
africana muluyensis Kocher, 1957: 178 N: MO
africana pardoi Baraud, 1981a: 133 N: MO
anatolica Reitter, 1890r: 381 A: TR
argentea Poda von Neuhaus, 1761: 20 (Scarabaeus) E: AL AU BE BH CR FR GE GR HU IT KO LU MC ME RO SB
SK SL SP SZ
ambigua Mulsant, 1842: 513
fissa Reitter, 1890r: 383
deflorata Mulsant, 1842: 512
farinosa Linnaeus, 1760: 138 (Scarabaeus)
glauca Mulsant, 1842: 512
griseola Rey, 1890a: 172
karamani Reitter, 1893d: 175
micans Mulsant, 1842: 512
rufolutea Mulsant, 1842: 512
sedicolor Mulsant, 1842: 512
sublutea Mulsant, 1842: 512
sulfurea Dufour, 1843: 64
viridula Mulsant, 1842: 512

220

SCARABAEIDAE

asteria Reitter, 1901c: 178 A: TD
attilioi Massa, 1979: 42 E: IT (Sicilia)
aulica aulica Linnaeus, 1767: 555 (Scarabaeus) N: AG MO
floralis A.G. Olivier, 1789: no 3: 69 (Scarabaeus)
regia Fabricius, 1787a: 22 (Melolontha)
sulphurea P.H. Lucas, 1846d: pl. 25 [HN]

aulica hispida Escalera, 1914a: 181 N: MO (Oujda)
bilineata Fabricius, 1801b: 178 (Melolontha) E: SP N: AG MO TU
bioscae Galante, 1978: 129 E: SP
brevipes S.I. Medvedev, 1952: 267 E: GG
brunnipes Bonelli, 1812: 156 E: AL AU BH BU CR FR GR IT PL SB SL SZ TR
pauperata Mulsant, 1842: 504 (Decamera)

caucasica Kolenati, 1846: 27 E: AB AR ST A: IN TR
chlorophana Erichson, 1847a: 710 E: PT SP
freyi Baraud, 1967b: 59
sulphuricolor Reitter, 1903a: 116

ciscaucasica S.I. Medvedev, 1952: 252 E: ST
coerulea Drury, 1773: 59 (Scarabaeus) E: FR SP SZ
argentata Geoffroy, 1785: 8 (Scarabaeus)
cosimii Sabatinelli, 1991b: 225
formosa Latreille, 1806: 116
imposita Buysson, 1906: 17
nigricollis Bedel, 1911a: 141
squamosa Villers, 1789: 30 (Scarabaeus)

colchica Petrovitz, 1967b: 339 A: TR
concolor Sharp, 1878b: 171 A: HP XIN
weisei Semenov, 1891: 327

cupulosa Peyerimhoff, 1940: 110 N: AG TU
cylindrica Reitter, 1903a: 123 E: AB A: TR
detrita Solsky, 1876a: 391 A: KI TD TM UZ
digitifera Peyerimhoff, 1940: 109 N: AG MO
dilutipes Reitter, 1890r: 383 E: AL BH BU CR HU RO SB UK
auriventris Apfelbeck, 1908: 530

dubia P. Rossi, 1790: 22 (Melolontha) E: IT
flavipes Germar, 1824: 130 E: AL BH GR ME
golovjankoi Jakobson, 1914: 4 E: BY UK
graminicola Fabricius, 1792b: 175 (Melolontha) E: BE BU BY CR CT CZ DE GE EN IT LA LT NL PL RO SB SK
SL UK
hakonensis Sawada, 1938b: 39 A: JA
herminiana Apfelbeck, 1908: 529 E: BH CR GR
hungarica Burmeister, 1844: 182 E: AL AU BH BY CR CZ FR GE HU MC ME PL RO SB SK SL SP UK
brunnescens Reitter, 1903a: 125

SCARABAEIDAE
christinae Baraud, 1967c: 196
nuda Erichson, 1847a: 714
subnuda Reitter, 1903a: 124

hyrcana S.I. Medvedev, 1952: 264 A: IN
korbi Petrovitz, 1958e: 160 E: SP
laconiae Petrovitz, 1958e: 161 E: GR (Pelopónnisos)
danieli Tesař, 1969a: 60

maremmana Leo, Liberto, Rattu & Sechi, 2010: 454 E: IT
mina Jakobson, 1914: 3 E: AB GG
minuta Panzer, 1789: 9 (Melolontha) E: IT
lepidota Illiger, 1803: 228
monticola O.G. Costa, 1839a: 1
oblongosquamulata Leoni, 1907: 188

moerens C.O. Waterhouse, 1875a: 100 A: JA
modestula S.I. Medvedev, 1952: 235
ohbayashii Nomura, 1968: 59

nigrina Reitter, 1885b: 378 E: AL GR
nova Pic, 1903a: 97
subaurina Reitter, 1903a: 125

paganettii J. Müller, 1907: 62 E: IT
parvula Krynicki, 1832: 127 E: BY CT EN KZ LA LT PL ST UK A: KZ
pollinosa Erichson, 1847a: 712 [HN]
pulverosa Küster, 1853: 63

pentheri Ganglbauer, 1906: 282 A: TR
peronii C.É. Blanchard, 1850: 72 N: AG MO TU
aulicoides Reitter, 1914b: 38
kobelti Reitter, 1891b: 33

pollinosa Krynicki, 1832: 127 E: AB AR GG ST A: TR
pilicollis Erichson, 1847a: 711

pontica Petrovitz, 1967b: 340 A: TR
pubicollis Küster, 1849a: 62 E: FR (Corse) IT (Sardegna)
puncticollis Gebler, 1832: 53 A: ES
rossica S.I. Medvedev, 1952: 260 E: CT ST
rungsi Kocher, 1957: 177 N: AG MO
schuberti Petrovitz, 1962b: 124 A: TR
shirakii Nomura, 1959b: 39 A: JA
stenolepis Apfelbeck, 1908: 529 E: AL BU MC ME
susiana Escalera, 1913a: 32 N: MO
uniformis Reitter, 1885b: 377 E: GR
walterrossii Sabatinelli, 1993a: 35 E: IT (Molise)
zaitzevi Jakobson, 1914: 5 E: BY CT EN KZ LA ST UK A: KZ

221

222

SCARABAEIDAE

subgenus Hyperis Dejean, 1833: 167 type species Hoplia eversmanni Faldermann, 1833 (=Hoplia paupera
Krynicki, 1832)
Xenoplia S.I. Medvedev, 1952: 229 type species Hoplia paupera Krynicki, 1832

kunzii W.L.E. Schmidt, 1840a: 73 E: TR A: TR
paupera Krynicki, 1832: 127 E: KZ ST A: KI KZ MG XIN
averini S.I. Medvedev, 1952: 229
eversmanni Faldermann, 1833: 50
kaszabi Endrődi, 1968b: 80
kuldshensis Balthasar, 1936b: 97
zaissanica Skopin, 1963: 142

subgenus Pseudohoplia Reitter, 1890r: 375 type species Hoplia gabriellina Fairmaire, 1887
Sinoplia S.I. Medvedev, 1952: 233 type species Hoplia bifasciata S.I. Medvedev, 1952 (=Hoplia gabriellina Fairmaire, 1887)

campestris Fairmaire, 1887b: 105 A: YUN
gabriellina Fairmaire, 1887b: 104 A: SCH YUN
bifasciata S.I. Medvedev, 1952: 233

shibatai makiharai Miyake, 1986c: 202 (Pseudohoplia) A: TAI
shibatai matsudai Miyake, 1986c: 202 (Pseudohoplia) A: TAI
shibatai shibatai Miyake, 1986c: 201 (Pseudohoplia) A: TAI
shibatai yushana Miyake, 1986c: 202 (Pseudohoplia) A: TAI
genus Hoplia, species incertae sedis
bisignata Gyllenhal, 1817: 69 A: NP
bomiensis Zeng, 1986: 273 A: XIZ
choui Miyake, 1986b: 203 A: TAI
chinensis Endrődi, 1952b: 60 A: FUJ
citrea Endrődi, 1952b: 58 A: YUN
citrinella Fairmaire, 1887b: 105 A: YUN
clotildae Sabatinelli, 1983a: 184 A: NP
coeruleosignata Moser, 1916a: 187 A: BT KA NP SD
schereri Tesař, 1969a: 57

coluzzii Sabatinelli, 1983a: 186 A: BT
elegantula White, 1844: 424 A: HKG
excellens Endrődi, 1952b: 55 A: FUJ
forsteri Tesař, 1969a: 56 A: NP
freudei bhutanica Sabatinelli, 1983a: 180 A: BT
freudei freudei Tesař, 1969a: 54 A: NP
fukiensis Endrődi, 1952b: 59 A: FUJ
fulvofemorata Moser, 1912a: 308 A: NP SD
gracilis Endrődi, 1952b: 57 A: FUJ
grisea Moser, 1912a: 313 A: NP SD UP
lahulensis Moser, 1919: 363

griseosparsa Moser, 1921: 72 A: GUA ORR

SCARABAEIDAE
gyirongensis Zeng, 1986: 273 A: XIZ
harpagon Fairmaire, 1887b: 104 A: YUN
hirsuta Moser, 1912a: 314 A: BT SD
indica Moser, 1912a: 310 A: SD
ingrata Fairmaire, 1888a: 18 A: FUJ
inornata Kobayashi, 1990a: 28 A: TAI
iridescens Zeng, 1986: 271 A: XIZ
klapperichi Endrődi, 1952b: 53 A: FUJ
koreana Moser, 1920: 27 A: NC
laetitiae Sabatinelli, 1983a: 182 A: BT
lama Endrődi, 1952b: 54 A: “Thibet”
latesuturata Fairmaire, 1900b: 620 (Ectinohoplia) A: SCH TAI
hayashii Miyake, 1986b: 204

lishana Miyake, 1986b: 205 A: TAI
mahayana Sabatinelli, 1983a: 200 A: BT SD
mediocris Fairmaire, 1887d: 315 A: BEI
minuscula Fairmaire, 1889a: 17 A: YUN
monticola Miyake, 1986b: 209 A: TAI
nakanei Miyake, 1986b: 207 A: TAI
nebulosa Fairmaire, 1889a: 6 A: JIX
nepalensis Tesař, 1969a: 59 A: NP
nengkaoshana alishana Kobayashi, 1990a: 29 A: TAI
nengkaoshana nengkaoshana Miyake, 1986b: 211 A: TAI
nigromaculata Moser, 1912a: 308 A: NP SD
nigrosparsa Moser, 1921: 70 A: YUN
ochraceoscutellata Moser, 1921: 70 A: YUN
ochreata Fairmaire, 1889a: 17 A: JIX
pictipes Fairmaire, 1889a: 15 (Ectinohoplia) A: SCH
platyca Zeng, 1986: 272 A: HUB
plebeja Moser, 1921: 69 A: YUN
polita Bates, 1891a: 14 A: HP KA PA
pulchra Miyake, 1986b: 207 A: TAI
rufopicta Fairmaire, 1889a: 16 A: SCH
sabatinellii A. Rey, 2006: 189 [RN] A: SD
argenteola Sabatinelli, 1983a: 193 [HN]

sabraechatilae Sabatinelli, 1983a: 192 A: BT
scheibei Balthasar, 1936c: 207 A: AF
scheini Endrődi, 1953a: 114 A: SCH
scutellaris C.O. Waterhouse, 1877c: 268 A: “N. China”
shimomurai Kobayashi, 1990a: 28 A: TAI
shirozui Kobayashi, 1990b: 21 A: TAI

223

224

SCARABAEIDAE

simillima Miyake, 1986b: 206 A: TAI
siningensis J. Frivaldszky, 1892: 122 A: QIN
squamacea White, 1844: 424 A: HKG
squamigera Hope, 1831: 24 A: NP
squamiventris Burmeister, 1855: 486 A: “Indien”
taipeienis Kobayashi & Chou, 2008: 71 A: TAI
taiwana Miyake, 1986b: 204 A: TAI
tesari Sabatinelli, 1983a: 188 A: BT
testudinis Sabatinelli, 1997a: 65 A: NP
tuberculicollis Moser, 1912a: 311 A: SD
tuberculifera Moser, 1924: 136 [RN] A: YUN
bituberculata Moser, 1920: 28 [HN]

tuberculifrons Moser, 1912a: 311 A: SD
ushijimai Kobayashi, 1990b: 19 A: TAI
validipes Fairmaire, 1889a: 17 A: SCH
versicolor Moser, 1921: 71 A: YUN
vestita Boheman, 1858: 53 A: HKG
vicina J. Frivaldszky, 1890: 199 A: SCH
virginioi Sabatinelli, 1983a: 190 A: BT
viridissima Brenske, 1894a: 34 A: BT NP SD
viridula gibbosa Sabatinelli, 1983a: 199 A: BT
viridula viridula Brenske, 1899: 177 A: NP SD ORR
vittata Nonfried, 1895: 285 A: GUA

tribe Leucopholini Burmeister, 1855

genus Asactopholis Brenske, 1894c: 276 type species Lepidiota gracilipes Sharp, 1876
Empectida Moser, 1914b: 154 type species Empectida vethi Moser, 1914

bicolor bicolor Sharp, 1876: 78 (Lepidiota) A: BT NP UP ORR
capucina Fairmaire, 1893d: 289 (Phytalus) A: YUN ORR
bituberculata Moser, 1908c: 334

microsquamosa Frey, 1972a: 119 (Holotrichia) A: BT NP SD UP
dehradunus Mittal, 1988: 159

genus Cyphochilus C.O. Waterhouse, 1867: 141 type species Melolontha candida A.G. Olivier, 1789
apicalis C.O. Waterhouse, 1867: 144 A: FUJ HAI HUN
candidus A.G. Olivier, 1789: no. 5: 15 (Melolontha) A: SD ORR
septentrionalis C.O. Waterhouse, 1867: 141

costulatus Bates, 1891b: 76 A: SCH ORR
crataceus crataceus Niijima & Kinoshita, 1923: 75 (Lepidiota) A: TAI
miwai Nakabayashi, 1939: 139

crataceus taipeiensis Kobayashi & C. K. Yu, 1993b: 350 A: TAI

SCARABAEIDAE

225

elongatus Brenske, 1894a: 80 A: JIX SHG
farinosus C.O. Waterhouse, 1867: 143 A: HAI HUN NC NE SC SHG
insulanus Moser, 1918: 242 A: TAI
marginalis Fairmaire, 1902c: 316 A: JIX
ochraceus Moser, 1915a: 132 A: SHN
pygidialis Nonfried, 1893: 332 A: NP ORR
testaceipes Fairmaire, 1902c: 316 A: JIX
unidentatus Nomura, 1977b: 102 A: TAI
genus Dasylepida Moser, 1913c: 287 type species Lepidiota nana Sharp, 1876 (=Dasylepida fissa Moser, 1913)
costata Nonfried, 1892d: 266 (Melolontha) A: CH
gononii Fairmaire, 1897e: 213 (Pseudolontha) A: JIX
ishigakiensis Niijima & Kinoshita, 1927: 23 (Brahmina) A: JA
nana Sharp, 1876: 76 (Lepidiota) A: CE TAI
fissa Moser, 1913c: 287

obscurata Fairmaire, 1878a: 99 (Schizonycha) A: CH
genus Dedalopterus Sabatinelli & Pontuale, 1998a: 65 type species Cyphochilus signatus Moser, 1908
fencli Zídek & Krajčík, 2007: 11 A: GUA
fujianensis Y.-W. Zhang, 1990b: 192 (Malaisius) A: FUJ
hainanensis Y.-W. Zhang, 2004: 334 (Malaisius) A: HAI
intermedius Y.-W. Zhang, 1990b: 190 (Malaisius) A: YUN
itohi Sabatinelli & Pontuale, 1998a: 70 A: SCH
melanodiscus Y.-W. Zhang, 1990b: 191 (Malaisius) A: GUI
signatus Moser, 1908d: 356 (Cyphochilus) A: YUN
pinae Y.-W. Zhang, 1990b: 198 (Malaisius)

genus Lepidiota Kirby, 1828a: 445 type species Melolontha stigma Fabricius, 1798
Eucirrus Melly, 1832: cl. IX, pl. 47 type species Eucirrus mellyi Melly, 1832

acuminata Moser, 1913c: 284 A: YUN ORR
albistigma Burmeister, 1855: 295 A: HP NP ORR
bimaculata W.W. Saunders, 1839: 176 (Melolontha) A: HAI KA NP SD YUN ORR
griffithii Hope, 1841b: 62 (Eucirrus)

crenulata Wiedemann, 1821: 132 (Melolontha) A: AF PA ORR
discedens Sharp, 1876: 73 A: SD YUN ORR
haasei Nonfried, 1906: 218
hispida Frey, 1969b: 113

hirsuta Brenske, 1893a: 46 A: CH
hispida Frey, 1969b: 113 A: CH
impluviata C.É. Blanchard, 1851: 157 A: “Inde sept.”
nonfriedi Brenske, 1893a: 42 A: HKG

226

SCARABAEIDAE

praecellens Bates, 1891b: 76 A: SCH
punctatipennis C.É. Blanchard, 1851: 157 A: SD ORR
richteri Brenske, 1893c: 44 A: “Himalaya”
rugosipennis C.É. Blanchard, 1851: 157 A: “Inde sept.”
sticticoptera C.É. Blanchard, 1851: 157 A: UP ORR
tridens Sharp, 1876: 74 A: SCH ORR
amoena Felsche, 1902: 360

genus Leucopholis Dejean, 1833: 160 type species Melolontha rorida Weber, 1801
crassa Brenske, 1893a: 58 A: SD ORR
lateralis Brenske, 1893a: 56 A: CH ORR
nudiventris Burmeister, 1855: 300 A: CH
genus Leucophorus Brenske, 1894a: 79 type species Leucophorus flabellatus Brenske, 1894
flabellatus Brenske, 1894a: 80 A: UP
genus Malaisius Arrow, 1941: 5 type species Malaisius mollis Arrow, 1941
similis C.-L. Li & P.-S. Yang, 1999a: 199 A: YUN
yunnanus Moser, 1908d: 357 (Cyphochilus) A: YUN

tribe Melolonthini Leach, 1819

genus Achranoxia Kraatz, 1888a: 208 type species Polyphylla koenigi Brenske, 1888
Ochranoxia Kraatz, 1888a: 208 type species Anoxia semiflava Kraatz, 1883

expallida Semenov, 1935: 281 (Ochranoxia) A: KZ
koenigi Brenske, 1888: 353 (Polyphylla) A: KZ TD TM UZ
planipalpis Reitter, 1902b: 278

peltzi Semenov & S.I. Medvedev, 1936: 115 (Ochranoxia) A: KZ UZ
semiflava Kraatz, 1883f: 357 (Anoxia) A: TM
sijesowi Reitter, 1913e: 657 (Ochranoxia) A: UZ
transaralica Semenov & S.I. Medvedev, 1936: 116 (Ochranoxia) A: KZ
turcomanica Nikolajev & Nikritin, 1973: 1875 (Ochranoxia) A: TM
varentzowi Semenov, 1897: 244 A: TM
genus Anoxia Laporte, 1832b: 407 type species Melolontha villosa Fabricius, 1781
subgenus Anoxia Laporte, 1832b: 407 type species Melolontha villosa Fabricius, 1781
Catalasis Dejean, 1833: 159 type species Melolontha villosa Fabricius, 1781

arenbergeri Petrovitz, 1971c: 183 A: TR
asiatica asiatica Desbrochers des Loges, 1872: 357 E: AR A: IQ SY TR
syriaca Brenske, 1894a: 24

asiatica montandoni Dewailly, 1945b: 77 E: RO
caphtor Petrovitz, 1971c: 184 E: GR (Kríti)

SCARABAEIDAE

227

cretica Kiesenwetter, 1858: 236 E: GR (Kríti)
kraatzi Reitter, 1890h: 175 A: TR
lodosi Baraud, 1990d: 297 A: TR
makrisi Keith, 2002b: 25 A: CY
maculiventris Reitter, 1890d: 106 A: IS LE SY
israelitica Petrovitz, 1971c: 184

maldesi Baraud, 1980b: 254 N: AL MO TU
naviauxi Baraud, 1990d: 299 A: LE
nicaeensis Baraud, 1990d: 300 A: TR
nigricolor Pic, 1905f: 177 A: TR
laterufa Pic, 1946: 99

pasiphae Reitter, 1890d: 107 E: GR MC
gracilis Reitter, 1890h: 176

pilosa Fabricius, 1792b: 156 (Melolontha) E: AB AR AU CR CZ GG HU IT PL RO SB SK ST UK A: IN KZ
cinerea Motschulsky, 1860i: 524 [=1861c: 423]
scutellaris scutellaris Mulsant, 1842: 422 E: AL BH CR FR GG IT ME SL SP ST N: MO TU
tunisia Heyden, 1886a: 56

scutellaris argentea Aliquò & Massa, 1976: 152 E: IT (Sicilia)
scutellaris naxiana Reitter, 1890d: 106 E: GR (Náxos)
scutellaris rumelica Apfelbeck, 1899: 291 E: BU GR MC
scutellaris sicula Motschulsky, 1860i: 524 [=1861c: 424] E: IT (Sicilia)
tristis Reitter, 1902b: 286 E: GG
tristis Shokhin, 2001: 449 [HN]

villosa tethys Reitter, 1890h: 176 E: GR
villosa villosa Fabricius, 1781: 496 (Melolontha) E: AL AU BE BU FR GE IT LU NL PT RO SB SP SZ UK
anceyi Pic, 1925: 1
cerealis Scopoli, 1786: 49 (Scarabaeus)
funebris Reitter, 1902b: 287
lanata Balthasar, 1931a: 57
pauliani Dewailly, 1945b: 60
pilosa Mulsant, 1842: 425 [HN]

subgenus Mesanoxia S.I. Medvedev, 1951a: 161 type species Melolontha australis Schönherr, 1817
Archianoxia Mikšič, 1958a: 6 type species Anoxia derelicta Desbrochers des Loges, 1874 (=Anoxia emarginata
Coquerel, 1860)

australis Schönherr, 1817: 169 (Melolontha) E: FR IT PT SP
hungarica Desbrochers des Loges, 1874: 83
occidentalis Fabricius, 1775: 32 (Melolontha)
occitanica Mulsant, 1842: 420
rosinae Reitter, 1891b: 36

cypria Zurcher, 1911: 254 A: CY IS
mavromoustakisi Mikšič, 1959a: 41

228

SCARABAEIDAE

desbrochersi Baraud, 1980b: 254 N: AL TU
emarginata Coquerel, 1860: 424 N: AL MO TU
derelicta Desbrochers des Loges, 1873: xli

hirta Reitter, 1890h: 175 E: GR
luteipilosa Desbrochers des Loges, 1874: 82 N: TU
matutinalis corsicana Sabatinelli, 1976: 150 E: FR (Corse) IT (Sardegna)
matutinalis matutinalis Laporte, 1832b: 407 E: AL CR IT ME SL
armerinae Ragusa, 1893b: 237
curzolensis J. Müller, 1938: 57
vespertina Mulsant, 1842: 418

matutinalis moltonii Sabatinelli, 1976: 153 E: IT (Isola Vulcano)
matutinalis sardoa Motschulsky, 1860i: 525 [=1861c: 425] E: IT (Sardegna)
matutinalis suturalis Reitter, 1890h: 174 E: BU GR
matutinalis thumseri Guerlach, 2011: 250 E: IT (Sardegna)
rattoi Escalera, 1906: 162 N: MO
theryi Kocher, 1952b: 77
zuercheri Reitter, 1909c: 119

reisseri Petrovitz, 1964d: 128 E: GR (Kríti)
subgenus Protanoxia S.I. Medvedev, 1951 type species Melolontha orientalis Krynicki, 1832
baraudi Keith, 2003c: 354 [RN] A: CY
cypria Baraud, 1989b: 280 [HN]

ciliciensis Baraud, 1989b: 282 A: TR
cingulata Marseul, 1868: 173 A: LE SY
kocheri Dewailly, 1957: 174 N: MO
laevimaculata Petrovitz, 1973b: 32 A: IS
orientalis Krynicki, 1832: 123 (Melolontha) E: AU BH BU CR GR HU IT MC RO SB SL ST TR UK A: IS LE
SY TR
meridionalis Reitter, 1890d: 105

rotroui Dewailly, 1957: 174 N: MO
smyrnensis Petrovitz, 1965b: 191 A: TR
genus Anoxoides Petrovitz, 1971d: 232 type species Anoxoides bytinskisalzi Petrovitz, 1971
bytinskisalzi Petrovitz, 1971d: 232 A: IS
genus Carlschoenherria Bezděk, nom. nov. [see new Acts] type species Melolontha sulcipennis Laporte, 1840
Schoenherria Burmeister, 1855: 417 [HN] type species Melolontha sulcipennis Laporte, 1840

brenskei Nonfried, 1906: 219 [DA] A: NW
genus Clypeolontha C.-L. Li & P.-S. Yang, 1999b: 394 type species Melolontha alboplagiata Brenske, 1898
alboplagiata alboplagiata Brenske, 1898b: 236 (Melolontha) A: BT SD

SCARABAEIDAE
genus Cryptotrogus Kraatz, 1888a: 206 type species Cryptotrogus weisei Kraatz, 1888
Meganoxia Reitter, 1902b: 280 type species Meganoxia orita Reitter, 1902

afghanus Balthasar, 1956b: 426 A: AF PA
gallagheri Sabatinelli & Pontuale, 1998b: 129 A: OM
kruppi Sabatinelli & Pontuale, 1998b: 127 A: SA
maljuzhenkoi Zaitzev, 1928: 397 (Cyphonoxia) E: AR
mesopotamicus Petrovitz, 1962b: 119 (Meganoxia) A: IQ
miksici Petrovitz, 1965b: 195 A: IN
niveus Hampe, 1852: 311 (Anoxia) E: AR
orita Reitter, 1902b: 280 (Meganoxia) E: AR
pajnii Mittal, 1979: 265 A: HP
weisei Kraatz, 1888a: 206 N: EG A: JO
rorifera Reitter, 1902b: 280 (Melolontha)

zarudnianus Semenov & S.I. Medvedev, 1936: 114 (Meganoxia) A: IN
genus Cyphonotus Fischer von Waldheim, 1824: 213 type species Scarabaeus testaceus Pallas, 1781
bicoloratus Petrovitz, 1962b: 119 A: IQ
testaceus Pallas, 1781: 19 (Scarabaeus) E: AR ST A: IN KZ PA TM UZ
anketeri Herbst, 1790: 43 (Melolontha)
caucasicus Jacquelin du Val, 1860: 54
glaucus Gmelin, 1790: 1583 (Scarabaeus)
maximus Reitter, 1895e: 205
integriceps Reitter, 1902b: 292
oryctoides Semenov, 1892c: 474

genus Cyphonoxia Reitter, 1889g: 276 type species Cyphonoxia praestabilis Reitter, 1889
brenskei Reitter, 1895d: 152 A: AF TM
buettikeri Sabatinelli & Pontuale, 1998b: 123 A: SA
glasunowi Semenov, 1897: 245 A: AF IN TM
haarlovi Petrovitz, 1955: 280 A: AF
kermanensis Semenov & S.I. Medvedev, 1936: 113 A: IN
kircheri Balthasar, 1930b: 59 N: AG LB MO
praestabilis Reitter, 1889g: 276 A: IN IQ IS
tatianae Semenov & S.I. Medvedev, 1936: 114 A: IN
zarudnyi Semenov & S.I. Medvedev, 1936: 113 A: IN
genus Exolontha Reitter, 1902b: 269 type species Melolontha umbraculata Burmeister, 1855
Parexolontha Chang, 1965b: 229 type species Exolontha tonkinensis Moser, 1913

bhutanensis Frey, 1975b: 238 A: BT (Schoenherria)
castanea Chang, 1965b: 227 A: HAI ORR

229

230

SCARABAEIDAE

chiangmaiensis laotica Li & Keith, 2012: 273 A: YUN ORR
fuliginosa Fairmaire, 1889a: 22 (Melolontha) A: SCH
laticauda Bates, 1891b: 77 (Melolontha) A: SCH
mandarina Sharp, 1876: 87 (Melolontha) A: CH ORR
obscura Chang, 1980b: 409 (Parexolontha) A: SCH
omeia Chang, 1965b: 226 A: CHQ SCH
rugulosa Y.-W. Zhang, 1993: 270 A: CHQ HUN
serrulata Gyllenhal, 1817: 73 (Melolontha) A: FUJ GUA HAI HKG HUN JIX ZHE ORR
manillarum C.É. Blanchard, 1851: 160 (Melolontha)

similis Chang, 1965b: 228 A: GUA GUX
tonkinensis Moser, 1913c: 292 A: YUN ORR
umbraculata Burmeister, 1855: 418 (Melolontha) A: CHQ FUJ GUX HKG HUN SCH
xizangensis Y.-W. Zhang, 1981: 347 (Parexolontha) A: XIZ
yaoshana Chang, 1965b: 227 A: GUX
genus Euranoxia Semenov, 1890b: 196 type species Euranoxia valida Semenov, 1890
valida Semenov, 1890b: 197 A: TD
genus Melolontha Fabricius, 1775: 31 type species Scarabaeus melolontha Linnaeus, 1758
Apropyga S.I. Medvedev, 1951a: 126 type species Melolontha aceris Faldermann, 1835
Hoplosternus Guérin-Méneville, 1838c: 63 type species Melolontha chinensis Guérin-Méneville, 1838
Ludibrius Gozis, 1886: 33 [RN] type species Scarabaeus melolontha Linnaeus, 1758
Oplosternus Guérin-Méneville, 1838c: 63 type species Melolontha chinensis Guérin-Méneville, 1838 [incorrect original
spelling]
Zhangia Bunalski, 2002: 408 type species Zhangia margaritae Bunalski, 2002 (=Hoplosternus maculatus Chang, 1983)

aceris Faldermann, 1835a: 270 E: AB AR GG ST
praeambula Kolenati, 1846: 24

aeneicollis Bates, 1891a: 16 A: HP PA
afflicta afflicta Ballion, 1871a: 343 A: UZ
zeravschanica Protsenko, 1974: 3

afflicta clypeata Reitter, 1887e: 542 A: KI TD UZ
afflicta hissarica S.I. Medvedev, 1951a: 129 A: TD
albida I. Frivalszkky, 1835: 258 E: GR TR A: IS SY TR
candidans Burmeister, 1855: 413
naxiana Reitter, 1887e: 538

albopruinosa Fairmaire, 1878a: 98 A: CE
anita Reitter, 1902b: 263 A: SY TR
borumandi Montreuil, 2012: 450 A: IN
chinensis Guérin-Méneville, 1838c: 63 A: GUA HUN JIX SCH ORR
heydeni Moser, 1913c: 288 (Hoplosternus)

ciliciensis Petrovitz, 1962b: 120 A: TR

SCARABAEIDAE

231

costipennis Fairmaire, 1889a: 19 A: SCH XIZ
tricostata Brenske, 1903: 335

cuprescens C.É. Blanchard, 1871: 811 A: HP KA SCH XIZ
davidis Fairmaire, 1878a: 97 A: CE
excisicauda Balthasar, 1936b: 95 A: LE
flabellata Sharp, 1876: 86 A: SD XIZ ORR
frater frater Arrow, 1913c: 400 A: JA
frater gobiensis Nomura, 1952a: 28 A: HEN NMO
furcicauda Ancey, 1881: 412 A: AF HP KA NP PA SD TD XIN XIZ ORR
bifurcata Brenske, 1896a: 164 (Hoplosternus)
insignis Semenov, 1890b: 197

gussakovskii S.I. Medvedev, 1945: 14 A: TD
guttigera Sharp, 1876: 87 A: AP SD ORR
hippocastani hippocastani Fabricius, 1801b: 162 E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE HU IR IT LA
LT LU MD NL NR NT PL RO SB SK SL SP ST SV SZ UK A: BEI ES KZ NE WS
albicans Stierlin, 1863: 58
aethiops Westhoff, 1884: 72
amasicola Westhoff, 1884: 68
coronata Mulsant, 1842: 415
discoidalis Dalla Torre, 1879: 113
festiva Westhoff, 1884: 73
fuscicollis Kraatz, 1885a: 69
metzleri Westhoff, 1884: 68
mulsanti Bedel, 1911a: 118 [RN]
nigricollis Mulsant, 1842: 415
nigripes Comolli, 1837: 24
picea Wanach, 1906: 233
rex Dalla Torre, 1879: 113
romana Reitter, 1887e: 535
schwarzi Kraatz, 1888b: 362
tibialis Mulsant, 1842: 415

hippocastani mongolica Ménétriés, 1854: 28 A: BEI GAN ES FE MG NE NO
baicalica Reitter, 1892a: 152

incana Motschulsky, 1854c: 46 (Hoplosternus) A: BEI FE GUI HEB HEI HUN JIN LIA NC SCH
indica Hope, 1831: 23 A: HP NP PA UP YUN ORR
nitidicollis C.É. Blanchard, 1851: 158 (Hoplosternus)

insulana Moser, 1918: 246 A: TAI (Hoplosternus)
japonica Burmeister, 1855: 420 A: CE HEI JA JIL
gotenbaensis Niijima & Kinoshita, 1923: 62
haroldi Moser, 1913c: 290 (Hoplosternus) [RN]
japonica Harold, 1875e: 291 (Hoplosternus) [HN]

232

SCARABAEIDAE

kraatzi Reitter, 1906a: 737 [RN] E: AB A: IN
brenskei Reitter, 1902b: 267
persica Reitter, 1902b: 267
tibialis Kraatz, 1882h: 245 [HN]

macrophylla Fischer von Waldheim, 1830: 186 A: “Persien”
maculata Chang, 1983b: 395 (Hoplosternus) A: YUN ORR
margaritae Bunalski, 2002: 409 (Zhangia)

mandarina Fairmaire, 1878a: 98 A: CH ORR
masafumii Nomura, 1952a: 29 A: JA (Okinawa-shotō: Ishigaki-jima)
medvedevi Kryzhanovskij, 1978: 133 A: UZ
melolontha Linnaeus, 1758: 351 (Scarabaeus) E: AL AU BE BH BU BY CR CT CZ DE EN FR GB GE GR HU IR IT
LA LS LT LU MC MD ME NL PL RO SB SK SL SP ST SV SZ TR UK A: TR
albida L. Redtenbacher, 1847b: 265 [HN]
asiatica Brenske, 1900a: 151
bipunctata Papp, 1943: 457
bioculata Papp, 1943: 457
colpopyga Petz, 1905: 100
discicollis Mulsant, 1842: 413
femoralis Kraatz, 1885a: 69
funesta Westhoff, 1884: 60
humeralis Westhoff, 1884: 61
luctuosa Westhoff, 1884: 61
lugubris Mulsant, 1842: 412
maialis Moll, 1785: 179 (Scarabaeus)
melanopus Westhoff, 1884: 60
nigra Kellner, 1877: 405
nigroapicalis Papp, 1943: 458
nigromarginalis Papp, 1943: 458
nigropygidialis Papp, 1943: 458
obscuripes Westhoff, 1884: 61
pulcherrima Dalla Torre, 1879: 113
redtenbacheri Dalla Torre, 1913: 272 [RN]
ruficeps Kraatz, 1885a: 69
ruficollis Mulsant, 1842: 413
scapularis Westhoff, 1884: 58
unicolor Papp, 1943: 457
unipunctata Papp, 1943: 458
vulgaris Fabricius, 1775: 32

minima Kobayashi, 1985b: 20 A: TAI
hualiensis Kobayashi & Chou, 2009: 49

nepalensis C.É. Blanchard, 1851: 158 (Hoplosternus) A: HP NP XIZ ORR

SCARABAEIDAE

233

papposa Illiger, 1803: 215 E: PT SP
fucata C.É. Blanchard, 1851: 160
hybrida T. Charpentier, 1825: 212
rufotestacea Kraatz, 1885a: 70

pectoralis Megerle von Mühlfeld, 1812: 21 E: AB AL AR AU BH BU CR CZ FR GE GG GR HU IT KO MC ME PL
RO SB SK SL ST TR UK
aceris Erichson, 1847a: 675 [HN]
albida Laporte, 1840: 131 [HN]
extorris Erichson, 1847a: 675
farinosa Kraatz, 1864: 11
fosca Reitter, 1902b: 268
fuscotestacea Kraatz, 1885a: 70
fuscotestacea Reitter, 1887e: 538 [HN]
ilsia Reitter, 1902b: 268
nigritula Kraatz, 1885a: 71
pulverea Mulsant, 1842: 410
rhenana Bach, 1845: 19
rufithorax Kraatz, 1885a: 70
satema Reitter, 1902b: 268
soror Marseul, 1879b: 474
spatulata Ballion, 1871a: 343
velebitica J. Müller, 1902: 460

permira Reitter, 1887e: 539 E: GG ST
pseudofurcicauda Keith, 2008d: 343 A: XIZ
reichenbachi Keith, 2008d: 347 A: XIZ
rufocrassa Fairmaire, 1889a: 20 A: SCH
sardiniensis Drumont, Muret, Hager & Penner, 1999: 188 E: IT (Sardegna)
satsumaensis satsumaensis Niijima & Kinoshita, 1923: 62 A: JA (Honshū)
kinoshitai Tesař, 1938: 165 (Hoplosternus)

satsumaensis shikokuana Nomura, 1977a: 78 A: JA (Shikoku)
sculpticollis Fairmaire, 1891d: x A: HUB
shanghaiana Brenske, 1896b: 203 A: CH
taihokuensis Niijima & Kinoshita, 1923: 65 A: TAI
taiwana Nomura, 1952a: 29

tamina Nomura, 1964a: 54 A: JA (Amami-Ôshima)
tarimensis Semenov, 1896a: 22 A: XIN
taygetana A. Rey, 1999a: 97 E: GR
tenuicauda Fairmaire, 1896a: 87 A: “Tibet”
virescens Brenske, 1896b: 202 A: BT HP NP PA SD XIZ
amplipennis Frey, 1975b: 238 (Melolontha)

wushana Nomura, 1977b: 103 A: TAI

234

SCARABAEIDAE

genus Microphylla Kraatz, 1890: 351 type species Anoxia paupera Hampe, 1852
paupera Hampe, 1852: 312 (Anoxia) E: AR ST A: IN TM
albella Reitter, 1902b: 288
xorema Reitter, 1902b: 288

genus Oligophylla Kraatz, 1894e: 289 type species Anoxia detrita Fairmaire, 1875
detrita Fairmaire, 1875a: 509 (Anoxia) N: AG TU
peyerimhoffi Dewailly & Baraud, 1982: 93 N: AG
genus Polyphylla Harris, 1841: 30 type species Melolontha variolosa Hentz, 1830
subgenus Centrasiobia Semenov, 1902d: 351 type species Polyphylla schestakowi Semenov, 1900
schestakowi Semenov, 1900d: 310 A: XIN
subgenus Grananoxia Brenske, 1890d: 198 type species Granida annamenis Fleutiaux, 1887
annamensis Fleutiaux, 1887: 63 (Granida) A: FUJ GUA HAI HKG ORR
nuda Petrovitz, 1964d: 129

dahnshuensis C.-L. Li & P.-S. Yang, 1997a: 114 A: TAI
subgenus Granida Motschulsky, 1862a: 8 type species Granida albolineata Motschulsky, 1861
albolineata Motschulsky, 1862a: 8 (Granida) A: JA
jessopi Dewailly, 1993: 12 A: GUX FUJ
minor Nomura, 1977b: 104 A: TAI YUN
parva Kobayashi & Chou, 2008: 74 A: TAI
schoenfeldti Brenske, 1890d: 198 A: JA
taiwana Sawada, 1950a: 6 (Granida) A: TAI
subgenus Gynexophylla S.I. Medvedev, 1951a: 86 type species Polyphylla laticollis Lewis, 1887
edentula Harold, 1878a: 71 (Granida) A: NP UP XIZ
brevicornis Petrovitz, 1970b: 20
brevicornis Y.-W. Zhang, 1981: 347 [HN]

exilis Y.-W. Zhang, 1984: 107 A: YUN
gracilicornis gracilicornis C.É. Blanchard, 1871: 811 (Melolontha) A: BEI CHQ GAN GUI HEB SCH SHX QIN
XIZ YUN
hirtifrons Reitter, 1899c: 202
mongola Fairmaire, 1888c: xvi

gracilicornis licenti Dewailly, 1948: 109 A: BEI SHX
intermedia Y.-W. Zhang, 1981: 349 A: WP XIZ
laticollis chinensis Fairmaire, 1888c: xvii A: BEI CHQ GAN HEB HEI SCH SHN XIZ ZHE
potanini Semenov, 1890b: 198
vacca Semenov, 1890b: 199

laticollis laticollis Lewis, 1887b: 231 A: JA
laticollis mandshurica Semenov, 1900c: 308 A: JIL LIA NC
lerestifi Guerlach, 2012: 45 A: BT NP
maculipennis Moser, 1919: 353 A: SCH XIZ YUN

SCARABAEIDAE

235

sikkimensis albosparsa Moser, 1919: 353 A: HP
sikkimensis sikkimensis Brenske, 1896b: 201 A: BT NP SD UP XIZ
subgenus Mesopolyphylla S.I. Medvedev, 1951a: 89 type species Melolontha irrorata Gebler, 1841
irrorata Gebler, 1841a: 372 (Melolontha) A: KI KZ XIN
tridentata Reitter, 1890i: 186 A: KI KZ TD TM UZ
subgenus Polyphylla Harris, 1841: 30 type species Melolontha variolosa Hentz, 1830
Macronoxia Crotch, 1873: 61 type species Melolontha occidentalis Linnaeus, 1767
Polylamina A. R. Hardy, 1974: 6 type species Polyphylla pubescens Cartwright, 1939

albertischulzi Kuntzen, 1933: 461 E: GR (Kríti)
boryi Brullé, 1832: 175 (Melolontha) E: BU CR GR ME TR A: TR
lesinae Reitter, 1890a: 21

davidis Fairmaire, 1888c: xvii A: CHQ FUJ GUI HUN JIA SCH ZHE
formosana Niijima & Kinoshita, 1923: 69 A: TAI
fullo algirana Brenske, 1896b: 200 N: AG TU
fullo fullo Linnaeus, 1758: 352 (Scarabaeus) E: AL AR AU BE BH BU BY CR CT CZ DE FR GB GE GG GR HU IT
KO LA LT LU MC MD ME NL PL RO SB SK SL SP ST SV SZ TR UK A: TR
luctuosa Mulsant, 1842: 407 (Melolontha)
marmorata Mulsant, 1842: 407 (Melolontha)

fullo macrocera Reitter, 1891b: 36 N: MO
hassi Hass & Reichenbach, 2014: 207 E: GR (Ródos)
maroccana descarpentriesi Dewailly & Baraud, 1982: 91 N: AG
maroccana maroccana Peyerimhoff, 1925a: 23 N: MO
naxiana Reitter, 1902b: 273 [RN] E: GR (Náxos)
nubecula Frey, 1962c: 614 A: FUJ GUA GUX ZHE
olivieri Laporte, 1840: 131 (Melolontha) E: AB AR GG GR ST TR A: IN IS TM TR SY
abchasica Motschulsky, 1845a: 57
caucasica Heyden, 1864: 319 [HN]

ploceki Tesař, 1944b: 339 A: ZHE
persica Brenske, 1902b: 375 A: IN TR
ragusae aliquoi Massa & Tassi, 1977: 271 E: IT (Sicilia)
ragusae ragusae Kraatz, 1882e: 82 [=1882b: 239] E: IT (Sicilia)
tonkinensis Dewailly, 1945a: 47 A: GUI ORR
turkmenoglui Petrovitz, 1965b: 193 A: TR
subgenus Xerasiobia S.I. Medvedev, 1951a: 100 type species Scarabaeus albus Pallas, 1773
adspersa Motschulsky, 1854b: 28 E: AB AR GG ST A: AF IN KI KZ ST TD TM TR UZ
araxidis König, 1901: 9
atrana Reitter, 1902b: 274
pulverea Ballion, 1871a: 342
turcestana Brenske, 1890c: 126

alba alba Pallas, 1773: 718 (Scarabaeus) E: ST UK A: KI KZ TM UZ
hololeuca Pallas, 1781: 19 (Scarabaeus)

236

SCARABAEIDAE

permagna Reitter, 1901a: 76

alba vicaria Semenov, 1900c: 307 A: GAN MG NMO XIN
sicardi Bedel, 1917: 363 N: MO
genus Pseudotocama Keith, 2008d: 339 type species Pseudotocama minima Keith, 2008
minima Keith, 2008d: 340 A: JIX
genus Sphodroxia Kraatz, 1890: 352 type species Melolontha mauritanica P.H. Lucas, 1846
algirica Ley, 1914: 55 N: AG
atripennis Pic, 1918c: 10 N: AG
maroccana Ley, 1923: 128 N: MO
mauritanica P.H. Lucas, 1846a: pl. 24 (Melolontha) N: AG
lucasi Coquerel, 1860: 423 (Anoxia)
septemfoliata Desbrochers des Loges, 1872: 356

quedenfeldti Brenske, 1890e: 331 (Polyphylla) N: MO
genus Tocama Reitter, 1902b: 265 type species Melolontha rubiginosa Fairmaire, 1889
formosana C.K. Yu, Kobayashi & Chu, 1998: 27 (Melolontha) A: TAI
hebeica Keith & Sabatinelli, 2012: 269 A: HEB HUB JIX
laevipennis C.É. Blanchard, 1851: 158 (Hoplosternus) A: FUJ GUA GUX HAI ORR
squamulata Frey, 1969b: 114 (Hoplosternus)

rubiginosa Fairmaire, 1889a: 21 (Melolontha) A: FUJ GUA HEB HUN JIA JIX ZHE
albidiventris Fairmaire, 1889a: 21 (Melolontha)

similis C.-L. Li & C.-C. Wang, 2015a: 105 A: YUN
tonkinensis Moser, 1913c: 290 (Hoplosternus) A: GUI GUX HUN SHN YUN ORR
pygidialis Moser, 1915d: 589 (Hoplosternus)
atra Keith, 2006f: 225
reichenbachi Keith, 2007f: 338

varimaculata C.-L. Li & P.-S. Yang, 2015a: 103 A: YUN
zhangia C.-C. Wang & C.-L. Li, 2015a: 104 A: YUN
genus Tricholontha Nomura, 1952a: 26 type species Tricholontha papagena Nomura, 1952
papagena Nomura, 1952a: 26 A: JA (Okinawa-shotō)

tribe Pachydemini Burmeister, 1855

genus Alaia Petrovitz, 1980: 631 type species Alaia sexdentata Petrovitz, 1980
sexdentata Petrovitz, 1980: 631 A: IN
genus Asiactenius Nikolajev, 2000: 137 type species Asiactenius afghanus Nikolajev, 2000
afghanus Nikolajev, 2000: 138 A: AF
mirabilis Nikolajev, 2000: 139 A: AF

SCARABAEIDAE

237

reitteri S.I. Medvedev, 1952: 78 (Hemictenius) A: KZ
tekkensis Reitter, 1889b: 105 (Pachydema) A: AF IN TM UZ
tokgajevi S.I. Medvedev, 1962: 603 (Hemictenius) A: IN TM
genus Atanyproctus Petrovitz, 1954: 272 type species Atanyproctus simplicitarsis Petrovitz, 1954
afghanus Petrovitz, 1968f: 244 A: AF
alexandri Petrovitz, 1965d: 687 A: AF
ernae Petrovitz, 1980: 628 A: IN
miksici Petrovitz, 1965d: 688 A: AF
opacipennis Petrovitz, 1968f: 245 A: IN
simplicitarsis Petrovitz, 1954: 273 A: IN
genus Brenskiella Berg, 1898: 17 [RN] type species Pachychaeta flavomicans Brenske, 1897
Pachychaeta Brenske, 1897d: 362 [HN] type species Pachychaeta flavomicans Brenske, 1897

flavomicans Brenske, 1897d: 362 (Pachychaeta) A: IS
genus Buettikeria Sabatinelli & Pontuale, 1998b: 140 type species Buettikeria graingeri Sabatinelli &
Pontuale, 1998
echinocephala Nikolajev, 2003b: 117 A: AE
graingeri Sabatinelli & Pontuale, 1998b: 141 A: AE SA
genus Canudema M. Lacroix, 1994: 154 type species Canudema socotrae M. Lacroix, 1994
homhil Král, Sehnal & Bezděk, 2012: 155 A: YE (Suqutra)
socotrae M. Lacroix, 1994: 155 A: YE (Suqutra)
genus Ceramida Baraud, 1987a: 126 type species Melolontha longitarsis Illiger, 1803
adusta Kraatz, 1882a: 20 (Elaphocera) E: SP
abderramani Escalera, 1923: 403 (Elaphocera)
certa Báguena, 1955a: 135 (Elaphocera)

bedeaui Erichson, 1840: 37 (Elaphocera) E: PT SP
brancoi Baraud, 1975b: 61 (Elaphocera) E: PT
brandeiroi Flach, 1906: 119 (Elaphocera) E: PT
championi Reitter, 1902b: 97 (Elaphocera) E: SP
onubensis Escalera, 1923: 400 (Elaphocera)
stenostoma Báguena, 1955a: 136 (Elaphocera)

cobosi Báguena, 1955a: 139 (Elaphocera) E: SP
dinizi Branco, 1981: 139 (Elaphocera) E: PT
isabellae López-Colón, 1993b: 313 E: SP (Huelva)
jiennensis Sánchez García, Pérez Gordillo, Cabrera Fernández, Romo Bedate, Cortázar Hurtado, Pla Benítez
& López-Colón, 2010: 31 E: SP (Andalucía)
longitarsis Illiger, 1803: 218 (Melolontha) E: PT

238

SCARABAEIDAE

baraudi Branco, 1981: 134 (Elaphocera)
transtagana Branco, 1981: 132 (Elaphocera)

luisae López-Colón & Rodríguez Arias, 1990: 291 E: SP
luna Sánchez García, Pérez Gordillo, Cabrera Fernández, Romo Bedate, Cortázar Hurtado, Pla Benítez &
López-Colón, 2010: 28 E: SP (Andalucía)
lusitanica Baraud, 1975b: 62 (Elaphocera) E: PT
malacensis Rambur, 1843: 343 (Elaphocera) E: SP
luctuosa Báguena, 1955a: 133 (Elaphocera)
mancinii Báguena, 1955a: 134 (Elaphocera)

mauritanica Rambur, 1843: 341 (Elaphocera) N: AG MO
bolivari Escalera, 1914a: 161 (Elaphocera)
numidica Rambur, 1843: 343 (Elaphocera)

moelleri Flach, 1906: 120 (Elaphocera) E: PT SP
gigas Serrano, 1985: 361 (Elaphocera)

tangeriana Kraatz, 1882a: 20 (Elaphocera) N: MO
zuzartei Branco, 1981: 138 (Elaphocera) E: PT
genus Ceramida, nomen dubium
ohausi Reitter, 1902b: 98 (Elaphocera) E: GR
genus Elaphocera Gené, 1836: 189 type species Elaphocera obscura Gené, 1836 (=Melolontha emarginata
Gyllenhal, 1817)
Elaphocerida Reitter, 1902b: 98 type species Melolontha emarginata Gyllenhal, 1817

affinis Schaufuss, 1874: 312 E: SP
alonsoi alonsoi López-Colón, 1992a: 158 E: SP (Valencia)
alonsoi semperei López-Colón, 1992a: 158 E: SP (Alicante)
ampla Báguena, 1955a: 147 E: SP
angusta Kraatz, 1882a: 23 E: SP
aristidis Raffray, 1871: 160 N: AG
autumnalis Motschulsky, 1860d: 139 E: SP
laufferi Flach, 1907: 17
pygmaea Escalera, 1923: 405

baguenae Mancini, 1926: 96 E: SP
torressalai Báguena, 1955a: 143

barbara antoniae Kocher, 1960: 235 N: MO
barbara barbara Rambur, 1843: 350 N: AG
funebris Fairmaire, 1879e: 249

barbara pardoi Escalera, 1931b: 715 N: MO
cacerensis López-Colón & Rodríguez Arias, 1986: 99 (Elaphocerida) E: SP
capdeboni Schaufuss, 1882a: 533 E: SP (Mallorca)
carteiensis Rambur, 1843: 356 E: SP

SCARABAEIDAE
granatensis Rambur, 1843: 349

christina Baraud, 1966b: 31 E: SP
churianensis Rambur, 1843: 355 E: SP
cretica Reitter, 1902b: 102 E: GR (Kríti)
dalmatina Motschulsky, 1860d: 141 E: CR
denticornis Reitter, 1902b: 102 E: MC
dilatata Erichson, 1840: 39 E: IT (Sardegna)
elongata Schaufuss, 1874: 311 E: GR MC
graeca Kraatz, 1882a: 27

emarginata Gyllenhal, 1817: 70 (Melolontha) E: IT (Sardegna)
sardoa Rambur, 1843: 352

erberi Kraatz, 1882a: 29 E: GR (Tinos)
ferreri López-Colón & Plaza Lama, 1989: 193 (Elaphocerida) E: SP
gibbifrons Reitter, 1902b: 105 E: GR (Attiki)
gracilis Waltl, 1838: 457 (Leptopus) E: BU GR MC TR
bysantica Rambur, 1843: 354

heydeni Kraatz, 1882a: 21 E: SP
ardoini Baraud, 1966b: 34
cobosi Baraud, 1966b: 33 [HN]
bicolorea Baraud, 1992: 561 [RN]
margaritae Blanco Villero, 1988: 283

hiemalis Erichson, 1840: 39 E: MC
raymondi Kraatz, 1882a: 28

hirticollis Kraatz, 1882a: 23 E: SP
hispalensis Rambur, 1843: 353 E: SP
baetica Sáez Bolaño, 1993: 65 (Elaphocerida)

ibicensis Escalera, 1926a: 160 E: SP (Baleares: Ibiza, Formentera, Cunillera)
insularis Fairmaire, 1877c: lxvi E: GR (Kríti)
maltzani Heyden, 1883b: 368

kosensis Keith & Trichas, 2007: 51 E: GR (Kós)
lajonquierei Baraud, 1966b: 29 E: SP
leoi Fancello & Cillo, 2014: 87 E: IT (Sardegna)
longicornis Serrano, 1985: 364 E: PT
martini López-Colón, 1993a: 280 E: SP
martorellii Fairmaire, 1879d: 241 E: SP
rubicunda Reitter, 1902b: 100

nigroflabellata Burmeister, 1855: 456 E: GR TR
nupcialis Ruiz & Ávila, 1995b: 244 E: SP
obscura erichsoni Kraatz, 1882a: 26 E: IT (Sardegna)
obscura obscura Gené, 1836: 189 E: IT (Sardegna)
ochsi Baraud, 1987a: 130 E: GR

239

240

SCARABAEIDAE

perezlopezi Ruiz, 1997: 360 E: SP
phungae Baraud, 1987a: 132 E: GR (Paros)
roessneri Keith, 2005d: 242 E: GR (Pelopónnisos)
schmidti Keith, 2005d: 243 E: GR (Chalkidiki)
segurensis Escalera, 1923: 408 E: SP
staudingeri Reitter, 1902b: 103 A: SY
sulcatula Fairmaire, 1884e: cxvii N: MO
abdelkrimi Escalera, 1934b: 129 (Elaphocerida)

suturalis Schaufuss, 1874: 312 E: GR (Ródos) A: SY TR
confusa Kraatz, 1882a: 30
nigrata Fairmaire, 1884b: 178
rhodana Marseul, 1878: 59

syriaca Kraatz, 1882a: 29 A: SY
tethys Reitter, 1902b: 101 E: SP
genus Europtron Marseul, 1867c: lxxxi type species Europtron gracile Marseul, 1867
atlanticum Bezděk, Král, Keith & M. Lacroix, 1999: 399 N: MO
confusum Marseul, 1878: 60 N: AG
ibericum Balthasar, 1929a: 41

gracile Marseul, 1867c: lxxxi N: AG
bayonnei Chobaut, 1896b: 413

genus Falsotoclinius Keith & Montreuil, 2004: 217 type species Otoclinius richteri Petrovitz, 1958
fragilis Petrovitz, 1980: 629 (Otoclinius) A: IN
nikodymi Keith, 2007d: 507 A: PA
richteri Petrovitz, 1958a: 6 (Otoclinius) A: IN
schaeuffelei Petrovitz, 1958a: 5 (Otoclinius) A: IN
sehnali Keith, 2007d: 505 A: PA
genus Hemictenius Reitter, 1897j: 223 type species Tanyproctus opacus Ballion, 1871
apterus Nikolajev, 1976a: 66 A: TD
bactrianus S.I. Medvedev, 1952: 78 A: TD UZ
badkhyzicus Nikolajev, 2000: 146 A: TM
comatus Nikolajev, 1975a: 148 A: TD
danilevskyi Gusakov, 2008: 3 A: TD
drescheri Reitter, 1906b: 34 A: TD
elongatus S.I. Medvedev, 1962: 606 A: TD
gracilipes Semenov, 1891: 317 (Pachydema) A: TM UZ
kabakovi Gusakov, 1998: 568 A: TD
klimenkoi Gusakov, 2004: 18 A: UZ
kryzhanovskii Nikolajev & Schukronajev, 1977: 157 A: TD

SCARABAEIDAE

241

latitarsis Reitter, 1891b: 24 (Tanyproctus) A: KZ UZ
lebedevi Reitter, 1908d: 216 A: KZ UZ
lopatini S.I. Medvedev, 1959: 874 A: TD
magnitarsis Reitter, 1897j: 223 A: AF
medvedevi Nikolajev, 1976b: 166 A: TM
merkli Gusakov, 2004: 23 A: TD
nigrociliatus Reitter, 1897j: 224 A: AF TD
nikolajevi Gusakov, 2004: 21 A: TD
ochripennis Reitter, 1902b: 136 A: TD
opacipes Reitter, 1902b: 135 A: TD
opacus Ballion, 1871a: 339 (Tanyproctus) A: TD TM UZ
albociliatus Reitter, 1902b: 134
nitidipes Reitter, 1902b: 135
scutellaris Reitter, 1902b: 136

petrovitzi Gusakov, 2003c: 69 A: AF
pygmaeus S.I. Medvedev, 1962: 604 A: TD
simplicitarsis Reitter, 1897j: 225 A: AF TD UZ
skomorokhovi Gusakov, 2004: 17 A: TM
smirnovi Gusakov, 2004: 13 A: TD
stackelbergi S.I. Medvedev, 1952: 81 A: TD
subpilosus S.I. Medvedev, 1959: 872 A: UZ
tarsalis S.I. Medvedev, 1962: 601 A: TD
talukanus Ivanova, 2007: 32 A: AF
tshistjakovae Gusakov, 1998: 569 A: TD
walteri Reitter, 1889b: 106 (Pachydema) A: TM
yuferevi Gusakov, 2004: 20 A: TM
zhantievi Gusakov, 2004: 15 A: TM
genus Jalalabadia Balthasar, 1967a: 111 type species Jalalabadia povolnyi Balthasar, 1967
povolnyi Balthasar, 1967a: 112 A: AF
genus Kryzhanovskia Nikolajev & Kabakov, 1977a: 85 type species Kryzhanovskia olegi Nikolajev &
Kabakov, 1977
cashmirensis Keith, 2006c: 425 A: KA
olegi Nikolajev & Kabakov, 1977a: 85 A: AF
genus Leptochristina Baraud & Branco, 1991: 176 type species Pachydemocera pubimargo Reitter, 1902
annamariae Baraud & Branco, 1991: 176 A: IQ
pubicavula Reitter, 1902b: 107 (Pachydemocera) A: IS
pubimargo Reitter, 1902b: 106 (Pachydemocera) A: SY TR
variabilis Petrovitz, 1962b: 123 (Pachydemocera) A: IQ

242

SCARABAEIDAE

genus Otoclinius Brenske, 1897c: 320 type species Otoclinius gracilipes Brenske, 1897
gracilipes Brenske, 1897c: 320 A: “Beluchistan”
genus Pachydema Laporte, 1832a: IX, pl. 37 type species Pachydema nigricans Laporte, 1832 (=Melolontha
hirticollis Fabricius, 1787)
Artia Rambur, 1843: 332 type species Artia cartaginensis Rambur, 1843
Dasysterna Rambur, 1843: 331 type species Dasysterna barbara Rambur, 1843
Flatipalpus Fairmaire, 1873b: 391 type species Flatipalpus albolanosus Fairmaire, 1873
Ootoma C.É. Blanchard, 1850: 120 type species Melolontha castanea Brullé, 1838
Oromia López-Colón, 1989c: 300 [HN] type species Melolontha obscura Brullé, 1838
Oromiella López-Colón, 1992c: 245 [RN] type species Melolontha obscura Brullé, 1838
Phlexis Erichson, 1841a: 172 type species Phlexis wagneri Erichson, 1841
Phygotoxeuma Brenske, 1897b: 118 type species Pachydema theryi Brenske, 1897
Physopalpus Heyden, 1900: 254 type species Pachydema rufina Fairmaire, 1879
Sparophysa Burmeister, 1855: 442 type species Pachydema bullata Burmeister, 1855

abeillei Fairmaire, 1881a: 84 A: SY
adusta Karsch, 1881: 46 N: EG
albipilis Reitter, 1902b: 118 N: EG
albolanosa Fairmaire, 1873b: 391 (Flatipalpus) N: AG TU
ameliae López-Colón, 1986a: 89 N: CI (Gran Canaria)
keithi M. Lacroix, 2000: 2

amphicomella Peyerimhoff, 1943: 13 N: MO
angustipalpis Normand, 1951: 160 N: TU
anthracina Fairmaire, 1860c: 420 N: MO
decipiens Fairmaire, 1867: 396
maroccana Brenske, 1889: 333
nigropilosa M. Antoine, 1951: 313

antoinei Baraud, 1979e: 288 N: MO
ariasi Escalera, 1914a: 167 N: MO
autumnalis Normand, 1915: 254 N: TU
bertiae Baraud, 1982b: 235 N: TU
bipartita Brullé, 1839: 60 (Melolontha) N: CI
africana Brenske, 1897b: 117

buettikeri Sabatinelli & Pontuale, 1998b: 132 A: OM
bullata Burmeister, 1855: 442 N: EG
caesariata Normand, 1925: 314 N: TU
cambeforti Chavanon & Zirari, 1998: 329 N: MO
cartereaui Fairmaire, 1868a: 479 N: AG
monochroa Reitter, 1902b: 116

cartaginensis Rambur, 1843: 332 (Artia) N: TU
castanea Brullé, 1839: 60 (Melolontha) N: CI

SCARABAEIDAE
canariensis Rambur, 1843: 331 (Dasysterna)
macaronesica López-Colón, 1986a: 94

cinctipennis Peyerimhoff, 1949b: 286 N: MO
concinna Burmeister, 1855: 446 A: TR
confinalis Peyerimhoff, 1949b: 286 N: MO
conica Reitter, 1902b: 110 N: EG
curvipedes Escalera, 1914a: 165 N: MO
daoudii Chavanon & Labrique, 2013: 180 N: MO
decorosa Normand, 1936: 380 N: TU
demoflysi Normand, 1938b: 367 N: TU
doriae Fairmaire, 1875a: 508 N: LB TU
doumeti Mayet, 1887b: xliv N: TU
doursii P.H. Lucas, 1859c: xxxi N: AG
dubitalis Reitter, 1902b: 111 N: TU
emflusi Escalera, 1914a: 165 N: MO
eremicola Peyerimhoff, 1928b: 305 N: AG
fortunatorum Baraud, 1985b: 132 N: CI (Tenerife)
ferreri López-Colón, 1986a: 86

foveiceps Marseul, 1878: 60 N: AG
foveola P.H. Lucas, 1859a: 455 N: AG
fuscipennis Brullé, 1839: 61 (Melolontha) N: CI
getula Baraud, 1979e: 261 N: AG
girardi Baraud, 1982b: 236 N: MO
gomerae López-Colón, 1999: 1 N: CI (La Gomera)
gourvesi Baraud, 1985a: 625 N: MO
grandipalpis Reitter, 1902b: 108 N: AG EG LB TU
inflatipalpis Reitter, 1902b: 108

hingrati Chavanon & François, 2015: 2 N: MO
hirticollis Fabricius, 1787a: 21 (Melolontha) E: IT (Sicilia, Lampedusa) N: AG LB TU
barbara Rambur, 1843: 331 (Dasysterna)
nigricans Laporte, 1832b: IX, pl. 37
quedenfeldti Brenske, 1889: 332
spreta Fairmaire, 1860c: 419

hontoriai Escalera, 1914a: 162 N: MO
hornbeckii P.H. Lucas, 1859c: xxx N: AG MO
distinguenda Fairmaire, 1866a: 22

icositana Baraud, 1979e: 264 N: AG
immanipalpis Baraud, 1979e: 298 N: AG
immatura Burmeister, 1855: 443 N: EG LB
lessepsii P.H. Lucas, 1869a: 527

integra Wollaston, 1864: 197 (Ootoma) N: CI (Gran Canaria)

243

244
israelitica Crotch, 1872: 266 A: SI
jeannei Baraud, 1980: 260 N: CI (Tenerife)
lamotteae Miessen, 2000: 367 N: MO
lesnei Peyerimhoff, 1928a: 171 N: CI (Gran Canaria)
lethierryi P.H. Lucas, 1861: 101 N: AG
letourneuxi Pic, 1903b: 113

lucasi Reiche, 1860b: 644 [RN] N: AG
hirticollis P.H. Lucas, 1851: 523 (Dasysterna) [HN]

lucianae Peyerimhoff, 1925a: 24 N: AG
marraquensis Escalera, 1914a: 168 N: MO
marmottani Fairmaire, 1868a: 478 N: AG TU
megalops Micó, 2009: 42 N: CI (Gran Canaria)
menieri Baraud, 1985b: 138 N: CI
pierae López-Colón, 1986a: 92

mogadorica Escalera, 1914a: 164 N: MO
mozabensis Baraud, 1979e: 277 N: AG
nitidicollis Fairmaire, 1876b: 37 N: AG TU
nocturna Crotch, 1872: 266 A: SI
normandi Baraud, 1979e: 265 N: TU
navatteae Baraud, 1985b: 138 N: CI (Gran Canaria)
obscura Brullé, 1839: 61 (Melolontha) N: CI
obscurata Fairmaire, 1883b: 459 N: LB
magna Brenske, 1889: 334

obscurella Wollaston, 1864: 200 (Ootoma) N: CI (El Hierro)
oraniensis P.H. Lucas, 1869a: 525 N: AG
oromii López-Colón, 1986a: 87 N: CI (La Gomera)
otini Peyerimhoff, 1949b: 285 N: MO
palposa Reitter, 1902b: 109 A: SI
peltastes Marseul, 1879a: 99 (Tanyproctus) N: AG
theryi Brenske, 1897b: 118

peyerimhoffi Baraud, 1979e: 288 N: LB
phylloperthoides Reitter, 1902b: 111 N: TU
pilosa Walker, 1871: 12 A: SI
rebellis Normand, 1925: 314 N: TU
renaudi Chavanon & François, 2014: 237 N: MO
rubripennis P.H. Lucas, 1848: xlviii (Elaphocera) N: AG MO
rungsi Peyerimhoff, 1949b: 286 N: MO
sancta Crotch, 1872: 267 A: SI
schrammi fusciceps Peyerimhoff, 1949b: 288 N: MO
schrammi schrammi Peyerimhoff, 1949b: 287 N: MO

SCARABAEIDAE

SCARABAEIDAE

245

sicardi Baraud, 1979e: 265 N: TU
sinaitica Crotch, 1872: 266 A: SI
succidua Normand, 1951: 161 N: TU
suspiciosa Normand, 1951: 162 N: TU
tarsalis Reitter, 1902b: 117 N: AG
thysdritana Baraud, 1979e: 273 N: AG
tinerfensis Galante & Stebnicka, 1992: 230 N: CI (Tenerife)
unicolor P.H. Lucas, 1851: 525 (Dasysterna) N: AG EG TU
valdani P.H. Lucas, 1859c: xxxi N: AG
veneriata Normand, 1936: 379 N: AG A: JO
vestita Normand, 1949: 88 N: TU
volaki Roubal, 1945: 68 A: JO
wagneri Erichson, 1841a: 172 (Phlexis) N: AG
wollastoni Peyerimhoff, 1928a: 173 N: CI (Fuerteventura)
moralesi López-Colón, 1989a: 55

xanthochroa Fairmaire, 1879e: 247 N: TU
rufina Fairmaire, 1879e: 247

zohra Normand, 1951: 157 N: TU
genus Pachydemocera Reitter, 1902b: 105 type species Elaphocera lucidicollis Kraatz, 1882
lucidicollis Kraatz, 1882a: 31 (Elaphocera) E: GR (Ródos) A: IS SY
genus Phalangonyx Reitter, 1889g: 277 type species Phalangonyx coniceps Reitter, 1889
bibatillatus Petrovitz, 1955: 283 (Tanyproctus) A: AF
buettikeri Sabatinelli & Pontuale, 1998b: 137 A: AE SA
coniceps Reitter, 1889g: 277 A: IN TM
hadhramauticus Decelle, 1982: 377 A: YE
hormoziaensis Petrovitz, 1980: 623 (Tanyproctus) A: IN
loebli Baraud, 1991c: 289 (Otoclinius)

irakanus Arrow, 1932: 196 A: IQ
mesopotamicus S.I. Medvedev, 1952: 64 (Tanyproctus) A: IQ
mirzayani Petrovitz, 1968f: 243 (Tanyproctus) A: IN
semenovi S.I. Medvedev, 1952: 63 (Tanyproctus) A: IN IQ
genus Pseudopachydema Balthasar, 1930a: 39 type species Pseudopachydema caucasica Balthasar,
1930
Protoctenius S.I. Medvedev, 1952: 88 type species Protoctenius bogatshevi S.I. Medvedev, 1952

caucasica Balthasar, 1930a: 39 E: AB
bogatshevi S.I. Medvedev, 1952: 88 (Protoctenius)

medvedevi Iablokoff-Khnzorian, 1971: 182 E: AR

246

SCARABAEIDAE

genus Socotraproctus Král, Sehnal & Bezděk, 2012: 160 type species Socotraproctus haghier Král, Sehnal &
Bezděk, 2012
haghier Král, Sehnal & Bezděk, 2012: 161 A: YE (Suqutra)
genus Tanyproctoides Petrovitz, 1971d: 229 type species Tanyproctoides arabicus Petrovitz, 1971 (=Phalangonyx
arabicus Arrow, 1932)
subgenus Odontiellus Keith & Montreuil, 2004: 221 type species Tanyproctoides silfverbergi Keith, 2001
freyi Petrovitz, 1968f: 242 (Tanyproctus) A: IQ
gormicus Montreuil & Keith, 2013: 66 A: IN
mirzayansinus Keith & Montreuil, 2004: 222 A: IN
silfverbergi Keith, 2002c: 143 A: IQ SA
subgenus Tanyproctoides Petrovitz, 1971d: 229 type species Tanyproctoides arabicus Petrovitz, 1971
(=Phalangonyx arabicus Arrow, 1932)
arabicus Arrow, 1932: 196 (Phalangonyx) A: KU QA SA
arabicus Petrovitz, 1971d: 229 [HN]

genus Tanyproctus Ménétriés, 1832: 185 type species Tanyproctus persicus Ménétriés, 1832
subgenus Brachydema Fairmaire, 1881a: 86 type species Brachydema lamberti Fairmaire, 1884
adanensis Reitter, 1908c: 129 A: TR
aphodioides Fairmaire, 1866b: 257 (Pachydema) A: TR
beskindensis Reitter, 1902b: 124 A: LE
carcelii Fairmaire, 1884b: 178 (Brachydema) A: SY
demaisoni Reitter, 1902b: 122 A: TR
kraatzi Reitter, 1902b: 124 A: SY
dichrous Reitter, 1902b: 124

lamberti Fairmaire, 1884b: 177 (Brachydema) A: LE
latimanus Reitter, 1902b: 125 A: IQ SY TR
malatiensis Reitter, 1902b: 125

longipes Burmeister, 1855: 445 (Pachydema) A: SY
novicius Reitter, 1902b: 124 A: LE
pilimargo Reitter, 1902b: 126 A: LE TR
pygidialis Reitter, 1902b: 126 A: LE
riparius Petrovitz, 1963c: 264 A: TR
rugosulus Fairmaire, 1892a: 148 A: LE
sinuatifrons Fairmaire, 1881a: 85 (Pachydema) A: LE
unicolor Motschulsky, 1860d: 134 A: TR
verryi Fairmaire, 1884b: 177 (Brachydema) A: LE
acrossoides Reitter, 1902b: 126

yunnanicus Keith, 2006c: 429 A: YUN
subgenus Tanyproctocera Reitter, 1902b: 119 type species Pachydema festiva Burmeister, 1855
crinitus Petrovitz, 1971d: 226 A: IS

SCARABAEIDAE
delesserti Reiche & Saulcy, 1856: 376 (Pachydema) A: SY
festivus Burmeister, 1855: 445 (Pachydema) A: LE SY
coyei Marseul, 1876: 379 (Pachydema)
brenskei Reitter, 1902b: 120

israeliticus Petrovitz, 1971d: 227 A: IS
kriecheldorffi Reitter, 1909b: 80 A: TR
lanatus Chevrolat, 1873: 203 (Pachydema) A: SY
ledereri Reiche, 1861a: 3 (Pachydema) A: IS LE SY
rufus Reitter, 1902b: 120

minutus Petrovitz, 1973b: 30 A: IS
nabataeus Petrovitz, 1973b: 29 A: IS
portusus Reitter, 1902b: 120 A: TR
pumilus Petrovitz, 1973b: 32 A: IS
saulcyi Reiche & Saulcy, 1856: 378 (Pachydema) A: IQ IS JO SY
sinaiticus Heyden, 1900: 253 (Pachydema) A: IS SI
syriacus S.I. Medvedev, 1952: 39 A: SY
subgenus Tanyproctus Ménétriés, 1832: 185 type species Tanyproctus persicus Ménétriés, 1832
araxis Reitter, 1902b: 132 E: AB AR
arher Bezděk, Sehnal & Král, 2013: 192 A: YE (Suqutra)
armeniacus S.I. Medvedev, 1952: 58 A: TR
batangicus Keith, 2009c: 82 A: SCH
beludschistanus Petrovitz, 1968f: 241 A: IN
bicuspidatus Peyerimhoff, 1926b: 219 (Pachydema) N: MO
brevipennis Motschulsky, 1860d: 137 E: GG
bucharicus Reitter, 1897j: 221 (Pachydema) A: AF TD UZ
subciliatus Reitter, 1902b: 128
suturiferus Reitter, 1897j: 222 (Pachydema)

canui M. Lacroix, 1999c: 92 A: YE (Suqutra)
carbonarius Faldermann, 1835a: 273 E: AB AR A: IN
cariensis Petrovitz, 1971d: 225 A: TR
confinis Motschulsky, 1860d: 138 E: AB AR A: IN
ordubadensis Reitter, 1902b: 132

davidis Fairmaire, 1886: 324 A: YUN
dechambrei M. Lacroix, 1996: 93 A: SCH
eghtedari Petrovitz, 1980: 621 A: IN
eversmanni Reitter, 1902b: 131 E: “Ural”
farsensis Keith, 2009c: 85 A: IN
fastus Petrovitz, 1968g: 476 A: TR
francottei Keith & M. Lacroix, 1999: 473 A: YUN
nocturnus Král, 1999b: 478

ganglbaueri Brenske, 1897b: 115 (Pachydema) A: IN

247

248
holzschuhi Petrovitz, 1973b: 27 A: TR
indescriptus Baraud, 1979e: 303 A: SI
inflatus Motschulsky, 1860d: 136 E: AB GG A: IN
iranicus Petrovitz, 1968f: 240 A: IN
jizu Král, 1999b: 477 A: YUN
jordaniacus Baraud, 1990e: 300 A: JO
kabulensis Petrovitz, 1967e: 486 A: AF
keithi Král, Sehnal & Bezděk, 2012: 166 A: YE (Suqutra)
kermanshahensis Petrovitz, 1980: 625 A: IN
kindermanni Reiche, 1861a: 2 (Pachydema) A: IS TR
kurdistanus Reitter, 1902b: 131 A: “Kurdistan: Edgin”
lacroixi Král, Sehnal & Bezděk, 2012: 172 A: YE (Suqutra)
lydiensis Petrovitz, 1971e: 582 A: TR
mourzinei Keith & M. Lacroix, 1999: 474 A: YUN
diurnus Král, 1999b: 474

muchei Petrovitz, 1962b: 121 A: SY
nitidus Petrovitz, 1968f: 241 A: IN
opacipennis Petrovitz, 1968g: 477 A: TR
ortospanaensis Petrovitz, 1965d: 689 A: AF
ovatus Motschulsky, 1860d: 136 E: AB AR GG A: TR
pallidus Petrovitz, 1962b: 122 A: IQ
pamphilus Petrovitz, 1967b: 333 A: TR
parallelus Petrovitz, 1968f: 238 A: IN
parvus Chang & Luo, 1981: 55 A: JIA
paulusi Petrovitz, 1980: 627 A: LE
persicola Reitter, 1909b: 81 A: IN
persicus Ménétriés, 1832: 185 E: AB GG A: IN
brevipennis Motschulsky, 1860d: 137
subcylindricus Motschulsky, 1860d: 138

poggii Keith, 2009c: 84 A: YUN
puncticeps C.O. Waterhouse, 1881b: 471 (Pachydema) A: YE (Suqutra)
reichii Rambur, 1843: 332 (Dasysterna) E: GR A: TR
graecus Motschulsky, 1860d: 139 (Dasysterna)
olivieri C.É. Blanchard, 1851: 163

rubicundus Reitter, 1902b: 129 A: AR
rufidens Marseul, 1879b: 474 (Elaphocera) E: AB AR GG A: IN
samai Piattella & Sabatinelli, 1996: 363 E: GR (Aykathonisi)
sanxiaensis Chang & Li, 1997: 770 A: CHQ SCH
satanas Reitter, 1902b: 130 E: AB AR A: IN
sichuanicus Keith, 2006c: 426 A: SCH
similis Petrovitz, 1962b: 123 A: SY

SCARABAEIDAE

249

SCARABAEIDAE

speculator Petrovitz, 1963c: 264 A: TR
tuniseus Baraud, 1979e: 303 N: TU
turanicus Reitter, 1900b: 52 A: AF TM
varians Petrovitz, 1980: 622 A: IN
vedicus Kalashian, 1999: 80 A: AR
waziristanicus Keith, 2009c: 79 A: PA
wraniki Král, Sehnal & Bezděk, 2012: 178 A: YE (Suqutra)
xisangensis Chang, 1987: 73 A: XIZ
yunnanus Keith & M. Lacroix, 2003: 38 A: YUN
zartoshti Petrovitz, 1980: 626 A: IN
subgenus Tetraproctus Iablokoff-Khnzorian, 1953: 149 type species Tanyproctus antennatus IablokoffKhnzorian, 1953
antennatus Iablokoff-Khnzorian, 1953: 149 E: AR

tribe Pachypodini Erichson, 1840

genus Pachypus Dejean, 1821: 57 type species Scarabaeus excavatus Fabricius, 1792 (=Scarabaeus candidae V.
Petagna, 1787)
Caelodera Dejean, 1833: 159 type species Scarabaeus excavatus Fabricius, 1792
Pachypus Audinet-Serville, 1825: 366 [HN] type species Melolontha candidae Petagna, 1787

caesus Erichson, 1840: 35 E: IT (Sicilia)
intermedius Ragusa, 1893a: 205
siculus Laporte, 1840: 129

candidae V. Petagna, 1787: 3 (Melolontha) E: FR (Corse) IT
cornutus A.G. Olivier, 1789: no 5: 20 (Melolontha)
erichsoni Reitter, 1899a: 41
excavatus Fabricius, 1792a: 31 (Scarabaeus)
impressus Erichson, 1840: 33

demoflysi Normand, 1936: 376 N: AG TU
melonii Sparacio, 2008: 5 E: IT (Sardegna)
sardiniensis Guerlach, Bazzato & Cillo, 2013: 74 E: IT (Sardegna)

tribe Rhizotrogini Burmeister, 1855

genus Aethiaratrogus Decelle, 1982: 384 type species Rhizotrogus clypeatus C.É. Blanchard, 1850
marqueti Montreuil, 2003: 486 A: YE
yemenicus Decelle, 1982: 385 A: YE
genus Amadotrogus Reitter, 1902b: 248 type species Rhizotrogus quercanus Burmeister, 1855
grassii Mainardi, 1902: 105 (Rhizotrogus) E: IT
insubricus Burmeister, 1855: 383 (Rhizotrogus) E: FR IT SZ
fiorii Brenske, 1893c: 190 (Rhizotrogus)

oertzeni Brenske, 1886: 201 (Rhizotrogus) E: GR

250

SCARABAEIDAE

patruelis Reiche, 1862b: 541 (Rhizotrogus) E: SP
bolivari Martínez y Sáez, 1873: 64 (Rhizotrogus)
catalaunicus Báguena, 1956b: 30 (Amphimallon)
lajonquieri Baraud, 1970b: 483 (Rhizotrogus)

quercanus Burmeister, 1855: 395 (Rhizotrogus) E: FR IT
sassariensis Perris, 1869: 14 E: IT
tarsalis Reiche, 1862b: 542 (Rhizotrogus) E: IT
truncatus anatolicus Keith, 2006a: 134 A: TR
truncatus truncatus Brenske, 1886: 202 (Rhizotrogus) A: LE
vicinus Mulsant, 1842: 438 (Rhizotrogus) E: FR IT SP
pallipennis Mulsant, 1842: 438 (Rhizotrogus)
rugifrons Burmeister, 1855: 384 (Rhizotrogus)

genus Amphimallon Latreille, 1825: 371 type species Scarabaeus solstitialis Linnaeus, 1758
Amphimallina Reitter, 1905b: 203 type species Amphimallina jenrichi Reitter, 1905 (=Melolontha volgensis Fischer von
Waldheim, 1823)
Amphimallum Agassiz, 1846: 52 type species Scarabaeus solsticialis Linnaeus, 1758
Amphimallus Mulsant, 1842: 440 type species Scarabaeus solsticialis Linnaeus, 1758
Erytrotrogus Iablokoff-Khnzorian, 1955: 28 type species Amphimallon medvedevi Iablokoff-Khnzorian, 1955
(=Rhizotrogus vernalis, Brullé, 1832)
Microdonta Hope, 1837: 105 type species Melolontha pini Fabricius, 1792 (=Melolontha pini A.G. Olivier, 1789)
Zantheumia Stephens, 1829b: 115 type species Scarabaeus solstitialis Linnaeus, 1758

adanense Montreuil, 2000: 263 A: TR
alatavicum S.I. Medvedev, 1951a: 459 A: KZ
altaicum Mannerheim, 1825: 29 (Melolontha) E: BU CT GG GR RO ST UK A: ES KZ WS
suwortzewi Semenov, 1891: 319 (Monotropus)

altifrons Baraud, 1971b: 38 (Rhizotrogus) [DA] N: MO
amphibolum Peyerimhoff, 1949b: 283 (Rhizotrogus) [DA] N: MO
arianae Fairmaire, 1879a: 4 (Rhizotrogus) [=1879e: 253] E: GR (Kríti, Náxos)
arnoldii S.I. Medvedev, 1951a: 496 (Amphimallina) E: AB
assimile Herbst, 1790: 83 (Melolontha) E: AU BH BU BY CR CZ FR GE HU IT KO MC ME PL RO SB SK SL SP SZ
UK
aprilinum Duftschmid, 1805: 90 (Melolontha)
fulvicolle Erichson, 1847a: 689
neapolitanum Reitter, 1902b: 247 (Rhizotrogus)
obscurum Brenske, 1890a: 13

atrum atrum Herbst, 1790: 84 (Melolontha) E: AU BE FR GE GR IT LU PT SB SL SP SZ
fuscum A.G. Olivier, 1789: no 5: 19 (Melolontha)

atrum nomadicum Reiche, 1862b: 543 (Rhizotrogus) E: SP
besnardi Keith, 2009a: 353 N: TU
brucki Fairmaire, 1879e: 253 E: GR

SCARABAEIDAE
burmeisteri Brenske, 1886: 207 E: AU BU CR FR GE HU IT SB SL SZ
cantabricum Heyden, 1870a: 116 E: PT SP
felicitanum Reitter, 1902b: 245
trichroum Reitter, 1907f: 486

circassicum Brenske, 1894a: 59 E: GG
circumligatum Peyerimhoff, 1949a: 26 (Rhizotrogus) N: MO
crinitum Brenske, 1894a: 60 A: “Turkestan”
dalmatinum Brenske, 1894a: 58 E: AL BH CR GR ME SL
evorense Reitter, 1913e: 656 E: PT
fallenii Gyllenhal, 1817: 85 (Melolontha) A: BE DE GB GE NL SV
fissiceps Fairmaire, 1860c: 442 (Rhizotrogus) N: MO
obtusilobum Fairmaire, 1879e: 251 (Rhizotrogus)

furvum furvum Germar, 1817: 215 (Melolontha) E: CR
furvum graecum Keith, 2010b: 113 E: GR
fuscum Scopoli, 1786: 53 (Scarabaeus) E: CR IT SL SZ
logesi Mulsant, 1872: 214
nebrodense Ragusa, 1881: 64 (Rhizotrogus)
sirentense Leoni, 1906: 63 (Rhizotrogus)

gianfranceschii Luigioni, 1931: 509 E: IT (Puglia)
helenae Iablokoff-Khnzorian, 1983: 46 (Amphimallina) [DA] E: AR
insculptum Brenske, 1889: 328 (Rhizotrogus) N: MO
javeti Stierlin, 1864a: 147 (Rhizotrogus) E: IT
jeannae Montreuil, 2000: 266 N: MO
jeannei Baraud, 1971e: 75 (Monotropus) E: GR
jedlickai Balthasar, 1936b: 94 A: SY
julieni Baraud, 1972: 160 (Rhizotrogus) [DA] N: MO
kalliesi Montreuil, 2013: 138 A: TR
keithi Montreuil, 2002: 24 A: TR
krali Montreuil, 2002: 25 E: GR (Ródos)
leuthneri Reitter, 1902b: 238 A: IN SY TR
litigiosum Fairmaire, 1860c: 441 (Rhizotrogus) N: MO
galleti Baraud, 1970b: 487 (Rhizotrogus)
semivillosum Fairmaire, 1883d: cix (Rhizotrogus)

lusitanicum Gyllenhal, 1817: 89 (Melolontha) E: PT SP
angulicolle Fairmaire, 1859: clii (Monotropus)

maevae Montreuil, 1999: 108 E: GR
majale bonadonai R. Paulian, 1959a: 215 (Rhizotrogus) E: FR (Alpes du Sud)
majale majale Razoumowsky, 1789: 249 (Melolontha) E: BE FR GE LU PT SL SP SZ NARi
korbi Reitter, 1894g: 302
pellitulum Reitter, 1902b: 248
rufescens Latreille, 1804b: 188 (Melolontha)

251

252

SCARABAEIDAE

semirufum Gyllenhal, 1817: 91 (Melolontha)

maniense Montreuil, 2000: 266 E: GR
menori Báguena, 1955b: 291 E: PT SP
melillanum Baraud, 1972: 158 (Rhizotrogus) [DA] E: MO
montanum Strassen, 1954: 279 E: IT (Sardegna)
mussardi M. Antoine, 1960: 213 (Rhizotrogus) N: MO
naceyroi Mulsant, 1859: 189 [=1860: 254] E: SP
nigripenne Reitter, 1902b: 238 E: TR A: TR
nigrum ebeninum Baraud, 1973b: 44 (Rhizotrogus) E: SP
nigrum nigrum Waltl, 1835: 68 (Melolontha) E: PT SP
flavicorne C.É. Blanchard, 1851: 144 (Rhizotrogus)

obscurum Reiche, 1864: 240 (Rhizotrogus) N: AG
albipile Reitter, 1902b: 245
brunneum Fairmaire, 1871a: 378 (Rhizotrogus)

occidentale Petrovitz, 1964d: 127 E: PT
ochraceum Knoch, 1801: 90 (Melolontha) E: AB AL AR AU CR FR GR IT MC PL PT SB SK SP ST SZ A: CY KZ TR
irtishense Nikolajev, 1979d: 173 (Rhizotrogus)
matutinale Nonveiller, 1963: 171
parumsetosum S.I. Medvedev, 1951a: 472

pardoi Baraud, 1971a: 23 (Rhizotrogus) N: MO
peropacum Reitter, 1911b: 53 E: PT
peropacum Baraud, 1972: 162 (Rhizotrogus) [HN]

pini A.G. Olivier, 1789: no 5: 18 (Melolontha) E: FR IT SP SZ N: MO
bicolor Mulsant, 1842: 443
ustulatipenne Mulsant, 1842: 443

pseudomajale Sabatinelli, 1977a: 38 E: IT (Calabria, Sicilia)
pygiale Mulsant, 1846: suppl. 9 E: FR PT SP
roris Baraud, 1981d: 273 E: PT SP
ruficorne Fabricius, 1775: 33 (Melolontha) E: BE BY CT CZ FR GE LU NL PL SZ UK
marginatum Herbst, 1783: 14 (Melolontha)
paganum A.G. Olivier, 1789: no 5: 82 (Melolontha)

safiense Montreuil, 2000: 265 N: MO
sainzii Graëlls, 1858: 65 (Rhizotrogus) E: PT SP
scutellare P.H. Lucas, 1847b: 284 (Rhizotrogus) [DA] E: MA N: AG MO TU
cristatifrons Fairmaire, 1883e: cxxiv
lobatum Fairmaire, 1860c: 440 (Rhizotrogus)
marginiceps Fairmaire, 1860c: 440 (Rhizotrogus)
parallelum Fairmaire, 1860c: 440 (Rhizotrogus)
warioni Marseul, 1876: 66 (Rhizotrogus)

seidlitzi Brenske, 1891a: 57 (Rhizotrogus) E: PT SP
trisinuatum Reitter, 1902b: 242

semenovi S.I. Medvedev, 1951a: 475 A: KI

SCARABAEIDAE

253

sogdianum Nikolajev, 2001: 104 A: UZ
solstitiale grossatum Reitter, 1902b: 235 E: ST UK
solstitiale orientale Reitter, 1902b: 235 E: TR A: TR
solstitiale pictum Reitter, 1902b: 235 E: GR
solstitiale setosum Reitter, 1902b: 235 E: AB AR GG ST A: IN SY TR
solstitiale sibiricum Reitter, 1902b: 234 A: ES FE KZ MG WS
solstitiale silvestrii Schatzmayr, 1936: 19 E: GR (Aegean Islands)
solstitiale simplicissimum J. Müller, 1902: 457 (Rhizotrogus) E: AL ME
solstitiale solstitiale Linnaeus, 1758: 351 (Scarabaeus) E: AL AN AU BE BH BU BY CR CT CZ DE EN FI FR GB GE
GR HU IR KO KZ LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP SV SZ UK A: KI KZ NEI QIN
TD UZ XIN XIZ
aurantiacum Mulsant, 1842: 447
autumnale Geoffroy, 1785: 6 (Scarabaeus)
fulvicolle Mulsant, 1842: 447
laterale Mulsant, 1842: 447
mesasiaticum S.I. Medvedev, 1951a: 470
montivagum Halbherr, 1892: 24
pineticola Graëlls, 1858: 63 (Rhizotrogus)
subsulcatum Faldermann, 1835a: 277 (Rhizotrogus)
suturale Mulsant, 1842: 447

solstitiale tropicum Gyllenhal, 1817: 86 (Melolontha) E: FR IT SZ
spartanum Brenske, 1884: 77 (Rhizotrogus) E: GR
subcristatum Fairmaire, 1879e: 253 (Rhizotrogus) [DA] N: AG
subparallelum Escalera, 1913a: 30 (Rhizotrogus) [DA] N: MO
suturale P.H. Lucas, 1859b: xvii (Rhizotrogus) N: AG MO
theryi Peyerimhoff, 1949b: 284 (Rhizotrogus) N: MO
vernale vernale Brullé, 1832: 176 (Rhizotrogus) E: AB AL AR BU GG GR MC RO SB TR A: IN IQ TR
medvedevi Iablokoff-Khnzorian, 1955: 27
schiraziense Reitter, 1902b: 237
torrulosum Waltl, 1838: column 456 (Rhizotrogus)

vernale persicum Petrovitz, 1970c: 40 A: IN
verticale Burmeister, 1855: 389 (Rhizotrogus) E: GR A: TR
vitalei Luigioni, 1932a: 131 E: IT (Sicilia)
vivesi Baraud, 1967a: 39 (Rhizotrogus) E: SP
volgense Fischer von Waldheim, 1823: pl. XXVIII [=1824: 213] (Melolontha) E: AB KZ ST UK A: ES KZ
caspicum Ménétriés, 1832: 184 (Melolontha)
jenrichi Reitter, 1905b: 203 (Amphimallina)

vulpecula Peyerimhoff, 1931a: 6 (Rhizotrogus) N: MO
zorni Montreuil, 2013: 137 A: TR
genus Amphimallon, nomen dubium
limbatipenne A. Villa & G.B. Villa, 1833: 34 (Melolontha) E: IT

254
genus Aplidia Hope, 1837: 101 type species Melolontha transversa Fabricius, 1801
Haplidia Harold, 1869g: 1171 type species Melolontha transversa Fabricius, 1801

aegyptiaca Kraatz, 1882b: 40 (Haplidia) N: EG A: CY
akbesiana Petrovitz, 1971e: 578 (Haplidia) E: SP A: SY TR
andreasmuelleri Keith, 2000b: 564 (Haplidia) A: TR
armeniaca Baraud, 1990c: 160 (Haplidia) E: AR A: TR
attenuata Reiche, 1862b: 543 [DA] E: IT (Sardegna) A: CY
baraudi Sabatinelli, 1991a: 45 (Haplidia) A: JO
baudii Kraatz, 1882b: 38 (Haplidia) E: GR (Ródos) A: CY
besucheti Baraud, 1988b: 571 (Haplidia) A: TR
bodemeyeri Reitter, 1909b: 82 (Haplidia) A: TR
caesariana Reitter, 1902b: 187 (Haplidia) A: IS TR
chaifensis Kraatz, 1882b: 38 (Haplidia) A: IS SY
chloes Keith & Lo Cascio, 2005: 139 (Haplidia) E: GR (Nisyros)
ciliciensis Baraud, 1988b: 565 (Haplidia) A: TR
claudii Baraud, 1988b: 575 (Haplidia) A: TR
cypria Kraatz, 1882b: 41 (Haplidia) A: CY
dilatata Reitter, 1902b: 185 (Haplidia) A: SY TR
endroedii Baraud, 1988b: 576 (Haplidia) A: IS JO LE
etrusca Kraatz, 1882b: 34 (Haplidia) E: IT
fissa Burmeister, 1855: 345 A: IS LE SY
graeca Kraatz, 1882b: 35 (Haplidia) E: GR
heinzorum Baraud, 1990c: 159 (Haplidia) A: TR
hirticollis Burmeister, 1855: 344 E: IT MA
houryae Montreuil, 2005: 130 (Haplidia) A: LE
indaryi Montreuil, 2005: 129 (Haplidia) A: LE
iranica Petrovitz, 1970c: 39 (Haplidia) A: IN IQ TR
janczyki Baraud, 1988b: 561 (Haplidia) A: SY
joannis Baraud, 1988b: 571 (Haplidia) A: LE SY
lizleri Keith, 2000a: 381 (Haplidia) A: SY TR
lodosi Baraud, 1988b: 573 (Haplidia) A: TR
massai Baraud, 1975c: 195 (Haplidia) E: IT (Sicilia)
migliaccioi Baraud, 1988b: 558 (Haplidia) A: TR
montreuili Keith, 2000b: 567 (Haplidia) A: CY
nididula Kraatz, 1882b: 40 (Haplidia) N: EG A: IS SY TR
taurica Petrovitz, 1971e: 577 (Haplidia)

pamphyliensis Baraud, 1988b: 573 (Haplidia) A: TR
petrovitzi lassallei Keith, 2000a: 382 (Haplidia) A: TR
petrovitzi petrovitzi Baraud, 1988b: 572 (Haplidia) A: IS SY
pilicollis pilicollis Petrovitz, 1967b: 336 (Haplidia) A: SY TR
pilicollis temperei Baraud, 1988b: 571 (Haplidia) A: LE SY

SCARABAEIDAE

SCARABAEIDAE

255

preissi Keith, 2000b: 565 (Haplidia) A: CY
pruinosa Baudi di Selve, 1870: 74 A: CY
pubiventris Kraatz, 1882b: 40 (Haplidia) A: TR
schatzmayri Petrovitz, 1958f: 204 (Haplidia)

rugicollis Petrovitz, 1967b: 335 (Haplidia) A: TR
sparsepunctata Petrovitz, 1967b: 337 (Haplidia) A: TR
tarsensis Kraatz, 1882b: 37 (Haplidia) A: IN LE SY TR
transversa cretica Petrovitz, 1971d: 231 (Haplidia) E: GR (Kríti)
transversa peloponnisica Petrovitz, 1971d: 231 (Haplidia) E: GR (Pelopónnisos)
transversa transversa Fabricius, 1801b: 163 (Melolontha) E: AL AU BH BU CR FR GR IT MC ME RO SB SL SZ TR
A: TR
lateriventris Reitter, 1902b: 185 (Haplidia)
quercina Burmeister, 1855: 344

turcica Kraatz, 1882b: 36 (Haplidia) E: TR
vagepunctata Kraatz, 1882b: 36 (Haplidia) A: TR
heydeni Kraatz, 1882b: 36 (Haplidia)

villigera Burmeister, 1855: 344 E: IT (Sicilia)
villosicollis Kraatz, 1882b: 39 (Haplidia) A: IS LE SY
leuthneri Petrovitz, 1971d: 230 (Haplidia)

werneri Keith, 2000a: 380 (Haplidia) A: SY TR
wewalkai Petrovitz, 1971e: 576 (Haplidia) A: TR
genus Asophrops Petrovitz, 1961c: 115 type species Asophrops jettmari Petrovitz, 1961
jettmari Petrovitz, 1961c: 115 A: PA
medvedevi Nikolajev & Kabakov, 1980: 224 (Lasiopsis) A: AF
punjabensis Arrow, 1948: 53 (Lachnosterna) A: PA UP ORR
rudolfpetrovitzi Keith, 2007b: 291 A: PA
staveni Keith, 2007b: 293 A: PA
genus Brachyllus Brenske, 1896a: 162 type species Brachyllus ulcerosus Brenske, 1896
bhutanensis Keith, 2005c: 28 A: BT
cechovskyi Sabatinelli & Pontuale, 1997: 454 A: NP
deuveianus Keith, 2003b: 307 A: YUN
frontalis Brenske, 1892b: 83 (Brahmina) A: NP SD ORR
itohi mantillerii Keith, 2003b: 309 A: YUN
rougeriei Keith, 2003b: 309 A: YUN
siniaevi Keith, 1999b: 284 A: YUN ORR
sprecherae Keith, 2006b: 421 A: AP
svetlanae Keith, 2006e: 55 A: GUX
ulcerosus Brenske, 1896a: 162 A: NP SD
lacroixi Keith, 1999b: 281

256

SCARABAEIDAE

genus Brahmina C.É. Blanchard, 1851: 140 type species Brahmina calva C.É Blanchard, 1851 (=Ancylonycha
cribricollis Kollar & L. Redtenbacher, 1844)
Coniotrogus Brenske, 1903: 333 type species Coniotrogus abdominalis Brenske, 1902
Pseudolontha Fairmaire, 1897e: 211 type species Pseudolontha canaliculata Fairmaire, 1897
Rhizocolax Motschulsky, 1860e: 130 type species Rhizocolax conspersus Motschulsky, 1860 (=Rhizotrogus sedakovii
Mannerheim, 1849)

subgenus Anoxiella Reitter, 1902b: 178 type species Brahmina senescens J. Frivaldszky, 1890
Schistocometa Brenske, 1903: 334 type species Brahmina brenskei Reitter, 1900 (=Rhizotrogus potanini Semenov, 1891)

potanini Semenov, 1891: 318 (Rhizotrogus) A: GAN HEB NE SCH QIN
brenskei Reitter, 1900d: 158

senescens J. Frivaldszky, 1890: 200 A: GAN
subgenus Brahmina C.É. Blanchard, 1851: 140 type species Brahmina calva C.É Blanchard, 1851 (=Ancylonycha
cribricollis Kollar & L. Redtenbacher, 1844)
Rhizocolax Motschulsky, 1860e: 130 type species Rhizocolax conspersus Motschulsky, 1860 (=Rhizotrogus sedakovii
Mannerheim, 1849)

amurensis Brenske, 1892b: 95 A: BEI FE HEI
crenicollis Motschulsky, 1854d: 64 (Ancylonycha) A: FE HEI NC NE NO SC
cribricollis Kollar & L. Redtenbacher, 1844: 524 (Ancylonycha) A: BT CH HP KA PA ORR
calva C.É. Blanchard, 1851: 140

darcisi Reitter, 1902b: 181 A: NE NO QIN SC
mandarina Reitter, 1902b: 181 A: NE
sedakovii Mannerheim, 1849: 237 (Rhizotrogus) A: ES FE HEB HEI MG
castanipes Brenske, 1892b: 89
conspersa Motschulsky, 1860e: 136 (Rhizocolax)
grandipunctata Murayama, 1941: 40
intermedia Mannerheim, 1849: 238 (Rhizotrogus)
sibirica Brenske, 1892b: 89

serricollis Motschulsky, 1854c: 46 (Ancylonycha) A: BEI
subgenus Brahminella S.I. Medvedev, 1951a: 239 type species Melolontha agnella Faldermann, 1835
agnella agnella Faldermann, 1835b: 375 (Melolontha) A: ES FE GAN HEI MG NMO
lederi Reitter, 1902b: 169 (Lasiopsis)

agnella gobica Endrődi, 1964: 428 A: MG
faldermanni Kraatz, 1892b: 309 A: BEI FE HEB HEI LIA
rubetra Faldermann, 1835b: 376 (Melolontha) A: LIA NC NO SC
genus Brahmina, species incertae sedis
abdominalis Brenske, 1903: 334 (Coniotrogus) A: QIN
adaequata Moser, 1910b: 475 A: SD
braeti Brenske, 1896a: 161 A: BT NP SD ORR
brevipilosa Moser, 1918b: 320 A: CH (“Singley”)
canaliculata Fairmaire, 1897e: 212 (Pseudolontha) A: SCH
cardoni Brenske, 1892b: 108 A: SD

SCARABAEIDAE
cariniclypeata Nomura, 1977b: 99 A: TAI
carinifrons Moser, 1910b: 474 (Holotrichia) A: SD
clypealis Moser, 1915a: 142 A: “Tibet (Poo)”
coriacea Hope, 1831: 23 (Melolontha) A: HP NP UP
gradatifrons Bates, 1891a: 15 (Lachnosterna)

cribriceps Moser, 1915a: 141 A: HUB
crinicollis Burmeister, 1855: 364 A: BT HP PA ORR
cuprea Mittal & Pajni, 1977: 86 A: UP
cylindrica Gyllenhal, 1817: 83 (Melolontha) A: “India or.”
elongata Moser, 1913a: 66 A: “Himalaya”
excisiceps Moser, 1915a: 136 A: CH (“Ho-chan”) SC
extraria Keith, 2009d: 350 A: NP
flabellata Brenske, 1892b: 104 A: SD
glabella Nikolajev & Kabakov, 1980: 225 (Lasiopsis) A: AF PA
itohi Kobayashi, 1998: 212 [RN] A: TAI
macrophylla Nomura, 1977b: 98 [HN]

jubata Frey, 1969c: 523 A: NP XIZ
kurseongana Moser, 1924: 132 A: SD
latispina Kobayashi, 1987a: 17 A: TAI
lutea Moser, 1913a: 66 A: “Himalaya”
monticola Kobayashi, 1993: 72 A: TAI
nomurai Itoh, 1989: 13 [RN] A: TAI
bituberculata Nomura, 1977b: 97 [HN]

nuda Moser, 1915a: 140 A: FUJ
parvula Moser, 1915a: 137 A: YUN
plagiatula Brenske, 1897d: 351 A: SD
pseudoglabella Keith, 2008f: 491 A: NP
pubiventris Kobayashi, 1993: 74 A: TAI
pulchella Motschulsky, 1854c: 46 (Rhizotrogus) A: BEI
ruficollis Moser, 1915a: 139 A: YUN
rugifrons Moser, 1918b: 319 A: YUN
rugosicollis Frey, 1971a: 130 A: UP
ruida Y.-W. Zhang & Wang, 1997a: 212 A: SHX
sakishimana Nomura, 1965: 3 A: JA (Iriomote-shima)
sculpticollis Frey, 1969c: 524 A: NP
serricollis Motschulsky, 1854c: 46 A: BEI
shibatai Kobayashi, 1987a: 18 A: TAI
sikkimensis Brenske, 1892b: 113 A: SD
simplex Frey, 1972a: 115 A: FUJ
sulcifrons Moser, 1913d: 344 A: SD
taitungensis Nomura, 1977b: 98 A: TAI

257

258

SCARABAEIDAE

tuberculifrons Moser, 1926: 204 A: KA
turcestana Brenske, 1892b: 94 A: “Turkestan”
wutaiensis Y.-W. Zhang & Wang, 1997a: 212 A: SHX
yabloniana Murayama, 1941: 40 A: HEI
yunnana Moser, 1915a: 138 A: YUN
genus Bunbunius Nomura, 1970: 65 type species Holotrichia reticulata Murayama, 1941
abditus Keith, 2008e: 66 A: HUN
bicallosifrons Frey, 1972b: 165 (Holotrichia) A: FUJ GUA
csorbai Keith, 2008e: 66 A: NP
diversifrons Fairmaire, 1891f: cxcix (Rhizotrogus) A: HUB
gamdoensis Keith & J. Li, 2013: 21 A: XIZ
hangayi Keith, 2008e: 62 A: SCH
hartmanni Keith, 2009d: 354 A: NP
liukueiensis Kobayashi, 1986a: 14 (Holotrichia) A: TAI
opacipennis Nomura, 1970: 66 (Pollaplonyx) A: TAI
privus Keith, 2005c: 31 A: GUI
prunellus Arrow, 1946a: 29 (Holotrichia) A: YUN ORR
pseudoopacipennis Keith, 2008e: 60 A: TAI
puchneri hubeicus Keith, 2008e: 65 A: HUB
puchneri puchneri Keith, 2008e: 65 A: GAN SCH
quadrihamis Sehnal, 2013b: 650 A: SCH
reticulatus Murayama, 1941: 39 (Holotrichia) A: FUJ HEI HUN GUA GUX SC SCH
sinensis Keith, 2005c: 30 A: GAN HUB SHA
turnai Keith, 2008e: 63 A: SCH
genus Butozania Mikšič 1955: 514 type species Butozania macedonica Mikšič, 1955 (=Rhizotrogus bulgaricus
Kantardzhieva-Minkova, 1953)
bulgarica Kantardzhieva-Minkova, 1953: 44 (Rhizotrogus) E: BU MC
macedonica Mikšič, 1955: 516

genus Chilotrogus Reitter, 1905b: 202 type species Chilotrogus panotrogoides Reitter, 1905
Articephala Moser, 1920: 26 type species Articephala planifrons Moser, 1920 (=Lachnosterna stoliczkae Sharp, 1878)

batillarius Bates, 1891a: 15 (Lachnosterna) A: HP PA
densepunctatus Keith, 2007a: 47 A: NP
fabriziae Keith, 2007a: 45 A: SD
farsensis Keith, 2003a: 92 A: IN
heinzi Keith, 2005a: 275 A: PA
himachali Mittal, 1988: 158 (Articephala) A: HP
kalabi Keith, 2005a: 277 A: PA
lahauli Chandra, 1991: 132 (Articephala) A: HP

SCARABAEIDAE

259

merkli Keith, 2005a: 278 A: PA
nadaii Keith, 2007a: 49 A: NP
ottomerkli Keith, 2007a: 48 A: NP
panotrogoides Reitter, 1905b: 203 A: IN
phasmellus Biggs, 1945: 110 (Articephala)

schmidti Keith, 2007a: 46 A: NP
stoliczkae Sharp, 1878b: 172 (Lachnosterna) A: HP NP PA UP
fulgidus Brenske, 1897d: 351 (Brahmina)
planiceps Frey, 1973a: 232 (Holotrichia)
planifrons Moser, 1920: 27 (Articephala)
popei Frey, 1963: 552 (Brahmina)

genus Chioneosoma Kraatz, 1891e: 354 type species Rhizotrogus komarovi Brenske, 1886
subgenus Aleucolomus Reitter, 1902b: 192 type species Melolontha porosa Fischer von Waldheim, 1823
Aleucoschema Semenov, 1935: 281 type species Melolontha vulpina Gyllenhal, 1817

demetrii Semenov, 1902a: 214 (Rhizotrogus) A: KZ UZ
porosum Fischer von Waldheim, 1823: pl. XXVIII [=1824: 213] (Melolontha) E: AB KZ ST A: IN KI KZ TD TM UZ
akinini Brenske, 1888: 352 (Rhizotrogus)
holsericeum Ménétriés, 1849: 64 (Ancylonycha)
nitidurum Reitter, 1902b: 193

reitteri Brenske, 1887b: 232 (Rhizotrogus) A: MG QIN XIN XIZ
bilobum Sharp, 1878b: 173 (Rhizotrogus)
eoum Semenov, 1902a: 215 (Rhizotrogus)

shestoperovi Semenov & S.I. Medvedev, 1936: 108 A: KI KZ XIN
subporosum Reitter, 1902b: 194 A: KZ TD UZ
ferganense Semenov, 1902a: 215 (Rhizotrogus)

tedschenense Semenov, 1895b: 198 (Rhizotrogus) A: KZ TM
vulpinum Gyllenhal, 1817: 81 (Melolontha) E: KZ ST UK A: IN KZ
ochraceum Motschulsky, 1860i: 523 (Rhizotrogus)
persicum Brenske, 1888: 359 (Rhizotrogus)
thoracicum Krynicki, 1829: 192 (Ancylonycha)

subgenus Chioneosoma Kraatz, 1891e: 354 type species Rhizotrogus komarowi Brenske, 1886
komarowi Brenske, 1886: 195 (Rhizotrogus) A: KZ TD TM UZ
nix Semenov & S.I. Medvedev, 1936: 112 A: UZ
subgenus Chionotrogus Reitter, 1902b: 190 type species Melolontha pulverea Knoch, 1801
Leucoschesis Semenov, 1935: 280 type species Rhizotrogus jakowlewi Semenov, 1895

astrachanicum Semenov, 1902a: 207 (Rhizotrogus) E: KZ ST A: KZ
bucharicum Semenov & S.I. Medvedev, 1936: 112 A: UZ
candidum Semenov, 1902a: 208 (Rhizotrogus) A: TM
kraatzi Brenske, 1886: 198 (Rhizotrogus)

casacorum Semenov, 1902a: 205 (Rhizotrogus) A: KZ

260

SCARABAEIDAE

glazunowi Semenov, 1895b: 208 (Rhizotrogus) A: TM
jakowlewi Semenov, 1895b: 201 (Rhizotrogus) A: UZ
kizilkumense Semenov, 1895b: 201 (Rhizotrogus) A: KZ UZ
kokujewi Semenov, 1895d: 251 (Rhizotrogus) A: UZ
lopatini S.I. Medvedev, 1962: 599 A: TD
molare Reitter, 1902b: 191 A: TM
mucidum Semenov & S.I. Medvedev, 1936: 110 A: KZ UZ
niveum Semenov & S.I. Medvedev, 1936: 208 A: TM
obenbergeri Semenov & S.I. Medvedev, 1936: 109 A: KZ
parfentjevi S.I. Medvedev, 1966: 851 A: KZ TM
peetzi Semenov, 1899c: 647 (Rhizotrogus) A: TM
pulvereum Knoch, 1801: 88 (Melolontha) E: KZ ST UK A: KZ TM UZ
macrophyllum Fischer von Waldheim, 1844: 49 (Cyphonotus)
thoracicum Zubkov, 1833: 323 (Melolontha)

rimskii Semenov & S.I. Medvedev, 1936: 110 A: KZ
senex Semenov, 1902a: 204 (Rhizotrogus) A: KI KZ UZ
sequensi Reitter, 1902b: 190 A: TM
turcomanum Brenske, 1888: 352 (Rhizotrogus) A: TM
subgenus Leucolasium Semenov, 1935: 281 type species Rhizotrogus gorilla Brenske, 1886
Leucoschema Semenov, 1935: 281 type species Rhizotrogus rostowtzowi Semenov, 1899

aralense Semenov & S.I. Medvedev, 1936: 111 A: KZ
arnoldii S.I. Medvedev, 1951a: 430 A: TD
baadeni Brenske, 1886: 196 (Rhizotrogus) A: TM
gorilla gorilla Brenske, 1886: 197 (Rhizotrogus) A: UZ TM
gorilla transoxiana Semenov, 1899c: 643 (Rhizotrogus) A: KZ
rostowtzowi Semenov, 1899c: 644 (Rhizotrogus) A: TM
tschitscherini Semenov, 1895a: 128 (Rhizotrogus) A: TM UZ
genus Chioneosoma, species incertae sedis,
corporaali Balthasar, 1935g: 321 A: “Karakorum”
genus Chioneosoma, nomina dubia
farinosum Krynicki, 1829: 192 (Cyphonotus) A: TM “Transkaspien”
hispidum Ballion, 1871a: 341 (Rhizotrogus) E: ST
genus Dasytrogus Reitter, 1902b: 230 type species Amphimallon transcaspicus Brenske, 1886
subgenus Cyprotrogus Nikolajev, 2004a: 60 type species Ancylonycha nitens Baudi di Selve, 1870
nitens Baudi di Selve, 1870: 75 (Ancylonycha) A: CY
subgenus Leucotrogus Nikolajev, 2004a: 48 type species Amphimallon glabripennis Ballion, 1871
afghanus Petrovitz, 1967e: 483 (Madotrogus) A: AF
aruktavicus Nikolajev, 1976a: 65 (Madotrogus) A: TD
glabripennis Ballion, 1871a: 341 (Amphimallon) A: KI KZ TD TM UZ

SCARABAEIDAE

261

kabakovi Nikolajev, 1976d: 696 (Madotrogus) A: AF
makarovi Gusakov, 2004: 11 (Madotrogus) A: TM
muelleri Balthasar, 1956b: 430 (Amphimallon) A: AF
pakistanus Keith, 2010a: 77 (Madotrogus) A: PA
palpalis Moser, 1924: 133 (Madotrogus) A: HP
petrovitzi Nikolajev & Kabakov, 1977b: 647 (Madotrogus) A: AF
pumilio Balthasar, 1956b: 431 (Amphimallon) A: AF
tadzhikorum Nikolajev, 1975c: 1261 (Madotrogus) A: AF TM UZ
subgenus Dasytrogus Reitter, 1902b: 230 type species Amphimallon transcaspicus Brenske, 1886
Madotrogus Reitter, 1902b: 251 type species Rhizotrogus glabricollis Reitter, 1888

diurnus Nikolajev, 2004a: 54 (Madotrogus) A: TM
ferganensis Protsenko, 1962: 100 (Amphimallon) A: KI TD UZ
glabricollis Reitter, 1888a: 68 (Rhizotrogus) E: AR A: IN KZ TM TR UZ
perses Zaitzev, 1928: 396 (Madotrogus)
varentzowi Semenov, 1895d: 252 (Rhizotrogus)

kabulicus Nikolajev & Kabakov, 1977b: 647 (Madotrogus) A: AF
kirgisicus Semenov, 1895b: 203 (Rhizotrogus) A: KI KZ UZ
kuhitangicus Nikolajev, 1976b: 166 (Madotrogus) A: TM UZ
transcaspicus Brenske, 1886: 200 (Amphimallon) A: AF KZ TM UZ
tekkensis Brenske, 1888: 355 (Amphimallon)

subgenus Micramphimallon S.I. Medvedev, 1951a: 486 type species Amphimallon mesopotamicum S.I.
Medvedev, 1951
baal Petrovitz, 1968g: 480 (Amphimallon) A: IQ
beludschistanus Brenske, 1897c: 318 (Rhizotrogus) A: “Beludschistan”
girardi Keith, 2002a: 14 (Madotrogus) A: IQ
jubatus Reitter, 1890g: 169 (Rhizotrogus) E: AB AR GG A: TR
obscuricollis Zaitzev, 1927: 378 (Amphimallon)

laghmanicus Nikolajev & Kabakov, 1977b: 647 (Madotrogus) A: AF
mesopotamicus S.I. Medvedev, 1951a: 487 (Amphimallon) A: IQ
nikolajevi Sabatinelli & Katbeh-Bader, 2009: 427 (Madotrogus) A: JO
problematicus Bunalski, 1993: 87 (Madotrogus) A: IQ
sinaicus Baraud, 1975c: 194 (Eriotrogus) A: IS SI
zembi Keith, 2002a: 15 (Madotrogus) A: KU
genus Eotrichia S.I. Medvedev, 1951a: 308 type species Holotrichia titanis Reitter, 1902 (=Lachnosterna
n­ iponensis Lewis, 1895)
biehli Brenske, 1892d: 170 (Holotrichia) A: SD
kurseongana Brenske, 1896a: 157 (Holotrichia)

braeti Brenske, 1896a: 159 (Holotrichia) A: BT SD ORR
brevispina Chang, 1964: 258 (Holotrichia) A: YUN
capitalis Chang, 1964: 257 (Holotrichia) A: YUN

262

SCARABAEIDAE

cavifrons Brenske, 1892d: 167 (Holotrichia) A: NP SD
cheni Chang, 1964: 256 (Holotrichia) A: YUN
costipennis Moser, 1912b: 421 (Holotrichia) A: CH
eberti Keith, 2008f: 493 (Holotrichia) A: NP
femoralis Chang, 1964: 257 (Holotrichia) A: YUN
frontalis Brenske, 1892d: 167 (Holotrichia) A: SD ORR
fulgida Brenske, 1896a: 159 (Holotrichia)

glabricollis Brenske, 1896a: 157 (Holotrichia) A: BT SD ORR
hainanensis Chang, 1964: 258 (Holotrichia) A: HAI ORR
impressicollis Moser, 1910b: 468 (Holotrichia) A: SHN
laticollis Moser, 1921: 62 (Holotrichia) A: SD
longipennis C.É. Blanchard, 1851: 140 (Ancylonycha) A: BT HP UP “Thibet” ORR
brevicollis Burmeister, 1855: 314 (Ancylonycha) [HN]
simlana Brenske, 1897d: 346 (Holotrichia)
tereticollis Burmeister, 1855: 538 (Ancylonycha) [RN]

maxima Chang, 1964: 258 (Holotrichia) A: CHQ SCH
nepalensis Frey, 1972a: 120 (Holotrichia) A: NP
niponensis Lewis, 1895c: 398 (Lachnosterna) A: FE HEB HEI NC SC SCH SHA
titanis Reitter, 1902b: 178 (Holotrichia)

protracta Bates, 1891b: 76 (Lachnosterna) A: SCH
scrobiculata Brenske, 1892d: 174 (Holotrichia) A: SD YUN
sericina Frey, 1971b: 213 (Holotrichia) A: SD
setosifrons Khan & Ghai, 1982a: 36 (Holotrichia) A: HP
sikkimensis Brenske, 1892d: 169 (Holotrichia) A: BT HP NP SD
staudingeri Brenske, 1892d: 166 (Holotrichia) A: SD ORR
nobilissima Brenske, 1894a: 20 (Holotrichia)

subiridea Fairmaire, 1891f: ccii (Lachnosterna) A: HUB
taiwana Nomura, 1977b: 90 (Holotrichia) A: TAI
tarsalis Chang, 1964: 257 (Holotrichia) A: CE NE NO FE
tsignata Chang, 1964: 258 (Holotrichia) A: YUN
yatungensis Chang, 1964: 259 (Holotrichia) A: XIZ
genus Firminus Coca-Abia, 2003: 451 type species Rhizotrogus punicus Burmeister, 1855
alsasuanus Reitter, 1902b: 206 (Rhizotrogus) E: SP
baudii Brenske, 1886: 202 (Rhizotrogus) E: IT
bellieri Reiche, 1862a: 295 (Rhizotrogus) E: FR (Corse) IT (Sardegna)
ciliatus ciliatus Reiche, 1862b: 539 (Rhizotrogus) E: IT (Sicilia)
proximus Brenske, 1886: 203 (Rhizotrogus)

ciliatus vexillis Reitter, 1902b: 202 (Rhizotrogus) E: IT
fossulatus Mulsant & Rey, 1859a: 228 [=1859b: 164] (Rhizotrogus) E: FR (Corse) IT (Sardegna)
lautiusculus Schaufuss, 1864: 675 (Rhizotrogus) E: AL AU BH CR GR ME SL

SCARABAEIDAE

263

maltensis Rössner & Zorn, 2013: 243 E: MA (Ghawdex)
massai Arnone, Lo Cascio & Grita, 2014: 341 E: IT (Isole Eolie)
procerus Baudi di Selve, 1870: 73 (Rhizotrogus) E: IT SL
punicus Burmeister, 1855: 378 (Rhizotrogus) N: AG MO TU
densaticollis Fairmaire, 1884c: lxi (Rhizotrogus)
eburneicollis Raffray, 1871: 160 (Rhizotrogus)
lamellatus Reitter, 1902b: 217 (Rhizotrogus)

putoni Reitter, 1902b: 214 (Rhizotrogus) N: AG
genus Geotrogus Guérin-Méneville, 1842: 7 type species Rhizotrogus magagnoscii Guérin-Méneville, 1842
Apterogyna Reitter, 1902b: 210 [HN] type species Rhizotrogus euphytus Buquet, 1840
Pseudoapterogyna Escalera, 1914a: 175 [RN] type species Rhizotrogus euphytus Buquet, 1840

araneipes Fairmaire, 1860c: 428 (Rhizotrogus) N: AG MO TU
barbarus P.H. Lucas, 1847b: 281 (Rhizotrogus) N: AG MO
fastidiosus Fairmaire, 1860c: 436 (Rhizotrogus)
reflexus C.É. Blanchard, 1851: 147 (Rhizotrogus)
reichii Mulsant & Rey, 1871a: 586 [=1871c: 330] (Rhizotrogus)
truncatipennis P.H. Lucas, 1847b: 283 (Rhizotrogus)

bedeli Reitter, 1902b: 218 (Rhizotrogus) N: AG
bleicherii Fairmaire, 1879e: 250 (Rhizotrogus) N: AG MO
capito Fairmaire, 1882c: clxxvii (Rhizotrogus) N: LB TU
cretei Ancey, 1888: [no page] (Rhizotrogus)
minutus Brenske, 1889: 329 (Rhizotrogus)

cludtsi Keith, 2005b: 137 (Pseudoapterogyna) N: LB
crassus Fairmaire, 1860c: 433 (Rhizotrogus) N: AG
cribripennis Peyerimhoff, 1949a: 24 (Rhizotrogus) N: AG
demoflysi Baraud, 1970b: 489 (Rhizotrogus) N: TU
dispar Buquet, 1840a: 171 (Rhizotrogus) N: AG TU
editorum Peyerimhoff, 1949a: 23 (Rhizotrogus) N: AG (Djurdjura)
elegans Brenske, 1890e: 328 (Rhizotrogus) N: AG TU
euphytus euphytus Buquet, 1840a: 171 (Rhizotrogus) N: AG TU
biskrensis Marseul, 1878: 65
dilutus Fairmaire, 1884c: lxiii (Rhizotrogus)
maculicollis Fairmaire, 1860c: 431 (Rhizotrogus) [HN]
tuniseus Fairmaire, 1884c: lxii (Rhizotrogus)

euphytus lamantiai Sparacio, 2014: 360 (Pseudoapterogyna) E: IT (Pantelleria)
gabalus Buquet, 1840a: 172 (Rhizotrogus) N: AG MO
asperiventris Fairmaire, 1867: 397 (Rhizotrogus)
humericavus Marseul, 1878: 64 (Rhizotrogus)
laticeps Marseul, 1878: 63 (Rhizotrogus)
melanocephalus Reitter, 1902b: 226

264
nitidiventris Fairmaire, 1867: 398 (Rhizotrogus)
punctiventris Fairmaire, 1860c: 426 (Rhizotrogus)
sinuatocollis Fairmaire, 1860c: 425 (Rhizotrogus)
translucens Marseul, 1878: 64 (Rhizotrogus)
triangulus Reitter, 1902b: 226

genei C.É. Blanchard, 1851: 147 (Rhizotrogus) E: FR (Corse) IT (Sardegna)
insularis Reiche, 1862a: 294 (Rhizotrogus)
sardous Burmeister, 1855: 371

gerardi aurasiacus Peyerimhoff, 1949a: 24 (Rhizotrogus) N: AG (Massif de l’Aurès)
gerardi gerardi Buquet, 1840a: 171 (Rhizotrogus) N: AG (litoral)
castanopterus Fairmaire, 1871a: 377 (Rhizotrogus)
guyoni P.H. Lucas, 1857b: lxxxvi (Rhizotrogus)
laeviscutatus Fairmaire, 1868a: 481 (Rhizotrogus)
lateritius Fairmaire, 1864: 640 (Rhizotrogus)
nitidicollis Fairmaire, 1864: 640 (Rhizotrogus)
polita Fairmaire, 1867: 398 (Rhizotrogus)
sparsinus Reitter, 1902b: 220 (Rhizotrogus)

gerardi henoni Fairmaire, 1860c: 430 (Rhizotrogus) N: AG (mountains of interior)
tenuispinus Fairmaire, 1867: 399 (Rhizotrogus)
thiebaulti Fairmaire, 1879e: 251 (Rhizotrogus)

gonoderus Fairmaire, 1860c: 427 (Rhizotrogus) N: AG MO TU
grossus cedretorum Peyerimhoff, 1949a: 23 (Rhizotrogus) N: AG (Forêt des Cèdres de Sgag)
grossus grossus C.É. Blanchard, 1851: 147 (Rhizotrogus) N: AG
cilipes Marseul, 1878: 65 (Apterogyna)
decoloratus Fairmaire, 1860c: 432 (Rhizotrogus)
pallidocinctus Raffray, 1873: 371 (Rhizotrogus)

grossus mesatlanticus Peyerimhoff, 1949a: 23 (Rhizotrogus) N: AG MO
holoxanthus Fairmaire, 1868a: 480 (Rhizotrogus) N: AG
ignobilis Reitter, 1902b: 223 (Rhizotrogus) N: AG
inflatus deserticola C.É. Blanchard, 1851: 147 (Rhizotrogus) N: AG MO
nubicollis Marseul, 1878: 64 (Rhizotrogus)
subopacus Fairmaire, 1860c: 427 (Rhizotrogus)

inflatus inflatus Buquet, 1840a: 171 (Rhizotrogus) N: AG
lepidulus Normand, 1932: 154 N: TU
magagnoscii Guérin-Méneville, 1842: 7 (Rhizotrogus) N: AG
majorinus Peyerimhoff, 1949a: 24 (Rhizotrogus) N: AG TU
michaelis Sparacio, 2014: 364 (Pseudoapterogyna) E: IT (Sicilia)
normandi aridulus Peyerimhoff, 1949a: 25 (Rhizotrogus) N: TU
normandi normandi Peyerimhoff, 1949a: 25 (Rhizotrogus) N: TU
numidicus P.H. Lucas, 1846d: pl. 25 (Rhizotrogus) N: AG
atlanticus Raffray, 1871: 160 (Rhizotrogus)
leucostethus Reitter, 1902b: 220 (Rhizotrogus)

SCARABAEIDAE

SCARABAEIDAE

265

nigratus Fairmaire, 1860c: 429 (Rhizotrogus)
oppositus Reitter, 1902b: 221 (Rhizotrogus)
pallidimargo Reitter, 1902b: 222 (Rhizotrogus)

olcesii Fairmaire, 1867: 396 (Rhizotrogus) N: MO
pellegrinensis Brenske, 1893d: 233 E: IT (Sicilia)
psilopus denigratus Peyerimhoff, 1949: 26 (Rhizotrogus) N: MO
psilopus psilopus Fairmaire, 1883d: cxix (Rhizotrogus) N: MO
prophettii Fairmaire, 1860c: 434 (Rhizotrogus) N: AG
lejeunii Fairmaire, 1860c: 434 (Rhizotrogus)

rectibasis Reitter, 1902b: 229 N: AG TU
sicelis C.É. Blanchard, 1851: 146 (Rhizotrogus) E: IT (Pantelleria, Sicilia, Ustica)
siculus Burmeister, 1855: 369

sordescens Fairmaire, 1871a: 375 (Rhizotrogus) N: MO
mscalicus Escalera, 1914a: 171 (Rhizotrogus)

stupidus Fairmaire, 1860c: 435 (Rhizotrogus) N: AG
syrtanus Normand, 1951: 156 (Rhizotrogus) N: TU
temperei Baraud, 1970b: 489 (Rhizotrogus) N: AG
tusculus Buquet, 1840a: 171 (Rhizotrogus) N: AG TU
amphytus Buqueta, 1840: 171 (Rhizotrogus)
fargesi Théry, 1900b: 33 (Rhizotrogus)
inundationis Raffray, 1871: 160 (Rhizotrogus)
obesus P.H. Lucas, 1847b: 282 (Rhizotrogus)
scabrosus Balthasar, 1930d: 97 (Rhizotrogus)

tynetis Baraud, 1972: 160 (Rhizotrogus) N: AG TU
unguicularis Normand, 1936: 378 N: TU
vorax Marseul, 1878: 65 (Apterogyna) E: IT (Lampedusa, Lampione) N: AG LB MO TU
quedenfeldti Brenske, 1890e: 326 (Rhizotrogus)
tripolitanus Fairmaire, 1884c: lxii (Rhizotrogus)

genus Glacilitarsus Matsumoto, 2014: 1 type species Holotrichia clypealis Brenske, 1896
clypealis Brenske, 1896a: 159 (Holotrichia) A: SD
genus Gnaphalostetha Reiche & Saulcy, 1856: 383 type species Gnaphalostetha bonvoisini Reiche &
Saulcy, 1856
bonvoisini Reiche & Saulcy, 1856: 354 A: IS JO
heinzi Keith, 2005f: 107 A: IS
hiemalis Baraud, 1990e: 296 (Lasiexis) A: JO
longepilosa Keith, 2005f: 108 A: “Arabie”
genus Holochelus Reitter, 1889g: 279 type species Holochelus subseriatus Reitter, 1889
subgenus Holochelus Reitter, 1889g: 279 type species Holochelus subseriatus Reitter, 1889
Eriotrogus Reitter, 1902b: 252 type species Amphimallon erivanicus Reitter, 1902

266

SCARABAEIDAE

Odontochelus Semenov, 1905b: 141 type species Holochelus lineolatus Reitter, 1902

aschhabadensis Nonveiller, 1965: 54 (Miltotrogus) E: ST A: KZ TM
costulatus I. Frivalszkky, 1835: 257 (Rhizotrogus) E: BU GR MC TR A: TR
erivanicus Reitter, 1902b: 253 (Amphimallon) E: AB AR GG A: TR
gracilis Nonveiller, 1965: 57 (Miltotrogus)

fallaciosus Nonveiller, 1965: 47 A: IN
fallax fallax Marseul, 1879b: 472 (Rhizotrogus) E: AB AL AR BU GG GR KO MC SB TR A: IN SY TM TR
bodemeyeri Brenske, 1900a: 150 (Rhizotrogus)
heydeni Brenske, 1891c: 216 (Rhizotrogus)

fallax mandator Reitter, 1902b: 200 (Rhizotrogus) A: SY
gigas Sabatinelli, 1977c: 103 (Miltotrogus) A: LE SY
lineolatus Reitter, 1902b: 195 A: TR
majusculus Nonveiller, 1965: 52 (Miltotrogus) E: “Caucasus” A: TR
nadaii Keith, 2003a: 89 A: IN
pilicollis pilicollis Gyllenhal, 1817: 89 (Melolontha) E: AU BH BU CR CZ GR HU MD RO SB SK TR UK A: SY TR
castaneus Gyllenhal, 1817: 90 (Melolontha)

pilicollis tauricola Petrovitz, 1968g: 480 (Miltotrogus) A: TR
pseudonadaii Keith, 2003a: 91 A: IN
setiventris Reitter, 1902b: 200 (Rhizotrogus) A: IS LE SY
subseriatus Reitter, 1889g: 279 E: ST UK (Krym)
uvarovi Semenov & S.I. Medvedev, 1936: 355 (Rhizotrogus) A: TM
subgenus Miltotrogus Reitter, 1902b: 196 type species Melolontha aequinoctialis Herbst, 1790
aequinoctialis Herbst, 1790: 62 (Melolontha) E: AB AU BH BU CR CT CZ GE GG GR HU IT KZ MC MD PL RO
SB SK SL ST SZ TR UK A: KZ TR
grandicornis Ballion, 1871a: 342 (Rhizotrogus)

angustifrons Nonveiller, 1965: 61 (Miltotrogus) A: SY
armeniacus Zaitzev, 1927: 396 (Rhizotrogus) A: TR
boudjelalae Baraud, 1987c: 359 (Miltotrogus) A: TR
brenskei Reitter, 1888a: 68 (Rhizotrogus) E: AB AR GG ST
intermedius S.I. Medvedev, 1951a: 364 (Rhizotrogus)
transcaucasicus S.I. Medvedev, 1951a: 363 (Rhizotrogus)

brussensis Nonveiller, 1965: 76 (Miltotrogus) A: TR
caucasicus Gyllenhal, 1817: 82 (Melolontha) A: “Caucasus”
escherichi Brenske, 1897a: 21 (Rhizotrogus) E: AR GR A: TR
fraxinicola Hagenbach, 1825: 488 (Melolontha) E: AL AU BH CR FR (Corse) GR IT ME SL A: TR
fusculus Nonveiller, 1965: 59 (Miltotrogus) A: IN
gradojevici Nonveiller, 1965: 98 E: BU GR A: TR
jarkae Sehnal & Nikodým, 2011: 2 E: GR
kerimi A. Rey, 1994: 242 (Miltotrogus) A: IN
kurdistanus Petrovitz, 1968g: 478 (Miltotrogus) A: TR
mimicus Reitter, 1902b: 201 (Rhizotrogus) A: SY TR

SCARABAEIDAE

267

nocturnus Nonveiller, 1958: 104 (Miltotrogus) E: AU BH CZ GG GR HU MC SB SK UK “Caucasus”
ovchinnikovi Nikolajev, 1998: 42 A: TD
parvus Nonveiller, 1965: 65 (Miltotrogus) A: IN
phrygicus Petrovitz, 1971e: 580 (Miltotrogus) A: TR
rapuzzii Keith, 2004: 476 A: SY
rusticus Faldermann, 1835a: 275 (Ancylonycha) E: AR RO A: TR
obenbergeri Nonveiller, 1965: 73 (Miltotrogus)

serrifunis Marseul, 1879b: 472 (Rhizotrogus) E: AB GG
syriacus Brenske, 1886: 204 (Rhizotrogus) A: LE SY
tataricus Faldermann, 1835a: 278 (Ancylonycha) E: AB AR GG TR ST A: TR
arcilabris Marseul, 1879b: 473 (Rhizotrogus)

tauricus C.É. Blanchard, 1851: 146 (Rhizotrogus) E: GR A: TR
vernus Germar, 1824: 127 (Melolontha) E: AL AU BH BU CR CZ GE GR HU IT KO MC MD ME RO SB SK SL ST
TR UK A: TR
vulpinus Burmeister, 1855: 376 (Rhizotrogus) E: MD RO ST UK
lanatus Fairmaire, 1860c: 436 (Rhizotrogus)

zimmermanni Nonveiller, 1965: 58 A: TR “Kurdistan”
zuvandi Qarjagdy, 1939: 86 (Rhizotrogus) E: AB
subgenus Pakistanotrogus Keith, 2006e: 48 type species Holochelus puchneri Keith, 2006
laszlonadaii Keith, 2008a: 342 [RN] A: PA
nadaii Keith, 2006e: 50 [HN]

nepalensis Keith, 2009d: 343 A: NP
nonveilleri Nikolajev & Kabakov, 1980: 227 (Miltotrogus) A: AF
pakistanus Keith, 2006e: 49 A: PA
puchneri Keith, 2006e: 48 A: KA PA
reichenbachi Keith, 2006e: 51 A: PA
genus Holotrichia Hope, 1837: 99 type species Melolontha serrata Fabricius, 1781
Ancylonycha Dejean, 1833: 160 type species Melolontha serrata Fabricius, 1781

acuticollis Frey, 1972a: 116 A: MG
amamiana Nomura, 1964a: 52 (Lachnosterna) A: JA (Amami-Ôshima)
anthracina Brenske, 1892d: 184 A: BT NP SD SHG ORR
cotesi Brenske, 1892d: 185

aritai Nomura, 1964b: 63 (Lachnosterna) A: JA (Iriomote-shima)
bhutanensis Frey, 1975b: 237 A: BT
calliglypta Bates, 1891b: 75 (Lachnosterna) A: SCH
cochinchina Nonfried, 1892e: 360 (Ancylonycha) A: GUX SCH YUN ORR
cocinchinae Brenske, 1892d: 185

consanguinea C.É. Blanchard, 1851: 139 A: PA UP ORR
convexopyga Moser, 1912b: 435 A: GAN HEB JA JIX SC
tametomo Nomura, 1969: 92 (Lachnosterna)

268

SCARABAEIDAE

danjoensis Miyake & Imasaka, 1982: 75 A: JA (Danjo)
diomphalia Bates, 1888b: 373 (Lachnosterna) A: BEI ES FE HEI JA JIL MG NC SC SHN
dolini Keith, 2005e: 541 A: JIX GUX
dubiosa Frey, 1971b: 217 A: SD ORR
dubitabilis Fairmaire, 1891e: cxxiii (Lachnosterna) A: KA
ernesti Reitter, 1902b: 177 A: FE HEB JA MG NC NO SC
ussuriensis S.I. Medvedev, 1951a: 305

fissa Brenske, 1894a: 71 A: UP ORR
gebleri Faldermann, 1835b: 374 (Melolontha) A: LIA NMO QIN
geilenkeuseri Reitter, 1902b: 175 A: CH
guandaoshana Kobayashi, 1990d: 73 A: TAI
hankowiensis Brenske, 1894a: 69 A: HUB SHG
shanghaiana Brenske, 1894a: 69

herwangshana Kobayashi, 1990d: 74 A: TAI
horishana Niijima & Kinoshita, 1923: 222 A: TAI
intermedia Brenske, 1894a: 64 A: CE FE NE ORR
impressa Burmeister, 1855: 314 (Ancylonycha) A: SD ORR
iridescens Moser, 1908c: 340 A: YUN ORR
iridipennis Brenske, 1892d: 177 A: “Himalaya” ORR
kiotonensis Brenske, 1894a: 68 A: FE JA NC NE SC
inelegans Lewis, 1895c: 396 (Lachnosterna)
waterhousei Brenske, 1894a: 68

koraiensis Murayama, 1937: 37 A: HEB HEI SC
kulzeri Frey, 1975b: 236 A: BT
kunmina Chang, 1965a: 45 A: YUN
kwatungensis Chang, 1965a: 46 A: FUJ
laevigata Chang, 1965a: 46 A: TAI YUN
lata Brenske, 1892d: 163 A: GUX HAI HKG SHG TAI ORR
pruinosa Niijima & Kinoshita, 1923: 51

loochooana loochooana Sawada, 1950b: 265 (Lachnosterna) A: JA (Ishigaki-jima)
loochooana okinawana Nomura, 1964a: 53 A: JA (Okinawa-shotō: Yogi, Okinoerabu-jima)
umebayashii Miyake, 1986a: 1

mizusawai Kobayashi, 1986a: 12 A: TAI
murzini Keith, 2006e: 56 A: SCH
nagpurensis Khan & Ghai, 1982a: 38 A: UP ORR
nigricollis Brenske, 1892d: 177 A: NP SD
rubricollis Moser, 1910b: 469

nubiliventris Bates, 1891a: 16 (Lachnosterna) A: HP
oblita Faldermann, 1835b: 459 (Ancylonycha) A: ANH BEI FE GAN HEB HEI HEN JIA SC SCH SHN
amplipennis Fairmaire, 1887b: 109 (Ancylonycha)

obscura Brenske, 1892d: 162 A: SD ORR

SCARABAEIDAE

269

occipitalis Bates, 1891a: 14 (Lachnosterna) A: HP
ochrogaster Bates, 1891a: 14 (Lachnosterna) A: HP SCH XIZ
omeia inexpectata Kobayashi, 1995b: 40 A: TAI
omeia omeia Chang, 1965a: 43 A: GAN SCH
ondulata Khan & Ghai, 1982a: 43 A: SD
ovata Chang, 1965a: 44 A: FUJ GUA GUX HAI HEN JIA JIX SHG
parallela Motschulsky, 1854d: 64 (Ancylonycha) A: ANH FE GAN GUI HEI HEN HUN JA NC SC SCH SHG
SHN ORR
morosa C.O. Waterhouse, 1875a: 104

picea C.O. Waterhouse, 1875a: 103 A: FE HEI JA NC SC
glossata Brenske, 1894a: 65
infantula S.I. Medvedev, 1951a: 308
izuensis Nomura, 1969: 73
rufopicea C.O. Waterhouse, 1875a: 103

pilifrons Moser, 1921: 63 A: JIX
pilipyga Chang, 1965a: 43 A: YUN
plumbea Hope, 1842: 60 [=1843a: 63, 1845: 8] A: GAN GUX TAI ZHE
problematica Brenske, 1899: 178 A: KA
protracta Bates, 1891b: 76 (Lachnosterna) A: SCH
pubifemorata Kobayashi, 1995b: 35 A: TAI
pulvinosa Sharp, 1878b: 172 (Lachnosterna) A: “Nordindien” ORR
rubida Chang, 1965a: 48 A: ZHE
rufescens Moser, 1912b: 433 A: HUB SCH
rufina Moser, 1912b: 433 A: GUA
sakurai Miyake & Yamaya, 1993: 35 A: NP
sauteri lutaoensis Kobayashi, 1990e: 84 A: TAI
sauteri sauteri Moser, 1913f: 436 A: GUX TAI
scutata Reitter, 1902b: 175 A: HKG
scutulata Dalla Torre, 1913: 206 [RN] A: HKG
scutata Moser, 1910b: 471 [HN]

semihirta Frey, 1971b: 210 A: UP
semiserrata Brenske, 1894a: 70 A: SHG
serrata Fabricius, 1781a: 35 (Melolontha) A: BT SD ORR
ceylonensis Moser, 1912b: 423
serraticollis Moser, 1912b: 423

seticollis Moser, 1913f: 426 A: NP SD
shibatai Nomura, 1977b: 91 A: TAI
excavata Miyake & Yamaya, 1993: 36

shishona Chang, 1965a: 44 A: YUN
sichotana Brenske, 1897d: 343 A: FE GAN HEI
amurensis Brenske, 1897d: 344

270

SCARABAEIDAE

similis S.I. Medvedev, 1951a: 299 A: GAN NO
simillima Moser, 1912b: 434 A: FUJ
stridulans Sharp, 1878b: 172 (Lachnosterna) A: PA
szechuanensis Chang, 1965a: 48 A: CHQ SCH
tokara Nakane, 1956a: 59 (Lachnosterna) A: JA (Tokara-rettō)
truncata Chang, 1965a: 47 A: JIA TAI
tuberculata Moser, 1908d: 358 A: GUA
ungulata Chang, 1965a: 47 A: FUJ
wangerbaoensis Y.-W. Zhang & Li, 1997: 779 A: CHQ SCH
yui Kobayashi, 1995b: 37 A: TAI
hualiensis Kobayashi, 1995b: 39
shizumui Kobayashi, 1995b: 39

yunnana Moser, 1912b: 428 A: YUN
genus Iranotrogus Nikolajev, 2006b: 53 type species Chioneosoma deserti Baraud, 1987
deserti Baraud, 1987b: 165 (Chioneosoma) A: IN
genus Lachnota Reitter, 1889g: 278 type species Melolontha henningii Fischer von Waldheim, 1823
henningii Fischer von Waldheim, 1823: pl. XXVIII [=1824: 213] (Melolontha) A: ES MG WS
genus Lasiexis Semenov & S.I. Medvedev, 1936: 107 type species Ancylonycha dilaticollis Ballion, 1871
dilaticollis Ballion, 1871a: 342 (Ancylonycha) A: AF IN KI KZ TD UZ
duplopunctata Reitter, 1902b: 180 (Brahmina)

glabrithoracica Keith, 2003a: 95 A: IN
kabulica Nikolajev & Kabakov, 1980: 226 (Lasiopsis) A: AF PA
vartiani Petrovitz, 1980: 617

kandaharica Nikolajev & Kabakov, 1980: 227 (Lasiopsis) A: AF
nurestanica Nikolajev & Kabakov, 1980: 227 (Lasiopsis) A: AF
pakistanus Keith, 2006e: 54 A: PA
persicola Petrovitz, 1980: 616 A: IN
primaeveris Semenov & S.I. Medvedev, 1936: 107 A: IN TM
reichenspergeri Balthasar, 1956b: 428 A: AF
genus Lasiopsis Erichson, 1847a: 658 type species Melolontha canina Zubkov, 1829
Askeptonycha Kraatz, 1883d: 154 type species Rhizotrogus sahlbergi Mannerheim, 1849
Brahminula S.I. Medvedev, 1951a: 246 type species Brahmina alatavica Semenov, 1925
Lasiobrahmina S.I. Medvedev, 1951a: 257 type species Lasiopsis duchoni Reitter, 1902

balgensis Murayama, 1941: 38 A: HEI
canina canina Zubkov, 1829: 156 (Melolontha) E: CT ST UK A: ES KZ WS
bergrothi Reitter, 1892c: 101
caucasica Semenov, 1900b: 124

SCARABAEIDAE
duchoni Reitter, 1902b: 168 A: KI KZ XIN
alatavica Semenov, 1925: 132 (Brahmina)

golovjankoi S.I. Medvedev, 1951a: 257 A: FE MG WS
sibirica S.I. Medvedev, 1951a: 259

koltzei Reitter, 1900b: 52 A: KI
kozlovi S.I. Medvedev, 1951a: 261 A: NEI NMO QIN XIN
kryzhanovskii Nikolajev & Kabakov, 1980: 223 A: AF
manchurica Murayama, 1941: 38 A: HEI NC SC
sahlbergi Mannerheim, 1849: 239 (Rhizotrogus) A: ES MG NO SC
bedeli Bergroth, 1892: 99 (Lachnota)
dahurica C.É. Blanchard, 1851: 144 (Rhizotrogus)

genus Lasiotropus Reitter, 1913e: 655 type species Lasiotropus poonensis Reitter, 1913
poonensis Reitter, 1913e: 656 A: “Tibet: Poo”
poonensis Frey, 1971a: 131 (Brahmina)

genus Medeotrogus Keith, 2001c: 76 type species Medeotrogus linnavuorii Keith, 2001
linnavuorii Keith, 2001c: 76 A: IN IQ
genus Megistophylla Burmesiter, 1855: 424 type species Megistophylla junghuhnii Burmeister, 1855
Hecatomnus Fairmaire, 1891f: ccii type species Hecatomnus grandicornis Fairmaire, 1891

grandicornis Fairmaire, 1891f: ccii (Hecatomnus) A: CHQ JIX HUN
genus Miridiba Reitter, 1902b: 170 type species Rhizotrogus trichophorus Fairmaire, 1891
subgenus Miridiba Reitter, 1902b: 170 type species Rhizotrogus trichophorus Fairmaire, 1891
Hippotrichia Arrow, 1948: 51 type species Hippotrichia hainana Arrow, 1948
Holotrochus Brenske, 1894a: 75 [HN] type species Holotrochus vestitus Brenske, 1894
Neodontocnema Arrow, 1948: 50 type species Ancylonycha coromandeliana C.É. Blanchard, 1851
Shangaia R. Lucas, 1920: 592 [RN] type species Holotrochus vestitus Brenske, 1894

abdominalis Hope, 1831: 23 (Melolontha) A: NP ORR
aequabilis Bates, 1891b: 75 (Lachnosterna) A: SCH
brancuccii Sabatinelli, 1983b: 211 (Neodontocnema) A: SD
castanea C.O. Waterhouse, 1875a: 104 (Holotrichia) A: NC NO SC JA YUN
frontalis Fairmaire, 1886: 325 (Rhizotrogus)

diversiceps Moser, 1913f: 445 (Holotrichia) A: SD
formosana Moser, 1910b: 470 (Holotrichia) A: LIA TAI
furcillata Keith & Sabatinelli, 2010: 340 A: NP
herteli Frey, 1971b: 220 (Holotrichia) A: “N.W. Indien”
hirsuta Itoh, 2001: 435 A: JA (Ishigaki-jima)
huesiotoi C.-L. Li & P.-S. Yang, 2015: 524 A: TAI (Green Island)
hybrida Moser, 1912b: 444 (Holotrichia) A: YUN XIZ ORR

271

272

SCARABAEIDAE

imitatrix Brenske, 1899: 178 (Holotrichia) A: SD
koreana Niijima & Kinoshita, 1923: 221 A: NC
kuraruana Nomura, 1977b: 89 A: TAI
newari Sabatinelli & Migliaccio, 1982: 107 (Neodontocnema) A: NP SD
sus Moser, 1912b: 443 (Holotrichia) A: HAI ORR
hainana Arrow, 1948: 51 (Hippotrichia)

taipei C.-C. Wang & C.-L. Li, 2015: 530 A: TAI
taoi C.-L. Li & C.-C. Wang, 2015b: 528 A: TAI (Lanyu)
trichophora Fairmaire, 1891f: cxcix (Rhizotrogus) A: BEI CHQ HEN HUB SCH SHG SHX ZHE
vestita Brenske, 1894a: 75 (Holotrochus)

wangi Y.-W. Zhang & Y. Li, 1997: 778 (Holotrichia) A: CHQ SCH
subgenus Pledina Reitter, 1902b: 173 type species Holotrichia sinensis Hope, 1842
sinensis Hope, 1842a: 60 [=1843a: 63, 1845: 8] (Holotrichia) A: HAI JIA NO TAI ZHE
cribellatus Fairmaire, 1891f: cc (Rhizotrogus)
sinae C.É. Blanchard, 1851: 139 (Ancylonycha)

genus Monotropus Erichson, 1847a: 658 type species Rhizotrogus nordmanni C.É. Blanchard, 1851
beauprei Pic, 1906b: 25 (Rhizotrogus) [DA] N: AG
brancoi Baraud, 1979a: 38 E: PT
fausti Semenov, 1899a: 95 E: AB ST
hirticollis P.H. Lucas, 1847b: 287 (Rhizotrogus) [DA] N: AG MO
humilis Marseul, 1878: 67 (Rhizotrogus) N: AG
laticollis Pérez Arcas, 1874: 126 (Rhizotrogus) E: SP
parvicollis Reitter, 1902b: 167

lusitanicus Baraud, 1976a: 80 E: PT
marquezi López-Colón & de la Puebla, 2014: 202 E: SP (Galicia)
nordmanni C.É. Blanchard, 1851: 142 (Rhizotrogus) E: RO ST UK
starcki Reitter, 1892f: 142

rotroui Peyerimhoff, 1931a: 6 (Rhizotrogus) [DA] N: AG MO
staudingeri Schaufuss, 1861: 48 E: SP
octus Báguena, 1960: 29

genus Onychosophrops Frey, 1972a: 114 type species Sophrops brevisetosa Frey, 1972
brevisetosus Frey, 1972a: 114 (Sophrops) A: FUJ
holosetosus Nomura, 1977b: 94 A: TAI
genus Panotrogus Reitter, 1902b: 187 type species Rhizotrogus myschenkovi Ballion, 1871
afghanus Petrovitz, 1965d: 691 A: AF IN PA
batillinus Bates, 1891a: 15 (Lachnosterna) A: HP
expansus Keith, 2003a: 96 A: PA
genieri Keith, 2006d: 436 A: NP

SCARABAEIDAE
grossepunctatus Keith, 2006e: 53 A: PA
hartmanni Keith, 2009d: 333 A: NP
heinzi Keith, 2001b: 10 A: AF PA
hirsutus Moser, 1913a: 52 (Holotrichia) A: BT HP SD
inexpectatus Keith, 2001b: 9 A: HP
kabulensis Keith, 2002d: 145 A: AF
kliri Keith, 2001b: 12 A: PA
miesseni miesseni Keith, 2009d: 336 A: NP
miesseni weiperti Keith, 2009d: 338 A: NP
myschenkovi Ballion, 187 1a: 341 (Rhizotrogus) A: AF IN KI KZ TD TM UZ
nadaii Keith, 2003a: 97 A: PA
pakistanus Keith, 2002d: 147 A: PA
pallidulus Gusakov, 2003a: 29 A: TD
petrovitzi Baraud, 1989a: 60 [RN] A: AF
afghanus Baraud, 1987b: 162 [HN]

petrovitzianus Keith, 2002d: 143 A: AF
pygmaeus Baraud, 1987b: 163 A: AF
schmidti Keith, 2006d: 435 A: NP UP
swatensis Keith, 2002d: 146 A: PA
genus Pectinichelus Ballion, 1871a: 340 type species Pectinichelus rhizotrogoides Ballion, 1871
subgenus Cryphaeobius Kraatz, 1882i: 313 type species Cryphaeobius brunneus Kraatz, 1882
brunneus Kraatz, 1882i: 313 A: KI KZ UZ TD
bucharicus Reitter, 1902b: 172 (Pectinichelus)

lopatini S.I. Medvedev, 1959: 869 A: TD (Pectinichelus)
simlanus Moser, 1913a: 63 (Brahmina) A: UP
subgenus Pectinichelus Ballion, 1871a: 340 type species Pectinichelus rhizotrogoides Ballion, 1871
ahrensi Keith, 2007a: 51 A: NP
chinensis Reitter, 1905b: 202 A: CH “Thibet”
emarginatus Keith & Sabatinelli, 2010: 339 A: PA
heinzi Keith, 2007c: 75 A: XIZ
kabakovi Nikolajev, 1976d: 695 A: AF
kopetzi Keith, 2009d: 348 A: NP
kryzhanovskii Nikolajev, 2004c: 871 A: AF
kryzhanovskii Nikolajev, 2007a: 52 [HN]

nasulatus Arrow, 1944c: 639 (Brahmina) A: XIZ
nepalensis Keith, 2007a: 54 A: NP
pakistanus Keith, 2003a: 98 A: PA
pommeranzi Keith, 2007a: 52 A: NP
rhizotrogoides Ballion, 1871a: 340 A: KI KZ TD UZ
hauseri Reitter, 1890p: 359 (Rhizocolax)

273

274

SCARABAEIDAE

medvedevi Protsenko, 1959: 174

sabatinellii Nikolajev, 2004c: 872 A: PA
schmidti Keith, 2007a: 53 A: NP
simikotensis Keith, 2009d: 346 A: NP
spinifer Keith, 2007a: 50 A: NP
tibetanus Arrow, 1944c: 640 (Brahmina) A: XIZ
tibetanicus Keith, 2007c: 75 A: XIZ
tuberculifrons Moser, 1926: 204 (Brahmina) A: KA
genus Pollaplonyx C.O. Waterhouse, 1875a: 105 type species Pollaplonyx flavidus C.O. Waterhouse, 1875
eriophorus Nomura, 1970: 64 A: TAI
flavidus C.O. Waterhouse, 1875a: 105 A: JA
genus Pseudopanotrogus Petrovitz, 1969b: 217 type species Pseudopanotrogus kashmirensis Petrovitz, 1969
(=Brahmina donckieri Brenske, 1892)
brevipilosus Keith, 2009d: 340 A: NP
donckieri Brenske, 1892b: 86 (Brahmina) A: KA NP SD
clypealis Moser, 1915a: 142 (Brahmina)
kashmirensis Petrovitz, 1969b: 217
nepalensis Frey, 1969c: 525 (Panotrogus)

extrarius Keith, 2005c: 26 A: NP ORR
holotrichius Keith, 2009d: 342 A: NP
kuluensis Moser, 1919: 352 (Brahmina) A: HP
pubescens Arrow, 1921a: 10 (Brahmina)

lassallei Keith, 2010c: 242 A: HP
obscurus Keith, 2007a: 44 A: NP
pilifer Moser, 1915a: 143 (Brahmina) A: BT “Tibet: Poo”
pusillus Arrow, 1921a: 11 (Brahmina) A: KA UP
rosettae Frey, 1971a: 129 (Brahmina) A: BT UP ORR
genus Pseudosymmachia Dalla Torre, 1913: 224 [RN] type species Symmachia chinensis Brenske, 1892
(=Metabolus impressifrons Fairmaire, 1887)
Ablotemus Paulsen & Smith, 2003: 254 [RN] type species Metabolus tumidifrons Fairmaire, 1887
Metabolus Fairmaire, 1887b: 107 [HN] type species Metabolus tumidifrons Fairmaire, 1887
Symmachia Brenske, 1892c: 151 [HN] type species Symmachia chinensis Brenske, 1892 (=Metabolus impressifrons
Fairmaire, 1887)

babai Kobayashi, 1990c: 55 (Metabolus) A: TAI
brevicollis Moser, 1915a: 147 (Metabolus) A: HUB
brevispina Nomura, 1977b: 100 (Metabolus) A: TAI
callosiceps Frey, 1972a: 110 (Metabolus) A: FUJ
costata Gu & Y.-W. Zhang, 1995: 10 (Metabolus) A: YUN

SCARABAEIDAE

275

clypeata Y.-W. Zhang, 1993: 275 (Metabolus) A: CHQ SCH
excisa Frey, 1972a: 110 (Metabolus) A: FUJ
flavescens Brenske, 1892c: 153 (Metabolus) A: BEI GAN
formosana Niijima & Kinoshita, 1927: 21 (Lasiopsis) A: TAI
fukiensis Frey, 1972a: 108 (Metabolus) A: FUJ
glabrosa Y.-W. Zhang, 1992a: 432 (Metabolus) A: HUN
impressifrons Fairmaire, 1887d: 316 (Metabolus) A: BEI JA NC SC
chinensis Brenske, 1892c: 151 (Symmachia)
niponicus Lewis, 1895c: 399 (Rhizotrogus)

kashmirensis Moser, 1915a: 148 (Metabolus) A: KA
longiuscula Y.-W. Zhang, 1993: 275 (Metabolus) A: HUB
manifestes manifestes Kobayashi, 1988a: 14 (Metabolus) A: TAI
manifestes nantouensis Kobayashi, 1988a: 15 (Metabolus) A: TAI
montana Kobayashi, 1987a: 15 (Metabolus) A: TAI
nitididorsis Kobayashi, 1995b: 40 (Metabolus) A: TAI
setifrons Moser, 1915a: 146 (Metabolus) A: YUN
setifrons Moser, 1915: 146 (Metabolus) A: YUN
similaris Y.-W. Zhang, 1992a: 432 (Metabolus) A: HUN
similis Kobayashi, 1987a: 15 (Metabolus) A: TAI
thibetana Moser, 1914a: 71 (Metabolus) A: “Thibet (Poo)”
tuberculifrons Nomura, 1977b: 100 (Metabolus) A: TAI
tumidifrons Fairmaire, 1887b: 107 (Metabolus) A: HEB ZHE
wulaiensis Kobayashi, 1988a: 13 (Metabolus) A: TAI
yunnanesis Gu & Y.-W. Zhang, 1995: 10 A: YUN
genus Pseudotrematodes Jacquelin du Val, 1860: 57 type species Rhizotrogus frivaldszkyi Ménétriés,
1836
frivaldszkyi Ménétriés, 1836a: 150 (Rhizotrogus) E: BU GR MC SB TR A: SY TR
carbonarius C.É. Blanchard, 1851: 148 (Trematodes)

genus Rhizotrogus Latreille, 1825: 371 type species Melolontha aestiva A.G. Olivier, 1789
aestivus A.G. Olivier, 1789: no 5: 17 (Melolontha) E: AB AL AN AR AU BE BH BU CR CT CZ FR GE GG GR HU IT
KZ LU MC MD NL PT RO SB SK SL SP ST SZ TR UK A: IN KZ SY TR
bimaculatus Herbst, 1790: 80 (Melolontha)
castaneipennis Pic, 1926b: 9
faldermanni Reiche, 1862b: 540
gracilis Frölich, 1792: 96 (Melolontha)
inanis Brahm, 1790: 85 (Melolontha)
incertus Mulsant, 1842: 428
lividigaster Mulsant, 1842: 429
maculicollis Zubkov, 1833: 322 (Melolontha) [HN]

276
nudicollis Dalla Torre, 1879: 12
subvittatus Mulsant, 1842: 429

alcaidei Baraud, 1971f: 472 N: MO
alicantinus Dieck, 1870: 118 E: SP
almeriensis Baraud, 1970b: 485 E: SP
angelesae Galante, 1981: 248 E: PT SP
balloyi M. Antoine, 1953: 217 N: MO
flavopennatus Baraud, 1973b: 45

breviceps Fairmaire, 1891f: cc A: HUB
camerosensis Báguena, 1955b: 285 E: SP
parcepunctatus Báguena, 1955b: 286
pilicauda Báguena, 1955b: 286
turolensis Báguena, 1955b: 289

camusi M. Antoine, 1960: 215 N: MO
carduorum Erichson, 1841a: 173 N: AG MO TU
batnensis Fairmaire, 1876b: 37
ignavus Fairmaire, 1860c: 437
maroccanus Brenske, 1889: 327
serraticollis P.H. Lucas, 1847b: 284
similis Marseul, 1878: 68
sternalis Marseul, 1878: 66
tingis Baraud, 1971f: 480

cariosicollis Fairmaire, 1873a: 336 N: AG MO
chevrolati Graëlls, 1858: 61 E: PT SP N: MO
bicoloripennis Baraud, 1975c: 196
plagiatus Marseul, 1878: 67

cicatricosus Mulsant, 1842: 433 E: FR GE IT SP SZ
auriculatus Báguena, 1955b: 289
parvicornis Baraud, 1972: 161
praecox Leoni, 1906: 62
rubidus Mulsant, 1842: 434

coiffaiti Baraud, 1979a: 35 E: PT
creticus Brenske, 1891c: 218 E: GR (Kríti)
flavicans C.É. Blanchard, 1851: 145 E: PT SP
viedmai Báguena, 1959: 213

gracilis Marseul, 1878: 68 N: AG
modestus Fairmaire, 1879a: 4 [=1879b: 254]

granatensis Báguena, 1955b: 283 E: SP
corduensis Báguena, 1955b: 284
floritae Compte, 1986: 72
tarifesis Báguena, 1955b: 284

SCARABAEIDAE

SCARABAEIDAE
guruguensis Baraud, 1971f: 480 N: MO
ifranensis Baraud, 1971f: 473 N: MO
iglesiasi Báguena, 1955b: 291 E: PT SP
impressifrons Fairmaire, 1891f: cc A: BEI JIA
latesulcatus Fairmaire, 1889a: 18 A: SCH
maculicollis A. Villa & G.B. Villa, 1833 (Melolontha) E: FR GE IT SP SZ
collaris Mulsant, 1842: 431
lineicollis Mulsant, 1842: 431
pallidifrons Mulsant, 1842: 431
thoracicus Mulsant, 1842: 431
vitticollis Mulsant, 1842: 431

marginipes Mulsant, 1842: 435 E: CR FR GE IT PT SL SP SZ
curtus Marseul, 1878: 66
foveolatus Bach, 1850: 16
geniculatus Chevrolat, 1865: 351
granulifer Rosenhauer, 1856: 119
pallidus Mulsant, 1842: 436
piemontanus Brenske, 1891b: 178
signatus Mulsant, 1842: 436
variolatus Fairmaire, 1880b: 239

mascarauxi Desbrochers des Loges, 1896: 166 E: FR PT SP
fernandesi Baraud, 1966a: 19
sulcifrons Báguena, 1955b: 290
zuzartei Baraud & Branco, 1982: 92

monticola C.É. Blanchard, 1851: 145 E: PT SP
cobosi Báguena, 1956a: 3
jeannei Baraud, 1970b: 486
monochromus Báguena, 1955b: 286
zapaterii Martínez y Sáez, 1873: 62

neglectus Pérez Arcas, 1865a: 27 E: PT SP
nevadensis Reitter, 1902b: 205 E: SP
pallescens Fairmaire, 1866a: 23 N: AG MO TU
pilidorsis Reitter, 1902b: 215
serripes Reitter, 1902b: 243 (Amphimallon)
vicarius Bedel, 1902: 212

pallidipennis C.É. Blanchard, 1851: 146 E: IT (Pantelleria) PT SP N: AG MO TU
anachoreta Rosenhauer, 1856: 120
areolus Reitter, 1902b: 203
gulosus Fairmaire, 1860c: 438
lepidus Schaufuss, 1869: 16
menorcanus Reitter, 1902b: 241 (Amphimallon)

277

278
occidentalis Escalera, 1914a: 172
phidias Reitter, 1902b: 216
signatitarsis Chevrolat, 1866: 102
vexator Schaufuss, 1869: 17

parvulus Rosenhauer, 1856: 122 E: SP
carthagenae Fairmaire, 1880b: 239
hidalgoi Pérez Arcas, 1872: 99
toletanus Báguena, 1955b: 288

pilosissimus Escalera, 1914a: 169 N: MO
ribbei Reitter, 1908a: 24 E: SP
romanoi Sabatinelli, 1975: 85 E: IT (Sicilia)
rosalesi Fairmaire, 1862: 549 E: SP
carmonensis Báguena, 1960: 30
epistomalis Báguena, 1955b: 289
gougeleti Fairmaire, 1869: 10

schaufussi Brenske, 1891c: 218 A: SY
siculus Baraud, 1970b: 479 E: IT (Calabria, Sicilia)
stigmaticollis Fairmaire, 1871a: 376 N: AG
subemarginatus Reiche, 1862b: 541 E: PT SP
rosinae Brenske, 1894b: 272
tornosii Pérez Arcas, 1872: 98

subsinuatus Báguena, 1955b: 287 E: SP
villiersi Baraud, 1970b: 480 E: PT SP
genus Rhizotrogus, nomen dubium
ballioni Brenske, 1888: 357 A: “Turkestan”
genus Sophrops Fairmaire, 1887b: 106 type species Sophrops parviceps Fairmaire, 1887
Eretusa Reitter, 1902b: 174 type species Ancylonycha nigra L. Redtenbacher, 1868
Microtrichia Brenske, 1900d: 343 type species Phytalus eurystomus Burmeister, 1855
Noanda Reitter, 1902b: 174 type species Holotrichia standfussi Brenske, 1892

acalcarius Gu & Z. Zhang, 1996: 23 A: GUA
acutangularis Moser, 1918: 243 (Microtrichia) A: JIX
analesetalis Gu & Z. Zhang, 1992: 257 A: YUN
brevisetalis Gu & Z. Zhang, 1992: 258 A: HUB
brunneus Moser, 1915d: 585 (Microtrichia) A: YUN
burmanicus Moser, 1913a: 67 (Brahmina) A: BT ORR
cantonensis Petrovitz, 1969b: 215 A: GUA
cardoni Brenske, 1892b: 108 (Brahmina) A: NP SD ORR
cephalotes Burmeister, 1855: 352 (Phytalus) A: GUX YUN ORR
chinensis Brenske, 1892b: 101 (Brahmina) A: CH ORR
coronarius Gu & Z. Zhang, 1992: 257 A: SCH YUN

SCARABAEIDAE

SCARABAEIDAE
cotesi Brenske, 1892b: 100 (Brahmina) A: NP SD
defreinai Frey, 1973a: 236 A: UP
formosanus Moser, 1913c: 273 (Microtrichia) A: TAI
hauseri Balthasar, 1932b: 20 A: YUN
heteropyga Moser, 1913a: 57 (Microtrichia) A: SD ORR
heydeni Brenske, 1892b: 108 (Brahmina) A: FE HEI LIA NC SC
himalayicus Brenske, 1892b: 113 (Brahmina) A: SD
kawadai kawadai Nomura, 1959b: 48 (Microtrichia) A: JA (Amami-Ôshima)
kawadai okinawaensis Nomura, 1977a: 77 A: JA (Okinawa-shotō)
konishii konishii Nomura, 1970: 67 A: JA (Iriomote-shima)
konishii yonaguniensis Nomura, 1970: 68 A: JA (Yonaguni-shima)
latus Frey, 1972a: 113 A: FUJ
latiusculus Nomura, 1977b: 93 A: TAI
longiflabella Gu & Y.-W. Zhang, 1995: 9 A: HUB
luridipennis Moser, 1915a: 134 (Microtrichia) A: YUN
parviceps Fairmaire, 1887b: 106 A: YUN
peronosporus Gu & Z. Zhang, 1992: 256 A: GUA HUN SCH
pexicollis Fairmaire, 1886: 325 (Schizonycha) A: YUN
planicollis Burmeister, 1855: 352 (Phytalus) A: HKG JIX ZHE SE ORR
niger L. Redtenbacher, 1868: 67 (Ancylonycha)

pruinosipyga Gu & Z. Zhang, 1996: 25 A: ANH
purkynei Balthasar, 1932b: 21 A: YUN
roeri Frey, 1972a: 112 A: FUJ
rugatus Moser, 1915d: 588 (Microtrichia) A: YUN
rugipennis Frey, 1972a: 111 (Metabolus) A: FUJ
setosus Brenske, 1892b: 99 (Brahmina) A: SD
stenocorpus Gu & Z. Zhang, 1996: 24 A: SCH
striatus Brenske, 1892b: 112 (Brahmina) A: CH NC SC
subrugatus Moser, 1921: 67 (Microtrichia) A: GUA
subrugosus Moser, 1915a: 133 (Microtrichia) A: JIX
taiwanus Nomura, 1977b: 93 A: TAI
takatoshii Itoh, 1990: 9 A: JA (Ryukyus: Miyako-jima)
thoracicus Brenske, 1892b: 109 (Brahmina) A: SD ORR
umbilicatus Frey, 1973a: 235 A: BT
yangbiensis Gu & Z. Zhang, 1996: 26 A: YUN
genus Stenosophrops Nomura, 1977b: 95 type species Stenosophrops longicornis Nomura, 1977
convexopyga Nomura, 1977b: 95 A: TAI
fuscicollis Nomura, 1977b: 96 A: TAI
longicornis Nomura, 1977b: 95 A: TAI
shukshanus Kobayashi, 1990c: 54 A: TAI
tuberculatus Kobayashi, 1986b: 9 A: TAI

279

280

SCARABAEIDAE

genus Tosevskiana Pavićević, 1985: 334 type species Tosevskiana inexpectata Pavićević, 1985
inexpectata Pavićević, 1985: 335 E: MC
machackovae Sehnal, 2004: 128 E: GR
sithoniensis Král, 1998: 41 (Amphimallon) E: GR
genus Trematodes Faldermann, 1835b: 372 type species Scarabaeus tenebrioides Pallas, 1781
grandis Semenov, 1902c: 346 A: NMO QIN
paratenebrioides J. K. Li, 1992: 77 A: JIL
potanini Semenov, 1902c: 345 A: GAN NMO SHX
tenebrioides Pallas, 1781: 9 (Scarabaeus) A: BEI ES HEB HEI JA LIA MG NEI
pallasii Faldermann, 1835b: 373

genus Xanthotrogus Reitter, 1902b: 212 type species Rhizotrogus sieversi Reitter, 1896
subgenus Xanthotrogus Reitter, 1902b: 212 type species Rhizotrogus sieversi Reitter, 1896
cariniclypeatus Keith, 2003a: 93 A: IN
fortis Reitter, 1902b: 212 (Rhizotrogus) A: UZ
leonhardi Reitter, 1909c: 119 (Rhizotrogus) A: IN
peratratus Reitter, 1909c: 119 (Rhizotrogus) A: IN
semnanensis Keith, 2003a: 94 A: IN
sieversi Reitter, 1896b: 33 (Rhizotrogus) A: IN TM
badkhyzicus Nikolajev, 2004: 40

tanyproctoides Reitter, 1906b: 34 (Amphimallon) A: IN TM
bicolor Petrovitz, 1970c: 40 (Madotrogus)

tadzhikorum Nikolajev, 2008b: 24 A: AF TD UZ
tekkensis Nikolajev, 2007d: 49 A: TM
subgenus Xestotrogus Reitter, 1902b: 211 type species Rhizotrogus validus Kraatz, 1884
validus Kraatz, 1884b: 233 (Rhizotrogus) A: IQ TR

tribe Schizonychini Burmeister, 1855

genus Canuschiza M. Lacroix, 1999: 87 type species Canuschiza insularis M. Lacroix, 1999
adah Sehnal, Král & Bezděk, 2014: 143 A: YE (Suqutra)
croton Sehnal, Král & Bezděk, 2014: 145 A: YE (Suqutra)
dracaena Sehnal, Král & Bezděk, 2014: 149 A: YE (Suqutra)
firmihin Sehnal, Král & Bezděk, 2014: 151 A: YE (Suqutra)
hagher Sehnal, Král & Bezděk, 2014: 155 A: YE (Suqutra)
insularis M. Lacroix, 1999: 89 A: YE (Suqutra)
jatropha Sehnal, Král & Bezděk, 2014: 161 A: YE (Suqutra)
minuta M. Lacroix, 1999: 88 A: YE (Suqutra)
skand Sehnal, Král & Bezděk, 2014: 163 A: YE (Suqutra)
genus Idionycha Arrow, 1932: 195 type species Idionycha excisa Arrow, 1932 (=Brahmina verticalis Moser, 1915)

281

SCARABAEIDAE
verticalis Moser, 1915a: 135 (Brahmina) A: HP NP ORR
excisa Arrow, 1932: 195

genus Redotus Fairmaire, 1860c: 443 type species Redotus rufulus Fairmaire, 1860
rufulus Fairmaire, 1860c: 444 N: AG
genus Schizonycha Dejean, 1833: 161 type species Scarabaeus globator Fabricius, 1781
algirina Fairmaire, 1876b: 38 N: AG MO TU
angustata yemenensis Arrow, 1944a: 198 A: OM SA YE
buettikeri Sabatinelli & Pontuale, 1998a: 115 A: AE OM SA
dumonti Peyerimhoff, 1926a: 106 N: AL MO TU
flavicornis Brenske, 1898c: 361 N: EG A: SA YE AFR
africana Burmeister, 1855: 269 [HN]
arabica Arrow, 1944a: 196
nigrofusca Arrow, 1900: 24

fuscescens C.É. Blanchard, 1851: 151 A: IN NP ORR
kadleci Bezděk & Sehnal, 2014: 38 A: YE
kocheri Baraud, 1980b: 257 N: MO
layeti Peyerimhoff, 1935: 147 N: AG
mucorea Fairmaire, 1887b: 108 [DA] A: YUN
nilotica C.É. Blanchard, 1851: 150 N: EG AFR
pygidialis Arrow, 1944a: 196 A: SA YE
ruficollis Fabricius, 1781: 39 (Melolontha) A: HP NP PA UP ORR
xanthodera C.É. Blanchard, 1851: 151

scorteccii Decelle, 1982: 382 A: OM YE
tenebrosa Fairmaire, 1887b: 108 [DA] A: YUN
genus Schizonycha, nomen dubium
diehli Muche, 1961: 107 A: SA

tribe Sericini Kirby, 1837

genus Ablaberoides C.É. Blanchard, 1850: 102 type species Ablaberoides aeneus C.É. Blanchard,
1850
Bielliana Baraud, 1980b: 61 type species Holoschiza abyssinica Brenske, 1902
Holoschiza van Lansberge, 1886b: 97 type species Holoschiza dentilabris van Lansberge, 1886
Trochaloserica Brenske, 1900c: 82 type species Trochaloserica festiva Brenske, 1902

abyssinicus Brenske, 1902a: 21 (Holoschiza) N: EG MO A: OM SA YE AFR
freyi Baraud, 1980b: 62 (Bielliana)
maroccanus Baraud, 1980b: 65 (Bielliana)
saatiensis Frey, 1973b: 244
somalicola Frey, 1969a: 77
yemenicus Baraud, 1980b: 62 (Bielliana)

282

SCARABAEIDAE

naviauxi Baraud, 1980b: 64 (Bielliana) A: SA YE
dhofarensis D. Ahrens, 2000e: 191 A: OM
genus Amiserica Nomura, 1974: 81 type species Amiserica rufidula Nomura, 1974
argentata Frey, 1975b: 223 (Serica) A: BT
babai Kobayashi, 1988b: 53 A: TAI
breviflabellata D. Ahrens, 2004a: 136 A: SD
costulata Frey, 1969c: 518 (Lasioserica) A: NP
flavolucida D. Ahrens, 2003b: 67 A: AP ORR
krausei D. Ahrens, 2004a: 130 A: NP
loi Kobayashi, 1991c: 215 A: TAI
longiflabellata D. Ahrens, 2004a: 134 A: SD
omeiensis D. Ahrens, 2003b: 71 A: SCH
pardalis Arrow, 1946a: 8 (Serica) A: YUN ORR
patibilis D. Ahrens, 2004a: 137 A: SD
recurva D. Ahrens & Fabrizi, 2009b: 258 A: AP
rufidula Nomura, 1974: 84 A: TAI
shizumui Kobayashi, 1980: 17 A: TAI
sparsesetosa D. Ahrens, 1999b: 239 A: AP SD
surda D. Ahrens, 2004a: 139 A: NP
taplejungensis D. Ahrens, 2004a: 133 A: NP
genus Anomalophylla Reitter, 1887b: 231 type species Anomalophylla tristicula Reitter, 1887
Melaserica Brenske, 1897e: 421 type species Melaserica thibetana Brenske, 1897 (=Anomalophylla tristicula Reitter,
1887)
Xorema Reitter, 1902b: 147 type species Anomalophylla thibetana Reitter, 1902 (=Anomalophylla tristicula Reitter, 1887)

bicolor D. Ahrens, 2005d: 22 A: SCH
dongchuanensis D. Ahrens, 2005d: 51 A: YUN
ganhaiziensis D. Ahrens, 2005d: 50 A: YUN
hispidulosa D. Ahrens, 2005d: 31 A: GAN
huashanica D. Ahrens, 2005d: 41 A: SHA
kangdingensis D. Ahrens, 2005d: 44 A: SCH
kozlovi S.I. Medvedev, 1952: 126 A: BEI NIN
liciata D. Ahrens, 2005d: 25 A: YUN
majori D. Ahrens, 2005d: 33 A: GAN
mandhatensis D. Ahrens, 2004a: 141 A: XIZ
mawi Arrow, 1946b: 274 (Microserica) A: SCH
morula D. Ahrens, 2005d: 35 A: YUN
moupinea Fairmaire, 1891b: cxcviii A: SCH
moxiensis D. Ahrens, 2005d: 57 A: SCH
obsuripennis D. Ahrens, 2005d: 20 A: SCH

SCARABAEIDAE

283

plagipennis D. Ahrens, 2005d: 59 A: YUN
qinlingensis D. Ahrens, 2005d: 53 A: SHA
stoetzneri D. Ahrens, 2005d: 29 A: SCH
subcarinata D. Ahrens, 2005d: 18 A: SCH YUN
subfastuosa D. Ahrens, 2005d: 27 A: SCH YUN
tristicula Reitter, 1887b: 232 A: GAN MG QIN SCH XIZ YUN
discoidalis Fairmaire, 1897f: 244 (Homaloplia)
thibetana Brenske, 1897g: 421 (Melaserica)
thibetana Reitter, 1902b: 147 [HN]

tsangpoana D. Ahrens, 2005d: 13 A: XIZ
vidua D. Ahrens, 2005d: 38 A: SCH
wulingshanica D. Ahrens, 2005d: 47 A: HUN
genus Apotriodonta Baraud, 1962c: 8 type species Triodonta hispanica Baraud, 1962
hispanica Baraud, 1962a: 192 E: SP
genus Archeohomaloplia Nikolajev, 1982: 286 type species Archeohomaloplia potanini Nikolajev, 1982
(=Homaloplia abbreviata Fairmaire, 1897)
Melanomaladera Miyake & Yamaya, 2001: 38 type species Melanomaladera yunnana Miyake & Yamaya, 2001

abbreviata Fairmaire, 1897f: 244 (Homaloplia) A: SCH
potanini Nikolajev, 1982: 286

frolovi D. Ahrens, 2011: 124 A: SCH
ganhaiziensis D. Ahrens, 2011: 131 A: YUN
hebashana D. Ahrens, 2011: 123 A: YUN
kalabi D. Ahrens, 2011: 128 A: SCH
medvedevi Nikolajev, 1982: 288 A: SCH
mingi D. Ahrens, 2011: 129 A: SCH
nikolajevi D. Ahrens, 2011: 125 A: SCH
safraneki D. Ahrens, 2011: 133 A: XIZ
yaregongensis D. Ahrens, 2011: 127 A: XIZ
yunnana Miyake & Yamaya, 2001: 38 (Melanomaladera) A: YUN
genus Autoserica Brenske, 1897e: 377 type species Serica piceorufa Fairmaire, 1888
comosa Brenske, 1898a: 301 [DA] A: SD
piceorufa Fairmaire, 1888b: 118 (Serica) A: BEI
umbilicata Brenske, 1898a: 303 [DA] A: SD
genus Calloserica Brenske, 1894a: 10 type species Calloserica tigrina Brenske, 1894
autumnalis D. Ahrens, 1999a: 193 A: NP
barabiseana D. Ahrens, 1999a: 194 A: NP
begnasia D. Ahrens, 1999a: 186 A: NP

284
bertiae D. Ahrens, 2000g: 816 A: SD XIZ
brendelli D. Ahrens, 1999a: 186 A: NP
cambeforti D. Ahrens, 2000g: 820 A: SD
capillata D. Ahrens, 2005e: 225 A: NP
chiplingensis D. Ahrens, 1999a: 192 A: NP
delectabilis D. Ahrens, 2000g: 819 A: NP
gosaikundensis D. Ahrens, 1999a: 195 A: NP
hingstoni D. Ahrens, 1999a: 192 A: XIZ
indrai D. Ahrens, 2004a: 57 A: XIZ
lachungensis D. Ahrens, 2000g: 817 A: SD
langtangica D. Ahrens, 1999a: 188 A: NP
manangensis D. Ahrens, 2005e: 228 A: NP
poggii D. Ahrens, 1995b: 137 A: NP
raksensis D. Ahrens, 2004a: 59 A: NP
rupthangensis D. Ahrens, 2004a: 58 A: NP
tigrina Brenske, 1894a: 10 A: SD
trisuliensis D. Ahrens, 1999a: 190 A: NP
zhangmuensis Liu & D. Ahrens, 2014b: 159 A: XIZ
genus Chrysoserica Brenske, 1897d: 355 type species Serica auricoma Brenske, 1896
angoris D. Ahrens, 2001a: 143 A: SD
auricoma Brenske, 1896a: 154 (Serica) A: SD NP YUN ORR
stebnickae D. Ahrens, 2001a: 140 A: HP NP PA
genus Eumaladera Nomura, 1967: 53 type species Maladera nitididorsis Nomura, 1967
fushanica D. Ahrens, 2002a: 75 A: TAI
gravida D. Ahrens, 2006b: 34 [RN] A: TAI
nitididorsis Kobayashi, 1991c: 215 (Amiserica) [HN]

latitibia Nomura, 1974: 106 (Maladera) A: TAI
nitididorsis nitididorsis Nomura, 1967: 54 (Maladera) A: JA
nitididorsis ootsuboi Kobayashi, 2009c: 55 A: JA (Tokunoshima Isl.)
nitidiceps Nomura, 1967: 55 (Maladera) A: TAI
opaciventris Moser, 1915c: 355 (Autoserica) A: SC
coreana J.I. Kim & A.Y. Kim, 2003b: 83 (Maladera)

paralatitibia D. Ahrens, 2002a: 78 A: TAI
planiuscula Nomura, 1974: 105 (Maladera) A: TAI
puliensis D. Ahrens, 2002a: 78 A: TAI
subrugata Moser, 1926: 198 (Autoserica) A: BEI
yakushimana Kobayashi, Kusui & Imasaka, 2006: 74 (Maladera) A: JA
genus Euserica Reitter, 1896e: 182 type species Melolontha mutata Gyllenhal, 1817

SCARABAEIDAE

SCARABAEIDAE
alcaidei Baraud, 1965a: 82 N: MO
alonsoi Ruiz & Ávila, 1993: 116 N: MO
brancobaraudi Ruiz & Ávila, 1995a: 97 N: MO
cambeloi Ruiz & Ávila, 1993: 112 N: MO
cobosi Baraud, 1965a: 87 N: MO
lucipeta Baraud, 1965a: 77 E: FR PT SP
mamorensis Baraud, 1965a: 86 N: MO
monstruosa Baraud & Branco, 1985: 628 N: MO
monticola Baraud, 1965a: 84 N: MO
mulsanti Brenske, 1902a: 61 E: SP
ariasi Brenske, 1897e: 364 [HN]

mutata Gyllenhal, 1817: 93 (Melolontha) E: PT SP
ariasi Mulsant & Rey, 1871a: 599 (Serica)

paenibaeticae Galante, 1987: 348 E: SP
pauliani López-Colón, 1988b: 257 E: SP
segurana Brenske, 1897e: 364 (Serica) E: SP
villarreali Baraud, 1975a: 42 E: SP
genus Gastromaladera Nomura, 1973c: 119 type species Gastroserica major Nomura, 1959
koyamai Hirasawa, 1985: 13 A: TAI
major Nomura, 1959b: 40 (Gastroserica) A: JA
nitida Nomura, 1974: 93 A: TAI
taitungensis Kobayashi, 1991a: 48 A: TAI
genus Gastroserica Brenske, 1897e: 412 type species Serica marginalis Brenske, 1894
angustula Brenske, 1897e: 416 A: GUI
asulcata D. Ahrens, 2000c: 79 A: GUI GUX JIX SCH YUN ORR
bicolor Niijima & Kinoshita, 1923: 36 A: TAI
bilyi D. Ahrens, 2000c: 113 A: YUN ORR
brevicornis Lewis, 1895c: 393 (Serica) A: JA
damingshanica Liu & D. Ahrens, 2014a: 103 A: GUX
fanjingensis D. Ahrens, 2000c: 104 A: GUI GUX SCH
fengduana Liu & D. Ahrens, 2014a: 93 A: SCH
guangdongensis D. Ahrens, 2000c: 90 A: FUJ GUA GUX
guizhouana D. Ahrens, 2000c: 89 A: GUI GUX HUN SCH
haoyui Liu & D. Ahrens, 2014a: 95 A: ZHE
haucki D. Ahrens, 2000c: 110 A: YUN ORR
herzi Heyden, 1887a: 264 (Serica) A: FUJ GUI GUX HEI HUN JIX NC SC SCH ZHE
higonia Lewis, 1895c: 394 (Serica) A: JA
hubeiana D. Ahrens, 2000c: 94 A: HUB SCH ZHE
impressicollis Fairmaire, 1891f: cxcvi (Serica) A: GUI GUX HUB JIX SCH

285

286

SCARABAEIDAE

jinxiuensis Liu & D. Ahrens, 2014a: 97 A: GUX
kabakovi D. Ahrens, 2002b: 66 A: YUN ORR
kucerai D. Ahrens & Pacholátko, 2003: 2 A: GAN GUX HUB SCH
liboensis Liu & D. Ahrens, 2014a: 99 A: GUI
marginalis Brenske, 1894a: 10 (Serica) A: FUJ GUA GUI GUX HAI HKG HUB HUN JIX SCH SHG SHN TAI
ZHE ORR
puncticollis Brenske, 1897e: 413

nigrofasciata Liu, D. Ahrens, Bai & X.-K. Yang, 2011: 32 A: FUJ GUI GUX
nikodymi D. Ahrens, 2000c: 102 A: FUJ HUN JIX
patkaiensis D. Ahrens, 2000c: 108 A: YUN ORR
shaanxiana D. Ahrens & Pacholátko, 2003: 4 A: SHA
sichuana D. Ahrens, 2000c: 95 A: SCH
similaris Kobayashi, 2015a: 27 A: TAI
sulcata Brenske, 1897e: 414 A: GUI GUX HUB HUN
wenzhui Liu & D. Ahrens, 2014a: 101 A: GUX
yingi D. Ahrens & Pacholátko, 2007: 137 A: GUX HAI YUN ORR
yunnanesis Liu, D. Ahrens, Bai & X.-K. Yang, 2011: 26 A: YUN
genus Gynaecoserica Brenske, 1897e: 356 type species Serica pellecta Brenske, 1896
Chaetoserica Brenske, 1897e: 355 type species Serica cymosa Brenske, 1896
Paragynaecoserica Khan & Ghai, 1982b: 61 type species Paragynaecoserica pubescens Khan & Ghai, 1982 (=Serica
cymosa Brenske, 1896)

alma D. Ahrens & Fabrizi, 2009a: 1511 A: YUN
amara D. Ahrens & Fabrizi, 2009a: 1522 A: YUN
aniniensis D. Ahrens & Fabrizi, 2011: 152 A: AP
annuliforceps D. Ahrens & Fabrizi, 2009b: 256 A: AP
arunachalensis D. Ahrens & Fabrizi, 2009a: 1524 A: AP
bocaki D. Ahrens & Fabrizi, 2009a: 1529 A: YUN
bomdilana D. Ahrens & Fabrizi, 2009a: 1531 A: AP
cymosa Brenske, 1896a: 154 (Serica) A: BT SD ORR
pubescens Khan & Ghai, 1982b: 63 (Paragynaecoserica)

dirangensis D. Ahrens & Fabrizi, 2009a: 1539 A: AP
exilis D. Ahrens & Fabrizi, 2009a: 1540 A: AP
gogonaica D. Ahrens, 1999a: 184 A: BT
hani Liu & D. Ahrens, 2014b: 160 A: XIZ
latesquamosa Frey, 1975b: 230 (Lasioserica) A: BT
lohitensis D. Ahrens & Fabrizi, 2009a: 1555 A: SCH XIZ ORR
lubosi D. Ahrens & Fabrizi, 2009a: 1559 A: AP
marginipes Brenske, 1896a: 155 (Serica) A: BT NP SD
obliqua D. Ahrens & Fabrizi, 2009a: 1570 A: YUN

SCARABAEIDAE

287

pellecta Brenske, 1896a: 154 (Serica) A: BT NP SD ORR
minutula Brenske, 1894a: 50 (Serica) [HN]

perdita D. Ahrens, 2004a: 105 A: SD
singhikensis D. Ahrens, 2004a: 109 A: SD
tawangensis D. Ahrens & Fabrizi, 2009b: 257 A: AP
tumba D. Ahrens, 2004a: 107 A: SD
variipennis variipennis Moser, 1916a: 153 (Lasioserica) A: NP SD ORR
variipennis wuermlii Frey, 1975b: 231 (Lasioserica) A: BT
victori D. Ahrens & Fabrizi, 2011: 151 A: BT
vogleri D. Ahrens & Fabrizi, 2009a: 1580 A: AP
yigongensis Liu & D. Ahrens, 2014b: 160 A: XIZ
genus Hellaserica Baraud & Nicolas, 1966: 119 type species Homaloplia elongata Reitter, 1887
elongata Reitter, 1887a: 138 (Homaloplia) E: GR
genus Hemiserica Brenske, 1894a: 57 type species Hemiserica nasuta Brenske, 1894
Rhynchosymmela Frey, 1974: 120 type species Rhynchosymmela pallida Frey, 1974 (=Hemiserica nasuta Brenske, 1894)

nasuta Brenske, 1894a: 57 A: UP ORR
pallida Arrow, 1945: 119
pallida Frey, 1974: 120 (Rhynchosymmela) [HN]

nasutella D. Ahrens, 2004a: 344 A: NP SD ORR
genus Hetamius Fairmaire, 1893g: cxlvii [=1893a: 123] type species Hetamius demaisoni Fairmaire, 1893
demaisoni Fairmaire, 1893g: cxlvii [=1893a: 123] N: AG EG
genus Hoplomaladera Nomura, 1974: 82 type species Hoplomaladera shibatai Nomura, 1974
hualiensis Kobayashi, 2001: 337 A: TAI
kurosawai Kobayashi, 2001: 337 A: TAI
monticola anmashana Kobayashi, 2001: 336 A: TAI
monticola monticola Kobayashi, 1991c: 219 A: TAI
saitoi Kobayashi, 1975: 9 A: TAI
shibatai Nomura, 1974: 92 A: TAI
taitungensis Kobayashi, 2015a: 30 A: TAI
genus Hymenochelus Reitter, 1890k: 263 type species Hymenoplia distincta Uhagon y Vedia, 1876
distinctus Uhagon y Vedia, 1876: 51 (Hymenoplia) E: PT SP
genus Hymenoplia Eschscholtz, 1830: 65 type species Melolontha strigosa Illiger, 1803
algirica Reitter, 1890k: 261 N: AG MO TU
antoinei Kocher, 1952a: 49 N: MO

288
arragonica Escalera, 1927: 119 E: SP
atlantea Baraud, 1985a: 503 N: MO
atlasica Escalera, 1914a: 180 N: MO
azrouensis Escalera, 1934a: 14 N: MO
castiliana Reitter, 1890k: 261 E: SP
castilianissima Báguena, 1954: 39 E: SP
chevrolati Mulsant, 1842: 471 E: FR SP
lugdunensis Dalla Torre, 1912: 66
omissa Escalera, 1925: 431
ramburi Heyden, 1875: 379

cinerascens Rosenhauer, 1856: 124 E: SP
clypealis Reitter, 1902b: 157 E: SP
carpetana Báguena, 1954: 34

escalerai Báguena, 1954: 42 E: SP
fulvescens Escalera, 1934a: 9 N: MO
fulvipennis C.É. Blanchard, 1850: 74 E: SP
mateui Báguena, 1954: 40
onubensis Báguena, 1954: 24

galaica Báguena, 1954: 35 E: SP
heydeni Reitter, 1890k: 262 N: AG MO
hirsutissima Báguena, 1956c: 64 N: MO
illigeri Pérez Arcas, 1874: 120 E: PT
jayenensis Báguena, 1954: 26 E: SP
ketamensis ketamensis Escalera, 1934a: 13 N: MO
ketamensis reymondi Kocher, 1952a: 49 N: MO
lata Heyden, 1870b: 180 E: SP
lecerfi Escalera, 1934a: 12 N: MO
lineolata C.É. Blanchard, 1850: 74 E: SP
lucusensis Escalera, 1934a: 10 N: MO
mairei Escalera, 1934a: 10 N: MO
meknesensis Escalera, 1934a: 7 N: MO
miegii Graëlls, 1858: 68 E: SP
fulvolineata Escalera, 1926b: 359
granatensis Escalera, 1926b: 358
korbi Reitter, 1890k: 262
moroderi Escalera, 1927: 118
nigripennis Báguena, 1954: 44

minuta Báguena, 1955b: 280 E: SP
oudjdensis Escalera, 1934a: 15 N: MO
pardoalcaidei López-Colón & de la Puebla, 2014: 197 E: SP (Madrid)
pilosissima Báguena, 1956c: 63 N: MO

SCARABAEIDAE

SCARABAEIDAE
pseudocinerascens Báguena, 1954: 25 E: SP
riffensis Escalera, 1925: 433 N: MO
roulleaui Báguena, 1956c: 62 N: MO
rufinoides Báguena, 1956c: 61 N: MO
rugosa Báguena, 1956c: 62 N: MO
rugulosa Mulsant, 1842: 473 E: PT SP
costulata Graëlls, 1858: 66
estrellana Heyden, 1870b: 182

sefroensis Báguena, 1956c: 60 N: MO
sericea Escalera, 1925: 432 N: MO
fezensis Escalera, 1934a: 8

sicula C.É. Blanchard, 1850: 74 E: IT
strigosa Illiger, 1803: 220 (Melolontha) E: PT SP
angusta Heyden, 1870b: 181
bifrons Eschscholtz, 1830: 65
cristata Graëlls, 1858: 67
discolor Báguena, 1955b: 280
hungarica C.É. Blanchard, 1850: 74

subcinerascens Escalera, 1925: 430 N: MO
theryi Escalera, 1934a: 14 N: MO
ungulata Peyerimhoff, 1949b: 280 N: MO
vulpecula Reitter, 1890k: 263 N: AG MO
zibana Peyerimhoff, 1949b: 280 N: MO
genus Lasioserica Brenske, 1896a: 155 type species Serica nobilis Brenske, 1894
Orchiserica Miyake & Yamaya, 2001: 38 type species Lasioserica brevipilosa Moser, 1919

antennalis Nomura, 1974: 83 [DA] A: TAI
beibengana Liu & D. Ahrens, 2014b: 161 A: XIZ
bispinosa D. Ahrens & Fabrizi, 2009b: 258 A: AP
braeti Brenske, 1896a: 155 A: SD
breviclypeata D. Ahrens, 1999a: 232 A: BT
brevipilosa Moser, 1919: 332 A: YUN
bumthangana D. Ahrens, 1999a: 225 A: BT
chitreana D. Ahrens, 1999a: 226 A: NP
dekensis D. Ahrens, 1999a: 231 A: SD
dolakhana D. Ahrens, 2004a: 116 A: NP
dolangsae D. Ahrens, 2004a: 117 A: NP
dragon Miyake & Yamaya, 2001: 36 [DA] A: YUN
godavariensis D. Ahrens, 1999a: 220 A: NP
hamifer D. Ahrens & Fabrizi, 2011: 153 A: BT
imminuta D. Ahrens & Fabrizi, 2011: 155 A: BT

289

290

SCARABAEIDAE

ilamensis D. Ahrens, 2000a: 6 A: NP
kuatunica D. Ahrens, 1996: 25 A: FUJ
kulbei D. Ahrens, 1999a: 218 A: NP
maculata bhutanica D. Ahrens, 1996: 19 A: BT
maculata galadrielae D. Ahrens, 1996: 18 A: AP SD
maculata jiriana D. Ahrens, 1996: 18 A: NP
maculata maculata Brenske, 1894a: 49 (Serica) A: NP UP
pilosa Moser, 1919: 332
umbrina C.É. Blanchard, 1850: 77 (Omaloplia)

modikholae D. Ahrens, 1996: 7 A: NP
nepalensis D. Ahrens, 1996: 5 A: NP
nobilis Brenske, 1894a: 48 (Serica) A: SD
calva Brenske, 1896a: 155

nudosa D. Ahrens, 1996: 15 A: NP
orlovi D. Ahrens, 2004a: 119 A: NP
pacholatkoi D. Ahrens, 2000a: 17 A: BT XIZ
pilosella Brenske, 1896a: 155 A: SD
piloselloida D. Ahrens, 1999a: 223 A: NP
pseudopilosella D. Ahrens, 1996: 12 A: NP
sabatinelii D. Ahrens, 1996: 7 A: NP
sikkimensis D. Ahrens, 1996: 4 A: SD
silkae D. Ahrens, 1996: 16 A: NP
smithi D. Ahrens, 2005c: 773 A: SD
soror D. Ahrens, 2004a: 121 A: SD
thoracica Brenske, 1898a: 311 A: SD
schereri Frey, 1962b: 132

tuberculiventris Moser, 1915a: 118 A: SHN
wittmeri D. Ahrens, 1999a: 217 A: NP
genus Leuroserica Arrow, 1946b: 281 type species Leuroserica fulgida Arrow, 1946
fulgida Arrow, 1946b: 281 A: SD
stemmleri Frey, 1975b: 225 (Microserica) A: AP BT
genus Maladera Mulsant & Rey, 1871a: 599 [=1871c: 343] type species Scarabaeus holosericeus Scopoli, 1772
subgenus Amaladera Reitter, 1896e: 188 type species Serica euphorbiae Burmeister, 1855
cobosi Baraud, 1964: 33 [DA] E: SP
espagnoli Baraud, 1964: 32 [DA] E: SP
euphorbiae Burmeister, 1855: 177 (Serica) E: ST A: AF KZ TD TM UZ
fusca Ballion, 1871a: 339 (Serica)

golovjankoi S.I. Medvedev, 1952: 147 A: TD TM

SCARABAEIDAE
longipennis Verdú, Mico & Galante, 1997: 116 [DA] E: SP
subgenus Aserica Lewis, 1895c: 394 type species Autoserica secreta Brenske, 1897
aureola Murayama, 1938: 19 [DA] A: NC SC TAI
chinensis Arrow, 1946b: 268 [DA] A: FUJ
cruralis Frey, 1972b: 170 (Autoserica) A: FUJ
secreta horaiana Nomura, 1974: 112 (Maladera) A: TAI
secreta secreta Brenske, 1897e: 431 (Autoserica) A: JA
subgenus Cephaloserica Brenske, 1900c: 79 type species Serica carinirostris Brenske, 1896
affinis C.É. Blanchard, 1850: 80 (Omaloplia) A: NP PA UP AFRi ORR
calcuttae Brenske, 1898a: 246 (Autoserica)
lahulensis Moser, 1919: 330 (Autoserica)
moreli Coquerel, 1866: 350 (Serica)

castanea Arrow, 1913c: 398 (Autoserica) E: GGi A: FE JA NC SC TRi NARi
korgei Petrovitz, 1967b: 332 (Serica)

cardoni Brenske, 1896a: 153 (Serica) A: AF NP PA SD UP SCH ORR
bhutanensis Frey, 1975b: 229 (Cephaloserica)
carinata Khan & Ghai, 1980: 24 (Autoserica)
nuristanica Petrovitz, 1965d: 684 (Serica)
tsienluana Brenske, 1897e: 394 (Serica)

formosae Brenske, 1898a: 210 (Autoserica) A: TAI
fusca Frey, 1972b: 170 (Autoserica) A: FUJ TAI
hiekei Frey, 1972b: 169 (Autoserica) A: FUJ
hongkongica Brenske, 1898a: 208 (Autoserica) A: HKG
insanabilis Brenske, 1894a: 37 (Autoserica) N: LB TU A: AE AF IN IS JO KA NP OM PA QA SA UP YE ORR
adjuncta Brenske, 1897e: 381 (Autoserica)
esfandiarii Petrovitz, 1970c: 38 (Autoserica)
matrida Argaman, 1986: 68 (Maladera)

iridescens C.É. Blanchard, 1850: 77 (Omaloplia) A: NP PA UP ORR
forsteri Frey, 1965a: 89 (Autoserica)
globus Burmeister, 1855: 165 (Serica)

lishana Miyake, 1989: 37 A: TAI
masumotoi Nomura, 1974: 112 A: TAI
nagporeana Brenske, 1898a: 241 (Autoserica) A: AF HP IN NP PA UP ORR
ignorata Petrovitz, 1965d: 685 (Serica)

nakamurai Miyake, 1991: 8 A: TAI
nanshanchiana Nomura, 1974: 111 (Maladera) A: TAI
omanica D. Ahrens, 2000e: 205 (Autoserica) A: OM
perniciosa Brenske, 1898a: 336 (Autoserica) A: NP SCH YUN ORR
rubescens Moser, 1908c: 325

rufocuprea C.É. Blanchard, 1850: 77 (Omaloplia) A: PA UP ORR

291

292

SCARABAEIDAE

carinirostris Brenske, 1896a: 153 (Serica)
fatifera Brenske, 1898a: 298 (Autoserica)
truncata Mittal, 1976: 180

satoi Nomura, 1961b: 73 (Maladera) A: JA
sauteri Moser, 1918: 213 (Autoserica) A: TAI
verticalis Fairmaire, 1888b: 118 (Serica) A: BEI HEB
koreana J.I. Kim & A.Y. Kim, 2003b: 90

subgenus Cycloserica Reitter, 1896e: 181 type species Cycloserica excisiceps Reitter, 1896
Brachyserica Brenske, 1897e: 386 type species Serica arenicola Solsky, 1876
Leucoserica Reitter, 1896e: 181 type species Serica arenicola Solsky, 1876

arenicola Solsky, 1876a: 394 (Serica) E: ST A: KZ TM UZ
diluta Reitter, 1896e: 185

caspia Faldermann, 1836: 370 (Omaloplia) E: KZ ST A: KZ
caspica Reitter, 1896e: 188

excisiceps Reitter, 1896e: 184 A: TM UZ
deserta L. N. Medvedev, 1956: 86

kerleyi D. Ahrens, 2004a: 337 A: NP SD
mechiana D. Ahrens, 2004a: 338 A: NP
paraquinquidens D. Ahrens, 2004a: 335 A: NP
quinquidens Brenske, 1896a: 153 (Serica) A: NP PA UP ORR
subgenus Eusericula Reitter, 1902b: 142 type species Serica modesta Fairmaire, 1881
ahrensi Keith, 2005g: 94 A: IN
farsensis Petrovitz, 1980: 615 A: IN
frischi Keith, 2010d: 161 A: IN
girardi Keith & D. Ahrens, 2002a: 196 A: IN
hivae Montreuil & Keith, 2009: 60 A: IN
hodkovae Nikodým & Král, 1998: 348 A: IN
jiraskovae Sehnal, 2008: 197 A: IN
mirzayansi Montreuil & Keith, 2009: 61 A: IN
modesta Fairmaire, 1881a: 83 (Serica) A: IS SI SY
pertusa Fairmaire, 1881a: 82 (Serica)

mofidii Montreuil & Keith, 2009: 62 A: IN
sinaica D. Ahrens, 2000d: 86 A: SI
subgenus Macroserica S.I. Medvedev, 1952: 139 type species Serica punctatissima Faldermann, 1835
allemandi Keith, 1998: 288 (Maladera) A: TR
angusta Baraud, 1990e: 300 (Maladera) A: TR
apfelbecki Petrovitz, 1969c: 392 (Maladera) A: AL BU GR MC
armeniaca Reitter, 1896e: 187 (Maladera) A: IN TR
attaliensis Petrovitz, 1969c: 396 (Maladera) A: TR
baluchestanica Petrovitz, 1971d: 225 (Maladera) A: IN
besucheti Baraud, 1990e: 297 (Maladera) A: TR

SCARABAEIDAE

293

bruschii Sabatinelli, 1977d: 107 (Maladera) A: TR
cerrutii Sabatinelli, 1977d: 111 (Maladera) A: CY
dentipenis Sehnal & Simandl, 2008: 496 A: CY
fusconitens Fairmaire, 1892b: 147 (Serica) A: SY
villiersi Petrovitz, 1969c: 391 (Maladera)

graeca Petrovitz, 1969c: 390 (Maladera) E: GR
iraqensis Keith, 2000b: 567 (Maladera) A: IN IQ
krueperi Petrovitz, 1969c: 388 (Maladera) E: GR
lodosi Baraud, 1975c: 191 (Maladera) A: TR
loebli Baraud, 1990e: 298 (Maladera) A: TR
lyciensis Petrovitz, 1969c: 395 (Maladera) A: TR
miliouensis Keith & Miessen, 2009: 179 A: CY
ollivieri Keith, 1998: 287 (Maladera) A: TR
phoenicica Petrovitz, 1969c: 394 (Maladera) A: LE
punctatissima Faldermann, 1835a: 279 (Serica) E: AB AR ST A: IN IS GG LE TR TM
elata Küster, 1849b: 39 (Serica)
iberica Kolenati, 1846: 26 (Omaloplia)
rugosa C.É. Blanchard, 1850: 77 (Omaloplia)

seleuciensis Petrovitz, 1969c: 397 (Maladera) A: TR
syriaca jeraschiensis Baraud, 1990e: 300 (Maladera) A: JO
syriaca syriaca Petrovitz, 1969c: 394 (Maladera) A: IS SY
taurica Petrovitz, 1969c: 389 (Maladera) A: TR
vingnai Sabatinelli, 1977b: 77 (Maladera) A: TR
wewalkai Petrovitz, 1969cc: 393 (Maladera) A: TR
subgenus Maladera Mulsant & Rey, 1871a: 599 [=1871c: 343] type species Scarabaeus holosericeus Scopoli, 1772
furcillata Brenske, 1897e: 400 (Autoserica) A: HUB
holosericea Scopoli, 1772: 77 (Scarabaeus) E: AL AU BE BH BU BY CR CT CZ DE FR GE HU IT LA LU MD ME
NL PL RO SB SK SL SP ST SV SZ UK A: FE JA HEI KZ WS
berolinensis Herbst, 1786: 155 (Melolontha)
fumosa Geoffroy, 1785: 11 (Scarabaeus)
fusca Mulsant, 1842: 462 (Omaloplia)
lamellata Geoffroy, 1785: 11 (Scarabaeus)
pellucida Schönherr, 1817: 179 (Melolontha)
pellucidula Sulzer, 1776: 18 (Scarabaeus)
sulzeri Fuessly, 1775: 3 (Scarabaeus)
variabilis Fabricius, 1777: 210 (Melolontha)

motschulskyi Brenske, 1897e: 370 (Serica) A: “Korea”
renardi Ballion, 1871a: 339 (Serica) A: FE HEI JA MG
nakayamai Murayama, 1938: 16 (Serica)
sibirica Brenske, 1897e: 372 (Serica)

schoenfeldti Murayama, 1937: 33 (Serica) A: NC SC

294

SCARABAEIDAE

spissigrada Brenske, 1897e: 430 (Serica) A: FE JA
kurentzovi Kalinina, 1978: 53

subgenus Omaladera Reitter, 1896e: 188 type species Amaladera diffinis Reitter, 1896 (=Serica orientalis
Motschulsky, 1858)
cariniceps Moser, 1915c: 341 (Autoserica) A: JA NC SC
clypeata Fairmaire, 1887b: 109 (Serica) A: YUN
dierli Frey, 1969c: 522 (Cephaloserica) A: NP PA UP
emmrichi D. Ahrens, 2004a: 227 A: NP UP
gardneri D. Ahrens, 2004a: 216 A: PA UP
himalayica Brenske, 1896a: 152 (Autoserica) A: SD
franklinmuelleri Moser, 1925: 56 (Autoserica)
spatiosa Brenske, 1898a: 377 (Autoserica)

himalayica immunda D. Ahrens, 2004a: 222 A: NP
himalayica incola D. Ahrens, 2004a: 223 A: NP
himalayica thakkholae D. Ahrens, 2004a: 225 A: NP
himalayica thimphuensis D. Ahrens, 2004a: 220 A: BT
joachimi D. Ahrens, 2004a: 229 A: NP
kawaii Kobayashi, 2010: 68 A: JA (Iriomote Isl.)
lignicolor Fairmaire, 1887b: 110 (Serica) A: FUJ HUB NC SC SCH TAI ZHE
fusiana Murayama, 1934: 35 (Aserica)
laboriosa Brenske, 1897e: 399 (Autoserica)
tenebrosa Frey, 1972b: 166 (Serica)

orientalis Motschulsky, 1858a: 33 (Serica) A: BEI FE GAN GUI HEB HEI HUB JA MG NC SC TAI
cavifrons Reitter, 1896e: 188
diffinis Reitter, 1896e: 188
famelica Brenske, 1897e: 391 (Serica)
pekingensis Brenske, 1897e: 366 (Serica)

oshimana Nomura, 1962: 38 A: JA
okinawana Nomura, 1964a: 51
sakishimana Nomura, 1964a: 51

simlana Brenske, 1898a: 302 (Autoserica) A: HP NP PA UP
sprecherae D. Ahrens, 2004a: 214 A: AP BT
stevensi D. Ahrens, 2004a: 231 A: SD
yasutoshii Nomura, 1974: 109 A: TAI
genus Maladera, species incertae sedis
allopruinosa D. Ahrens, 1998: 42 (Serica) [RN] A: NP SD ORR
pruinosa Burmeister, 1855: 165 (Serica) [HN]

amamiana Nomura, 1959b: 40 A: JA
bagmatiensis D. Ahrens, 2004a: 242 A: NP
bengalensis Brenske, 1898a: 250 (Autoserica) A: NP UP ORR
bhutanensis Frey, 1975b: 224 (Autoserica) A: BT SD

SCARABAEIDAE
lata Frey, 1975b: 224 (Lasioserica)

botrytibia Nomura, 1974: 111 A: TAI
brevistylis D. Ahrens, 2004a: 249 A: NP
chiruwae D. Ahrens, 2004a: 300 A: NP
conspicua D. Ahrens, 2004a: 293 A: SD
consularis D. Ahrens & Fabrizi, 2009b: 269 A: AP
davidis Brenske, 1898a: 213 (Autoserica) A: SCH
detersa Erichson, 1834: 239 (Serica) A: CH
diversipes Moser, 1915c: 346 (Autoserica) A: CH
duvivieri Brenske, 1896a: 152 (Serica) A: SD
ejimai Miyake & Imasaka, 1987: 137 A: JA
fatigata D. Ahrens, 2004a: 292 A: NP
ferruginea Kollar & L. Redtenbacher, 1844: 525 (Serica) A: KA NP PA UP
ferruginea C.É. Blanchard, 1850: 81 (Omaloplia) [HN]

festina Brenske, 1898a: 296 (Autoserica) A: SD
flammea Brenske, 1898a: 209 (Autoserica) A: HKG
fumosa Brenske, 1898a: 306 A: UP ORR
pilula Sharp, 1903: 470 (Serica)
tristicula Moser, 1915b: 157 (Autoserica)

fuscipes Moser, 1915c: 353 (Autoserica) A: JIX ORR
futschauana Brenske, 1897e: 404 (Autoserica) A: FUJ
gibbiventris Brenske, 1897e: 396 (Autoserica) A: HUB TAI
gopaldharae D. Ahrens, 2004a: 286 A: SD
gorkhae D. Ahrens, 2004a: 297 A: NP
haldwaniensis D. Ahrens, 2004a: 195 A: SD UP
harmonica Brenske, 1897e: 405 (Autoserica) A: FUJ
hayashii Hirasawa, 1991: 176 A: TAI
hiemalis Kobayashi, 2014b: 38 A: JA (Ishigaki-jima)
holzschuhi D. Ahrens, 2004a: 257 A: UP
horii Kobayashi, 2010: 71 A: JA (Ishikagi-jima)
howdeni D. Ahrens, 2003a: 5 A: TAI
imasakai Miyake & Yamaya, 1995: 34 A: JA
imbella Reitter, 1898b: 119 (Maladera) A: “Turkestan”
impubis D. Ahrens, 2004a: 324 A: SD
impressithorax Nomura, 1973c: 130 A: JA (Amami-Ôshima)
inadai Kobayashi, 2010: 72 A: JA (Amami-Ôshima Isl.)
infuscata Moser, 1915c: 340 (Autoserica) A: NC SC SCH SHN
koreana Moser, 1919: 330 (Autoserica)
okamotoi Murayama, 1938: 18 (Aserica)

invenusta Moser, 1918: 212 (Autoserica) A: TAI
japonica japonica Motschulsky, 1861a: 15 (Serica) A: JA

295

296
japonica korai Miyake & Yamaya, 1995: 35 A: NC
kamiyai Sawada, 1937: 27 (Serica) A: JA
kawaharai Kobayashi, 2010: 67 A: JA (Okinawa-shotō)
kobayashii Nomura, 1974: 106 A: TAI
krali D. Ahrens, 2004a: 301 A: SD
kreyenbergi Moser, 1918: 211 (Autoserica) A: SHN TAI
kubotoi Nomura & Kobayashi, 1979: 7 A: TAI
kumei Kobayashi, 1990e: 84 A: TAI
kunigami Kobayashi, Kusui & Imasaka, 2006: 69 A: JA
kusuii Miyake, 1986: 2 A: TAI
lanae Kobayashi, 2012: 21 A: TAI
levis Frey, 1972b: 169 (Serica) A: FUJ
liotibia Nomura, 1974: 110 A: FUJ TAI
liukueiensis Kobayashi, 1985a: 12 A: TAI
lugubris Brenske, 1896a: 152 (Serica) A: NP PA UP ORR
lukjanovitschi S.I. Medvedev, 1966: 1576 A: MG
kaszabi Frey, 1967: 145

maedai Nomura, 1974: 108 A: TAI
marginella Hope, 1831: 24 (Serica) A: HP KA NP SD UP ORR
semirufa Brenske, 1898a: 320 (Microserica)

merkli D. Ahrens, 2004a: 269 A: PA UP
mirabilis Brenske, 1894a: 41 (Serica) A: SHG
modestula Brenske, 1902a: 62 (Autoserica) [RN] A: NP SD XIZ ORR
modesta Brenske, 1896a: 153 (Serica) [HN]

mussooriensis D. Ahrens, 2004a: 328 A: UP
nomurai Hirasawa, 1991: 175 A: TAI
obscurata Moser, 1915c: 354 (Autoserica) A: YUN
okinawaensis Kobayashi, 1978: 14 A: JA
okinoerabuana Kobayashi, 1978: 15 A: JA (Okinoerabu-jima) TAI
hiranoi Miyake, 1986a: 3

opaca Frey, 1975b: 229 (Cephaloserica) A: BT NP SD ORR
opacifrons Fairmaire, 1891f: cxcv (Serica) A: HUB TAI
opima Nomura, 1967: 52 (Maladera) A: TAI
paris D. Ahrens, 2004a: 306 A: SD
parva Moser, 1908c: 326 (Cephaloserica) [DA] A: TAI ORR
kojimai Miyake, 1991: 9 (Cephaloserica)

phuntsholingensis D. Ahrens, 2004a: 197 A: BT
piceola Moser, 1915c: 352 (Autoserica) A: YUN
pokharae D. Ahrens, 2004a: 240 A: NP UP
polunini D. Ahrens, 2004a: 264 A: NP
poonensis Khan & Ghai, 1980: 25 (Autoserica) A: PA

SCARABAEIDAE

SCARABAEIDAE
prenai D. Ahrens, 2004a: 316 A: AP BT SD
raptiensis D. Ahrens, 2004a: 193 A: NP UP ORR
rufodorsata Fairmaire, 1888a: 19 (Homaloplia) A: FUJ
rufoplagiata Fairmaire, 1893e: 305 (Omaloplia) A: BT NP SD YUN ORR
birmanica Brenske, 1898a: 343 (Autoserica)

rufotestacea Moser, 1915c: 342 (Autoserica) A: YUN
rustica Brenske, 1896a: 153 (Serica) A: UP ORR
saitoi Niijima & Kinoshita, 1927: 79 (Autoserica) A: TAI ORR
schenklingi Moser, 1918: 214 (Autoserica) A: IN KA NP ORR
schereri Frey, 1975a: 183 (Autoserica) A: SD
senta Brenske, 1897e: 405 (Autoserica) A: SHG TAI
sericella Brenske, 1898a: 335 (Autoserica) A: BT NP SD ORR
setosa Brenske, 1896a: 153 (Serica) A: NP UP ORR
cinerea Brenske, 1898a: 279 (Autoserica)

shihzitouensis Kobayashi, 2002: 316 A: TAI
shouchiana Kobayashi & C. K. Yu, 1997: 179 A: TAI
sikkimensis Brenske, 1898a: 305 (Autoserica) A: NP SD ORR
siniaevi D. Ahrens, 2004a: 284 A: AP BT SD XIZ ORR
sinica Hope, 1842: 60 [=1843a: 63, 1845: 9] (Serica) A: ZHE TAI
siwakiliana D. Ahrens, 2004a: 322 A: NP UP
spatulata D. Ahrens, 2006c: 415 A: NP
spei D. Ahrens, 2004a: 299 A: SD
spinifemorata Kobayashi, 1993: 69 A: TAI
straminea Moser, 1915b: 156 (Autoserica) A: JIX
stridula Brenske, 1897e: 401 (Autoserica) A: SHN TAI ORR
subspinosa Brenske, 1898a: 243 (Autoserica) A: NP UP ORR
taiwana Nomura, 1974: 109 A: TAI ORR
taiyal Kobayashi, 2002: 315 A: TAI
takaoana Niijima & Kinoshita, 1927: 11, 80 (Autoserica) A: JA
taoyuanensis Kobayashi, 1991c: 212 (Maladera) A: TAI
thomsoni Brenske, 1894a: 55 (Serica) A: HP KA NP PA UP ORR
assamensis Brenske, 1898a: 333 (Autoserica)
phthisica Brenske, 1902a: 69 (Cephaloserica)

tienchihna Kobayashi, 1988b: 60 A: TAI
tienhsiangensis Kobyayshi, 2007b: 24 A: TAI
tokunoshimana Kobayashi, Kusui & Imasaka, 2006: 72 A: JA
trichotibialis D. Ahrens, 2004a: 311 A: NP SD
tridentipes Nomura, 1974: 107 A: TAI
tumida D. Ahrens, 2004a: 282 A: NP PA UP
uhligi D. Ahrens, 2004a: 246 A: SD
umbratica Brenske, 1898a: 303 (Autoserica) A: SD

297

298

SCARABAEIDAE

ventralis Brenske, 1897e: 397 (Autoserica) A: JIX
watanabei Kobayashi, 2002: 313 A: TAI
weigeli D. Ahrens, 2004c: 212 A: NP
wolfgangdierli D. Ahrens, 2004: 295 [RN] A: NP
dierli Frey, 1972c: 188 (Autoserica) [HN]

yakushimana Kobayashi, Kusui & Imasaka, 2006: 71 A: JA
yaeyamana Nomura, 1963a: 37 A: TAI
genus Microserica Brenske, 1894a: 52 type species Microserica quadrimaculata Brenske, 1894
arunensis D. Ahrens, 1998: 45 A: NP
bhutanensis Frey, 1975b: 227 A: BT ORR
bisignata Nomura, 1974: 99 A: TAI
cechovskyi D. Ahrens, 1999b: 243 A: NP
diversicornis Moser, 1915c: 387 A: YUN
fukiensis Frey, 1972b: 174 (Gastroserica) A: FUJ TAI
inornata Nomura, 1974: 98

gandakiensis D. Ahrens, 1998: 39 A: NP
hainana Brenske, 1898a: 216 (Microtrichia) A: HAI
hispidula Frey, 1975b: 226 A: BT
hiulca Brenske, 1897e: 419 A: GUX JIX
interrogator Arrow, 1946b: 267 A: NP UP
lucens D. Ahrens & Fabrizi, 2009b: 267 A: AP
marginata Brenske, 1896a: 154 (Serica) A: NP SD
problematica Arrow, 1946b: 269 (Aserica)
unctiuscula Brenske, 1898a: 304 (Autoserica)

martensi D. Ahrens, 1998: 43 A: NP
myagdiana D. Ahrens, 1998: 46 A: NP UP
nigropicta Fairmaire, 1891f: cxcvii (Serica) A: HUB
nikkonensis Brenske, 1897e: 435 A: JA
nitidipyga Nomura, 1974: 99 A: FUJ TAI
formosana Kobayashi, 1991c: 217 (Gastroserica)

pedongensis D. Ahrens, 1998: 50 A: SD
pruinosa Hope, 1831: 24 (Serica) A: HP NP UP XIZ
subaenea C.É. Blanchard, 1850: 79 (Omaloplia)

roeri Frey, 1972b: 171 A: FUJ
schawalleri D. Ahrens, 1998: 43 A: NP
schulzei D. Ahrens, 1998: 32 A: NP
sigillata Brenske, 1897e: 417 A: FUJ
steelei D. Ahrens, 2004a: 185 A: XIZ ORR
truncata Brenske, 1898a: 305 (Autoserica) A: SD
cribriceps Moser, 1915b: 183

SCARABAEIDAE
varians Moser, 1915c: 392 A: YUN
viridicollis Arrow, 1913b: 192 A: SD ORR
genus Neomaladera Baraud, 1965a: 90 type species Brachyphylla barbara P.H. Lucas, 1846
barbara P.H. Lucas, 1846d: pl. 25 (Brachyphylla) N: AG MO TU
lanata Escalera, 1925: 428 (Serica)
mecheriensis Pic, 1898c: 97 (Serica)
pardoi Cobos, 1961: 91 (Euserica)
pilicollis Burmeister, 1855: 178 (Serica)
substriata Küster, 1849b: 44 (Homaloplia)
susiana Escalera, 1913a: 31 (Euserica)

grandiclava Escalera, 1925: 422 (Serica) N: MO
labriquei Keith, 2012: 53 N: MO
longitarsis Escalera, 1925: 424 (Serica) N: MO
iridescens Escalera, 1925: 427 (Serica)

neglecta Baraud, 1965a: 96 N: MO
rugosula Escalera, 1914a: 177 (Euserica) N: MO
teluetica Escalera, 1914a: 176 (Euserica) N: MO
genus Neoserica Brenske, 1894a: 44 type species Serica ursina Brenske, 1894
abnormoides D. Ahrens, Liu, Fabrizi, Bai & X.-K. Yang, 2014: 65 A: ZHE ORR
ailaoshanica Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 78 A: YUN
anonyma Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 63 A: YUN
arunachalana D. Ahrens & Fabrizi, 2009b: 259 A: AP
bairailingshanica D. Ahrens, Liu, Fabrizi, Bai & X.-K. Yang, 2014: 72 A: YUN
baishuiensis D. Ahrens, Fabrizi & Liu, 2015: 1124 A: SCH YUN
baoshana D. Ahrens, Fabrizi & Liu, 2015: 1099 A: YUN
biuncinata D. Ahrens, Fabrizi & Liu, 2015: 1108 A: SCH
calvoides Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 60 A: YUN
cardamomensis D. Ahrens, Liu, Fabrizi, Bai & X.-K. Yang, 2014: 68 A: YUN ORR
changrae D. Ahrens, 2004a: 166 A: BT
chayuensis Liu & D. Ahrens, 2014b: 164 A: XIZ
crenatolineata D. Ahrens & Fabrizi, 2009b: 262 A: AP
curvipenis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014a: 110 A: SCH YUN
daweishanica D. Ahrens, Fabrizi & Liu, 2014: 90 A: YUN
dundai D. Ahrens, Fabrizi & Liu, 2015: 1119 A: SCH
emeishanensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014a: 108 A: SCH
euyunnanica D. Ahrens, Liu, Fabrizi, Bai & X.-K. Yang, 2014: 44 A: YUN
exoleta D. Ahrens & Fabrizi, 2009b: 264 A: AP
fanjingshanica D. Ahrens, 2003c: 203 A: GUI
fukiensis Frey, 1972b: 176 (Lasioserica) A: FUJ

299

300
gaoligongshanica D. Ahrens, Fabrizi & Liu, 2014: 88 A: YUN
ganhaiziana D. Ahrens, Fabrizi & Liu, 2015: 1106 A: SCH YUN
garlangensis D. Ahrens, 2004a: 157 [DA] A: NP
guangpingensis D. Ahrens, Fabrizi & Liu, 2014: 80 A: YUN
gulinqingensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 61 A: YUN
heishuiana D. Ahrens, Fabrizi & Liu, 2015: 1096 A: YUN
huangi D. Ahrens, Liu, Fabrizi, Bai & X.-K. Yang, 2014: 75 A: XIZ
igori D. Ahrens, Fabrizi & Liu, 2014: 98 A: YUN
incompta D. Ahrens & Fabrizi, 2009b: 265 A: AP
inops D. Ahrens & Fabrizi, 2009b: 263 A: AP
inspergata D. Ahrens & Fabrizi, 2009b: 262 A: AP
jiangxiensis D. Ahrens, Liu, Fabrizi, Bai & X.-K. Yang, 2014: 46 A: GUX JIX
jinggangshanica D. Ahrens, 2003c: 209 A: JIX
jiulongensis D. Ahrens, Fabrizi & Liu, 2014: 92 A: SCH
kanphantensis D. Ahrens, Liu, Fabrizi, Bai & X.-K. Yang, 2014: 47 A: YUN ORR
kaskiensis D. Ahrens, 2004a: 163 [DA] A: GUI
kereni D. Ahrens, Fabrizi & Liu, 2015: 1098 A: SCH YUN
koelkebecki Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 74 A: SC
kuaichangensis D. Ahrens, 2003c: 191 A: GUI
kunmingensis D. Ahrens, Fabrizi & Liu, 2015: 1101 A: GUI YUN
lateriuncinata D. Ahrens, Fabrizi & Liu, 2015: 1126 A: SCH
leiboensis D. Ahrens, Fabrizi & Liu, 2015: 1128 A: SCH
lenangensis D. Ahrens & Fabrizi, 2009b: 260 A: XIZ
liangi Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 58 A: YUN
lincangensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014a: 102 A: YUN
ludingensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014a: 95 A: SCH
lushuiana Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014a: 106 A: YUN
luxiensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 76 A: YUN
luzhouana D. Ahrens, Fabrizi & Liu, 2015: 1121 A: GUI SCH
matura D. Ahrens, 2004a: 160 [DA] A: NP
menghaiensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 56 A: GUX YUN
mengi Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 72 A: YUN
ningyuanensis D. Ahrens, Fabrizi & Liu, 2015: 1094 A: SCH YUN
nykli D. Ahrens, Fabrizi & Liu, 2015: 1110 A: SCH
obscura C.É. Blanchard, 1850: 79 (Omaloplia) [DA] A: CH
parausta D. Ahrens, Fabrizi & Liu, 2015: 1103 A: YUN
parursina D. Ahrens, 2003c: 189 A: SHN
pilosula Moser, 1915c: 377 A: YUN
plurilamellata D. Ahrens, Fabrizi & Liu, 2014: 82 A: YUN
probsti D. Ahrens, 2004a: 159 [DA] A: NP ORR
pseudovulpes D. Ahrens, Fabrizi & Liu, 2015: 1113 A: SCH
pubiforceps D. Ahrens, 2004a: 165 [DA] A: NP SD ORR

SCARABAEIDAE

SCARABAEIDAE

301

rangshuiensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014a: 104 A: GUI
rubellula D. Ahrens, Fabrizi & Liu, 2015: 1104 A: HUB
rutilans D. Ahrens & Fabrizi, 2009b: 261 A: XIZ
ruzickai D. Ahrens, Fabrizi & Liu, 2015: 1089 A: YUN
sangangana D. Ahrens, 2003c: 206 A: FUJ JIX
sapaensis D. Ahrens, Fabrizi & Liu, 2014: 94 A: YUN ORR
septemfoliata Moser, 1915c: 381 A: YUN
shennongjiaensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014a: 100 A: GUI HEN HUB SHX YUN
shibingensis D. Ahrens, 2003c: 198 A: GUI
shinkaisiensis D. Ahrens, Fabrizi & Liu, 2015: 1091 A: SCH
sichuanica D. Ahrens, Fabrizi & Liu, 2015: 1117 A: SCH
silvestris Brenske, 1902a: 61 [DA] A: ZHE
taipingensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014: 70 A: SHA
tianeana Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014a: 98 A: GUX
trifida D. Ahrens, Liu, Fabrizi, Bai & X.-K. Yang, 2014: 50 A: YUN
unicolor Frey, 1972b: 175 (Lasioserica) A: FUJ
ursina Brenske, 1894a: 49 (Serica) A: SHG
usta D. Ahrens, Fabrizi & Liu, 2015: 1115 A: SCH
weibaoshanica Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014a: 96 A: YUN
weishanensis D. Ahrens, Fabrizi & Liu, 2015: 1092 A: YUN
weishanica D. Ahrens, Fabrizi & Liu, 2014: 74 A: YUN
xiaguanensis D. Ahrens, Fabrizi & Liu, 2015: 1122 A: YUN
xizangensis Liu & D. Ahrens, 2014b: 165 A: XIZ
yangjiapingensis D. Ahrens, Fabrizi & Liu, 2015: 1087 A: YUN
yanzigouensi D. Ahrens, Fabrizi & Liu, 2014: 77 A: SCH
yaoi D. Ahrens, Liu, Fabrizi, Bai & X.-K. Yang, 2014: 34 A: GUX YUN
yingjiangensis D. Ahrens, Liu, Fabrizi, Bai & X.-K. Yang, 2014: 52 A: YUN
yongkangensis Liu & D. Ahrens, 2015: 2389 A: ZHE
zheijangensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 66 A: FUJ HUB YUN ZHE
zhibenshanica Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 67 A: YUN
zongyuani Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014b: 54 A: SCH
genus Neosericania Miyake & Yamaya, 1994: 37 type species Neosericania excisoclypeata Miyake & Yamaya,
1994
excisoclypeata Miyake & Yamaya, 1994: 37 A: TAI
haboensis D. Ahrens, 2002a: 66 A: TAI
nitida Kobayashi, 1985a: 10 (Lasioserica) A: TAI
genus Nepaloserica Frey, 1965a: 88 type species Nepaloserica procera Frey, 1965
archolabrata D. Ahrens & Sabatinelli, 1996: 238 A: NP XIZ
baehri D. Ahrens & Sabatinelli, 1996: 214 A: NP

302
baglungensis D. Ahrens, 2012: 311 A: NP
barbara D. Ahrens & Sabatinelli, 1996: 232 A: SD
brevipes D. Ahrens & Sabatinelli, 1996: 231 A:SD
bruschii D. Ahrens & Sabatinelli, 1996: 222 A: NP
fabriziae D. Ahrens, 1999a: 207 A: SD
ganeshi D. Ahrens, 1999a: 201 A: NP XIZ
goomensis D. Ahrens, 1999a: 199 A: SD
hartmanni D. Ahrens, 1999a: 203 A: NP
helambuensis D. Ahrens & Sabatinelli, 1996: 221 A: NP
induwae D. Ahrens, 1999a: 211 A: NP
jumlaica D. Ahrens, 1999a: 204 A: NP
lamjungi D. Ahrens, 1999a: 205 A: NP
longispina D. Ahrens, 1999a: 212 A: NP
manasluensis D. Ahrens, 2004a: 69 A: NP
migliaccioi D. Ahrens & Sabatinelli, 1996: 218 A: NP
muelleri muelleri D. Ahrens & Sabatinelli, 1996: 224 A: NP
muelleri tuberculata D. Ahrens, 2004a: 68 A: NP
mustangia D. Ahrens & Sabatinelli, 1996: 226 A: NP
nielamuensis Liu & D. Ahrens, 2014b: 163 A: XIZ
phulcokiensis D. Ahrens & Sabatinelli, 1996: 236 A: NP
perrecondita D. Ahrens, 2004a: 66 A: NP
pewaensis D. Ahrens, 2012: 313 A: NP
procera procera Frey, 1965a: 88 A: NP XIZ
procera rufescens Frey, 1965b: 182 A: NP
richardsonae D. Ahrens, 2012: 308 A: NP
rufobrunnea D. Ahrens, 1999a: 201 A: BT
sankhuwasabhae D. Ahrens & Sabatinelli, 1996: 230 A: NP
schmidti D. Ahrens & Sabatinelli, 1996: 215 A: NP
similis Frey, 1969c: 520 A: NP
telbrungensis D. Ahrens, 1999a: 208 A: NP
thimphui D. Ahrens & Sabatinelli, 1996: 234 A: BT
vignai D. Ahrens & Sabatinelli, 1996: 212 A: NP
vilya D. Ahrens & Sabatinelli, 1996: 227 A: NP
yeti D. Ahrens, 1999a: 213 A: NP
genus Nipponoserica Nomura, 1973c: 120 type species Serica similis Lewis, 1895
Pseudomaladera Nikolajev, 1980: 40 [HN] type species Serica koltzei Reitter, 1897

babai Kobayashi, 1991a: 47 A: TAI
chinensis Moser, 1915b: 144 (Serica) A: SHN
daisensis Sawada, 1937: 24 (Serica) A: JA
lewisi Chapin, 1938: 68 (Serica)

SCARABAEIDAE

SCARABAEIDAE
dahongshanica D. Ahrens, 2005a: 276 A: HUB
elliptica Murayama, 1938: 17 (Serica) A: SC
gomandana Nomura, 1976b: 187 A: JA
koltzei Reitter, 1897h: 214 (Serica) A: FE GAN HEI SC
opacicarina J.I. Kim & A.Y. Kim, 2003a: 76

kunitachiana Nomura, 1976b: 190 A: JA
laferi Nikolajev, 1980: 41 (Pseudomaladera) A: FE
peregrina Chapin, 1938: 68 (Serica) A: JA NARi
pubiventris Nomura, 1976b: 189 A: JA
similis Lewis, 1895c: 391 (Serica) A: JA
setiventris Nomura, 1976b: 188

sulciventris D. Ahrens, 2004b: 9 A: HUB SCH SHA
takeuchii Hirasawa, 1991: 171 [DA] A: TAI
genus Omaloplia Schönherr, 1817: 178 type species Melolontha ruricola Fabricius, 1775
subgenus Omaloplia Schönherr, 1817: 178 type species Melolontha ruricola Fabricius, 1775
Brachyphylla Mulsant, 1842: 465 Melolontha ruricola Fabricius, 1775
Homaloplia Erichson, 1847: 700 type species Melolontha ruricola Fabricius, 1775

baraudi baraudi Galante, 1985: 500 (Homaloplia) A: IN TR
baraudi pontica Ádám, 1994b: 25 E: BU GR MC RO TR A: TR
caeca Baraud, 1965d: 420 (Homaloplia) E: BU GR MC ME SB
balcanica Ádám, 1994b: 23

cerrutii Sabatinelli, 1977d: 113 (Homaloplia) E: AL BU GR TR A: TR
corcyrae Baraud, 1965d: 407 (Homaloplia) E: AL BH BU CR GR KO MC ME RO
depilis J. Müller, 1910: 130 (Homaloplia) E: GR
diabolica Reitter, 1887a: 139 (Homaloplia) A: SY TR
ursina Fairmaire, 1892b: 147 (Homaloplia)

erebea Baraud, 1965d: 426 (Homaloplia) E: TR A: TR
erythroptera I. Frivaldszky, 1835: 260 E: AL BH BU CR GR HU MC MD ME RO SB TR UK
carbonaria C.É. Blanchard, 1850: 76 (Brachyphylla)
transsylvanica Bielz, 1850: 180 (Homaloplia)

gibbosa Baraud, 1965d: 422 (Homaloplia) E: AL GR MC ME TR A: TR
macedoniae Baraud, 1965d: 423 (Homaloplia)

gobbii Piattella & Sabatinelli, 1993: 196 (Homaloplia) E: GR
graeca Reitter, 1887a: 137 (Homaloplia) E: GR MC
attica Baraud, 1965d: 424 (Homaloplia)
oetaea Baraud, 1965d: 425 (Homaloplia)

hericia Chobaut, 1907: 175 (Homaloplia) E: BU FR GR RO TR
majuscula Baraud, 1965d: 417 (Homaloplia)
rozneri Ádám, 1994b: 27

illyrica Baraud, 1965d: 411 (Homaloplia) E: AL BH BU CR GR KO MC ME RO SB

303

304

SCARABAEIDAE

iris Reitter, 1887a: 139 (Homaloplia) E: BU GR MC ME RO SB
kiritschenkoi S.I. Medvedev, 1952: 163 (Homaloplia) E: UK
lonae Schatzmayr, 1923: 7 (Homaloplia) E: AL AU BH CR IT MC ME SL
minuta Brenske, 1887a: 137 (Homaloplia) E: GR
mutilata Fairmaire, 1892b: 147 (Homaloplia) A: TR
corpulenta J.R. Sahlberg, 1908: 64 (Homaloplia)

nigromarginata Herbst, 1786: 155 (Melolontha) A: AU DE CT CZ FR GE HU LA LT PL RO SB SK SL ST SZ UK
A: ES MG WS
occidentalis Baraud, 1965d: 415 (Homaloplia)

polita Baraud, 1965d: 423 (Homaloplia) E: GR
pubipennis Rössner & D. Ahrens, 2004: 34 E: GR
ruricola nicolasi Baraud, 1965d: 110 (Homaloplia) E: AL FR IT
epirota Baraud, 1965d: 410 (Homaloplia)

ruricola ruricola Fabricius, 1775: 38 (Melolontha) E: AB AU BE BH BU CR CZ FR GB GE GG GR HU IT KO LA
MC ME RO SB SK SL SP ST SZ TR A: IN
arnoldii S.I. Medvedev, 1952: 168 (Homaloplia)
atrata Geoffroy, 1785: 11 (Scarabaeus)
floricola Laicharting, 1781: 41 (Melolontha)
fritschi Reitter, 1905b: 201 (Homaloplia)
humeralis Fabricius, 1775: 40 (Melolontha)
intermedia Abeille de Perrin, 1895d: ccix (Homaloplia)
marginata Fuessly, 1775: 3 (Scarabaeus)
populi Gistel, 1857a: 68 (Homaloplia)
pruinosa Küster, 1849b: 42 (Homaloplia)
svanetica S.I. Medvedev & Dzambazishvilli, 1973: 457 (Homaloplia)
tergestina Baraud, 1965d: 407 (Homaloplia)

settorum Uliana, 2014: 2 A: TR
subgenus Acarina Baraud, 1965d: 395 type species Scarabaeus spireae Pallas, 1773
labrata Burmeister, 1855: 156 (Homaloplia) A: IN IQ IS JO LE SY TR
subsinuata Burmeister, 1855: 156 (Homaloplia)

ottomana Baraud, 1965d: 433 (Homaloplia) A: TR
spireae longiclava Baraud, 1965d: 434 (Homaloplia) A: GR
spireae spireae Pallas, 1773: 719 (Scarabaeus) E: AR AU BU CT CZ GG GR HU KZ RO SB SK ST UK A: IN KZ
TR WS
adulta Reitter, 1887a: 138 (Homaloplia)
alternata Küster, 1849b: 43 (Homaloplia)
hirta Gebler, 1830: 109
limbata Krynicki, 1832: 126 (Homaloplia)
puberula Gebler, 1847: 465
sieversi Reitter, 1897e: 124 (Homaloplia)
margaritacea Ádám, 1994b: 27 (Acarina)

SCARABAEIDAE

305

genus Oxyserica Brenske, 1900c: 54 type species Oxyserica pygidialis Brenske, 1900
Parvulomaladera D. Ahrens, 1995a: 45 type species Microserica simlana Brenske, 1902 (=Oxyserica pygidialis
Brenske, 1900)

bimaculata Hope, 1831: 24 (Serica) A: NP
janetscheki Frey, 1969c: 521 (Microserica)

brancuccii D. Ahrens, 2001b: 223 (Microserica) A: NP
darjeelingia Brenske, 1898a: 319 (Microserica) A: SD
hellmichi Frey, 1965a: 91 (Microserica) A: NP
kanchenjungae D. Ahrens, 1995a: 51 (Microserica) A: NP
kurseongana Moser, 1915b: 182 (Microserica) A: SD
longefoliata Frey, 1965a: 92 (Microserica) A: NP XIZ
pygidialis annapurnae D. Ahrens, 1995a: 46 (Microserica) A: NP XIZ
pygidialis pygidialis Brenske, 1900c: 54 A: NP PA UP
simlana Brenske, 1902a: 62 (Microserica)

varia Frey, 1975b: 226 (Microserica) A: BT
genus Pachyserica Brenske, 1897e: 420 type species Pachyserica rubrobasalis Brenske, 1897
albosignata Moser, 1915a: 114 (Serica) A: FUJ SHN
albosquamosa Brenske, 1898a: 356 A: AP SD UP
ambiversa D. Ahrens, 2004a: 43 A: NP
bituberculata D. Ahrens, 2006a: 525 A: XIZ ORR
cipingensis D. Ahrens, 2006a: 505 A: JIA
conspersa D. Ahrens, 2006a: 509 A: GUI
darjeelingensis D. Ahrens, 2004a: 51 A: SD
gracilis D. Ahrens, 2004a: 35 A: NP UP
himalayensis D. Ahrens, 2004a: 37 A: BT NP SD
horishana Niijima & Kinoshita, 1927: 3, 78 (Serica) A: TAI
huangliangensis D. Ahrens, 2006a: 512 A: YUN
jendeki D. Ahrens, 2004a: 45 A: SD ORR
marmorata C.É. Blanchard, 1850: 77 (Omaloplia) A: NP UP
minax D. Ahrens & Fabrizi, 2009b: 255 A: XIZ
nantouensis Kobayashi & C. K. Yu, 1993c: 357 A: TAI
similis Kobayashi & C. K. Yu, 1993c: 358

nepalica D. Ahrens, 2004a: 39 A: NP
numensis D. Ahrens, 2004a: 41 A: NP
olafi D. Ahrens, 2004a: 32 A: NP SD
pellingensis D. Ahrens, 2004a: 52 A: SD
rubrobasalis Brenske, 1897e: 420 A: FUJ JIA
squamifera Frey, 1972b: 162 A: FUJ
stabilis D. Ahrens, 2004a: 47 A: SD
yanoi Nomura, 1959b: 41 A: JA

306
genus Paramaladera Nomura, 1974: 83 type species Paramaladera major Nomura, 1974
aserrata Kobayashi & Nomura, 1979: 9 A: TAI
cariniprinceps Kobayashi & Nomura, 1979: 8
wulaiana Kobayashi, 1980: 19

kiyoyamai Nomura, 1974: 100 A: TAI
babai Kobayashi, 1991a: 49

major Nomura, 1974: 100 A: TAI
makiharai Nomura, 1974: 100 A: TAI
masumotoi Hirasawa, 1991: 173 A: TAI
pishana Kobayashi, 1991a: 50 A: TAI
rufofusca Nomura, 1974: 100 A: TAI
simillima Kobayashi, 1985a: 11 A: TAI
genus Paraserica Reitter, 1896e: 183 type species Serica grisea Motschulsky, 1866
grisea Motschulsky, 1866: 171 (Serica) A: JA
taiwana Nomura & Kobayashi, 1979: 11 A: TAI
genus Paratriodonta Baraud, 1962c: 7 type species Melolontha morio Fabricius, 1792
algirica Reitter, 1889h: 285 (Triodonta) N: AG
alicantina Reitter, 1890c: 68 (Triodonta) E: SP
atlantis Baraud, 1962d: 171 (Triodonta) N: MO
cinctipennis P.H. Lucas, 1846d: pl. 25 (Hymenoplia) E: IT N: AG TU
puberula Erichson, 1847a: 703 (Triodonta)
pumila Burmeister, 1855: 151 (Triodonta)

demoflysi Normand, 1949: 87 (Triodontella) N: TU
kocheri Baraud, 1962d: 169 (Triodonta) N: MO
lentula Normand, 1949: 86 (Triodontella) N: AG
maroccana Brenske, 1889: 325 (Triodonta) N: MO
maura Baraud, 1962d: 172

morio Fabricius, 1792b: 178 (Melolontha) N: AG TU
aterrima P.H. Lucas, 1847b: 292 (Hymenoplia)

mskalica Escalera, 1914a: 179 (Triodontella) N: MO
normandi Baraud, 1962d: 177 (Triodonta) N: TU
ochroptera Erichson, 1841a: 173 (Omaloplia) N: AG MO
olivieri C.É. Blanchard, 1850: 75 (Omaloplia) N: AG EG LB A: IS SY
delicatula Fairmaire, 1881a: 83 (Serica)

proboscidea Illiger, 1802a: 169 (Melolontha) N: MO
romana Brenske, 1890b: 85 (Triodonta) E: IT
solitudinis Baraud, 1985a: 625 N: AG
temperei Baraud, 1962c: 50 N: AG
tripolitana Brenske, 1889: 326 (Triodonta) N: LB TU A: IS

SCARABAEIDAE

SCARABAEIDAE

307

unguicularis Erichson, 1841a: 174 (Omaloplia) N: AG MO
vicina Baraud, 1962c: 56 N: AG
genus Pseudomaladera Nomura, 1974: 82 type species Pseudomaladera nitidifrons Nomura, 1974
nitidifrons Nomura, 1974: 96 A: TAI
genus Pseudosericania Kobayashi, 1980: 15 type species Pseudosericania gibiventris Kobayashi, 1980
gibiventris Kobayashi, 1980: 15 A: TAI
makiharai Hirasawa, 1991: 172 A: TAI
nitididorsis Nomura, 1974: 89 (Nipponoserica) A: TAI
quadrifoliata Nomura & Kobayashi, 1979: 10 (Nipponoserica) A: TAI
genus Serica W.S. Macleay, 1819: 146 type species Scarabaeus brunneus Linnaeus, 1758
subgenus Serica W.S. Macleay, 1819: 146 type species Scarabaeus brunneus Linnaeus, 1758
Ophthalmoserica Brenske, 1897e: 356 type species Serica thibetana Brenske, 1897
Podoserica Breit, 1912: 202 type species Podoserica reitteri Breit, 1912
Trichoserica Reitter, 1896e: 181 type species Trichoserica fulvopubens Reitter, 1896

almorae D. Ahrens, 1999b: 280 A: UP
angustatotibialis D. Ahrens, 1999b: 303 A: BT SD
arborea D. Ahrens, 1999b: 210 A: NP
assequens D. Ahrens & Fabrizi, 2009b: 249 A: AP
basumtsoensis D. Ahrens, 2005b: 62 A: XIZ
becvari D. Ahrens, 2005b: 70 A: YUN
behluhdinensis D. Ahrens, 2005b: 90 A: SCH
benesi D. Ahrens, 2005b: 28 A: GAN QIN SCH SHA
bhaktai D. Ahrens, 1999b: 294 A: NP
bidentata D. Ahrens, 1999b: 292 A: NP
bidigitata D. Ahrens, 2000f: 287 A: NP
bolm D. Ahrens, 2005b: 44 A: YUN
boops C.O. Waterhouse, 1875a: 101 A: JA
niijimai Kontkanen, 1956: 60

brevitarsis brevitarsis Nomura, 1972: 111 A: JA
brevitarsis rectipes Nomura, 1972: 111 A: JA
brunnea Linnaeus, 1758: 352 (Scarabaeus) E: AL AU BE BH BU BY CR CT CZ DE FI FR GB GE HU IR IT LA LT
LU MC MD NL NR NT PL RO SB SK SL SP ST SV SZ UK A: ES KZ MG WS
berardii Luigioni, 1929: 72
erythrocephala V. Petagna, 1787: 5 (Melolontha)
fulva De Geer, 1774: 277 (Scarabaeus)
fulvescens Geoffroy, 1785: 10 (Scarabaeus)

chasilakhae D. Ahrens, 1999b: 296 A: BT
chautarana D. Ahrens, 2005b: 122 A: NP

308

SCARABAEIDAE

chuttana D. Ahrens, 1999b: 278 A: NP
daliangshanica D. Ahrens, 2005b: 35 A: SCH
daliensis D. Ahrens, 2005b: 78 A: YUN ORR
degenensis D. Ahrens, 2005b: 55 A: YUN
doiinthanonensis D. Ahrens, 2005b: 73 A: YUN ORR
eberti Frey, 1965a: 90 (Ophthalmoserica) A: BT NP UP
echigoana Nakane & Baba, 1960: 6 (Ophthalmoserica) A: JA
emeishanica D. Ahrens, 2005b: 38 A: HUB SCH
exhausta D. Ahrens & Fabrizi, 2011: 141 A: BT
filitarsata D. Ahrens, 1999b: 301 A: BT
foobowana Sawada, 1937: 20 A: JA
fulvopubens Reitter, 1896e: 184 (Trichoserica) A: FE
furcata D. Ahrens, 2005b: 68 A: SCH YUN
fusifemorata Nomura, 1974: 90, 91 A: TAI
gansuensis Miyake & Yamaya, 2001: 35 A: GAN
gonggashanica D. Ahrens, 2005b: 37 A: SCH
grahami D. Ahrens, 2005b: 33 A: SCH
guidoi D. Ahrens, 1999b: 268 A: UP NP SD XIZ
heishuensis D. Ahrens, 2005b: 98 A: YUN
heydeni Reitter, 1896e: 184 (Trichoserica) A: GAN HUB QIN SCH SHA
reitteri Breit, 1912: 202

hirsuta J.I. Kim & A.Y. Kim, 2003a: 74 A: SC
incognita D. Ahrens, 1999b: 282 A: NP
incurvata Nomura, 1971: 67 (Trichoserica) A: JA
inexspectata Kontkanen, 1956: 60 (Ophthalmoserica) A: JA
nitidiceps Nomura, 1971: 69 (Trichoserica)
pilosa Nomura, 1971: 66 (Trichoserica)

jaegeri D. Ahrens, 1999b: 273 A: NP
kalabi D. Ahrens, 2005b: 48 A: SCH
kangdingensis D. Ahrens, 2005b: 46 A: SCH
karafutoensis honshuensis Nomura, 1972: 130 A: JA
karafutoensis karafutoensis Niijima & Kinoshita, 1923: 219 A: FE (Kuril’skie ostrova, Sakhalin) JA
khajiaris Mittal, 1988: 157 A: AP BT KA NP PA SD UP XIZ
khasiana Moser, 1918: 210 (Ophthalmoserica) A: NP UP ORR
kingdoni D. Ahrens, 1999b: 300 A: GAN SCH XIZ YUN
kumaonensis D. Ahrens, 1999b: 291 A: UP
lama D. Ahrens, 1999b: 260 A: SD
leigongshanica D. Ahrens, 2005b: 31 A: GUI
litangensis D. Ahrens, 2005b: 51 A: SCH
meiguensis D. Ahrens, 2005b: 42 A: SCH
mianningensis D. Ahrens, 2005b: 80 A: SCH

SCARABAEIDAE
minshanica D. Ahrens, 2005b: 87 A: GAN
montreuili D. Ahrens, 2005b: 59 A: GAN QIN SCH XIZ YUN
moupinensis Fairmaire, 1889a: 18 (Homaloplia) A: SCH
muliensis D. Ahrens, 2005b: 66 A: SCH
mureensis D. Ahrens, 1999b: 271 A: NP SD
narya D. Ahrens, 1999b: 298 A: SD
nebulosa D. Ahrens, 1999b: 259 A: NP
nepalensis Frey, 1969c: 519 (Ophthalmoserica) A: NP XIZ
nigromaculosa Fairmaire, 1891f: cxcvi A: SCH
nigrovariata Lewis, 1895c: 392 A: JA
nigripennis Sawada, 1937: 15

nipponica Nomura, 1959b: 46 (Ophthalmoserica) A: JA
nitididorsis nitididorsis Nomura, 1971: 70 (Trichoserica) A: JA
nitididorsis opacidorsis Nomura, 1972: 127 A: JA
olivacea Brenske, 1896a: 155 A: SD
opaciclypealis D. Ahrens, 1999b: 253 A: NP
opacithorax Nomura, 1974: 90 A: TAI
ovata Nomura, 1971: 70 (Trichoserica) A: JA
pelelaensis D. Ahrens & Fabrizi, 2011: 140 A: BT
pigrans D. Ahrens & Fabrizi, 2009b: 251 A: AP
planifrons Nomura, 1972: 123 [RN] A: JA
opacifrons Nomura, 1971: 68 (Trichoserica)

plutenkoi D. Ahrens, 2005b: 24 A: HUB SHA
polita Gebler, 1832: 53 (Omaloplia) A: ES FE
lutea J.I. Kim & A.Y. Kim, 2003a: 75

pommeranzi D. Ahrens, 1999b: 305 A: NP
proclivis D. Ahrens, 1999b: 257 A: NP
puetzi D. Ahrens, 2005b: 94 A: SCH SHA
qinlingshanica D. Ahrens, 2005b: 40 A: SCH SHA
ramosa D. Ahrens, 1999b: 284 A: NP UP
ratcliffei D. Ahrens, 2005b: 89 A: SCH
rectidens D. Ahrens & Fabrizi, 2009b: 250 A: AP BT XIZ
ribbei D. Ahrens, 1999b: 275 A: SD
rosinae kurosawai Nomura, 1959b: 45 (Ophthalmoserica) A: JA
rosinae rosinae Pic, 1904b: 33 A: FE HEB HEI LIA
sawadai Nomura, 1959b: 43 (Ophthalmoserica) A: JA
takagii Sawada, 1937: 18
unicolor Sawada, 1937: 18

schoenmanni D. Ahrens, 2005b: 53 A: SCH YUN
septentrionalis Murayama, 1935: 7 A: NC
shaanxiensis D. Ahrens, 2005b: 26 A: GAN SCH SHA

309

310

SCARABAEIDAE

sherpa Sabatinelli & Migliaccio, 1982: 106 (Trichoserica) A: NP
shokhini D. Ahrens, 2005b: 83 A: SCH
sudhausi D. Ahrens, 2005b: 57 A: SHA
taibashanica D. Ahrens, 2005b: 85 A: SHA
tayanpingensis D. Ahrens, 2005b: 96 A: SCH
thibetana Brenske, 1897e: 395 A: BT KA HP NP PA SD UP XIZ ORR
umbrinella Brenske, 1898a: 288

tokejii Nomura, 1959b: 47 (Ophthalmoserica) A: JA
tomiensis D. Ahrens & Fabrizi, 2009b: 252 A: XIZ
tongluana D. Ahrens, 1999b: 308 A: NP SD
trichofemorata Nomura, 1959b: 44 (Ophthalmoserica) A: JA
tropdeana D. Ahrens, 1999b: 255 A: XIZ
tukucheana D. Ahrens, 1999b: 276 A: NP SD
weiperti D. Ahrens, 2004c: 211 A: NP
wrzecionkoi D. Ahrens & Fabrizi, 2009b: 251 A: XIZ
xichangensis D. Ahrens, 2005b: 82 A: SCH
yaogiensis D. Ahrens, 2005b: 50 A: SCH
yoshidai Nomura, 1959b: 45 (Ophthalmoserica) A: JA
yulongshanica D. Ahrens, 2005b: 64 A: YUN
zerchei D. Ahrens, 2005b: 92 A: SCH
subgenus Taiwanoserica Nomura, 1974: 82 type species Taiwanoserica elongata Nomura, 1974
anmashana Kobayashi, 1993: 71 (Taiwanoserica) A: TAI
bihluhensis Kobayashi & C. Yu, 2000: 126 (Taiwanoserica) A: TAI
chengtuensis D. Ahrens, 2009: 287 A: SCH
chunlinlii D. Ahrens, 2002a: 58 (Taiwanoserica) A: TAI
elongata Nomura, 1974: 85 (Taiwanoserica) A: TAI
gracilipes Nomura, 1974: 86 (Taiwanoserica) A: TAI
kubotai Kobayashi, 1983a: 16 (Taiwanoserica) A: TAI
lishana Nomura, 1974: 85 (Taiwanoserica) A: TAI
monticola Kobayashi & C. K. Yu, 2000: 126 (Taiwanoserica) A: TAI
niitakana Sawada, 1939a: 49 A: TAI
nitidipes Nomura, 1974: 86 (Taiwanoserica) A: TAI
ovata Nomura, 1974: 86, 89 (Taiwanoserica) A: TAI
pubisterna Nomura, 1974: 85 (Taiwanoserica) A: TAI
shinanshana Kobayashi, 1988b: 53 (Taiwanoserica) A: TAI
sigipinensis D. Ahrens, 2009: 288 A: SCH
simillima Kobayashi, 1983a: 19 (Taiwanoserica) A: TAI
sinuosa Kobayashi & C. K. Yu, 2000: 126 (Taiwanoserica) A: TAI
suturalis Nomura, 1974: 86 (Taiwanoserica) A: TAI
suzukii Kobayashi, 1983a: 17 (Taiwanoserica) A: TAI
taiyal Kobayashi & C. K. Yu, 2000: 126 (Taiwanoserica) A: TAI

SCARABAEIDAE
variegata Nomura, 1974: 85 (Taiwanoserica) A: TAI
wenchuanensis D. Ahrens, 2009: 289 A: SCH
yui Kobayashi, 1993: 70 (Taiwanoserica) A: TAI
genus Serica, species incertae sedis
albisetis D. Ahrens, 2005b: 118 A: GUX
baiyungshanica D. Ahrens, 2005b: 111 A: YUN
bicornis D. Ahrens, 2005b: 105 A: YUN
brevitarsis Kobayashi & C. K. Yu, 1993c: 355 (Pachyserica) A: TAI
brunnescens Frey, 1972b: 166 A: FUJ
calva Frey, 1972b: 173 (Trichoserica) A: FUJ
dathei D. Ahrens, 2005b: 113 A: FUJ
davidkrali D. Ahrens & Fabrizi, 2011: 149 A: YUN
delicta Brenske, 1897e: 372 A: FE
detersa Erichson, 1834: 239 A: CH ORR
deuvei D. Ahrens, 2005b: 107 A: YUN
erectosetosa D. Ahrens, 1999b: 315 A: NP XIZ
exigua Brenske, 1894a: 37 A: SHG
excisa Frey, 1972b: 172 (Trichoserica) A: FUJ
falcifera D. Ahrens & Fabrizi, 2009b: 254 A: AP
fashengi Liu & D. Ahrens, 2014b: 167 A: XIZ
feisintsiensis D. Ahrens, 2007: 10 A: CH (“Fei Sin Tsi pass”)
formosana Moser, 1915c: 338 A: TAI
brevipilosa Kobayashi, 1985a: 13 (Maladera)
curvifemora Nomura, 1974: 107 (Maladera)

hirtella D. Ahrens, 2005b: 109 A: YUN
jindrai D. Ahrens, 2007: 14 A: GUI SCH
karnaliensis D. Ahrens, 1999b: 319 A: NP XIZ
klapperichi Frey, 1972b: 173 (Trichoserica) A: FUJ
koshiana D. Ahrens, 1999b: 321 A: NP
lalashana Kobayashi, 1983a: 15 A: TAI
lijiagensis D. Ahrens, 2005b: 103 A: SCH YUN
longula Frey, 1972b: 165 A: FUJ
maculosa Moser, 1915a: 113 A: SHN ORR
nanjiangava D. Ahrens, 2005b: 116 A: SCH
nigrobrunnea Moser, 1926: 198 A: SCH
nigroguttata Brenske, 1897e: 389 A: FUJ HKG JIX SCH SHG SHN TAI ORR
nitens Moser, 1915a: 117 A: FUJ
ovatula Fairmaire, 1891f: cxcv A: HEB HAI SCH
palaea D. Ahrens, 2004a: 26 A: SD
pallida Burmeister, 1855: 167 A: HKG
panchaseana D. Ahrens, 2004a: 28 A: NP

311

312

SCARABAEIDAE

parasquamosa D. Ahrens, 2007: 24 A: FUJ HUN SCH
pilumna D. Ahrens & Fabrizi, 2009b: 253 A: AP
pulvinosa Frey, 1972b: 168 A: FUJ HUB TAI ZHE
albosquamosa Frey, 1972b: 163
taiwana Nomura, 1974: 97

salebrosa Brenske, 1897e: 428 A: JA
serripes Moser, 1915c: 339 A: ZHE
silviae D. Ahrens, 2009: 290 A: SCH
sinuaticeps Moser, 1915a: 115 A: SHN YUN
somathangana D. Ahrens, 1999b: 317 A: NP
squamosa D. Ahrens, 2007: 27 A: FUJ JIX
sticta D. Ahrens & Fabrizi, 2009b: 253 A: AP BT
subtruncata Fairmaire, 1887b: 109 A: YUN
trapezicollis D. Ahrens, 2005b: 114 A: SCH
tryznai D. Ahrens, 2007: 31 A: GUI SCH
wrasei D. Ahrens, 2007: 34 A: HUB
yadongensis Liu & D. Ahrens, 2014b: 168 A: XIZ
genus Sericania Motschulsky, 1860e: 256 [RN] type species Sericania fuscolineata Motschulsky, 1860
Mesoserica Matsumura, 1911: 118 [HN] type species Mesoserica sachalinensis Matsumura, 1911

alternata Sawada, 1938a: 8 A: JA
angulata Lewis, 1895c: 392 (Serica) A: JA
awana Nomura, 1976b: 168 A: JA
babaulti D. Ahrens, 2004a: 79 A: UP
besucheti D. Ahrens, 2004a: 91 A: PA
bhojpurensis D. Ahrens, 2004a: 102 A: NP
carinata Brenske, 1898a: 217 A: HKG
chikuzensis Sawada, 1938c: 70 A: JA
hayashii Nomura, 1959b: 42

costulata Moser, 1915b: 148 (Serica) A: NP PA UP
dispar D. Ahrens, 2004a: 82 A: PA
dubiosa D. Ahrens, 2004a: 94 A: KA
elongata Nomura, 1976b: 171 A: JA
fuscolineata ezoensis Nomura, 1976b: 172 A: JA
fuscolineata fulgida Niijima & Kinoshita, 1927: 14 A: JA
lineata Sawada, 1939a: 48
testacea Sawada, 1938a: 8

fuscolineata fuscolineata Motschulsky, 1860e: 137 A: FE HEI JA LIA NC SC
nipponensis Nomura, 1976b: 172

fuscolineata minuscula Nomura, 1976b: 171 A: JA
galloisi Niijima & Kinoshita, 1927: 82 A: JA

SCARABAEIDAE
aikyoi Sawada, 1955: 573
babai Nomura, 1976b: 169

gilgitensis D. Ahrens, 2004a: 96 A: PA
hazarensis D. Ahrens, 2004a: 90 A: PA
heinzi D. Ahrens, 2004a: 76 A: PA
hidana Niijima & Kinoshita, 1923: 218 A: JA
kamiyai Sawada, 1938a: 18

kadowakii Nakane, 1983b: 141 A: JA
kashmirensis Moser, 1919: 331 (Neoserica) A: KA
khagana D. Ahrens, 2004a: 93 A: PA
khandbariensis D. Ahrens & Fabrizi, 2011: 150 A: NP
kirai Sawada, 1938c: 69 A: JA
kleebergi D. Ahrens, 2004a: 101 A: NP
kobayashii Nomura, 1976b: 168 A: JA
koryoensis Murayama, 1935: 8 A: SC
laeticula Sharp, 1878b: 172 (Serica) A: PA
latisulcata Murayama, 1941: 36 A: SC
lewisi Arrow, 1913c: 397 A: JA
loebli D. Ahrens, 2004a: 87 A: PA
mara D. Ahrens, 2004a: 99 A: NP
marginata Nomura, 1973a: 17 A: JA
suturalis Nomura, 1976b: 171

matusitai Sawada, 1938a: 12
mimica kompira Miyake & Sano, 1996: 117 A: JA
mimica mimica Lewis, 1895c: 390 A: JA TAI
matsuyamana Sawada, 1938c: 70
quinquefoliata Sawada, 1938a: 12
takaoana Sawada, 1938a: 13

miyakei Nomura, 1960b: 60 A: JA
nepalensis Frey, 1965a: 89 (Autoserica) A: NP
ohirai Sawada, 1960: 9 A: JA
akitai H. Hirasawa, 1987: 13

ohtakei Sawada, 1955: 565 A: JA
hirosei Miyake & Nakamura, 1995: 9
imadatei Sawada, 1955: 569

opaca Nomura, 1973a: 16 A: JA
hiranoi Miyake, 1988: 77

pacis D. Ahrens, 2004a: 78 A: PA
piattellai D. Ahrens, 2004a: 95 A: KA PA
poonchensis D. Ahrens, 2004a: 83 A: PA
quadrifoliata Lewis, 1895a: 393 (Serica) A: JA

313

314

SCARABAEIDAE

shinanoensis Niijima & Kinoshita, 1923: 219

sachalinensis Matsumura, 1911: 118 A: FE (Kuril’skie ostrova, Sakhalin) JA
kurilensis S.I. Medvedev, 1952: 121

serripes Nomura, 1973a: 14 A: JA
ohtsukai Miyake, 1988: 78

shikokuana Nakane, 1954: 12 A: JA
sinuata Nomura, 1959b: 42 A: JA
tohokuensis Sawada, 1955: 571 A: JA
yamauchii mochizukii Kobayashi, 2014a: 33 A: JA (Shikoku)
yamauchii yamauchii Sawada, 1938a: 8 A: FE HEI JA LIA SC
hasegawai Murayama, 1941: 37
ussuriensis S.I. Medvedev, 1952: 105 (Serica)

yamayai Kobayashi & Fujioka, 2008: 70 A: JA
yanoi Kobayashi, 2014a: 34 A: JA (Shikoku)
genus Sphaerotrochalus Brenske, 1900c: 82 type species Pseudotrochalus boehmi G. Quedenfeldt, 1888
somalicola Frey, 1960: 633 (Trochalus) A: OM SA YE AFR
genus Tetraserica D. Ahrens, 2004a: 168 type species Neoserica gestroi Brenske, 1898
anhuaensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 115 A: FUJ HKG HUN JIX
brahmaputrae D. Ahrens, 2004a: 172 A: BT NP SD ORR
changjiangensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 92 A: HAI
changshouensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 112 A: HUB SCH
crenulata D. Ahrens & Fabrizi, 2009b: 267 A: AP
damaidiensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 96 A: FUJ GUX YUN ORR
daqingshanica Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 88 A: GUX
disoccupata D. Ahrens, 2004a: 175 A: SD
ferrugata C.É. Blanchard, 1850: 78 (Omaloplia) A: HP NP UP
alcocki Brenske, 1898a: 304 (Autoserica)

fikaceki Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 92 A: HAI
graciliforceps Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 102 A: YUN
hilaris D. Ahrens & Fabrizi, 2009b: 266 A: AP
jinghongensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 100 A: YUN
leishanica Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 116 A: GUI GUX
liangheensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 103 A: YUN
linaoshanica Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 113 A: GUX
longipenis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 98 A: GUI GUX SCH YUN
longzhouensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 97 A: GUX
maoershanensis D. Ahrens, Liu & Fabrizi, 2014: 91 A: GUX
mengeana Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 110 A: YUN
menglongensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 102 A: YUN

SCARABAEIDAE
pingjiangensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 105 A: GUX HUB HUN YUN ORR
ruiliana Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 115 A: YUN
ruiliensis D. Ahrens, Liu & Fabrizi, 2014: 113 A: YUN
rungbongensis D. Ahrens, 2004a: 171 A: SD
schneideri D. Ahrens, 2004a: 173 A: SD
sculptilis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 89 A: HUB YUN
shangsiensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 112 A: GUI GUX
shunbiensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 97 A: YUN
sigulianshanica Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 94 A: CHQ GAN SCH SHA
tianchiensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 102 A: HAI
tonkinensis Moser, 1908c: 328 (Neoserica) A: GUX ORR
wandingensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 107 A: YUN
wangtongensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 89 A: GUA HKG JIX
xichouensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 109 A: YUN
yaoanica Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 119 A: FUJ GUA GUX HUN JIX ORR
yaoquensis Liu, Fabrizi, Bai, X.-K. Yang & D. Ahrens, 2014c: 98 A: YUN
genus Trichomaladera Nomura, 1974: 82 type species Trichomaladera elongata Nomura, 1974
elongata Nomura, 1974: 93 A: TAI
infortunata D. Ahrens, 2002a: 72 A: TAI
rufofusca Nomura & Kobayashi, 1979: 9 A: TAI
yasutoshii Kobayashi, 1991c: 211 A: TAI
yui Kobayashi, 1993: 68 A: TAI
genus Triodontella Reitter, 1919b: 221 [RN] type species Serica aquila Laporte, 1840
Triodonta Mulsant, 1842: 468 [HN] type species Serica aquila Laporte, 1840

alni alni C.É. Blanchard, 1850: 83 (Triodonta) E: IT (Sardegna)
alni luteipes Fairmaire, 1881c: xxvi (Serica) E: IT (Sardegna)
aquila Laporte, 1840: 148 (Serica) E: FR SZ
noctua Mulsant, 1842: 469 (Triodonta)

asiatica Brenske, 1890b: 84 (Triodonta) A: TR
brunneipennis J.R. Sahlberg, 1908: 65 (Triodonta) A: TR
bucculenta Baraud, 1962b: 181 [=1962c: 12] (Triodonta) E: FR IT
castilliana Baraud, 1961: 19 (Triodonta) E: PT SP
corsica Baraud & Schaefer, 1959: 170 (Triodonta) E: FR (Corse)
cribellata Fairmaire, 1859a: 277 (Triodonta) E: FR (Corse)
dalmatica Baraud, 1962c: 13 (Triodonta) E: AL BH BU CR GR KO MC ME RO SB TR
bulgarica Bunalski, 2001: 82

demelti Petrovitz, 1963c: 263 (Triodonta) A: TR
difformipes Fairmaire, 1892a: 146 (Triodonta) A: SY TR
brignolii Sabatinelli, 1977b: 81 (Triodonta)

315

316

SCARABAEIDAE

dispar Fairmaire, 1892b: 146 (Triodonta) A: SY
delagrangei Pic, 1898c: 97 (Triodonta)

eggeri A. Rey, 1999b: 122 (Triodonta) A: TR
flavimana Burmeister, 1855: 151 (Triodonta) E: GG A: SY TR
sieversi Reitter, 1889h: 283 (Triodonta)

judaica C.É. Blanchard, 1850: 75 (Triodonta) A: IQ
lusitanica Brenske, 1894a: 7 (Triodonta)

lajonquierei Baraud, 1961: 17 (Triodonta) E: SP
lateristria Reitter, 1889h: 283 (Triodonta) A: TR
mimula Leo & Fancello, 2007: 266 E: IT (Sardegna)
nitidula P. Rossi, 1790: 23 (Melolontha) E: AU IT SL SZ
sericans Gyllenhal, 1817: 186 (Melolontha) [RN]
sericea Bonelli, 1812: 158 (Melolontha)

preissi Keith, 2001d: 157 A: TR
raymondi Perris, 1869: 15 (Triodonta) E: IT (Sardegna)
reitteri Brenske, 1890b: 84 (Triodonta) N: AG
ruizi Pérez-López & Hernández-Ruiz, 1994: 74 E: SP
sardoa Baraud, 1962c: 12 (Triodonta) E: IT (Sardegna)
tunisia Brenske, 1890b: 84 (Triodonta) [DA] E: TU
zuzartei Branco, 1978: 389 (Triodonta) E: PT
genus Trioserica Moser, 1922: 111 type species Trioserica lepichaeta Moser, 1922
Sinoserica Miyake & Yamaya, 2001: 40 type species Sinoserica maculipennis Miyake & Yamaya, 2001

alternata Nomura, 1974: 94 (Selaserica) A: TAI
antennalis Nomura, 1974: 95 (Selaserica) A: TAI
macrophthalma Moser, 1918: 219 (Neoserica) A: TAI
rufocastanea Kobayashi, 1980: 18 (Selaserica)

maculipennis Miyake & Yamaya, 2001: 40 (Sinoserica) A: YUN
taipeiensis Kobayashi, 2013: 78 A: TAI
tarsata Brenske, 1894a: 11 (Serica) A: BT ORR
genus Trochaloschema Reitter, 1896e: 182 type species Serica iris Semenov, 1893
armeniacum Brenske, 1897b: 388 A: TD
medusa Reitter, 1902b: 141

chikatunovi Nikolajev, 1987: 174 A: TD
iris Semenov, 1893c: 495 (Serica) A: TD
kanevskajae Nikolajev, 1987: 174 A: TD
kryzhanovskii Nikolajev, 1987: 173 A: TD
lopatini Nikolajev, 1987: 174 A: TD
medvedevi medvedevi Nikolajev, 1987: 173 A: TD
medvedevi vachshianum Nikolajev, 1987: 174 A: TD
michailovi Nikolajev, 1981: 123 A: TD

317

SCARABAEIDAE
ravshani Ivanova & Pak, 2012: 21 A: TD
ruginota Reitter, 1896e: 186 A: KI
saryhissorica Janushev, 1973: 139 A: TD
shukronajevi Nikolajev, 1987: 173 A: TD
tavildarense Nikolajev, 2002: 104 A: TD
valentini Novikov, 1999: 33 A: TD
genus Xenoserica D. Ahrens, 2005a: 273 type species Nipponoserica sindhensis D. Ahrens, 2000
brachyptera D. Ahrens, 2005a: 275 A: NP
matthiasi Shrestha, Eberle & D. Ahrens, 2012: 383 A: NP
pindarensis D. Ahrens, 2000b: 27 (Nipponoserica) A: UP
selaensis D. Ahrens & Fabrizi, 2009b: 255 A: AP
sindhensis D. Ahrens, 2000b: 26 (Nipponoserica) A: KA

subfamily Rutelinae W.S. Macleay, 1819
tribe Adoretini Burmeister, 1844
subtribe Adoretina Burmeister, 1844

genus Adoretus Dejean, 1833: 157 type species Melolontha nigrifrons Steven, 1809
subgenus Adoretus Dejean, 1833: 157 type species Melolontha nigrifrons Steven, 1809
Prionadoretus Ohaus, 1914c: 512 type species Prionadoretus serridens Ohaus, 1914

abdolrezagharibi Petrovitz, 1980: 634 A: IN
aegrotus Burmeister, 1844: 470 N: AG A: SI “Arabien” AFR
millingeni Pic, 1905e: 153

afghanus Machatschke, 1958: 178 A: AF
atiqi Abdullah, 1968: 419 A: PA
baquari Abdulah, 1968: 421 A: PA
bicolor Brenske, 1893b: 142 A: NP SD ORR
bilobatus Lin, 1974: 190 A: YUN
bimarginatus Ohaus, 1914c: 480 A: SD UP ORR
brachypygus Burmeister, 1855: 530 A: “India”
caliginosus Burmeister, 1844: 471 A: HP PA SD ORR
bicolor Brenske, 1900b: 38
nudostriatus Ohaus, 1914c: 482

convexus Burmeister, 1855: 531 A: HKG ORR
coronatus Burmeister, 1855: 533 A: NP ORR
discolor Faldermann, 1835a: 267 (Trigonostoma) E: AB AR GG IN ST
drurei Pic, 1905h: 200 A: IQ
duvauceli C.É. Blanchard, 1851: 232 A: HP SD ORR
concolor C.É. Blanchard, 1851: 232

epipleuralis Arrow, 1914b: 601 A: CH ORR
falciungulatus Nomura, 1965: 7 A: JA (Iriomote-jima, Ishigaki-jima)
fatehi Abdullah, 1968: 420 A: PA

318

SCARABAEIDAE

flavus Arrow, 1917: 315 A: IN PA ORR
gandolphei Guérin-Méneville, 1859: 185 N: AG MO TU
fuscitarsis Reitter, 1903a: 32
infissidens Pic, 1922b: 17

garamas Peyerimhoff, 1921: 235 N: AG EG LB AFR
geyri Ohaus, 1917b: 4 N: AG MO AFR
surcoufi Peyerimhoff, 1921: 234

granulifrons Fairmaire, 1882b: 68 N: EG A: IS JO AFR
graniceps Reitter, 1889e: 268

hainanensis Keith & J. Li, 2009: 165 A: HAI
hirticollis Lin, 1974: 190 A: YUN
horticola Arrow, 1917: 344 A: UP
hybogeneius Ohaus, 1930: 153 N: AG MO AFR
iftikhari Abdullah, 1968: 422 A: PA
irakanus Ohaus, 1928: 401 A: IQ
iranicus Petrovitz, 1958a: 7 A: IN
ismaili Abdullah, 1968: 421 A: PA
jarkandus Ohaus, 1930: 153 A: XIN
ladakanus horni Balthasar, 1936c: 205 A: AF
ladakanus ladakanus Ohaus, 1914c: 484 A: AF KA PA
ladakanus nuristanicus Balthasar, 1936c: 206 A: AF
lasiopygus Burmeister, 1855: 530 A: BT NP SD UP ORR
cardoni Brenske, 1893b: 141

limbatus C.É. Blanchard, 1851: 233 A: KA ORR
castaneus Nonfried, 1892e: 362
flavovittatus Nonfried, 1892e: 362
plagiatus Burmeister, 1855: 531

lithobius Ohaus, 1914c: 485 A: HP PA UP ORR
ludmilae Sabatinelli & Rittner, 2015: 303 A: IS
machatschkei Petrovitz, 1965d: 692 A: AF
minutus Brenske, 1893b: 143 A: YUN ORR
naeemi Abdullah, 1968: 422 A: PA
nanschanchianus Kobayashi, 1982: 54 A: TAI
nigrifrons Steven, 1809: 41 (Melolontha) E: AB GG KZ ST A: AF IN KI KZ TD TM UZ XIN XIZ
pallidulus Motschulsky, 1860i: 522

peregrinus Ohaus, 1914c: 477 A: “Ostindien”
persicus Reitter, 1889e: 268 A: IN
petrovitzi Baraud, 1975c: 192 A: IN
peyerimhoffi Baraud, 1979c: 222 N: MO
progrediens Ohaus, 1914c: 489 A: “Nord-Indien” ORR

SCARABAEIDAE
pruinosus Ballion, 1871a: 345 A: AF KI KZ TD TM UZ
pullus Baudi di Selve, 1870: 76 A: CY
punjabensis Arrow, 1917: 344 A: PA
quadridens Marseul, 1878: 71 N: EG
rubenyani Kalashyan, 2002: 76 E: AR
runcinatus Lin, 1974: 189 A: YUN
rufifrons Reitter, 1903a: 32 A: MG XIN
kaszabi Endrődi, 1964: 429

saleemi Abdullah, 1968: 420 A: PA
scabiosus Petrovitz, 1980: 636 A: IN
senescens Walker, 1871: 12 N: EG
seriesetosus Arrow, 1917: 303 A: PA
serratipes Arrow, 1914b: 595 A: NP SD ORR
serridens Ohaus, 1914c: 513 (Prionadoretus) A: “Ost-Indien”
setifer Reitter, 1889e: 269 A: CH
sexdentatus Lin, 1974: 192 A: YUN
simplex Sharp, 1878b: 173 A: HP KA NP
sistanicus Petrovitz, 1970c: 41 A: IN
sterbae Reitter, 1909b: 82 A: IS TR
stoliczkae Ohaus, 1914c: 490 A: PA ORR
striatus Lin, 1974: 191 A: YUN
tufaili Abdullah, 1968: 419 A: PA
vilosicollis C.É. Blanchard, 1851: 321 A: “Arabia”
versutus Harold, 1869d: 124 A: NP UP AURi ORR
bangalorensis Brenske, 1900b: 38
insularis Fairmaire, 1897b: 105
vestitus Boheman, 1859: 119
vitiensis Nonfried, 1892d: 268

subgenus Chaetadoretus Ohaus, 1914c: 492 type species Adoretus timoriensis Ohaus, 1914
costopilosus Ohaus, 1914c: 506 A: HP NP SD UP ORR
cribratus White, 1844: 424 A: HKG SE ORR
parallelus Kraatz, 1895e: 252

formosanus formosanus Ohaus, 1914c: 503 A: TAI
formosanus sakishimanus Kobayashi, 1982: 53 A: JA (Sakishima)
hirsutus Ohaus, 1914c: 503 A: FUJ LIA NC SC TAI ORR
inornatus Heyden, 1889c: 669 A: SCH
lasius Ohaus, 1914c: 501 A: BT SD ORR
mizusawai Kobayashi, 1982: 51 A: TAI
nigrofuscus Kobayashi, 1991b: 65 A: YUN ORR
polyacanthus Ohaus, 1914c: 504 A: TAI ORR

319

320

SCARABAEIDAE

puberulus Motschulsky, 1854c: 46 A: BEI LIA
siemsseni Ohaus, 1935: 128 A: FUJ
testaceus Hope, 1831: 23 (Geniates) A: BT NP
assimilis Hope, 1831: 23 (Geniates)

tetracanthus Ohaus, 1930: 158 A: YUN
subgenus Gemadoretus Reitter, 1903a: 30 type species Adoretus clypeatus Burmeister, 1844
clypeatus Burmeister, 1844: 471 N: EG AFR
signatus Reitter, 1889e: 268

subgenus Lepadoretus Reitter, 1903a: 30 type species Adoretus tenuimaculatus C.O. Waterhouse, 1875
bhutanensis Frey, 1975c: 245 A: BT
kashmarensis Keith, 2001c: 77 A: IN
maniculus Ohaus, 1914c: 511 A: “Ostindien”
sinicus Burmeister, 1855: 532 A: FUJ GUA HAI HKG HUN JA JIA LIA NC SC TAI ZHE AURi NARi ORR
tenuimaculatus C.O. Waterhouse, 1875a: 112 A: CH GUI HUN JA NC LIA SC TAI ZHE AURi ORR
vastus Petrovitz, 1958a: 7 A: IN
genus Lissadoretus Arrow, 1917: 293 type species Lissadoretus pallidus Arrow, 1917
pallidus Arrow, 1917: 293 A: AF IN PA ORR
australis Petrovitz, 1958a: 8 (Pseudadoretus)

genus Pseudadoretus Semenov, 1889b: 202 type species Adoretus phthisicus Dohrn, 1882
Eremadoretus Semenov, 1912: 500 type species Pseudadoretus validus Semenov, 1889
Peradoretus Reitter, 1903a: 35 type species Adoretus koechlini Marseul, 1867
Pristadoretus Reitter, 1903a: 36 type species Adoretus fallax Semenov, 1890
Stichadoretus Reitter, 1903a: 34 type species Adoretus arabicus Reitter, 1903

dilutellus Semenov, 1889b: 203 A: KZ TD TM UZ
perdilutus Reitter, 1903a: 37

fallax Semenov, 1890a: 4 A: KZ TM UZ
koechlini Marseul, 1867c: lxxx (Adoretus) N: AG MO AFR
evanescens Marseul, 1878: 72 (Adoretus)

phthisicus Dohrn, 1882: 108 (Adoretus) A: KZ TD TM UZ
dilutellus Reitter, 1889e: 269 (Adoretus)

validus Semenov, 1889b: 203 A: KZ TD TM UZ
genus Rhampadoretus Ohaus, 1912: 418 type species Adoretus minutulus Brenske, 1895
suillus Arrow, 1917: 291 A: NP ORR

subtribe Adorrhinyptiina Arrow, 1917

genus Adorrhinyptia Arrow, 1917: 273 type species Rhyniptia dorsalis Burmeister, 1855
dorsalis Burmeister, 1855: 494 (Rhinyptia) A: NP PA ORR

321

SCARABAEIDAE
subtribe Pachyrhinadoretina Ohaus, 1912

genus Pachyrhinadoretus Ohaus, 1912: 509 type species Pachyrhinadoretus rugipennis Ohaus, 1912
frontatus Burmeister, 1855: 530 (Adoretus) A: NP ORR

subtribe Prodoretina Ohaus, 1912

genus Clipadoretus Ohaus, 1941: 112 type species Adoretus epistomalis Chobaut, 1899
dentatus Machatschke, 1965: 134 A: IN PA
epistomalis Chobaut, 1899: 38 (Adoretus) N: AG MO AFR
ehrenbergi Ohaus, 1912: 425 (Rhamphadoretus) A: “Arabia felix”
persicus Ohaus, 1941: 113 A: IN
quadridentatus Petrovitz, 1967e: 488 A: PA
genus Oxyadoretus Arrow, 1917: 281 type species Oxyadoretus nasutus Arrow, 1917
nasutus Arrow, 1917: 282 A: “India Orient.”

subtribe Trigonostomusina Ohaus, 1912

genus Epadoretus Reitter, 1903a: 35 type species Pseudadoretus reitteri Semenov, 1890
reitteri Semenov, 1890a: 3 (Pseudadoretus) A: KZ TD TM UZ
expallidus S.I. Medvedev, 1949: 328
semenovi S.I. Medvedev, 1949: 328

genus Phaeadoretus Reitter, 1903a: 31 type species Adoretus comptus Ménétriés, 1849
comptus Ménétriés, 1849: 62 (Adoretus) E: AB A: AE AF IN KI KZ TD TM UZ
iranicus iranicus Machatschke, 1960: 843 A: IN
iranicus nuristanicus Machatschke, 1960: 846 A: AF
lindemannae Machatschke, 1960: 847 A: IQ SY
shirasensis Petrovitz, 1970c: 42 A: IN
syriacus C.É. Blanchard, 1851: 232 (Adoretus) A: IN IQ SY

tribe Anomalini Streubel, 1839
subtribe Anisopliina Burmeister, 1844

genus Anisoplia Schönherr, 1817: 186 type species Scarabaeus agricola Poda von Neuhaus, 1761
subgenus Anisoplia Schönherr, 1817: 186 type species Scarabaeus agricola Poda von Neuhaus, 1761
Ammanisoplia S.I. Medvedev, 1949: 305 type species Anisoplia deserticola Fischer von Waldheim, 1824
Lasioplia S.I. Medvedev, 1949: 275 type species Scarabaeus villosus Goeze, 1777

abdita Baraud, 1991d: 324 A: TR
agnata Reitter, 1889d: 107 E: AB AR GG A: IN TR
anagnata Zaitzev, 1917: 25 [= Zaitzev, 1918a: 113]

agricola Poda von Neuhaus, 1761: 21 (Scarabaeus) E: AL AU BH BU BY CR CT CZ GR HU IT KO MC MD PL RO
SB SK SL ST SV TR UK A: ES KI KZ MG TR WS

322
aetolica Apfelbeck, 1909: 502
deleta Schilsky, 1888a: 186 [HN]
conjuncta Schilsky, 1888a: 186
crucifera Herbst, 1790: 100 (Melolontha)
cyathigera Scopoli, 1763: 4 (Scarabaeus)

alazanica Zaitzev, 1917: 32 [= Zaitzev, 1918a: 120] E: GG
andreae Baraud, 1991d: 324 E: GR
angorensis Petrovitz, 1971e: 583 A: TR
aprica Erichson, 1847a: 636 E: BU TR A: TR
armeniaca Kraatz, 1883b: 19 E: AR GG TR
baetica Erichson, 1847a: 636 E: PT SP
brenskei Reitter, 1889d: 106 E: BU CT GR MD RO ST UK A: KZ
balcanica Reitter, 1889d: 106
banatica Reitter, 1903a: 100

bromicola Germar, 1817: 216 (Melolontha) E: AU BH CR CZ FR HU IT SB SK SL UK
maculata Schilsky, 1888a: 186
nigra Schilsky, 1888a: 186 [HN]
scutellata Schilsky, 1888a: 186
variabilis Schilsky, 1888a: 186

bulgarica Apfelbeck, 1909: 503 E: BU RO SB
bureschi Sacharieva-Stoilova, 1958: 83 E: BH BU GR MC SB TR A: TR
clypealis Reitter, 1889d: 103 E: AR “Kaukasus”
depressa Erichson, 1847a: 635 E: PT SP
nigra Paulino de Oliveira, 1884: 562

deserticola Fischer von Waldheim, 1824: 216 E: AB BU GR MD RO SB ST UK A: KZ
dispar Erichson, 1847a: 642 E: BU GR RO A: TR
morio Kraatz, 1883b: 20

enucleator Baraud, 1991d: 338 A: TR
erichsoni Reitter, 1889d: 104 E: AL AU BH CR GE GR HU IT MC ME SB SL
faldermanni Reitter, 1883a: 100 [RN] E: GG ST A: IN TR
villosa Faldermann, 1835a: 270 [HN]

farraria antoniae Reitter, 1889d: 109 E: AB A: IN TR
pastuchovae Zaitzev, 1917: 30 [= Zaitzev, 1918a: 103]

farraria farraria Erichson, 1847a: 641 E: AB AR GG A: TR
succincta C.É. Blanchard, 1851: 175

flavipennis Brullé, 1832: 179 E: AL AU BH CR GE GR MC ME SB TR
adiecta Erichson, 1847a: 647
bouaei C.É. Blanchard, 1851: 176

hebes Reitter, 1903a: 104 E: MC GR A:TR
hebrothracica Král, 1996a: 183 E: BU
hirta Zaitzev, 1917: 27 [Zaitzev, 1918a: 115] E: GG

SCARABAEIDAE

SCARABAEIDAE
imitatrix Apfelbeck, 1909: 502 E: GR MC
insolita Baraud, 1991e: 366 E: GR
kiritshenkoi S.I. Medvedev, 1949: 284 E: ST (Kavkaz)
koenigi Reitter, 1895g: 309 A: TM
lanuginosa Erichson, 1847a: 637 E: GR A: TR
lata lamiensis Apfelbeck, 1909: 503 E: GR
borchmanni Balthasar, 1959a: 108
maderi Pilleri, 1954a: 47

lata lata Erichson, 1847a: 648 E: AL AU BH CR CZ GR HU MC MD PL RO SB SK SL UK
atra Schilsky, 1888a: 186
macedonica Apfelbeck, 1909: 503
signata Schilsky, 1888a: 186 [HN]

limbata Kraatz, 1886b: 225 E: AB AR GG A: TR
lodosi Baraud, 1990b: 149 A: TR
minor J. Müller, 1902: 463 E: BH CR
monticola adriatica J. Müller, 1957: 202 E: CR
monticola marginata Kraatz, 1883b: 20 E: IT (Sicilia)
monticola monticola Erichson, 1847a: 638 E: AU CR IT SB SL
neapolitana Reitter, 1889d: 105

monticola muelleri Mikšić, 1952: 389 E: CR
noahi Petrovitz, 1973a: 11 A: TR
parva Kraatz, 1883b: 21 E: AB AR ST
persiaca Biczók, 1940: 30 A: IN TR
egregia Petrovitz, 1968g: 481
paupercula Petrovitz, 1971e: 584

phoenissa Zaitzev, 1917: 13 [= Zaitzev, 1918a: 101] A: SY
pubipennis C.É. Blanchard, 1851: 174 E: AL CR GR ME
agriniona Reitter, 1903a: 105
nasalis Reitter, 1903a: 105
valida Kraatz, 1883b: 18

reitteriana Semenov, 1903c: 99 E: AB AR A: TR
remota Reitter, 1889d: 103 E: FR SP
latenigra Pic, 1944a: 1
mulsanti Pic, 1944a: 1
noesskei Hänel, 1939: 13
perroudi Pic, 1944a: 1
reyi Pic, 1944a: 1
theresae Pic, 1944a: 1
weberi Reitter, 1898c: 342

sabatinellii Baraud, 1991e: 379 E: IT
seleucidis Baraud, 1991e: 358 A: TR

323

324
signata akbesiana Baraud, 1991e: 360 A: IS JO SY TR
signata signata Faldermann, 1835a: 268 E: AB AR GG ST A: IN TR
sila Zaitzev, 1917: 25 [= Zaitzev, 1918a: 113] E: GG
taocha Zaitzev, 1917: 28 [= Zaitzev, 1918a: 116] E: TR
tempestiva Erichson, 1847a: 643 E: AL BH CR CZ FR GR HU IT ME SB SK SL SZ UK
agricola Stephens, 1830: 227 [HN]
ambigua Mulsant, 1842: 487
connexa Mulsant, 1842: 486
deleta Mulsant, 1842: 486
evanida Mulsant, 1842: 487
excutellata Mulsant, 1842: 487
fasciata Mulsant, 1842: 487
georgi Schatzmayr, 1923: 7
graminivora Sainte-Claire Deville, 1914: 488
hungarica Reitter, 1903a: 104
immarginata Mulsant, 1842: 486
interrupta Mulsant, 1842: 486
maura Mulsant, 1842: 486
meridiana C. Rey, 1890a: 171
quadrimaculata Mulsant, 1842: 486
scutellaris Mulsant, 1842: 486
sicula Reitter, 1889d: 109
unipunctata Mulsant, 1842: 486

tenebralis Burmeister, 1844: 225 A: SY TR
thessalica Reitter, 1889d: 111 E: BU GR MC RO TR
tunneri Petrovitz, 1971e: 583 A: TR
turcomana Zaitzev, 1917: 13 [= Zaitzev, 1918a: 101] A: IN TM
ungulata Baraud, 1991e: 361 A: ST E: GG
venusta Baraud, 1991e: 361 E: AR
villosa Goeze, 1777: 74 (Scarabaeus) E: BE CZ FR GE IT NL PL SP SK SZ UK
curvipunctata Mulsant, 1842: 490
defectiva Mulsant, 1842: 490
donovani Marsham, 1802: 44
obscura Mulsant, 1842: 490
punctum Mulsant, 1842: 490
quadrata Mulsant, 1842: 490
simplicifrons Reitter, 1889d: 104
subarcuata Mulsant, 1842: 490
sycophanta Mulsant, 1842: 490
trimaculata Mulsant, 1842: 490
unicolor Mulsant, 1842: 490

SCARABAEIDAE

SCARABAEIDAE

325

villica Mulsant & Rey, 1871c: 386

zwickii Fischer von Waldheim, 1824: 216 E: CT ST UK A: KZ
scytha Motschulsky, 1849: 115

subgenus Autanisoplia S.I. Medvedev, 1949: 265 type species Melolontha austriaca Herbst, 1783
austriaca austriaca Herbst, 1783: 16 (Melolontha) E: AU BU CT CZ GE GR HU KO MC MD RO SB SK SL ST
TR UK
balcanica Mikšić, 1959c: 98 [RN] [HN]
bulgarica Sacharieva-Stoilova, 1954: 209 [HN]
bulgaricola Machatschke, 1957: 194 [RN]
miksici Baraud, 1991d: 313 [RN]

austriaca hordearia Petrovitz, 1959c: 127 E: GR (Ródos)
austriaca kurdistana Reitter, 1889d: 108 E: AB AR A: TR
austriaca major Reitter, 1889d: 108 E: AB GG ST A: IN
austriaca persica Petrovitz, 1980: 633 A: IN TR
merkli Baraud, 1991d: 316 A: TR
tritici Burmeister, 1855: 492 E: GR
subgenus Pilleriana Baraud, 1991d: 318 type species Anisoplia campicola Ménétriés, 1832
campicola Ménétriés, 1832: 187 E: CT ST UK A: ES IN KZ TR WS
hauseri Reitter, 1898b: 120 A: KI NW TD XIN
genus Anisoplia, nomina dubia
arvicola Fabricius, 1781: 42 (Melolontha) A: “Sibiria”
apicalis Mulsant, 1842: 494 E: FR
bifasciata Mulsant, 1842: 493 E: FR
bipunctata Mulsant, 1842: 493 E: FR
disjuncta Mulsant, 1842: 493 E: FR
fallax Mulsant, 1842: 493 E: FR
funerea Mulsant, 1842: 494 E: FR
laeta Mulsant, 1842: 494 E: FR
mariettii Osculati, 1844: 72 A: TR
unifasciata Mulsant, 1842: 493 E: FR
genus Anthoplia S.I. Medvedev, 1949: 273 type species Melolontha floricola Fabricius, 1787
floricola Fabricius, 1787a: 22 (Melolontha) E: PT SP N: AG LB MO TU
lusitanica Mikšić, 1954b: 21 (Anisoplia)
nigripennis Paulino de Oliveira, 1884: 563 (Anisoplia)

genus Brancoplia Baraud, 1986: 332 type species Anisoplia leucaspis Laporte, 1840
klapperichi Petrovitz, 1971e: 587 (Anisoplia) A: IS JO
leucaspis leucaspis Laporte, 1840: 151 (Anisoplia) E: AB AR ST A: IN IQ IS SY TM TR
leucaspis vseteckai Pilleri, 1951: 235 (Anisoplia) A: LE

326

SCARABAEIDAE

mesopotamica Pilleri, 1954b: 392 (Anisoplia) A: IQ IS LE SY TR
babylonica Petrovitz, 1973b: 34 (Anisoplia) [RN]
mesopotamica Petrovitz, 1971e: 585 (Anisoplia) [HN]

pumila Marseul, 1878: 68 (Anisoplia) N: EG A: IS JO LE
sabulicola Reitter, 1889: 101 (Anisoplia)

genus Chaetopteroplia S.I. Medvedev, 1949: 258 type species Melolontha segetum Herbst, 1783
aegyptica Machatschke, 1961: 620 (Anisoplia) N: EG
bidens Pilleri, 1948: 435 (Anisoplia) A: SY
cuprina Machatschke, 1961: 621 (Anisoplia) A: LE
gracilis Machatschke, 1961: 624 (Anisoplia) A: LE
inculta Erichson, 1847a: 634 (Anisoplia) A: SY
libanensis Machatschke, 1961: 623 (Anisoplia) A: LE
muelleri Pilleri, 1954b: 393 (Anisoplia) A: LE SY TR
naviauxi Baraud, 1984c: 248 (Anisoplia) A: IQ
obenbergeri Všetečka, 1941: 150 (Anisoplia) A: IS LE SY
petrovitzi Machatschke, 1971b: 295 (Anisoplia) A: TR
segetum balcanicola Machatschke, 1961: 639 (Anisoplia) E: BU GR MC
pontica Muche, 1963b: 195 (Anisoplia)

segetum cordofana Burmeister, 1855: 489 (Anisoplia) A: TR
curdistana Machatschke, 1961: 645 (Anisoplia)
ruficollis Kraatz, 1883b: 24 (Anisoplia)

segetum cretica Baraud, 1993: 25 A: GR (Kríti)
segetum croatica Machatschke, 1961: 639 (Anisoplia) E: CR
segetum griseovillosa Machatschke, 1961: 650 (Anisoplia) E: CT
segetum segetum Herbst, 1783: 15 (Melolontha) E: AU BE BY CT CZ FR GE HU KZ LU MD NL PL RO SB SK SL
ST SZ UK
campestris Herbst, 1783: 15 (Melolontha) [NO]
fruticola Fabricius, 1787a: 23 (Melolontha)
volhynica Roubal, 1931: 36 (Anisoplia)

segetum rasa Zoubkov, 1833: 324 (Anisoplia) A: “Turcménie”
segetum solitarium Muche, 1962b: 126 (Anisoplia) A: TR
segetum straminea Brullé, 1832: 178 (Anisoplia) E: AL BU MC GR
pirina Machatschke, 1961: 644 (Anisoplia)

segetum velutina Erichson, 1847a: 634 (Anisoplia) E: AB AR GG ST UK (Krym) GR (Sporádhes) A: TR
segetum zoubkovii Krynicki, 1832: 126 (Anisoplia) E: ST A: KZ
rufipes Burmeister, 1855: 489 (Anisoplia)

syriaca Burmeister, 1844: 220 (Anisoplia) A: IN IS SY TR
cintipennis Reitter, 1903a: 95 (Anisoplia)
nigripennis Reitter, 1903a: 95 (Anisoplia)

327

SCARABAEIDAE
genus Hemichaetoplia Baraud, 1986: 336 type species Trichius pallidipennis Gyllenhal, 1817
gossypiata Fairmaire, 1881a: 86 (Anisoplia) A: IS
ferruginipes Petrovitz, 1963c: 265 (Anisoplia)

lanata Escalera, 1914a: 184 (Anisoplia) N: MO
heterodoxa Peyerimhoff, 1949b: 290 (Anisoplia)

pallidipennis Gyllenhal, 1817: 40 (Trichius) E: IT (Sardegna) N: AG LB MO TU
hypocrita Burmeister, 1844: 218 (Anisoplia)
palleola Desbrochers des Loges, 1898: 75 (Anisoplia)
villosostriata Desbrochers des Loges, 1898: 75 (Anisoplia)

peyerimhoffi Kocher, 1958: 63 (Anisoplia) N: MO
peyerimhoffi Baraud, 1979c: 201 (Anisoplia) [HN]

genus Rhinyptia Burmeister, 1844: 227 type species Rhinyptia indica Burmeister, 1844
subgenus Pararhinyptia Machatschke, 1957: 182 type species Rhinyptia rostrata Burmeister, 1844
plana Walker, 1871: 12 A: “Arabia”
rostrata Burmeister, 1844: 229 N: EG AFR
tewfiki Pic, 1939b: 145 (Adoretus)

subgenus Rhinyptia Burmeister, 1844: 227 type species Rhinyptia indica Burmeister, 1844
indica Burmeister, 1844: 228 A: SD ORR
testacea Nonfried, 1891a: 230

nigrifrons Kraatz, 1895a: 105 A: PA
suturalis Kraatz, 1895a: 104 A: AF IN KA PA ORR
genus Tropiorhynchus C.É. Blanchard, 1851: 176 type species Anisoplia orientis Newman, 1838
orientis Newman, 1838c: 384 (Anisoplia) A: HP NP PA SD
podagricus Burmeister, 1844: 226 (Anisoplia) A: KA ORR

subtribe Anomalina Streubel, 1839

genus Adoretosoma C.É. Blanchard, 1851: 234 type species Adoretosoma elegans C.É. Blanchard, 1851
Euchrysinda Reitter, 1903a: 80 type species Phyllopertha chinensis L. Redtenbacher, 1867

atritarse atritarse Fairmaire, 1891d: xi (Phyllopertha) A: FUJ GUA GUI HUB HUN JIA JIX SCH SHG XIZ YUN ZHE
incostatum Fairmaire, 1891d: xi (Phyllopertha)
metallicum Arrow, 1899a: 266

bruschii Sabatinelli, 1991c: 396 A: BT NP SD
chinense L. Redtenbacher, 1868: 70 (Phyllopertha) A: GUA HKG ORR
chromaticum chromaticum Fairmaire, 1886: 327 (Phyllopertha) A: GUA GUI YUN
chromaticum pulchrum Machatschke, 1955a: 382 A: JIX
chromaticum setchuense Machatschke, 1955a: 382 A: SCH
elegans C.É. Blanchard, 1851: 234 A: FUJ GUA GUI GUX HKG HUB HUN JIA JIX SCH SHX YUN ZHE ORR
humerale Ohaus, 1905: 82

328

SCARABAEIDAE

tenuelimbatum Fairmaire, 1889a: 24 (Phyllopertha)

fairmairei Arrow, 1899a: 262 (Anomala) [RN] A: GUA HKG ORR
cinabarina Fairmaire, 1893e: 308 (Anomala) [HN]
crassipes Ohaus, 1905: 84

fulviventre C.É. Blanchard, 1851: 235 A: BT HP NP SD XIZ
fusipes Heller, 1923: 76 A: SCH
galerucina Arrow, 1917: 199 (Anomala) A: NP SD
nigripenne Ohaus, 1905: 83 A: XIZ ORR
lineatum Ohaus, 1905: 83

perplexum Machatschke, 1955a: 392 A: SCH YUN ORR
signaticolle Nonfried, 1893: 334 (Anomala) A: NP SD XIZ YUN ORR
singhikense Kacker, 1972: 96 (Anomala) A: SD
tonkinense Machatschke, 1955a: 385 A: HEN SCH YUN ORR
genus Anomala Samouelle, 1819: 191 type species Melolontha frischii Fabricius, 1775
Anomalepta Casey, 1915: 8 type species Anomala semilivida LeConte, 1879
Anomalopides E. Strand, 1928: 2 [RN] type species Anomala rhizotrogoides C.É. Blanchard, 1851
Anomalopus Casey, 1915: 40 [HN] type species Anomala rhizotrogoides C.É. Blanchard, 1851
Aprosterna Hope, 1836: 117 type species Mimela nigricans Kirby, 1825 (=Melolontha antiqua Gyllenhal, 1817)
Bifurcanomala J.I. Kim, 1998: 311 type species Melolontha aulax Wiedemann, 1823
Chejuanomala J.I. Kim, 1998: 312 type species Anomala quelparta Okamoto, 1924
Chrysoplethisa Reitter, 1903a: 56 type species Anomala octiescostata Burmeister, 1844
Dichomala Reitter, 1903a: 63 type species Melolontha devota P. Rossi, 1790
Diplomala Reitter, 1903a: 68 type species Anomala subvittata Reitter, 1903
Emphalena Reitter, 1903a: 67 type species Anomala exoleta Faldermann, 1835
Euchlora W.S. Macleay, 1819: 147 type species Melolontha viridis Fabricius, 1775
Euchronomala Reitter, 1903a: 65 type species Euchlora albopilosa Hope, 1839
Euporochlora Reitter, 1903a: 64 type species Melolontha viridis Fabricius, 1775
Euporomala Reitter, 1903a: 62 type species Anomala sieversi Heyden, 1887
Hemispilota Casey, 1915: 45 type species Melolontha lucicola Fabricius, 1798
Heteroplia Burmeister, 1844: 233 type species Melolontha elata Fabricius, 1792
Hybalonomala Reitter, 1903a: 81 type species Anomala bleusei Chobaut, 1896
Idiocnema Faldermann, 1835b: 377 type species Idiocnema sulcipennis Faldermann, 1835
Idiocnemina Reitter, 1903a: 67 type species Anomala gracilenta Reitter, 1903
Iliola Semenov & S.I. Medvedev, 1949: 198 type species Iliola pallens Semenov & S.I. Medvedev, 1949
Lamoana Casey, 1915: 48 type species Anomala villosella C.É. Blanchard, 1851
Nongoma Péringuey, 1902: 609 type species Anomala calcarata Arrow, 1899
Orphnomala Reitter, 1903a: 57 type species Anomala rufozonula Fairmaire, 1887
Paragematis Reitter, 1903a: 69 type species Anomala melanopa Reitter, 1903 (=Melolontha plebeja A.G. Olivier, 1789)
Paranomala Casey, 1915: 12 type species Melolontha binotata Gyllenhal, 1817

SCARABAEIDAE

329

Peripopillia H.J. Kolbe, 1894b: 208 type species Popillia oberthueri Kraatz, 1892 (=Anomala basalis GuérinMéneville, 1847)
Psammoscapheus Motschulsky, 1854b: 30 type species Psammoscapheus dilutus Motschulsky, 1854
Rhinoplia Burmeister, 1844: 232 type species Melolontha dorsalis Fabricius, 1775
Rhombonalia Casey, 1915: 5 type species Anomala cavifrons LeConte, 1868

acusigera Lin, 2002b: 420 A: FUJ GUA
acutangula Ohaus, 1914b: 201 A: GUA ORR
aegrota Arrow, 1917: 171 A: SD ORR
affinis Ganglbauer, 1882c: 245 E: GR (Sámos) A: TR
afghana Balthasar, 1968: 150 (Iliola) A: AF PA
agilis Arrow, 1917: 207 A: BT NP SD XIZ
albopilosa albopilosa Hope, 1839b: 70 (Euchlora) A: HEI JA JIL LIA NC SC
immarginata Reitter, 1903a: 65

albopilosa gracilis Schönfeldt, 1890: 171 A: JA (Liu Kiu)
horii Nomura, 1964b: 65

albopilosa sakishimana Nomura, 1964b: 65 A: JA (Ishigaki-jima, Miyake-jima, Iriomote-jima)
albopilosa trachypyga Bates, 1866: 344 (Euchlora) A: TAI
albopilosa yashiroi Sawada, 1950b: 307 A: JA (Okinawa-shotō)
albopilosa yonaguniana Nomura, 1964b: 65 A: JA (Yonaguni)
amarginata B. Zhang & Lin, 2008: 129 A: YUN SCH
amoena Frey, 1971a: 113 A: GUI JIA SCH SHN XIZ YUN ZHE
rufithorax Ohaus, 1933: 477 [HN]

anguliceps Arrow, 1917: 158 A: FUJ GUA GUI GUX SCH YUN ZHE ORR
corneola Lin, 2002b: 421

angusta Arrow, 1912a: 79 A: BT NP SD XIZ
latiungula Lin, 1981: 379

anthracina Arrow, 1912a: 76: BT SD
anthusa Ohaus, 1937b: 95 A: FUJ TAI
antiqua Gyllenhal, 1817: 196 (Melolontha) A: GUA GUI JIX NP AURi ORR
aerea Perty, 1831: 34 (Euchlora)
australasiae Blackburn, 1892b: 113
lugubris Wiedemann, 1823: 93 (Melolontha)
nigricans Kirby, 1825: 569 (Mimela)

artemida Prokofiev, 2015: 37 A: YUN ORR
atrovirens Lin, 2000a: 40 A: YUN
aulacoides Ohaus, 1915b: 327 A: TAI
poecilochalcea Ohaus, 1915b: 328

aulax Wiedemann, 1823: 93 (Melolontha) A: ANH FE FUJ GUA GUI GUX HAI HEI HKG HUB HUN JIL JIX LIA
NC SC SCH XIZ YUN ZHE ORR
costifera Reitter, 1895f: 209

330

SCARABAEIDAE

marginalis Newman, 1838c: 385

aureoflava Arrow, 1917: 190 A: NP SCH YUN ORR
ausonia Erichson, 1847a: 620 E: FR IT SP N: AG TU
cogina Reitter, 1903a: 76
neapolitana Reitter, 1903a: 75
sicula Ganglbauer, 1882c: 175
signaticollis Ohaus, 1915b: 312

australis Lin, 2002b: 421 A: FUJ GUA GUX HAI HUN
babai Kobayashi, 1987b: 24 A: TAI
badia Ohaus, 1925a: 127 A: TAI
barbellata Lin, 1996a: 168 A: HAI
barbellicauda Lin, 1999a: 171 A: YUN
bedeli Ohaus, 1914b: 198 A: ANH GUI YUN ORR
bella Arrow, 1917: 209 A: AP SD ORR
bengalensis C.É. Blanchard, 1851b: 182 A: BT NP UP ORR
pallidicollis C.É. Blanchard, 1851b: 181

biharensis Arrow, 1917: 166 A: NP ORR
bilineata Reitter, 1903a: 60 A: KZ
bilobata Arrow, 1912b: 329 A: NP SD ORR
bilunata Fairmaire, 1889b: 341 A: BT FUJ GUA GUI GUX NP SCH SD TAI YUN ORR
senooi Kobayashi, 1987b: 29 (Blitopertha)
sorortertia Prokofiev, 2013: 105
taitungensis Kobayashi, 1987b: 28 (Blitopertha)

bivirgulata Fairmaire, 1893e: 310 A: GUI YUN ORR
discimaculata R. Paulian, 1959b: 75

brancuccii Sabatinelli, 1991c: 399 A: NP
brevihirta Lin, 1996: 167 A: GUA
cantori Hope, 1839b: 73 (Euchlora) A: AP HP NP “Thibet” ORR
severa Burmeister, 1855: 504

capillula Lin, 1996a: 166 A: YUN
cariniventris Lin, 2002a: 353 A: GUA HAI
castaneoventris Bates, 1866: 344 (Euchlora) A: TAI
chamaeleon Fairmaire, 1887d: 317 A: BEI FE GAN HEB HEI JIL LIA MG NE NMO SHA SHN SHX TIA
dubia Ballion, 1871a: 344 [HN]

chloroderma Arrow, 1913c: 403 A: JA
chlorosoma Arrow, 1917: 231 A: HP NP ORR
choui Lin, 1999a: 168 A: YUN
chromicolor Burmeister, 1855: 505 A: FUJ
cinderella Arrow, 1917: 196 A: NP SD
hirticollis Frey, 1975d: 314

SCARABAEIDAE

331

clerica Arrow, 1917: 150 A: SD
collotra B. Zhang & Lin, 2008: 133 A: SCH YUN ORR
comma Arrow, 1917: 192 A: NP SD
concavifronta Lin, 1999a: 170 A: YUN
controversa Hope, 1842a: 62 [=1843a: 64, 1845: 14] A: GUA
corpulenta Motschulsky, 1854b: 28 A: ANH FUJ GAN GUI HEB HEI HEN HUB HUN JIA JIL JIX LIA NC NIN
NMO SC SCH SHA SHG SHN SHX ZHE
gottschei H.J. Kolbe, 1886: 190
pallidiventris Gautier des Cottes, 1870: 104 (Anisoplia)
planerae Fairmaire, 1891f: ccv

corrugata Bates, 1866: 343 A: FUJ GUA TAI
holosericioides Niijima & Kinoshita, 1927: 50

corvina Arrow, 1917: 214 [EA] A: SD
costulata Fairmaire, 1887b: 111 A: SCH YUN
coxalis Bates, 1891b: 77 A: ANH FUJ GUA GUI GUX HAI HUB HUN JIA JIX LIA SCH SHG SHX TAI YUN ZHE
ORR
fuscoviolacea Ohaus, 1917a: 7
heydeni J. Frivaldszky, 1892: 124 (Euchlora)
streptopyga Ohaus, 1915b: 329
viridirufa Ohaus, 1917a: 7

cpustulata cpustulata Hirayama, 1940: 62 A: JA
matsumurai Sawada, 1950b: 305
tokarana Nomura, 1961b: 74

crassipyga Benderitter, 1923b: 218 A: YUN “Thibet”
cruralis Fairmaire, 1887b: 110 A: YUN
cuprea Hope, 1839b: 72 (Euchlora) A: FE HEI JA JIL LIA NC NE
cupripes Hope, 1839b: 68 (Euchlora) A: YUN ORR
cyanipennis Lin, 1999a: 169 A: YUN ORR
cypriogastra Ohaus, 1938: 264 A: FUJ GUA HUN JIX TAI
daimiana Harold, 1877e: 354 A: JA
delavayi Fairmaire, 1886: 327 A: GUI YUN ORR
aulacina Frey, 1972d: 248
semiaurea Arrow, 1917: 205

deltoides Lin, 1999a: 170 A: YUN ORR
densa Arrow, 1917: 221 A: YUN ORR
dentifera Lin, 2002b: 422 A: ANH FUJ GUA GUI GUX HAI SCH YUN
devota P. Rossi, 1790: 19 (Melolontha) E: FR IT SP N: TU
apicalis Mulsant, 1842: 480 (Euchlora)
villosa C.É. Blanchard, 1851: 184
versicolor Mulsant, 1842: 480 (Euchlora)

332

SCARABAEIDAE

dimidiata barbata Burmeister, 1855: 505 A: AF HP KA PA UP
dimidiata dimidiata Hope, 1831: 23 (Euchlora) A: AP BT NP SD XIZ YUN ORR
sulcata Hope, 1839b: 72 (Euchlora)

diplopsyla Prokofiev, 2015: 44 A: YUN ORR
dorsalis Fabricius, 1775: 35 (Melolontha) A: AF BT HP NP PA SD UP ORR
centralis Nonfried, 1893: 335 [HN]
fraterna Burmeister, 1844: 232
imitatrix Nonfried, 1894: 12

dubia abchasica Motschulsky, 1854b: 29 A: AB AR GG ST
dubia dubia Scopoli, 1763: 3 (Scarabaeus) E: AL AU BE BH BU BY CR CT CZ DE EN FI FR GB GE HU IR IT LA
LT LU MC MD ME NL NR NT PL RO SB SK SL SP ST SV SZ UK A: TR WS NARi
aenea De Geer, 1774: 277 (Scarabaeus)
albanica Reitter, 1918a: 43
baudueri Preudhomme de Borre, 1881b: cxx
bicolor Schilsky, 1888a: 184
bicolor Schilsky, 1888a: 185 [HN]
caerulea A.G. Olivier, 1789: no 5: 36 (Melolontha)
caeruleocephala A.G. Olivier, 1789: no 5: 36 (Melolontha)
coerulescens Schilsky, 1888: 184
cuprea Westhoff, 1882: 154 [HN]
cyanicollis A. Villa & G.B. Villa, 1833: 34
cyanocephala Fabricius, 1801b: 169 (Melolontha)
fallax Schilsky, 1888a: 185
frischii Fabricius, 1775: 37 (Melolontha)
humeralis Schilsky, 1888a: 185 [HN]
incerta Mulsant, 1842: 476 (Euchlora)
janthina Gmelin, 1790: 1561 (Scarabaeus)
julii Fabricius, 1792b: 167 (Melolontha)
lutescens Schilsky, 1888a: 185
maculata Schilsky, 1888a: 185
maculipes Burmeister, 1844: 257
marginata Schilsky, 1888a: 184
micans Mulsant, 1842: 478 (Euchlora)
nigrita Fabricius, 1792b: 167 (Melolontha)
oblonga Fabricius, 1777: 209 (Melolontha)
ovata Burmeister, 1844: 257
pedemontana Tournier, 1865: 267
pygidialis Schilsky, 1888a: 184 [HN]
rubrocuprea Mulsant, 1842: 478 (Euchlora)
sublaevigata Motschulsky, 1854b: 29
sublucida Motschulsky, 1856: 81

SCARABAEIDAE
varians Mulsant, 1842: 476 (Euchlora) [HN]
virescens Schilsky, 1888a: 184
viridicuprea Mulsant, 1842: 478 (Euchlora)
viridis Schilsky, 1888a: 185 [HN]

ebenina Fairmaire, 1886: 328 A: GAN GUI HEB HUB HUN SCH SHA SHX
ochracea Benderitter, 1923a: 8

edentula amamiana Nomura, 1962: 39 A: JA (Amami-Ôshima)
edentula edentula Ohaus, 1925b: 128 A: GUA HKG TAI ORR
fusca Lin, 1985: 121

edentula okinawana Nomura, 1965: 6 A: JA (Okinawa-shotō)
edentula yaeyamana Nomura, 1965: 5 A: JA (Ishigaki-jima, Iriomote-jima)
egregia Gahan, 1896: 451 A: SA AFR
muchei Mikšić, 1960: 374

errans diluta Motschulsky, 1854b: 30 (Psammoscapheus) E: ST
plustschevskyi Reitter, 1894f: 239

errans errans Fabricius, 1775: 37 (Melolontha) E: AB BU KZ MD RO ST UK A: KZ
brunea Ménétriés, 1832: 186
circumcincta Reitter, 1896a: 34
desertorum Motschulsky, 1854b: 31 (Psammoscapheus)
praticola Fabricius, 1781: 42 (Melolontha)

erubescens Ohaus, 1925b: 129 A: TAI
esakii Sawada, 1950b: 273 A: JA
euops Arrow, 1917: 173 A: NP UP ORR
exoleta Faldermann, 1835b: 381 A: BEI GAN HEB HEI HEN LIA NMO QIN SHA SHN SHX
exoletoides Lin, 2000b: 121 A: HEN SHA
expansa expansa Bates, 1866: 343 (Euchlora) A: TAI
expansa lanshuensis Nomura, 1977a: 81 A: TAI (Lanshu)
expansa lutaoensis Nomura, 1977a: 82 A: TAI (Lutao)
fasciolata Ohaus, 1925b: 124 A: TAI
formosana Niijima & Kinoshita, 1927: 61 (Phyllopertha)

femoralis A.G. Olivier, 1789: no 5: 35 (Melolontha) A: “Indes Orientales”
ferruginea Marseul, 1867b: xxxvi N: AG MO
filigera Ohaus, 1933: 476 A: NP SD YUN ORR
flavilimbata Lin, 2002: 353 A: HAI
flavipunctuata Lin, 1999a: 168 A: YUN
flaviventris Arrow, 1912a: 74 A: BT NP SD XIZ
flavofasciata Arrow, 1912a: 72 A: BT NP SD ORR
flavofemorata Lin, 1989: 87 A: SHA
flavonotata Arrow, 1912a: 73 A: BT SD
flavopicta Arrow, 1912a: 75 A: BT NP SD
francottei Sabatinelli, 1997b: 244 A: YUN ORR

333

334
speciosa Lin, 1999a: 167

fukiensis Machatschke, 1955b: 501 A: FUJ
fuscicauda Lin, 1999a: 169 A: YUN ORR
fusitibia Lin, 1992: 521 A: YUN ORR
geniculata Motschulsky, 1866: 171 (Rhinoplia) A: HEI JA JIL LIA NE SC
pleurimargo Reitter, 1903a: 72

glabra Lin, 1981: 378 A: XIZ
gracilenta Reitter, 1903a: 67 A: SCH YUN
graminea Ohaus, 1905: 86 A: FUJ GUA GUX HUN ORR
grandis Hope, 1839b: 69 (Euchlora) A: “N. India” SD ORR
granulicauda Lin, 1989: 88 A: SCH SHA SHX
granuliformis Lin, 1996b: 310 A: YUN ORR
gudzenkoi Jakobson, 1903: xiii A: FE HEI JIL LIA NE
hemiseca B. Zhang & Lin, 2008: 130 A: YUN
hirsutoides Lin, 1996a: 159 A: ANH FUJ
hirsutula Nonfried, 1892a: 87 A: GUX HUN JIX ZHE
hirtidorsa Lin, 1999a: 167 A: YUN
hoplocosmeta Ohaus, 1925b: 125 A: TAI
ignicolor Reitter, 1903a: 73 A: HEI JIL LIA “Korea”
ignipes Lin, 1996b: 304 A: FUJ GUA GUX HAI
imperialis Arrow, 1899a: 264 A: YUN ORR
polychroma Ohaus, 1905: 85

inconcinna Bates, 1866: 343 A: FUJ TAI
chlorochalcea Ohaus, 1925b: 124
jonasi Ohaus, 1915b: 326
obscurior Ohaus, 1917a: 4

insulicola Lin, 2000a: 39 A: HAI
interrupta Lin, 2002b: 423 A: FUJ GUA
ishidai Niijima & Kinoshita, 1927: 52 A: TAI
itohi Miyake, 1987: 4 A: TAI
iwasei Miyake, 1994b: 144 A: YUN ORR
japonica Arrow, 1913c: 401 A: JP
borealis Arrow, 1913c: 402
izuensis Sawada, 1942b: 45

keithi Zorn, 2011: 311 A: YUN
kirgisica Borodin, 1915: 511 E: ST A: KI KZ
koreana J.I. Kim, 1997: 279 A: SC
kuatuna Machatschke, 1955b: 506 (Mimela) A: FUJ GUA GUX
laccata B. Zhang & Lin, 2008: 132 A: GUA YUN
laevisulcata Fairmaire, 1888a: 19 A: ANH FUJ ZHE JIX
holcoptera Fairmaire, 1889a: 26

SCARABAEIDAE

SCARABAEIDAE
latericostulata Lin, 1992: 522 A: SCH XIZ
lateripila Lin, 2002b: 423 A: FUJ
laticlypea Lin, 1999a: 166 A: YUN
leonfairmairei Zorn, 2005: 308 [RN] A: SCH
micans Fairmaire, 1897e: 214 [HN]

leptopoda Lin, 1992: 506 A: YUN ORR
libidinosa Ohaus, 1917a: 5 A: TAI
lignea Arrow, 1917: 202 A: YUN ORR
tongyaii Ohaus, 1938: 265

limbifera Ohaus, 1915b: 330 A: TAI
lineatopennis C.É. Blanchard, 1851: 191 A: BT HP NP PA SD UP XIZ ORR
tenuipes Lin, 1981: 380

loi Kobayashi, 1987b: 27 A: TAI
longicarcarata Lin, 2002b: 424 A: FUJ
longiclypea Lin, 1999a: 172 A: YUN ORR
longicornis Lin, 1999a: 168 A: YUN
longilamina Lin, 1999a: 171 A: YUN
lucens Ballion, 1871b: 155 [RN] A: FE (Kuril’skie ostrova, Sakhalin) JA
impressibasis Reitter, 1903a: 73
lucidula Motschulsky, 1866: 171 (Rhombonyx) [HN]
tokioensis Niijima & Kinoshita, 1923: 105

luculenta Erichson, 1847a: 620 A: ES FE HEB HEI JIL LIA MG NC NMO SHA SC TIA
daurica Mannerheim, 1849: 236
ruficornis Motschulsky, 1854b: 30
purpurea Niijima & Kinoshita, 1923: 100
rufofemorata Reitter, 1903a: 74

luniclypealis Lin, 1979a: 31 A: NP XIZ
machatschkei Frey, 1972d: 249 A: YUN
margina Lin, 1999a: 167 A: YUN
marginipennis Arrow, 1912a: 78 A: BT NP SD ORR
martinii Hope, 1839b: 70 (Euchlora) A: CH
masaakii Nomura, 1977a: 82 A: TAI
matzenaueri Reitter, 1918a: 42 E: AL ME
metonidia Reitter, 1903a: 61 A: KI KZ XIN
calliura Reitter, 1903a: 60

microda B. Zhang & Lin, 2008: 136 A: YUN
millestriga asticta Lin, 2002b: 424 A: FUJ GUA HEN HUB HUN
millestriga millestriga Bates, 1891b: 78 A: GUI SCH YUN
miyakona Nomura, 1977a: 79 A: JA
mizusawai Kobayashi, 1987b: 25 A: TAI
mongolica brevilimbata Lin, 1989: 89 A: ANH FUJ SCH SHA

335

336

SCARABAEIDAE

mongolica mongolica Faldermann, 1835b: 379 A: BEI FE HEB HEI JIL LIA MG NC NMO SC SHN
anomala Kraatz, 1879a: 234 (Mimela)
coerulea Ohaus, 1930: 147
subaurata Ballion, 1871a: 345

monogramma Zorn, 2011: 301 A: GUI GUX
montana Lin, 1996a: 160 A: JIX
multistriata Motschulsky, 1862a: 7 (Heteroplia) A: JA
puncticollis Harold, 1877e: 352

myriopila Lin, 2002: 354 A: HAI
nainitalii Shah, 1983: 403 A: UP
nepalensis Machatschke, 1966: 184 A: NP
nigricollis Lin, 1992: 522 A: SCH
nigripes Nonfried, 1892a: 88 A: FUJ GUA GUX ORR
nigrolineata Kobayashi, 1987b: 22 A: TAI
nigrovirens Reitter, 1894b: 46 A: TD UZ
sublucida Ballion, 1871a: 344 [HN]

niijimae Ohaus, 1925b: 126 A: JA (Ishigaki-jima)
nubeculosa nubeculosa Ohaus, 1905: 84 A: YUN ORR
nomurai Kobayashi, 1995a: 37 A: TAI
obliquipunctata Lin, 1989: 88 A: SHA
obliquisulcata Lin, 2002b: 425 A: FUJ GUA GUI GUX HAI HUN HUB JIX SHA ZHE
obscurata Reitter, 1903a: 65 A: FUJ ZHE
fuscipennis Reitter, 1903a: 65

obscuripes Fairmaire, 1892a: 97 N: EG AFR
femorata Reitter, 1903a: 70

ochii Miyake, 1987: 2 A: TAI
octiescostata Burmeister, 1844: 243 A: JA
ogloblini S.I. Medvedev, 1949: 171 A: FE HEI JIL LIA NC
oleaginea Ménétriés, 1836b: 181 A: ES
olivea Lin, 2002: 354 A: HAI
opaconigra Frey, 1972d: 247 A: FUJ
opalina Fairmaire, 1887b: 111 A: SCH XIZ YUN ORR
kambaitina Ohaus, 1937a: 3

osakana Sawada, 1942a: 40 A: JA
osmanlis C.É. Blanchard, 1851: 185 E: BU GR TR A: SY TR
haemorrhoidalis Reitter, 1903a: 78

oxiana Semenov, 1891: 321 A: AF KZ TD TM UZ
palaestina Pic, 1905c: 129 A: IS
pallens Semenov & S.I. Medvedev, 1949: 198 (Iliola) A: KZ
palleola Gyllenhal, 1817: 107 (Melolontha) A: “India orientalis”
parallela Benderitter, 1929: 107 A: YUN ORR

SCARABAEIDAE
perplexa diana B. Zhang & Lin, 2008: 134 A: YUN
perplexa perplexa Hope, 1839b: 70 (Euchlora) A: AP BT FUJ GUA HKG NP SD XIZ
herbea Lin, 1981: 378

phaeogastra Ohaus, 1925b: 129 A: TAI
pilicauda Lin, 2002: 354 A: HAI
pilifrons Lin, 1979a: 30 A: XIZ YUN
piliscutella Lin, 1981: 380 A: SCH XIZ YUN
pilosella Fairmaire, 1898b: 384 A: GUI HUB SCH
planicauda Lin, 1996a: 164 A: YUN
planicorna Lin, 1996b: 309 A: TAI
rufina Chujo, 1940: 76 [HN]

platypyga Fairmaire, 1893e: 309 A: BT NP SD XIZ YUN ORR
protea Arrow, 1912a: 80

plebeja plebeja A.G. Olivier, 1789: no 5: 25 (Melolontha) N: EG AFR
atrata Ohaus, 1914a: 154
femoralis Laporte, 1840: 136 (Euchlora) [HN]
melanopa Reitter, 1903a: 70
scutellaris Klug, 1835: 35

polita C.É. Blanchard, 1851: 182 [HN] A: BT HP NP PA UP ORR
polyanor Ohaus, 1917a: 4 A: TAI
pontualei Sabatinelli, 1997b: 247 A: SD
porrecta Zorn, 2011: 301 A: GUA HKG
potanini S.I. Medvedev, 1949: 143 A: GAN
praeclara R. Paulian, 1959b: 80 A: YUN ORR
praecoxalis Ohaus, 1914b: 204 A: GUI YUN ORR
praenitens Arrow, 1917: 171 A: NP ORR
profundisulcata Lin, 2002: 354 A: HAI
projecta Lin, 2002: 355 A: HAI
propinqua Arrow, 1912a: 77 A: BT NP SD UP XIZ ORR
picina Lin, 1981: 379

puncticlypea Lin, 1992: 522 A: YUN
pyroscelis Hope, 1841b: 66 (Mimela) A: BT NP UP “South China” ORR
fulgens C.É. Blanchard, 1851: 195
rufipes Burmeister, 1855: 506

pyxexcavata B. Zhang & Lin, 2008: 131 A: YUN
pyxofera B. Zhang & Lin, 2008: 135 A: ANH HUB JIX
quadripunctata A.G. Olivier, 1789: no 5: 82 (Melolontha) E: PT SP N: MO
pallidior Pic, 1905b: 105
profuga Erichson, 1847a: 617
tingitana C.É. Blanchard, 1851: 185
vagans Erichson, 1847a: 620

337

338

SCARABAEIDAE

quelparta Okamoto, 1924: 177 A: SC
raui Ohaus, 1914b: 206 A: GUX ORR
generosa Benderitter, 1929: 106
pygidialis Fairmaire, 1889b: 341 (Aprosterna) [HN]

rubripes rubripes Lin, 1996b: 302 A: GUA GUX HEI HAI ZHE ORR
rubripes virescens Lin, 1996b: 304 A: TAI
rufiventris Kollar & L. Redtenbacher, 1844: 526 A: BT GUI HEI HP KA NP SCH SD UP XIZ YUN ORR
iridiventris Fairmaire, 1878a: 102 (Aprosterna)
laevissima Burmeister, 1855: 506 (Aprosterna)
striolata C.É. Blanchard, 1851: 195

rufocuprea Motschulsky, 1861a: 14 A: FE (Kuril’skie ostrova, Sakhalin) HEI JA JIL LIA SC
hachijoensis Nomura, 1969: 74
motschulskyi Harold, 1877e: 351

rufopartita Fairmaire, 1889a: 27 A: GUI HUB JIX “Mongolei: Ho-chan”
colorata Reitter, 1903a: 58

rufozonula Fairmaire, 1887b: 112 A: SCH YUN ORR
fascipennis Reitter, 1903a: 57
bioculata Fairmaire, 1887b: 112

rugiclypea Lin, 1989: 89 A: FUJ GUA GUX HAI HUB HUN JIX SCH SHA SHX YUN
rugipennis C.É. Blanchard, 1851: 182 A: “Inde sept.”
rugosa Arrow, 1899a: 263 A: BT HP NP PA SD UP ORR
rugulipennis Lin, 1999a: 171 A: YUN
russiventris Fairmaire, 1893d: 290 A: FUJ GUA GUI GUX YUN ORR
lasiocaula Ohaus, 1914b: 212
shanica Arrow, 1917: 218

sagiens Ohaus, 1925a: 75 [RN] A: TAI
sagax Ohaus, 1917a: 6 [HN]

saitoi Kobayashi, 1995a: 35 A: TAI
sakaii Miyake, 1987: 1 A: TAI
sapa Miyake, 1994b: 145 A: GUX ORR
sauteri sauteri Ohaus, 1915b: 329 A: TAI
sauteri yanoi Nomura, 1959b: 50 A: JA (Ishigaki-jima, Iriomote-jima)
schereri Frey, 1965a: 93 A: YUN ORR
schoenfeldti Ohaus, 1915b: 322 A: JA
miyakensis Nomura, 1967: 57

semenovi S.I. Medvedev, 1949: 182 A: TM
semicastanea Fairmaire, 1888a: 21 A: ANH FUJ GUA GUX HKG HUN JIA JIX SHA SHG SHX ZHE
castaneipennis Fairmaire, 1891f: cciv (Aprosterna)

semiovalis Lin, 2002b: 425 A: FUJ GUA
sericipennis Lin, 2002b: 426 A: FUJ GUA HUN
shimenensis Lin, 2000a: 39 A: HUN

SCARABAEIDAE
shirakii Nomura, 1959b: 49 A: JA
sieversi Heyden, 1887a: 266 A: BEI FUJ GUI HEI HUB JIL LIA NC NE SC
atrocoerulea Reitter, 1903a: 63
subpurpurea Reitter, 1903a: 63

similopyga Miyake, 1987: 3 A: TAI
sinica Arrow, 1915: 231 [RN] A: GUA HKG
mimeloides Reitter, 1903a: 71 [HN]

siniopyga Ohaus, 1917a: 8 A: TAI
solida flexuosa Reitter, 1903a: 76 A: TR
solida solida Erichson, 1847a: 622 E: BU GR MC RO SB TR
amydra Ohaus, 1930: 147

spatuliformis Lin, 2002b: 422 A: FUJ
spiloptera Burmeister, 1855: 500 A: BT FUJ GUA GUI JIA JIX SCH SD ZHE ORR
densestrigosa Fairmaire, 1888a: 20

splendida Ménétriés, 1832: 185 E: AB GG ST A: IN
collaris Motschulsky, 1854b: 29 [HN]
turcomanica Harold, 1869d: 123 [RN]
variabilis Motschulsky, 1854b: 30 [HN]
viridisplendens Ménétriés, 1832: 185

stenodera Arrow, 1917: 172 A: BT ORR
straminea Semenov, 1891: 320 A: GAN SCH SHX
sublunalis Lin, 2002b: 426 A: GUA YUN
subpilosa Lin, 1996a: 162 A: HUN GUI
subtomentella Lin, 1996a: 163 A: YUN
subtrinata Lin, 1996a: 162 A: YUN
subvittata Reitter, 1903a: 68 A: FE
sulcipennis Faldermann, 1835b: 378 (Idiocnema) A: GUI HEB HEI HKG JIL LIA “China borealis”
idiocnema Burmeister, 1855: 497

taeniana B. Zhang & Lin, 2008: 128 A: YUN
taiwana Kobayashi, 1987b: 27 A: TAI
takasagoensis Sawada, 1941b: 51 (Phyllopertha) A: TAI
takeshii Sabatinelli, 1997b: 245 A: NP
tarowana Sawada, 1941b: 51 (Phyllopertha) A: TAI
testacea Hope, 1831: 24 A: BT NP UP
thoracica Zoubkov, 1833: 323 (Melolontha) A: KZ
transversa Lin, 1999a: 169 A: YUN
triancistris Lin, 1999a: 172 A: YUN
triangularis Schönfeldt, 1890: 171 A: JA
kyushuensis Nomura, 1964b: 68

trichophora Lin, 1996a: 160 A: HUN
trillesi R. Paulian, 1959b: 58 A: MAC ORR

339

340

SCARABAEIDAE

tristis Arrow, 1917: 158 A: SD PA UP
trivirgata Fairmaire, 1888a: 21 A: BT FUJ GAN GUI JIX NP SCH SHX YUN ORR
bifasciata Benderitter, 1929: 103
biguttata Frey, 1971a: 114

trochanterica Arrow, 1917: 238 A: AP BT NP SD
truncata chlorochelys Arrow, 1912b: 340 A: GUI NP YUN ORR
truncata truncata Bates, 1890: 244 A: GUI JIX SCH XIZ ORR
varicolor Gyllenhal, 1817: 114 (Melolontha) A: BT GUA GUI GUX NP SD TAI UP XIZ YUN ORR
variegata Hope, 1831: 24 A: BT NP SD
vestigator Arrow, 1917: 379 A: PA
virens Lin, 1996b: 307 A: FUJ GUA GUI GUX HAI HEN HUB HUN JIX SCH SHN SHX YUN ZHE ORR
viridana H.J. Kolbe, 1886: 192 (Euchlora) A: FE HEI JIL LIA SC NC
viridicostata Nonfried, 1892a: 86 A: ANH FUJ GUA GUI GUX HUB HUN JIX SCH YUN ZHE ORR
viridimicans Benderitter, 1929: 106 A: FUJ HKG ORR
viridilatera Arrow, 1917: 181 A: SD ORR
viridisericea Ohaus, 1905: 85 A: JIX ORR
tectiformis Frey, 1970b: 174

vitalisi Ohaus, 1914b: 207 A: FUJ GUA JIX ORR
vitis Fabricius, 1775: 37 (Melolontha) E: AL AU BH BU CR GR HU IT MD ME RO SB SK SL SZ UK
azurescens Reitter, 1903a: 77
cupreonitens Bau, 1883: 286
dichroa Reitter, 1903a: 77
lutea Schilsky, 1888a: 185 [HN]
metallica Burmeister, 1844: 256
signata Schilsky, 1888a: 185
signaticollis Burmeister, 1844: 256
variabilis Schilsky, 1888a: 185 [HN]
viridicollis Schilsky, 1888a: 185

vittata Gebler, 1841a: 372 A: KI KZ UZ XIN
circumdata Heyden, 1885a: 274
picticollis Ballion, 1878: 28

xantholoma Lin, 1981: 381 A: XIZ
xanthopleura Arrow, 1913c: 402 A: JA
xanthoptera C.É. Blanchard, 1851: 195 A: HP NP PA SD UP
prasinicollis Bates, 1891a: 18

yunnana Fairmaire, 1886: 328 A: GUI YUN ORR
semismaragdina Fairmaire, 1889a: 27

genus Asiopertha Machatschke, 1957: 131 type species Phyllopertha nazarena Marseul, 1878
fulvicornis Burmeister, 1855: 502 (Anomala) A: “Mesopotamien”
ganglbaueri Reitter, 1885c: 394 (Phyllopertha) A: IN SY TR
nazarena Marseul, 1878: 71 (Phyllopertha) A: IN IS TR

SCARABAEIDAE

341

genus Blitopertha Reitter, 1903a: 85 type species Melolontha lineata Fabricius, 1798
gracilipedes Baraud, 1990e: 295 N: MO
lineata Fabricius, 1798: 133 (Melolontha) E: SP N: AG MO TU
algirica Reiche, 1864: 241 (Phyllopertha)
lineigera Reitter, 1903a: 86
nigra Petrovitz, 1959d: 188
oberthurii Fairmaire, 1871a: 379 (Phyllopertha)

lineolata lineolata Fischer von Waldheim, 1824: 216 (Anisoplia) E: AL BU MC MD GR KO ME RO SB ST TR UK
arenaria Brullé, 1832: 177 (Anisoplia)
flavipennis Reitter, 1903a: 86
krueperi Brenske, 1884: 77 (Phyllopertha)
senticola Reitter, 1888c: 292 (Phyllopertha)

lineolata vanensis Shokhin, 2010: 40 A: TR
nigripennis Reitter, 1888c: 292 (Phyllopertha) E: AB AR GG ST A: CY IN IS JO LE SY TM TR
abdita Petrovitz, 1959d: 186
majuscula S.I. Medvedev, 1949: 222

genus Callistethus C.É. Blanchard, 1851: 198 type species Callistethus consularis C.É. Blanchard, 1851 (=Mimela
auronitens Hope, 1836)
Hadropopillia Kraatz, 1892a: 289 type species Popillia regina Newman, 1838
Pachystethus C.É. Blanchard, 1851: 201 type species Popillia vidua Newman, 1838
Poecilosticta Kraatz, 1892a: 290 type species Mimela variegata Walker, 1859
Spileuchlora Ohaus, 1903: 209 type species Spileuchlora waterstraati Ohaus, 1903
Spilota Burmeister, 1844: 266 type species Melolontha marginata Fabricius, 1792
Zaspilota Casey, 1915: 47 type species Anomala cupricollis Chevrolat, 1834

afghanus Balthasar, 1967a: 141 (Spilota) A: AF
amphilissus Arrow, 1917: 245 (Anomala) A: HP NP
anwari Abdullah & Roohi, 1969: 123 (Anomala) A: PA
ashrafii Abdullah & Roohi, 1968: 435 (Anomala) A: PA
auronitens Hope, 1836: 114 (Mimela) A: BT HP KA NP PA SCH SD UP XIZ YUN ORR
consularis C.É. Blanchard, 1851: 198

excisipennis Lin, 1981: 381 A: XIZ
formosanus Kobayashi, 1987b: 21 A: TAI
plagiicollis isidai Miyake, 1987: 5 A: TAI
plagiicollis plagiicollis Fairmaire, 1886: 329 (Spilota) A: ANH BEI FE FUJ GUA GUI GUX HEB HEI HEN HUB
HUN JIA JIX LIA MG NC SC SCH SHA SHX XIZ YUN ZHE ORR
impictus Bates, 1888b: 374 (Paraspilota)

pterygophorus Ohaus, 1903: 217 A: BT SD ORR
quadrii Abdullah & Roohi, 1968: 435 (Anomala) A: PA
shafqati Abdullah & Roohi, 1968: 439 (Anomala) A: PA
stoliczkae Sharp, 1878b: 173 (Anomala) A: BT HP KA NP PA SD SCH XIZ YUN
seminitidus Fairmaire, 1889a: 25

342

SCARABAEIDAE

tumidicauda tumidicauda Arrow, 1912a: 83 (Anomala) A: AP BT SD ORR
varius Newman, 1839: 365 (Popillia) A: NP ORR
popiliopsis Candèze, 1869: 43 (Anomala)

xanthonotus Arrow, 1917: 244 (Anomala) A: HP PA
yunusi Abdullah & Roohi, 1969: 124 (Anomala) A: PA
genus Choumala Kobayashi, 2008: 69 type species Choumala choui Kobayashi, 2008
choui Kobayashi, 2008: 70 A: TAI
genus Cyriopertha Reitter, 1903a: 91 type species Anisoplia glabra Gebler, 1841
subgenus Apleopertha Semenov, 1903g: 390 type species Phyllopertha reitteri Semenov, 1891
reitteri Semenov, 1891: 324 (Phyllopertha) A: TD “Turkestan”
subgenus Cyriopertha Reitter, 1903a: 91 type species Anisoplia glabra Gebler, 1841
glabra Gebler, 1841a: 372 (Anisoplia) A: KI KZ TD TM UZ WS XIN
asiatica Ballion, 1871a: 343 (Phyllopertha)
fulvibasis Reitter, 1903a: 91
monochroa Semenov, 1891: 325 (Phyllopertha)
multicolor Reitter, 1890l: 264 (Phyllopertha)

subgenus Pleopertha Reitter, 1903a: 92 type species Anisoplia arcuata Gebler, 1832
arcuata Gebler, 1832: 51 (Anisoplia) A: BEI ES FE GAN HEB HEI HEN JIL LIA MG NIN NMO SHA
arcuata C.É. Blanchard, 1851: 179 (Phyllopertha) [HN]
flavicollis Heyden, 1889c: 661 (Phyllopertha)
laeta Reitter, 1903a: 92 (Cyriopertha)
lunata Faldermann, 1835b: 383 (Anomala)

genus Exomala Reitter, 1903a: 88 type species Phyllopertha orientalis C.O. Waterhouse, 1875
subgenus Exomala Reitter, 1903a: 88 type species Phyllopertha orientalis C.O. Waterhouse, 1875
ohdaiensis Sawada, 1941b: 49 (Phyllopertha) A: JA
okiwaensis Ohaus, 1925b: 130 (Phyllopertha) A: JA (Okinawa-shotō)
orientalis C.O. Waterhouse, 1875a: 108 (Phyllopertha) A: JA HEI JIL LIA NC SC NARi ORR
flavipennis Reitter, 1903a: 89 (Blitopertha)
tanbaensis Niijima & Kinoshita, 1923: 128 (Phyllopertha)
xanthogastra Harold, 1881: 90 (Phyllopertha)

pallidipennis Reitter, 1903a: 89 (Blitopertha) A: FE HEI JIL LIA MG NC NE SC
nigrata Reitter, 1903a: 89 (Blitopertha)

subgenus Neoblitopertha Baraud, 1991b: 50 type species Melolontha camprestris Latreille, 1804
adriatica Petrovitz, 1968g: 485 (Blitopertha) E: AL GR ME
bileki Petrovitz, 1968g: 487 (Blitopertha) A: TR
campestris Latreille, 1804b: 195 (Melolontha) [NP] E: AN AU FR IT SL SP SZ
arenaria Laporte, 1840: 151 (Anisoplia) [HN]
circumcincta Mulsant, 1842: 497 (Phyllopertha)

SCARABAEIDAE
hispanica Báguena, 1955: 293
mulsanti Báguena, 1955: 293
occidentalis Mulsant, 1842: 497 (Phyllopertha)
sabulosa Mulsant, 1842: 497 (Phyllopertha)

leonii Luigioni, 1932b: 138 (Phyllopertha) E: IT
succincta Laporte, 1840: 151 (Anisoplia) E: AU FR IT SL SZ
abbreviata Mulsant, 1842: 496 (Phyllopertha)
cruciata Mulsant, 1842: 496 (Phyllopertha)
maculata Mulsant, 1842: 496 (Phyllopertha)
pauperata Mulsant, 1842: 496 (Phyllopertha)

subgenus Trichopertha Reitter, 1903a: 85 type species Anisoplia hirtella Brullé, 1832
hirtella Brullé, 1832: 178 (Anisoplia) E: AL AU BH BU CR GR KO MC ME RO SB TR A: TR
rumeliaca I. Frivaldszky, 1835: 260 (Anisoplia)

subgenus Taxipertha Baraud, 1991b: 51 type species Phyllopertha arenicola Mulsant & Pellet, 1870
arenicola Mulsant & Pellet, 1870: 29 (Phyllopertha) E: AB GG ST UK A: TR
pilosella Reitter, 1885c: 396 (Phyllopertha)

conspurcata Harold, 1878a: 71 (Phyllopertha) A: FE HEI JA JIL LIA NC NE SC
kiotoensis Niijima & Kinoshita, 1923: 126 (Phyllopertha)

mlokosiewiczi Zaitzev, 1917: 21 [= Zaitzev, 1918a: 109] (Blitopertha) E: GG
genus Hontoriella Escalera, 1914a: 186 type species Hontoriella elaphoceroides Escalera, 1914
elaphoceroides Escalera, 1914a: 186 N: MO
maroccana Ohaus, 1941: 111 (Trigonocnemis)

genus Hoplopus Laporte, 1832a: IX, pl. 20 type species Melolontha atriplicis Fabricius, 1787
Anisonchus Dejean, 1833: 157 type species Melolontha atriplicis Fabricius, 1787

atriplicis atriplicis Fabricius, 1787a: 19 (Melolontha) N: AG MO TU
atriplicis libyca Peyerimhoff, 1939b: 219 (Anomala) N: LB
bleusei Chobaut, 1896a: 388 N: AG
nigropunctatus Bleuse, 1912b: 376

marcens Peyerimhoff, 1939b: 219 (Anomala) N: LB TU
genus Megapertha Reitter, 1903a: 89 type species Phyllopertha massageta Kirsch, 1880
massageta Kirsch, 1880b: 164 (Phyllopertha) A: KZ TM UZ
bicolor Semenov, 1889b: 200 (Phyllopertha)
obscurata Semenov, 1891: 326 (Phyllopertha)
sarta Semenov, 1889b: 199 (Phyllopertha)

genus Melanopopillia Lin, 1980c: 300 type species Melanopopillia dinghuensis Lin, 1980
dinghuensis Lin, 1980c: 300 A: GUA
hainanensis Lin, 1980c: 301 A: HAI

343

344

SCARABAEIDAE

praefica Machatschke, 1971a: 199 (Callistethus) A: FUJ GUX ORR
genus Microlontha Petrovitz, 1967e: 484 type species Microlontha aphodioides Petrovitz, 1967
aphodioides Petrovitz, 1967e: 485 A: PA
genus Micropertha Baraud, 1991b: 53 type species Micropertha variabilis Ballion, 1871
variabilis Ballion, 1871a: 344 (Phyllopertha) A: KI KZ TD TM UZ
rufina Reitter, 1903a: 87 (Blitopertha)

genus Mimela Kirby, 1823: 101 type species Mimela chinensis Kirby, 1823
Amblomala Reitter, 1903a: 58 type species Melolontha aurata Fabricius, 1801
Eriomela Reitter, 1903a: 52 type species Mimela pomacea Bates, 1890
Paracrusis Newman, 1839: 366 type species Paracrusis cyanipes Newman, 1839
Paramimela Ohaus, 1915a: 88 type species Euchlora circumcincta Hope, 1842
Rhombonyx Hope, 1837: 106t ype species Melolontha holosericea Fabricius, 1787
Trimela Ohaus, 1924: 171 type species Mimela macassara Heller, 1896

ahrensi Wada, 2015a: 83 A: NP
anopunctata Burmeister, 1855: 500 (Anomala) A: YUN ORR
antiqua Ohaus, 1938: 268 A: SCH
aurata Fabricius, 1801b: 172 (Melolontha) E: AL AU BH BU CR GR HU MC RO SB SL SZ
auricollis Laporte, 1840: 136 (Euchlora)
cuprea Erichson, 1847a: 617 (Anomala) [HN]
meridionalis Mikšić, 1954c: 139

bicolor Hope, 1831: 23 A: AP NP SD XIZ YUN ORR
similis Hope, 1831: 23

bidentata Lin, 1966b: 144 A: GUX YUN ORR
bifoveolata Lin, 1966b: 145 A: JIX
bimaculata Lin, 1966b: 144 A: YUN
brancuccii Sabatinelli, 1994a: 59 A: NP
cariniventris Lin, 1990: 19 A: XIZ
chinensis Kirby, 1823: 107 A: FUJ GUA GUI GUX HAI HUN JIX SCH YUN ZHE ORR
stilbophora Wiedemann, 1823: 92 (Melolontha)

confucius confucius Hope, 1836: 112 A: ANH FUJ GUA GUI GUX HAI HEB HUB HUN JIX SCH SHX YUN ZHE
ORR
flexuosa Lin, 1966b: 144

confucius formosana Nomura & Kobayashi, 1976: 38 A: TAI
confucius ishigakiensis Sawada, 1950b: 302 (Anomala) A: JA (Ishigaki-jima)
confucius kurodai Nomura & Kobayashi, 1976: 39 A: TAI
costata Hope, 1839b: 73 (Euchlora) A: JA
cyanipes Newman, 1839: 366 (Paracrusis) A: GUX SCH YUN ORR
globosa Ohaus, 1905: 88

SCARABAEIDAE

345

dehaani Hope, 1839b: 71 (Euchlora) A: BT FUJ GUI GUX HUN JIX SCH SD YUN ORR
decipiens Hope, 1841b: 66
dulcissima Bates, 1891b: 78

dentifera Lin, 1990: 20 A: HAI
deretzi R. Paulian, 1959b: 108 A: YUN ORR
despumata Ohaus, 1915a: 95 A: YUN ORR
tonkinensis Ohaus, 1944: 87

difficilis C.O. Waterhouse, 1875a: 111 (Anomala) A: JA
lenzi Ohaus, 1915b: 323 (Anomala)

epipleurica Ohaus, 1930: 143 A: GUX YUN ORR
excisifemorata Lin, 1990: 20 A: HAI
excisipes Reitter, 1903a: 54 A: ANH FUJ GUA HEN HUB HUN JIA JIX SCH SHN SHX TAI ZHE
ferreroi Sabatinelli, 1994a: 60 A: GUX YUN ORR
flavilabris C.O. Waterhouse, 1875a: 110 (Anomala) A: FE (Kuril’skie ostrova, Sakhalin) JA
flavipes Lin, 1988b: 262 A: XIZ
flavocincta Lin, 1966b: 146 A: FUJ HUB HUN TAI ZHE
kitanoi Miyake, 1987: 7

flavomarginata Ohaus, 1915a: 97 A: NP SD
fukiensis Machatschke, 1955b: 502 A: ANH FUJ HUN JIX ZHE
fulgidivittata C.É. Blanchard, 1851: 197 A: AP BT SD ORR
furvipes Lin, 1990: 22 A: GUX
fusania Bates, 1888b: 375 A: FUJ HEB HEI HEN HUB HUN JIA JIL JIX LIA NC SC SCH SHN SHX TIA ZHE
plicicollis Arrow, 1908b: 247
sculpticollis Fairmaire, 1897e: 215 (Anomala)
viriditincta Fairmaire, 1891f: ccv

fusciventris Lin, 1990: 25 A: FUJ GUA
hauseri Ohaus, 1944: 85 A: GUA GUI GUX HUN SCH YUN ORR
heterochropus heterochropus C.É. Blanchard, 1851: 196 A: AP BT SD XIZ ORR
heterochropus hopei Burmeister, 1855: 507 [RN] A: NP
bruschii Sabatinelli, 1997b: 252
splendens Hope, 1831: 23 [HN]

hirtipyga Lin, 1966b: 145 A: YUN ORR
holosericea Fabricius, 1787a: 21 (Melolontha) A: BEI CT ES FE HEB HEI JIL LIA MG NC NMO QIN SC SHA
SHX WS
coerulea Ohaus, 1944: 83
japonica Machatschke, 1952: 358

horsfieldii Hope, 1836: 114 A: AP HP KA NP PA SD UP XIZ YUN ORR
vittata Kollar & L. Redtenbacher, 1844: 525 (Euchlora)

ignicauda Bates, 1866: 345 A: TAI
ignistriata Lin, 1990: 20 A: YUN ORR
inouei Nomura, 1967: 56 A: JA

346

SCARABAEIDAE

inscripta Nonfried, 1891a: 236 (Anomala) A: NP YUN ORR
iris Lin, 1990: 19 A: YUN ORR
junii calabrica Machatschke, 1957: 112 E: IT (Basilicata, Calabria)
junii gigliocola Machatschke, 1957: 113 E: IT (Isola del Giglio)
junii junii Duftschmid, 1805: 199 (Melolontha) E: AU CR FR IT SL SZ
cuprithorax Della Beffa, 1910: 103 (Anomala)
doublieri Mulsant, 1842: 483 (Anomala)
etrusca Erichson, 1847: 619
scutellaris Mulsant, 1842: 483 (Anomala)
signaticollis Erichson, 1847: 619
thoracica Mulsant, 1842: 483 (Anomala)

junii miksici Sparacio, 2003: 226 E: IT (Sicilia)
junii rugosula Fairmaire, 1859a: 276 (Anomala) E: FR (Corse) IT (Sardegna)
corsicana Machatschke, 1957: 112

laevicollis Lin, 1966b: 146 A: YUN
laevigata Arrow, 1908b: 241 A: BT SD ORR
leei Swederus, 1787: 188 (Scarabaeus) A: “India Orientalis” ORR
fastuosa Fabricius, 1792b: 168 (Melolontha)
sapphirina Parry, 1845: 85

lusitanica Ohaus, 1915b: 313 (Anomala) E: PT
luteipennis Motschulsky, 1853: 47 A: BEI
malaisei Ohaus, 1937a: 6 A: YUN ORR
malicolor Lin, 1990: 21 A: YUN
marginalis Arrow, 1908b: 242 A: BT SD ORR
nigritarsis Lin, 1990: 22 A: GUI YUN ORR
nubeculata Lin, 1990: 21 A: GUA HAI JIL
ohausi Arrow, 1908b: 243 A: SCH SD YUN ORR
opalina Ohaus, 1902: 50 A: FUJ GUA GUX SCH YUN ORR
pachygastra Burmeister, 1855: 507 A: XIZ
parva Lin, 1966b: 146 A: FUJ JIX
passerinii diana Lin, 1993: 105 A: YUN ORR
passerinii mediana Lin, 1993: 106 A: HUB SHX
passerinii oblonga Arrow, 1908b: 247 A: SCH
passerinii passerinii Hope, 1842c: 128 A: AP BT KA NP PA SCH SD XIZ ORR
limbata Burmeister, 1855: 508

passerinii pomacea Bates, 1890: 245 A: SCH
passerinii taiheizana Sawada, 1943a: 1 A: TAI
passerinii tienmusana Lin, 1993: 106 A: ZHE
peckinensis coreana Machatschke, 1952: 367 A: SC
peckinensis peckinensis Heyden, 1886d: 291 (Anomala) A: BEI HEB HEI JIL LIA NMO SHA SHX

SCARABAEIDAE

347

violata Reitter, 1903a: 56 (Anomala)

pectoralis C.É. Blanchard, 1851: 197 A: BT HP NP PA SCH UP XIZ YUN
laevisutula Lin, 1981: 383
rugosopunctata Fairmaire, 1889a: 26 (Anomala)

plicatulla Lin, 1990: 25 A: YUN ORR
pomicolor Ohaus, 1936: 149 A: YUN
princeps Hope, 1841b: 65 A: SD ORR
punctulata Lin, 1993: 104 A: GUX
pusilla Arrow, 1917: 378 A: UP
pyriformis Arrow, 1908b: 246 A: YUN ORR
expansa Lin, 1990: 24

rectangular Lin, 1990: 22 A: YUN ORR
repsimoides Ohaus, 1915a: 96 A: FUJ
rubrivirgata Lin, 1990: 23 A: YUN ORR
rugatipennis Graëlls, 1849: 620 (Anomala) E: PT SP
rugicauda Arrow, 1917: 119 A: SD
rugicollis Lin, 1966b: 147 A: HAI
ruyuanensis Lin, 1990: 24 A: GUA
sakishimana Nomura, 1973a: 19 A: JA (Iriomote shima)
sauteri Ohaus, 1938: 269 A: TAI
schneideri Ohaus, 1905: 89 A: SD ORR
seminigra Ohaus, 1908b: 640 A: FUJ GUA GUX HAI HUN JIX YUN ORR
sericea Ohaus, 1905: 88 A: NP SD
sericicollis Ohaus, 1944: 86 A: FUJ GUA GUX HUN JIX ORR
signaticollis Ohaus, 1902: 53 A: YUN ORR
siliguria Arrow, 1917: 188 (Anomala) A:NP SD UP ORR
calva Benderitter, 1927: 430 (Anomala)

specularis Ohaus, 1902: 49 A: FUJ GUA GUI GUX HAI SCH SHX ORR
splendens Gyllenhal, 1817: 110 (Melolontha) A: ANH BEI FUJ GUA GUI GUX HEB HEI HUB HUN JA JIL JIX LIA
NC SC SCH SHN SHX TAI YUN ZHE ORR
concolor C.É. Blanchard, 1851: 196
corusca Heyden, 1887a: 253
davidis Fairmaire, 1886: 330
foveolata Ohaus, 1944: 84
gaschkewitchii Motschulsky, 1858a: 32
lathamii Hope, 1836: 113
lucidula Hope, 1836: 113
murasaki Chûjô, 1940a: 76
simplex Bates, 1866: 345
takamurai Sawano & Kometani, 1939: 205

348

SCARABAEIDAE

sulcatula Ohaus, 1915a: 92 A: FUJ GUA GUI GUX HAI HUN JIX
klapperichi Machatschke, 1955b: 504

taiwana Sawada, 1943a: 2 A: TAI
takemurai Sawada, 1942a: 38 A: JA
testacea Lin, 1990: 25 A: XIZ
testaceipes testaceipes Motschulsky, 1861a: 14 (Rhombonyx) A: JA
testaceipes ussuriensis S.I. Medvedev, 1949: 104 (Rhombonyx) A: BEI FE HEI JIL LIA NC SC SHA
testaceipes yuasai Nomura, 1969: 73 (Anomala) A: JA
testaceoviridis C.É. Blanchard, 1851: 197 A: ANH BEI FUJ GUX HEB HEI HUB HUN JIA JIL JIX LIA SCH SHG
SHN SHX TAI ZHE
chryseis Bates, 1866: 345
rufofemoralis Fairmaire, 1887d: 319 (Spilota)
surigera Heyden, 1886d: 291 (Anomala)

vernicata Fairmaire, 1896a: 88 (Spilota) A: BT SD
biplagiata Ohaus, 1908b: 642

vittaticollis Burmeister, 1855: 506 A: BT NP SD XIZ ORR
werneri Sabatinelli, 1997b: 253 A: UP SD
xanthorrhoea Ohaus, 1902: 52 A: GUA GUX ORR
xutholoma Lin, 1993: 105 A: GUX
yonaguniensis Nomura, 1965: 5 A: JA
yunnana Ohaus, 1944: 87 A: YUN
genus Phyllopertha Stephens, 1830: 223 type species Scarabaeus horticola Linnaeus, 1758
abullosa Lin, 1988b: 262 A: XIZ
bifasciata Lin, 1966a: 83 A: BEI LIA
brevipilosa Lin, 1988b: 263 A: XIZ
chalcoides Ohaus, 1925b: 131 A: TAI
diversa C.O. Waterhouse, 1875a: 106 A: BEI HEB HEI HEN JA JIL LIA NC NMO SC SHA SHN SHX TIA ZHE
koreana Ohaus, 1915b: 325
maculicollis Reitter, 1903a: 84

euchroma Fairmaire, 1891f: cciv (Anomala) A: SCH
glabripennis S.I. Medvedev, 1949: 85 A: GAN SCH
horticola Linnaeus, 1758: 351 (Scarabaeus) E: AL AN AU BE BH BU BY CT CR CZ DE EN FI FR GB GE GR HU IR
IT LA LS LT LU MD NL NR NT PL RO SB SK SL SP ST SV SZ UK A: ES FE HEB HEI JIL KI KZ LIA MG NC NE
NO NW WP WS
adiophora Poda von Neuhaus, 1761: 20 (Scarabaeus)
cyanocephala Mulsant, 1842: 499
macularis Mulsant, 1842: 500
obscura Westhoff, 1882: 153
perrisi Mulsant, 1842: 500
rufiventris Westhoff, 1882: 154

SCARABAEIDAE

349

suturalis Stephens, 1839: 169
ustulatipennis A. Villa & G.B. Villa, 1833: 34 (Anisoplia)
viridicollis De Geer, 1774: 278 (Scarabaeus)

horticoloides Lin, 1965: 393 A: XIZ
humeralis Fairmaire, 1887b: 106 A: SCH YUN
cribricollis Fairmaire, 1887b: 105

intermixta Arrow, 1913c: 403 (Anomala) A: FE (Kunashir) JA NE
fuscata Niijima & Kinoshita, 1923: 129

irregularis C.O. Waterhouse, 1875a: 107 A: JA
latevittata Fairmaire, 1889a: 24 A: SCH
obscuricolor Fairmaire, 1891f: cciii A: HUB SCH
puncticollis Reitter, 1888c: 294 A: GAN QIN SCH
puncticollis Heyden, 1889c: 668 [HN]

punctigera Fairmaire, 1888a: 20 (Anomala) A: GAN GUI HEB HUB JIX NMO SCH SHA SHX
sexoculata Fairmaire, 1897e: 213 (Anomala)
tenuelimbata Fairmaire, 1891f: cciii [HN]

sublimbata Fairmaire, 1900b: 620 A: FUJ
suturata Fairmaire, 1887b: 106 A: SCH YUN
major Fairmaire, 1889a: 23

suzukii Sawada, 1943a: 2 A: TAI
taiwana Li C.-L. & P.-S. Yang, 1997b: 169 A: TAI
virgulata Fairmaire, 1889a: 24 A: SCH
wassuensis Frey, 1971a: 114 A: SCH
yangi Kobayashi & C.-L. Li, 1995: 31 A: TAI
zea Reitter, 1903a: 83 A: QIN
genus Proagopertha Reitter, 1903a: 50 type species Anomala acutisterna Fairmaire, 1878 (=Anomala lucidula
Faldermann, 1835)
lucidula Faldermann, 1835b: 380 (Anomala) A: ANH FE GAN HEB HEI HEN JIA JIL LIA NC NMO SC SCH SHA
SHN SHX
acutisterna Fairmaire, 1878a: 106 (Anomala)
aeneoflavida Reitter, 1913c: 212
starki Reitter, 1913c: 212

ohbayashii Nomura, 1965: 6 A: JA
pubicollis C.O. Waterhouse, 1875a: 111 (Anomala) A: JA
genus Pseudosinghala Heller, 1891b: 294 type species Pseudosinghala vorstmani Heller, 1891
dalmanni Gyllenhal, 1817: 109 (Melolontha) A: BEI FUJ GUA GUI GUX HEB HKG JIX SHG ORR
schaiblei Nonfried, 1892a: 89 (Singhala)
theicola Waga, 1842: 273 (Anisoplia)

transversa Burmeister, 1855: 513 (Phyllopertha) A: FUJ GUA GUI HUB SCH SD XIZ YUN ORR

350

SCARABAEIDAE

basipennis Fairmaire, 1889a: 28 (Singhala)
immaculata Fairmaire, 1893d: 291 (Singhala)

genus Spinanomala Ohaus, 1910a: 213 type species Spinanomala unispinosa Ohaus, 1910
dentipennis Lin, 1966a: 84 (Blitopertha) A: YUN ORR
hainanensis Lin, 1979c: 186 A: HAI
genus Trichanomala Arrow, 1917: 95 type species Popillia fimbriata Newman, 1838
dentipennis dentipennis Fairmaire, 1896a: 88 (Phyllopertha) A: SCH XIZ
fimbriata Newman, 1838b: 337 (Popillia) A: SD UP “East India”
rugicollis Lin, 1979c: 186 A: HAI
genus Trigonocnemis Kraatz, 1894b: 171 type species Trigonocnemis hauseri Kraatz, 1894
Ammogenia Semenov, 1896d: 341 type species Ammogenia lanuginosa Semenov, 1896

hauseri Kraatz, 1894b: 172 A: TM
lanuginosa Semenov, 1896d: 343 (Ammogenia) A: KZ TM UZ
varentzowi Semenov, 1896d: 345 (Ammogenia) A: TM
unguicularis Semenov, 1896e: 378 (Ammogenia)

subtribe Popilliina Ohaus, 1918

genus Callistopopillia Ohaus, 1903: 219 type species Anomala iris Candèze, 1869
davidis Fairmaire, 1878a: 101 (Callistethus) A: GUI GUX HUB SCH YUN “Thibet”
compressidens Fairmaire, 1887b: 112 (Callistethus)

iris Candèze, 1869: 43 (Anomala) A: KA NP SD XIZ
lurida Arrow, 1917: 93 A: NP “N. India”
scabrata Lin, 1999b: 280 A: XIZ
seperata Lin, 1999b: 280 A: XIZ
sulcipennis Lin, 1992: 523 A: SCH YUN
genus Dactylopopillia Arrow, 1917: 56 type species Popillia virescens Hope, 1831
blanchardi Ohaus, 1897: 348 (Popillia) A: “Himalaya” ORR
opacicollis Kraatz, 1892a: 272 (Popillia) A: NP ORR
virescens Hope, 1831: 23 (Popillia) A: BT NP SD
truncata Kollar & L. Redtenbacher, 1844: 527

genus Dicranoplia Reitter, 1903a: 42 type species Phyllopertha deserticola P.H. Lucas, 1859
demoflysi Baraud, 1989c: 318 N: TU
deserticola P.H. Lucas, 1859d: liii (Phyllopertha) N: AL MO TU
lucasii Fairmaire, 1871a: 379 (Anisoplia) [RN]

genus Glenopopillia Lin, 1980a: 77 type species Glenopopillia maculata Lin, 1980
maculata Lin, 1980a: 77 A: GUX

SCARABAEIDAE
nagaii Sabatinelli, 1997b: 249 (Callistethus) A: YUN ORR
rufipennis Lin, 1980a: 78 A: YUN
genus Ischnopopillia Kraatz, 1892a: 292 type species Popillia exarata Fairmaire, 1886
akbari Abdullah & Roohi, 1969: 124 (Anomala) A: PA
angusta Lin, 1992: 523 A: YUN
atrivaria Lin, 1981: 381 A: XIZ
atronitens Fairmaire, 1896a: 89 (Popillia) A: HUB SCH SHX
atronitens Machatschke, 1975: 6 [HN]

bidentata Lin, 1979b: 112 A: YUN
bivittata Lin, 1988b: 262 A: XIZ
brancuccii Sabatinelli, 1997b: 250 A: BT
chalconota Machatschke, 1975: 8 A: SD
consimilis Machatschke, 1975: 12 A: GUI
difficilis Fairmaire, 1888b: 119 (Popillia) A: BEI
erythroptera chinensis Machatschke, 1975: 11 A: JIX
erythroptera erythroptera Kraatz, 1892a: 295 A: SCH SD YUN
erythropteroides Machatschke, 1975: 14 A: GUI
exarata Fairmaire, 1886: 330 (Popillia) A: SCH XIZ YUN
cinnabarina Fairmaire, 1887b: 113 (Popillia)
cupreata Kraatz, 1892a: 293
suturella Machatschke, 1975: 10

festiva Arrow, 1917: 261 (Anomala) A: SD XIZ
flavomarginata Lin, 1965: 394 A: XIZ
fuscicollis Machatschke, 1975: 13 A: YUN ORR
ikrami Abdullah & Roohi, 1969: 125 (Anomala) A: PA
lateralis Hope, 1831: 24 (Anomala) A: BT NP SD XIZ
lineatocollis Kraatz, 1892a: 302
piliclunis Lin, 1965: 393
rugicollis Newman, 1838b: 337 (Popillia)

longicosta Lin, 1981: 382 A: XIZ
lyallpurensis Abdullah & Roohi, 1969: 124 (Anomala) A: PA
moorei Kraatz, 1892a: 294 A: HP NP UP
andrewesi Kraatz, 1897a: 332
flavipes Arrow, 1917: 266 (Anomala)

nana Lin, 1992: 523 A: YUN
phylloperthoides Fairmaire, 1887b: 114 (Popillia) A: YUN
purpuricollis Kraatz, 1897a: 331 A: GUI SCH YUN
pusilla Arrow, 1912a: 82 (Anomala) A: NP SD XIZ ORR
reflexa Lin, 1992: 524 A: SCH YUN
rhinosina Machatschke, 1975: 16 A: SCH XIZ YUN
robustipes Lin, 1981: 382 A: XIZ

351

352

SCARABAEIDAE

sulcatula Lin, 1981: 383 A: XIZ
genus Pharaonus C.É. Blanchard, 1851: 201 type species Popillia fasciculata Burmeister, 1844
subgenus Pharaonus C.É. Blanchard, 1851: 201 type species Popillia fasciculata Burmeister, 1844
adelphus Keith, Sabatinelli & Uliana, 2015: 168 A: IN
caucasicus Reitter, 1888c: 294 (Phyllopertha) E: AR A: TR
farsensis Keith, Sabatinelli & Uliana, 2015: 171 A: IN
fasciculatus Burmeister, 1844: 307 (Popillia) N: EG LB TU A: IS AFR
saharicus Král, 2012: 128 N: AG
varicoloreus Burmeister, 1844: 308 (Popillia) A: TR IN IQ JO KU SY
mesopotamica C.É. Blanchard, 1851: 178 (Phyllopertha)

subgenus Tamerlanius Keith, Sabatinelli & Uliana, 2015: 177 type species Phyllopertha lederi Reitter, 1888
lederi Reitter, 1888c: 293 (Phyllopertha) A: AF KZ TD TM UZ
ammodendri Semenov, 1889b: 201 (Phyllopertha)
turcmenius Ohaus, 1941: 114

semenowi Reitter, 1887d: 510 (Phyllopertha) A: KZ TD TM UZ
genus Popillia Dejean, 1821: 60 type species Trichius bipunctatus Fabricius, 1787
Calopopillia H.J. Kolbe, 1894b: 209 type species Popillia candezei Kraatz, 1892
Costapopillia Benderitter, 1922: 83 type species Costapopillia callewaerti Benderitter, 1922
Eupopillia H.J. Kolbe, 1894b: 209 type species Popillia callipyga Dohrn, 1876
Godschama Reitter, 1903a: 50 type species Popillia hexaspila Ancey, 1883
Metapopillia H.J. Kolbe, 1894b: 209 type species Popillia biimpressa H.J. Kolbe, 1894
Pseudopopillia H.J. Kolbe, 1894b: 209 type species Popillia phylloperthina H.J. Kolbe, 1894
Xenopopillia H.J. Kolbe, 1910: 80 type species Popillia ducatrix H.J. Kolbe, 1910

adamas Newman, 1838b: 337 A: “Ind. Orient.”
viridinitens Kraatz, 1892a: 302

anomaloides Kraatz, 1892a: 260 A: GUI SCH YUN ORR
barbellata Lin, 1988a: 69 A: YUN
bhutanensis Frey, 1975c: 244 A: BT
bhutanica Sabatinelli, 1993b: 209 A: BT
birmanica Arrow, 1913a: 43 A: NP ORR
brancuccii Sabatinelli, 1993b: 205 A: BT
cerchnopyga Lin, 1981: 384 A: XIZ
laticlypealis Lin, 1981: 384

cerinimaculata Lin, 1980b: 196 A: YUN ORR
clypealis Ohaus, 1897: 347 A: KA UP
comma Niijima & Matsumura, 1923: 139 A: JA (Okinawa-shotō)
cribricollis Ohaus, 1938: 271 A: GUI GUX HUB SCH YUN ORR
cupricollis Hope, 1831: 23 A: AP HP KA NP SD UP ORR
caschmirensis Kollar & L. Redtenbacher, 1844: 528

SCARABAEIDAE

353

formosa Hope, 1831: 23
hilaris Burmeister, 1855: 510
smaragdula Hope, 1831: 23
suturata Newman, 1841: 42

curtipennis Lin, 1980b: 197 A: YUN ORR
cyanea Hope, 1831: 23 A: AP GUI HP KA NP PA SCH SD UP XIZ YUN ORR
beryllina Hope, 1831: 23
concolor Laporte, 1840: 138

daliensis Lin, 1980b: 197 A: YUN
difficilis Newman, 1838b: 337 A: “Ind. orient.”
dilutipennis Fairmaire, 1889b: 342 A: ANH FUJ GUA GUX HKG SC TAI YUN ZHE ORR
formosana Arrow, 1913a: 45
quelpartiana Ohaus, 1924: 172

fallaciosa Fairmaire, 1889a: 29 A: GUI SCH YUN
feae Kraatz, 1892a: 269 A: AP NP SD ORR
fimbripes Lin, 1988b: 263 A: XIZ
flavofasciata Kraatz, 1892a: 271 A: YUN ORR
flavosellata Fairmaire, 1886: 331 A: ANH BEI FE GUI HEI HUB HUN JIA JIL JIX LIA NC SC SCH SHG SHN SHX
XIZ YUN ZHE ORR
atra Kraatz, 1892a: 245
atrocaerulea Bates, 1888b: 376
bicolor Kraatz, 1892a: 245
biplagiata Kraatz, 1892a: 245
chrysitis Kraatz, 1892a: 244
circumducta Kraatz, 1892a: 245
inconstans Fairmaire, 1889a: 28
losczyi J. Frivaldszky, 1890: 201
schoenfeldti Kraatz, 1892a: 245
sellata Kraatz, 1892a: 245
strigiventris Kraatz, 1892a: 245
viridicollis Kraatz, 1892a: 245

flexuosa Lin, 1988a: 71 [=1988b: 264] A: XIZ
fukiensis Machatschke, 1955b: 508 A: FUJ GUA GUI GUX JIX ZHE
gedongensis Lin, 1988b: 264 A: XIZ
gemma Newman, 1839: 366 A: BT NP SD ORR
nitidicollis Gory, 1844: 100

hainanensis Lin, 1988a: 65 A: HAI
hirta Lin, 1987b: 225 A: XIZ
hirtipyga Lin, 1987b: 226 A: NP XIZ
histeroidea Gyllenhal, 1817: 66 (Rutela) A: ANH FUJ GUA GUI GUX HAI HUB HUN JIX SCH YUN ZHE ORR
coerulea Boheman, 1858: 55

354

SCARABAEIDAE

discipennis Fairmaire, 1886: 332
unicolor Fairmaire, 1889a: 23

impressipyga Ohaus, 1897: 381 A: SD ORR
insularis Lewis, 1895a: 403 A: TAI
circumducta Kraatz, 1897a: 327 [HN]

japonica Newman, 1838c: 337 A: FE (Kuril’skie ostrova, Sakhalin) JA NARi
plicatipennis Burmeister, 1844: 300

laevicollis Kraatz, 1892a: 275 A: BT SD ORR
laevis Burmeister, 1855: 511 A: CH “Thibet”
clara Arrow, 1913a: 49

laeviscutula Lin, 1980b: 198 A: SCH
laevistriata Arrow, 1913a: 42 A: BT ORR
lasiopyga Lin, 1987a: 198 A: SD
laticlypealis Lin, 1981: 384 A: XIZ
latimaculata Nomura, 1970: 70 A: TAI
leptotarsa Lin, 1980b: 198 A: GAN GUI HUB HUN SCH XIZ YUN
lewisi Arrow, 1913a: 45 A: JA
limbatipennis Lin, 1988a: 67 A: YUN
linpingi Sabatinelli, 1994b: 338 A: NP
livida Lin, 1988a: 65 A: TAI
lucida Newman, 1838c: 337 A: “Ind. Orient.”
maclellandi Hope, 1842a: 60 [=1843a: 62, 1845: 8] A: XIZ ORR
marginicollis Hope, 1831: 23 A: GUX NP SD UP YUN ORR
atrata Kraatz, 1892a: 266
purpuricollis Kraatz, 1892a: 266

melanoloma Lin, 1988a: 67 A: GUX
metallicollis Fairmaire, 1887b: 113 A: GUI SCH YUN
miniatipennis Fairmaire, 1889a: 23 A: GUI
minuta Hope, 1831: 23 A: NP SD XIZ
mongolica Arrow, 1913a: 43 A: FUJ GUA GUI GUX HKG JIA JIX SHN TAI YUN ORR
mutans Newman, 1838c: 337 A: ANH BEI FE FUJ GAN GUA GUI GUX HAI HEB HEI HEN HUB HUN JIA JIL JIX
LIA NC NIN NMO SC SCH SHA SHN SHX TAI YUN ZHE “Ind. Orient.”
indigonacea Motschulsky, 1854c: 47
relucens C.É. Blanchard, 1851: 199

nitida Hope, 1831: 23 A: NP XIZ ORR
trichiopyga Lin, 1981: 386

nottrotti Kraatz, 1892a: 273 A: AP SD ORR
oviformis Lin, 1988a: 68 A: SCH YUN
patricia Arrow, 1917: 75 A: YUN ORR
bothynoma Ohaus, 1938: 270

petrarcai Sabatinelli, 1994b: 328 A: BT

SCARABAEIDAE

355

piattellai Sabatinelli, 1994b: 335 A: NP
pilicollis Kraatz, 1892a: 278 A: AP NP SD ORR
pilifera Lin, 1988a: 70 A: SCH
pilosa Arrow, 1913a: 51 A: NP UP
plagicollis Kraatz, 1897a: 329 A: SCH
pui Lin, 1988a: 66 A: GUA
puncticollis Kraatz, 1897a: 329 A: SD
pustulata Fairmaire, 1887b: 114 A: ANH FUJ GUA GUI GUX HUB HUN JIA JIX SCH SHN SHX YUN ZHE ORR
brunnipennis Kraatz, 1892a: 250
cupricollis Kraatz, 1892a: 250 [HN]
impustulata Kraatz, 1892a: 250

quadriguttata Fabricius, 1787b: 377 (Trichius) A: ANH BEI FE FUJ GAN GUA GUI GUX HEB HEI HEN HUB
HUN JIA JIL JIX LIA NC NIN NMO QIN SC SCH SHA SHG SHN SHX TAI YUN ZHE
biguttatus Fabricius, 1794: 499 (Trichius)
bogdanowi Ballion, 1871a: 345
castanoptera Hope, 1842a: 60 [=1843a: 63, 1845: 8]
chinensis J. Frivaldszky, 1890: 201
dichroa C.É. Blanchard, 1851: 200
frivaldszkyi Kraatz, 1892a: 249
purpurascens Kraatz, 1892a: 249
ruficollis Kraatz, 1892a: 254
sordida Kraatz, 1892a: 249
straminipennis Kraatz, 1892a: 258
uchidai Niijima & Kinoshita, 1923: 138

rotundata Lin, 1988a: 66 A: HAI
rubescens Lin, 1981: 384 A: XIZ
rubripes Lin, 1981: 385 A: XIZ
sabatinellii Lin, 2003: 195 A: GUX
sanmenensis Lin, 1980b: 198 A: GUX
sauteri Ohaus, 1917a: 2 A: TAI
scabricollis Lin, 1988b: 264 A: XIZ
semiaenea Kraatz, 1892a: 251 A: FUJ GUX HKG ORR
aenea Kraatz, 1892a: 302 [HN]
cupricollis Kraatz, 1892a: 302 [HN]
simoni Kraatz, 1892a: 252
tesari Sabatinelli, 1984b: 169

semicuprea Kraatz, 1892a: 256 A: SD “China interior”
spediceipennis Lin, 1987a: 199

sichuanensis Lin, 1988a: 39 [RN] A: HUB SCH
concolor Lin, 1980b: 197 [HN]

sikkimensis Lin, 1987a: 199 A: BT SD

356

SCARABAEIDAE

concolor Kraatz, 1897a: 328 [HN]
unicolor Frey, 1975c: 243 [HN]

simlana Arrow, 1913a: 52 A: HP NP
splendidicollis Fairmaire, 1886: 331 A: YUN
strumifera Lin, 1981: 385 A: XIZ
subquadrata Kraatz, 1892a: 259 A: GUI GUX HAI HUB HUN JIX SCH ZHE ORR
sulcata Kollar & L. Redtenbacher, 1844: 527 A: KA NP UP XIZ
cupripennis Kraatz, 1892a: 276
scutellaris C.É. Blanchard, 1851: 199
testaceipennis Kraatz, 1892a: 276

suturalis Lin, 1980b: 199 A: YUN
taiwana Arrow, 1913a: 44 A: TAI
tibidens Lin, 1980b: 199 A: SCH
transversa Lin, 1988a: 68 A: FUJ
varicollis Lin, 1988a: 70 A: SCH YUN
vignai Sabatinelli, 1994b: 341 A: BT
viridula Kraatz, 1892a: 255 A: SCH YUN
wittmeri Frey, 1975c: 243 A: BT
zerchei Sabatinelli, 1994b: 346 A: BT
genus Pseudoblitopertha Keith, 2000a: 384 type species Pseudoblitopertha inexpectata Keith, 2000
inexpectata Keith, 2000a: 385 A: TR
genus Spilopopillia Kraatz, 1892a: 181 type species Popillia sexguttata Fairmaire, 1887
quadriguttata Lin, 1988b: 265 A: XIZ
sexguttata Fairmaire, 1887b: 115 (Popillia) A: SCH SD YUN ORR
sexmaculata Kraatz, 1892a: 261 (Popillia) A: FUJ GUA GUX HKG HUN SCH ORR
cantonensis Ohaus, 1908a: 201

genus Tribopertha Reitter, 1903a: 43 type species Phyllopertha aegyptiaca C.É. Blanchard, 1851
aegyptiaca C.É. Blanchard, 1851: 179 (Phyllopertha) N: AG EG LY MO TU A: IS
colonnellii Piattella & Sabatinelli, 1992b: 337 N: AG
quedenfeldti Reitter, 1890g: 170 (Phyllopertha) N: LY TU
genus Trichopopillia Ohaus, 1897: 353 type species Gnatholabis dorsalis Kraatz, 1892
dorsalis nilotica Kraatz, 1892a: 299 (Gnatholabis) N: EG

tribe Rutelini W.S. Macleay, 1819

genus Cyphelytra C.O. Waterhouse, 1875b: 367 type species Cyphelytra ochracea C.O. Waterhouse, 1875
ochracea C.O. Waterhouse, 1875b: 367 A: SD ORR

SCARABAEIDAE

357

genus Dicaulocephalus Gestro, 1888a: 623 type species Dicaulocephalus feae Gestro, 1888
fruhstorferi Felsche, 1901: 314 [=1902: 359] A: GUX HAI YUN ORR
genus Didrepanephorus Wood-Mason, 1878: 422 type species Didrepanephorus bifalcifer Wood-Mason, 1878
nishiyamai Muramoto, 2006: 58 A: GUX
yunnanus yunnanus Ohaus, 1911: 334 (Fruhstorferia) A: HAI GUI GUX YUN
genus Kibakoganea Nagai, 1984: 25 type species Fruhstorferia sexmaculata Kraatz, 1900
formosana Kurosawa & Kobayashi, 1975: 213 (Fruhstorferia) A: TAI
fujiokai ushizana Nagai, 2004: 154 A: HAI
kawaii Muramoto, 2005: 27 A: HAI
kraatzi reichenbachi Muramoto, 2005: 30 A: GUX HAI
sinica Bouchard, 2005: 166 A: GUX HAI
tamdaoensis Miyake & Muramoto, 1992: 21 A: YUN ORR
vernicata Benderitter, 1929: 102 (Fruhstorferia) A: YUN ORR
genus Parastasia Westwood, 1842f: 204 type species Parastasia canaliculata Westwood, 1842
Barymorpha Guérin-Méneville, 1843b: 41 type species Barymorpha bimaculata Guérin-Méneville, 1843
Echmatophorus C.O. Waterhouse, 1895: 158 type species Echmatophorus pascoei C.O. Waterhouse, 1895
Ohkubous Sawada, 1938d: 73 type species Ohkubous quadridentatus Sawada, 1938 (=Parastasia ferrieri Nonfried, 1895)
Polymoechus LeConte, 1856: 24 type species Polymoechus brevipes LeConte, 1856
Urleta Westwood, 1875: 237 type species Urleta ometoides Westwood, 1875 (=Parastasia dimidiata Erichson, 1845)

akebono Wada, 2015b: 87 A: AP
alternata Arrow, 1899b: 489 A: BT SD “Himalaya” ORR
birmana Arrow, 1899b: 481 A: YUN ORR
canaliculata Westwood, 1842f: 204 A: TAI ORR
bipuncata Westwood, 1842g: 304
rubrotessellata C.É. Blanchard, 1851: 217
yukioi Wada, 1989: 3

ferrieri ferrieri Nonfried, 1895: 289 A: JA LIA SC
boninensis Nakane, 1983b: 142 (Ohkubous)
ichikawai Nakane, 1983b: 142 (Ohkubous)
quadridentata Sawada, 1938d: 73 (Ohkubous)
sakishimana Nomura, 1965: 4
tokarana Nomura, 1964a: 56

ferrieri formosana Ohaus, 1925b: 122 A: TAI
hainanensis Wada, 2009: 84 A: HAI
indica Ohaus, 1898: 9 A: BT SD ORR
fruhstorferi Ohaus, 1902: 57

oberthueri ishigakiana Nomura, 1964a: 55 A: JA (Ishigaki-jima)

358

SCARABAEIDAE

rufopicta Westwood, 1842a: 53 A: BT NP SD “Himalaya” ORR
anthracina Ohaus, 1902: 56

genus Peperonota Westwood, 1847d: 296 type species Peperonota harringtoni Westwood, 1847
harringtoni Westwood, 1847d: 298 A: BT SD ORR
bicornis Ohaus, 1913: 506

subfamily Dynastinae W.S. Macleay, 1819
tribe Cyclocephalini Laporte, 1840

genus Peltonotus Burmeister, 1847: 75 type species Peltonotus morio Burmeister, 1847
morio Burmeister, 1847: 75 A: BT NP SD ORR
sawaii Miyake, 2000: 112

nasutus Arrow, 1910b: 155 A: GUI GUX YUN ORR

tribe Dynastini W.S. Macleay, 1819

genus Chalcosoma Hope, 1837: 86 type species Scarabaeus atlas Linnaeus, 1758
atlas Linnaeus, 1758: 345 (Scarabaeus) A: NP ORR
hector Reiche, 1859a: 18
hesperus Erichson, 1834: 238 (Dynastes)
kirbii Hope, 1831: 23 (Dynastes)
phidias C.É. Blanchard, 1853: 107

chiron A.G. Olivier, 1789: no 3: 18 (Scarabaeus) A: NP ORR
belangeri Guérin-Méneville, 1833a: 485 (Scarabaeus)
caucasus Fabricius, 1801a: 10 (Geotrupes)

genus Eupatorus Burmeister, 1847: 268 type species Dynastes hardwickii Hope, 1831
Alcidosoma Laporte, 1867: 113 type species Alcidosoma siamense Laporte, 1867

gracilicornis Arrow, 1908c: 351 A: GUX HAI HUB HUN SCH YUN ORR
hardwickii Hope, 1831: 22 (Dynastes) A: BT GUA GUI GUX KA NP SD UP XIZ YUN ORR
atkinsoni Nonfried, 1890b: 89
childreni Hope, 1831: 22 (Dynastes)
cantori Hope, 1843e: 365 (Dynastes)
niger Arrow, 1910a: 269

siamensis hirokii Yamaya, 2013: 99 A: HAI
sukkiti Miyashita & Arnaud, 1997: 2 A: YUN ORR
genus Trypoxylus Minck, 1920: 216 type species Scarabaeus dichotomus Linnaeus, 1771
Xyloscaptes Prell, 1934: 58 type species Xylotrupes davidis Deyrolle, 1878

davidis Deyrolle, 1878: 106 (Xylotrupes) A: FUJ GAN HUN JIX ZHE
dichotomus dichotomus Linnaeus, 1771: 529 (Scarabaeus) A: ANH BEI CHQ FUJ GAN GUA GUI GUX HAI HEB
HEI HEN HKG HUB HUN JIA JIL JIX LIA NC SC SCH SD SHA SHN SZE YUN ZHE ORR

359

SCARABAEIDAE
politus Prell, 1934: 58

dichotomus septentrionalis Kôno, 1931a: 160 A: JA
dichotomus shennongjii Takeuchi, 2014: 149 A: HUB
dichotomus takarai Kusui, 1976: 52 (Allomyrina) A: JA (Ryukyus)
inchachina Kusui, 1976: 52 (Allomyrina)

dichotomus tsunobosonis Kôno, 1931a: 159 A: TAI
dichotomus xizangensis J. Li, 2015a: 48 (Allomyrina) A: XIZ
genus Xylotrupes Hope, 1837: 19 type species Scarabaeus gideon Linnaeus, 1767
Endebius van Lansberge, 1880: cxxii type species Xylotrupes florensis van Lansberge, 1879

mniszechii hainanianus Rowland, 2006a: 16 A: HAI
mniszechii mniszechii J. Thomson, 1859: 18 A: BT HP KA NP PA SD UP ORR
socrates Schaufuss, 1863a: 60

mniszechii tonkinensis Minck, 1920: 217 A: ANH FUJ GUA GUI GUX HKG MAC SCH TAI YUN XIZ ZHE ORR
bourgini R. Paulian, 1945: 196
kaszabi Endrődi, 1951b: 250
siamensis Minck, 1920: 217

philippinensis peregrinus Rowland, 2006b: 13 A: TAI

tribe Oryctini Mulsant, 1842

genus Blabephorus Fairmaire, 1898b: 382 type species Blabephorus pinguis Fairmaire, 1898
pinguis Fairmaire, 1898b: 383 A: GUA GUI GUX HAI TAI XIZ YUN ORR
genus Dichodontus Burmeister, 1847: 217 type species Dichodontus coronatus Burmeister, 1847
coronatus Burmeister, 1847: 218 A: TAI ORR
angulatus Prell, 1912: 103
hexagonus Ritsema, 1882: 167

genus Hoploryctoderus Prell, 1933: 67 [RN] type species Enoplus tridens Reiche, 1860
Enoplus Reiche, 1860c: 277 [HN]

chinensis Voirin, 1996: 226 A: CH (“Mts Yuning”)
genus Oryctes Hellwig, 1798: 100 type species Scarabaeus nasicornis Linnaeus, 1758
subgenus Eremoryctes Semenov & S.I. Medvedev, 1932a: 487 type species Oryctes ata Semenov &
S.I. Medvedev, 1932
ata Semenov & S.I. Medvedev, 1932a: 487 A: IN KZ TM
subgenus Oryctes Hellwig, 1798: 100 type species Scarabaeus nasicornis Linnaeus, 1758
nasicornis afganistanicus Endrődi, 1938: 88 A: AF
afganicus Endrődi, 1938: 60

nasicornis chersonensis Minck, 1915a: 7 E: UK
nasicornis corniculatus A. Villa & G.B. Villa, 1833: 34 E: CR FR IT SL SZ

360

SCARABAEIDAE

laevigatus Heer, 1841: 534

nasicornis edithae Endrődi, 1938: 90 A: TIA
nasicornis grypus Illiger, 1803: 212 (Geotrupes) E: FR (Corse) IT MA PT SP N: AG MO TU
continuus Minck, 1914: 9
nasutus Mulsant, 1842: 374
ragusae Riggio, 1882: 17
siculus Kollar, 1836: 335
simus Mulsant, 1842: 374

nasicornis hindenburgi Minck, 1915b: 533 A: IN
montanus Minck, 1915b: 534

nasicornis holdhausi Minck, 1914: 8 E: AU BY CR CZ HU MD RO SB SK SL UK
nasicornis illigeri Minck, 1915b: 535 A: IN HP KA PA
nasicornis kuntzeni Minck, 1914: 10 E: AL BH BU CR GR KO MC ME RO SB SL TR A: CY IS LE SI SY TR
mediterraneus Minck, 1915a: 16

nasicornis latipennis Motschulsky, 1845a: 57 E: AB AR GG ST A: IN
caspicus Minck, 1915a: 10

nasicornis mariei Bourgin, 1949: 88 E: FR (Ardèche)
nasicornis nasicornis Linnaeus, 1758: 346 (Scarabaeus) E: AU BE DE FI FR GE LU NL NR NT PL SV SZ
tuberculatus Mulsant, 1842: 377

nasicornis ondrejanus Minck, 1918: 20 E: AU CZ
nasicornis polonicus Minck, 1918: 23 E: BY CT EN LA LT PL SK ST UK
nasicornis przevalskii Semenov & S.I. Medvedev, 1932a: 485 A: XIN
nasicornis punctipennis Motschulsky, 1860i: 520 E: KZ ST
kazakorum S.I. Medvedev, 1960: 75

nasicornis shiraticus Endrődi & Petrovitz, 1974: 8 A: IN
nasicornis transcaspicus Endrődi, 1938: 86 A: IN TM
zarudnyi Semenov & S.I. Medvedev, 1932a: 486

nasicornis turcestanicus Minck, 1914: 10 A: KI KZ TD UZ XIN
pumilus Minck, 1918: 22

prolixus Wollaston, 1864: 202 N: CI
subgenus Rykanes Minck, 1919: 59 type species Scarabaeus rhinoceros Linnaeus, 1758
Macroryctes Semenov & S.I. Medvedev, 1932a: 482 type species Scarabaeus rhinoceros Linnaeus, 1758

gnu Mohnike, 1874: 261 A: HAI ORR
trituberculatus van Lansberge, 1879: cliii

hisamatsui Nagai, 2002b: 45 A: JA
rhinoceros Linnaeus, 1758: 346 (Scarabaeus) A: AEi GUA GUX HAI HKG JAi PA TAI YUN “Korea” AFRi AURi
ORR
subgenus Rykanoryctes Minck, 1919: 59 type species Scarabaeus monoceros A.G. Olivier, 1789
Hovoryctes Prell, 1934: 60 type species Oryctes pyrrhus Burmeister, 1847

agamemnon agamemnon Burmeister, 1847: 201 N: EG AFR
agamemnon arabicus Fairmaire, 1896b: 256 N: TUi A: AE KU OM QA SA

361

SCARABAEIDAE
agamemnon matthiesseni Reitter, 1907c: 205 A: AF IN PA
desertorum Arrow, 1910a: 276
persicus Endrődi, 1938: 61

agamemnon sinaicus Walker, 1871: 13 A: IS SI
boas Fabricius, 1775: 8 (Scarabaeus) A: SA YE AFR
ferrugineus Thunberg, 1818: 396 (Geotrupes)
senegalensis Klug, 1835: 35

elegans Prell, 1914: 210 A: AE BA IN IQ KU OM PA QA SA YE
piesbergeni Bodemeyer, 1916: 111

richteri Petrovitz, 1958a: 10 A: IN
sahariensis Bruneau de Miré, 1960: 248 N: AG EG AFR
vicinus Gahan, 1900: 11 A: YE (Suqutra)
genus Trichogomphus Burmeister, 1847: 219 type species Scarabaeus milo A.G. Olivier, 1789
martabani Guérin-Méneville, 1833a: 484 (Oryctes) A: GUA GUX SCH SD XIZ YUN ORR
tonkinensis Fairmaire, 1893e: 313

mongol Arrow, 1908c: 347 A: CHQ FUJ GUA GUI GUX HAI HEB HKG HUB HUN JIX SCH TAI YUN ZHE ORR
robustus Arrow, 1930: 542 A: FUJ GUX YUN ORR
rongi Dechambre & Drumont, 2000: 399 A: FUJ

tribe Pentodontini Mulsant, 1842

genus Alissonotum Arrow, 1908c: 322 type species Scarabaeus piceus Fabricius, 1775
amploides Endrődi, 1973a: 317 (Heteronychus) A: GUX
binodulum Fairmaire, 1891e: cxxix (Phileurus) A: AP HP KA NP
crassum Arrow, 1908c: 323 A: PA TAI YUN ORR
cribratellum Fairmaire, 1893e: 313 (Heteronychus) A: GUA GUX HAI JA XIZ YUN ORR
elongatum Arrow, 1910a: 299 A: AP ORR
impressicolle Arrow, 1908c: 323 A: FUJ GUA GUI GUX JA SD TAI ORR
parvum Chang, 1984: 52 A: SCH YUN
pauperum Burmeister, 1847: 94 (Heteronychus) A: FUJ GUA GUX JA TAI ORR
interruptum Fairmaire, 1893e: 312 (Heteronychus)
monator Reiche, 1859a: 8 (Heteronychus)
simplex C.O. Waterhouse, 1884a: 370 (Heteronychus)

piceum Fabricius, 1775: 14 (Scarabaeus) A: HP MAC PA SD UP ORR
detractum Walker, 1859a: 54 (Phileurus)

simile Arrow, 1910a: 300 A: HP NP PA ORR
yamayai Drumont, 2013: 89 A: GUA GUX YUN ORR
genus Calicnemis Laporte, 1832a: ix, pl. 7 type species Calicnemis latreillii Laporte, 1832
Colorhinus Erichson, 1841a: 171 type species Colorhinus obesus Erichson, 1841

latreillii Laporte, 1832a: x, pl. 7 E: FR IT SP

362

SCARABAEIDAE

truncatifrons sardiniensis Leo, 1985: 421 E: IT (Sardegna)
truncatifrons truncatifrons Laporte: 1832a: ix, pl. 7 (Pachypus) E: FR PT SP N: AG MO TU
atlantica Mosconi, 1996: 704
bahilloi López-Colón, 2003: 2
bercedoi López-Colón, 2004: 73
obesa Erichson, 1841a: 171 (Colorhinus)

genus Coptognathus Burmeister, 1847: 184 type species Coptognathus crassipes Burmeister, 1847
Macrator Reitter, 1895d: 151 type species Macrator hauseri Reitter, 1895 (=Xylotrupes attilla Ménétriés, 1849)

attilla Ménétriés, 1849: 61 (Xylotrupes) A: AF KZ TM UZ
hauseri Reitter, 1895d: 151 (Macrator)

crassipes Burmeister, 1847: 184 N: EG LB
curtipennis Fairmaire, 1883b: 459
lefranci Mulsant & Godart, 1866: 448
villiersi Dechambre, 1984: 73 (Phyllognathus)

genus Dipelicus Hope, 1845: 7 type species Dipelicus cantori Hope, 1842
Camelonotus Fairmaire, 1883a: 14 [RN] type species Scarabaeus daedalus Fabricius, 1792
Horonotus Burmeister, 1847: 178 type species Scarabaeus daedalus Fabricius, 1792
Neodipelicus Ritsema, 1887: 215 type species Dipelicus nasutus Bates, 1877
Palmerstonia Blackburn, 1888: 855 type species Palmerstonia bovilli Blackburn, 1888

deiphobus loi Kobayashi, 1995c: 148 A: TAI
genus Eutyctus Semenov, 1889: 208 type species Eutyctus deserti Semenov, 1889
deserti Semenov, 1889b: 210 A: KZ TM
genus Haplosoma Semenov, 1889b: 205 type species Haplosoma ordinatum Semenov, 1889
ordinatum Semenov, 1889b: 205 A: KZ TM UZ
aralense Semenov, 1889b: 206

genus Heteronychus Dejean, 1833: 152 type species Geotrupes cricetus Hausmann, 1807
annulatus Bates, 1891a: 19 A: HP KA PA UP ORR
curtipennis Fairmaire, 1891e: cxxiv (Phileurus)

curtulus Fairmaire, 1887b: 115 A: YUN ORR
digitiformis Chang, 1983a: 298 A: GUI GUX YUN
intermedius Chang, 1983a: 299 A: GUX XIZ YUN
licas Klug, 1835: 35 (Geotrupes) N: EG A: IS SI AFR
corvinus Klug, 1855: 657
curtipennis Fairmaire, 1894c: 315
mashunus Péringuey, 1902: 519

lioderes L. Redtenbacher, 1868: 75 A: GUA GUX HP NP SCH SD UP XIZ YUN ORR

SCARABAEIDAE

363

flavopilosus Prell, 1914: 198
poropygus Bates, 1891a: 19

parumpunctatus Burmeister, 1847: 95 N: EG A: SY AFR
punctolineatus Fairmaire, 1893e: 313 A: GUX TAI ORR
sacchari Arrow, 1908c: 329 A: PA ORR
sublaevis Fairmaire, 1891e: cxxiii (Phileurus) A: KA UP XIZ ORR
robustus Arrow, 1910a: 296

tesari Endrődi, 1968c: 161 A: NMO
genus Microryctes Arrow, 1908c: 324 type species Microryctes kanarensis Arrow, 1908
confinis Y.-W. Zhang, 1991: 184 A: XIZ YUN ORR
hainanensis Y.-W. Zhang, 2002: 331 A: HAI
monodon Fairmaire, 1893e: 313 (Heteronychus) A: GUX XIZ YUN ORR
silvestrei Drumont, Dechambre, Yamaya & Verdugo, 2011: 128 A: GUX YUN ORR
genus Musurgus Vauloger, 1898: 286 type species Musurgus stridens Vauloger, 1898
sabulicola Dechambre & Pope, 1980: 123 A: SA YE
stridens Vauloger, 1898: 286 N: AG MO
genus Papuana Arrow, 1911a: 156 type species Horonotus woodlarkianus Montrouzier, 1855
philippinica Arrow, 1937: 47 A: TAI ORR
genus Pentodon Hope, 1837: 92 type species Scarabaeus punctatus Villers, 1789
algerinus algerinus Fuessly, 1778: 40 (Scarabaeus) E: FR IT SP PT N: AG EG LB MO TU
balearicus Kraatz, 1882c: 60
bispinosus Küster, 1852: 36
cribratellus Fairmaire, 1893g: cxlvi (Heteronychus) [HN]
puncticollis Burmeister, 1847: 104

algerinus bispinifrons Reitter, 1894b: 45 E: AB ST A: AF IN KZ NW TD TM UZ XIN
persicus Jakovlev, 1902b: 274
subcostatus Jakovlev, 1902a: 95

algerinus dispar Baudi di Selve, 1870: 76 E: AR GR (Kríti, Ródos) A: AE CY IN IQ IS JO KU OM QA SA SY TR YE
nasutus Petrovitz, 1964d: 131
pachypus Jakovlev, 1902b: 272
syriacus Kraatz, 1882c: 62

algerinus indicus Endrődi, 1967b: 180 A: PA ORR
bengalense Arrow, 1910a: 304 A: PA ORR
bidens bidens Pallas, 1771: 461 (Scarabaeus) E: AB RO ST UK A: AF KA KI KZ TD TM UZ WS XIN
affinis Ballion, 1871a: 346
conjunctus Jakovlev, 1902b: 269
devius Jakovlev, 1902b: 266

364

SCARABAEIDAE

dubius Ballion, 1871a: 345
emarginatus Küster, 1848: 42
humilis Ballion, 1871a: 346
jakovlevi S.I. Medvedev, 1960: 166
kazachstanicus S.I. Medvedev, 1960: 161
tardus Jakovlev, 1902b: 275
tumidus Jakovlev, 1902b: 276

bidens punctatus Villers, 1789: 40 (Scarabaeus) E: AL AU BH BU CR FR GR IT MA MC ME RO SL SP N: AG MO
castaneus Mulsant, 1842: 384
pimelioides A. Costa, 1853a: 3
punctulatus P. Rossi, 1790: 9 (Scarabaeus) [HN]
simplex Depoli, 1910: 306
testudinarius Ragusa, 1882b: 279

bidens sulcifrons Küster, 1848: 43 E: AB AR BU GG GR (Kríti) RO ST UK N: EG A: CY IN IQ IS JO SY TR
rugulosus Reitter, 1899a: 35
xyphias Baudi di Selve, 1870: 78

brachypterus Gusakov, 2004: 25 A: TM UZ
caminarius caminarius Faldermann, 1835a: 265 (Scarabaeus) A: IN IQ SY TR
anthracinus Reiche, 1861a: 2
kurdistanus Reitter, 1899a: 37
mandibularis Reitter, 1901b: 99

caminarius elatus Küster, 1847a: 60 E: AB AR GG ST A: IN
georgicus S.I. Medvedev, 1960: 138
semenowi Jakovlev, 1902b: 268

foveipennis Jakovlev, 1903a: 17 E: “Transcaucasie: Tarabany”
idiota gumariensis Petrovitz, 1961c: 117 A: PA
idiota idiota Herbst, 1789: 164 (Scarabaeus) E: AB AL AR AU BH BU CR CT CZ GG GR HU KZ MC MD ME RO
SB SK SL ST TR UK A: CY IN IQ IS LE TR XIN
bilobus Motschulsky, 1849: 110
brunneus Mulsant, 1842: 382
coniceps Reitter, 1894b: 45
curtus Motschulsky, 1849: 111
monodon Fabricius, 1792a: 20 (Scarabaeus)
ottomanus Reitter, 1899a: 36
reitteri Jakovlev, 1904c: xvi
subdilatatus Motschulsky, 1849: 109
talpa Herbst, 1789: 165 (Scarabaeus)
tauricus Motschulsky, 1849: 110
ventricosus Motschulsky, 1849: 111

idiota memnon Jakovlev, 1902b: 271 E: FR SP N: AG MO TU
insularis Y.-W. Zhang, 1991: 183 A: HKG

SCARABAEIDAE

365

kuwaitensis Endrődi & Al-Houty, 1985: 205 A: KU
minutus Reitter, 1887e: 527 A: AF IN KZ TM UZ XIN
parmatus Jakovlev, 1902a: 97
sieversi Reitter, 1899a: 30

quadridens bidentulus Fairmaire, 1887d: 319 (Heteronychus) A: BEI HUN JIA LIA NMO SHN TIA
jeholensis Niijima & Kinoshita, 1937: 21

quadridens distantidens Reitter, 1899a: 31 E: AR A: CY IN LE SY TR
quadridens mongolicus Motschulsky, 1849: 111 A: ANH GAN HEB HEI HEN HUB HUN JIA JIL LIA MG NIN
NMO QIN SHA SHN SHX XIN ZHE
gobicus Endrődi, 1965: 198
patruelis J. Frivaldszky, 1890: 202

quadridens quadridens Gebler, 1844: 100 (Scarabaeus) A: KZ TM UZ XIN XIZ
bullatus Jakovlev, 1902a: 92
distans Erichson, 1848b: 66
latifrons Reitter, 1899a: 34
nikolskii Jakovlev, 1901b: 181
pumilus Sharp, 1891: 53
semiermis Jakovlev, 1902a: 94
truncatus Sharp, 1891: 52

snegovaae Shokhin, 2015: 56 E: AB
topali Endrődi, 1974: 3 A: PA ORR
variolopunctatus deserti Heyden, 1900: 253 (Heteronychus) Ei: BH N: EG LB MO TU A: IS SA SI
milini Mikšić, 1962b: 23

variolopunctatus variolopunctatus Fairmaire, 1879c: 172 N: AG TU
pygidialis Kraatz, 1882c: 61

genus Phyllognathus Eschscholtz, 1830: 65 type species Scarabaeus silenus Fabricius, 1775 (=Scarabaeus
excavatus Forster, 1771)
Oryx Guérin-Méneville, 1838b: 80 [HN] type species not designated

dionysius Fabricius, 1792a: 20 (Scarabaeus) A: HP NP PA SD TAI UP ORR
haworthii Hope, 1831: 22 (Oryctes)
reductus Walker, 1859a: 54 (Xylotrupes)

excavatus Forster, 1771: 1 (Scarabaeus) E: AL BG BH CR FR GR IT MA MC ME PT SP ST TR UK (Krym) N: AG CI
EG LB MO TU A: AE CY IN IQ IS JO KU KZ LE OM SA SI SY TM TR YE
bacchus Mulsant, 1842: 380
cephalotes Laboulbène, 1860: 645 (Oryctes)
curvicornis Mulsant, 1842: 379
gibbifrons Mulsant, 1842: 380
hauseri Reitter, 1894b: 46
inermis Motschulsky, 1849: 112
maritimus L. Petagna, 1819: 25 (Scarabaeus)

366

SCARABAEIDAE

punctatostriatus Motschulsky, 1845a: 57
silenus Fabricius, 1775: 13 (Scarabaeus)
tuberculifer Leoni, 1910: 157

genus Podalgus Burmeister, 1847: 117 type species Podalgus cuniculus Burmeister, 1847
Crator Semenov, 1889b: 206 type species Crator infantulus Semenov, 1889
Vertumnus Reiche, 1859a: 10 [HN] type species Podalgus cuniculus Burmeister, 1847

cuniculus arabicus Fairmaire, 1895f: cccliii N: EG A: AE IS KU OM QA SA SI YE
cuniculus infantulus Semenov, 1889b: 207 (Crator) A: IN IQ KZ TD TM UZ
cuniculus reichei R. Paulian, 1954: 1167 N: AG LB MO TU AFR
genus Temnorhynchus Hope, 1837: 93 type species Geotrupes retusus Fabricius, 1781
Neotemnorrhynchus A. Janssens, 1949a: 16 type species Temnorrhynchus overlaeti Burgeon, 1947

baal Reiche & Saulcy, 1856: 369 E: GR N: AG EG LB A: CY IS LE SY TR YE AFR
coronatus Fabricius, 1781: 13 (Scarabaeus) A: CY SA YE AFR
agysimbanus Fairmaire, 1868b: 793
erectilobus Arrow, 1937: 50
faunus Péringuey, 1902: 538
spadiceus H.J. Kolbe, 1914: 368
sublaevepygus R. Paulian, 1946: 31
zanzibaricus H.J. Kolbe, 1887: 258

tribe Phileurini Burmeister, 1847

genus Eophileurus Arrow, 1908c: 332 type species Geotrupes planatus Wiedemann, 1823
subgenus Eophileurus Arrow, 1908c: 332 type species Geotrupes planatus Wiedemann, 1823
assamensis Fairmaire, 1898b: 384 (Trionychus) A: SD ORR
chinensis Faldermann, 1835b: 370 (Phileurus) A: ANH BT FE FUJ GAN GUA GUI GUX HAI HEB HEI HEN HUB
HUN JA JIA JIL JIX LIA NC NMO SC SHG SHN SHX TAI TIA YUN ZHE ORR
irregularis Prell, 1913: 122
morio Faldermann, 1835b: 371 (Phileurus)
okinawanus Nomura, 1964a: 57
poteli Fairmaire, 1898b: 384 (Trionychus)

confinis Prell, 1913: 114 A: SHG YUN ORR
decipiens Prell, 1913: 115
siamensis Arrow, 1914a: 261

forsteri Endrődi, 1971: 13 A: NP
heyrovskyi Král & Strnad, 1992 A: GUX ORR
katsurai Muramoto, 1995: 223

himalayanus Endrődi, 1976: 105 A: SD
montanus Prell, 1913: 113 A: SD
planatus Wiedemann, 1823: 5 (Geotrupes) A: AP SD UP ORR
platypterus Wiedemann, 1823: 5 (Geotrupes) A: NP PA SD UP ORR

367

SCARABAEIDAE
spatulicornis Lamant-Voirin, 1995: 150 A: CH (“Mts Yuning”)
spinosus Lamant-Voirin, 1995: 148 A: YUN ORR
subgenus Tetrodontus Y.-W. Zhang, 1991: 182 type species Eophileurus tibialis Y.-W. Zhang, 1991
tetraspermexitus Ratcliffe, 1988: 46 A: YUN ORR
tibialis Y.-W. Zhang, 1991: 182

genus Rhizoplatys Westwood, 1841a: 38 type species Rhizoplatys cribrarius Westwood, 1841
arabicus Dechambre, 2005: 301 A: OM

subfamily Cetoniinae Leach, 1815
tribe Cetoniini Leach, 1815
subtribe Cetoniina Leach, 1815

genus Aethiessa Burmeister, 1842: 405 type species Cetonia floralis Fabricius, 1787
Brachytricha Reiche, 1871: 84 type species Cetonia feralis Erichson, 1841

feralis Erichson, 1841a: 174 (Cetonia) N: AG TU
floralis atlantis Krajčík, 2013: 3 N: MO (Rabat, Agadir)
floralis barbara Gory & Percheron, 1833: 231 (Cetonia) N: MO
floralis floralis Fabricius, 1787a: 31 (Cetonia) E: MA SP N: AG EG LB TU A: IS SAi SI
aethiopica Burmeister, 1842: 411
aupicki Gory & Percheron, 1833: 232 (Cetonia)
deserticola Waltl, 1839: 68 (Cetonia)
divergens Bedel, 1889: 91
dogueraui Gory & Percheron, 1833: 230 (Cetonia)
dolorosa Mulsant, 1842: 570 (Cetonia)
elongata Gory & Percheron, 1833: 228 (Cetonia)
flavospila Motschulsky, 1849: 120 (Cetonia)
funerea Mulsant, 1842: 571 (Cetonia)
inhumata Gory & Percheron, 1833: 227 (Cetonia)
lefebvrei Mulsant, 1842: 571 (Cetonia)
leucospila Burmeister, 1842: 413
numisma Newman, 1838a: 168 (Cetonia)
refulgens Herbst, 1790: 250 (Cetonia)
stigmatica Mulsant, 1842: 571 (Cetonia)
viridiflua Motschulsky, 1849: 118 (Cetonia)
tenebrionis Gory & Percheron, 1833: 233 (Cetonia)

floralis squamosa Gory & Percheron, 1833: 232 (Cetonia) E: IT (Calabria, Sicilia)
martini Bedel, 1889: 91 N: AG MO
mesopotamica Burmeister, 1842: 414 E: AB AR GG A: IS JO LE SY TR
albocincta Kraatz, 1884b: 231
maera Reitter, 1899a: 53

rugipennis Burmeister, 1842: 417 E: AB AR A: AF IN IQ JO TM TR
bagdadensis Burmeister, 1842: 414

368

SCARABAEIDAE

doriae Reiche, 1871: 85 (Cetonia)
zarudnyi Kiseritzky, 1910: 281

szekessyi Brasavola de Massa, 1939: 161 A: KZ TM UZ
crinita Nikolajev, 1975b: 282

genus Anatona Burmeister, 1842: 503 type species Anatona flavoguttata Burmeister, 1842 (=Cetonia stillata
Newman, 1838)
Eumimela Kraatz, 1881: 264 type species Eumimela pygialis Kraatz, 1881 (=Anoplochilus castanopterus Burmeister, 1842)

castanoptera Burmeister, 1842: 509 (Anoplochilus) A: HP SD UP ORR
pygialis Kraatz, 1881: 264 (Eumimela)

stillata Newman, 1838a: 169 (Cetonia) A: HP UP ORR
ferruginipennis Kraatz, 1898a: 223
flavoguttata Burmeister, 1842: 504
lignea C.É. Blanchard, 1842: 8 (Cetonia)
pilicollis Kraatz, 1898a: 223

genus Cetonia Fabricius, 1775: 42 type species Scarabaeus auratus Linnaeus, 1758
subgenus Cetonia Fabricius, 1775: 42 type species Scarabaeus auratus Linnaeus, 1758
Tecinoa A. Costa, 1852: 12 type species Scarabaeus auratus Linnaeus, 1758

aeratula Reitter, 1891d: 56 E: AB AR GG A: IN TM
asiatica Gory & Percheron, 1833: 244 E: GR (Kríti)
oertzeni Reitter, 1885b: 379
scutellaris Reitter, 1891d: 55

aurata aurata Linnaeus, 1758: 352 (Scarabaeus) E: AL AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU
IR IT KO LA LS LT LU MC MD ME NL NR NT PL RO SB SK SL SP ST SV SZ TR UK A: ES FE IN KI KZ MG TR
UZ WS XIN
aenea Fuessly, 1778: 313
chlorochrysa Schröter, 1776: 14
cuprifulgens Mulsant, 1842: 563
maculata Scriba, 1790: 68
nobilis Schrank, 1798: 10
piligera Mulsant, 1842: 563
praeclara Mulsant, 1842: 563
prasiniventris Reitter, 1896f: 245
purpurata Heer, 1841: 551
rubifera Bourgin, 1943: 110
smaragda De Geer, 1774: 279 (Scarabaeus)
purpurata Heer, 1841: 551
pyrochroa Olsoufieff, 1916: 173
rhilensis Nedelkow, 1905: 430
tunicata Reitter, 1896f: 245

SCARABAEIDAE

369

uniformis Reitter, 1899a: 59
variabilis Preyssler, 1790: 51

aurata jingkelii Flutsch & Tauzin, 2009: 101 A: SCH
aurata pallida Drury, 1773: 71 (Scarabaeus) E: AB BH BU CR CT GR SB ST TR UK (Krym) A: IN IQ KZ LE TR
amasicola Reitter, 1899a: 61
angorensis Reitter, 1896f: 246
nigriventris Reitter, 1896f: 246
strigiventris Reitter, 1896f: 246
tatarica Reitter, 1896f: 246
undulata Reitter, 1896f: 245
violaceiventris Reitter, 1899a: 61

aurata pisana Heer, 1841: 552 E: FR GR IT PT SP SZ
bicolora Leoni, 1910: 185
bilucida Reitter, 1899a: 62
castanea A. Costa, 1852: 27
coerulescens Leoni, 1910: 183
corsicana Heller, 1900: 56
cyanicollis Reitter, 1899a: 62
delfieui Le Comte, 1904: 94
devagnerii Leoni, 1910: 184
elegans Leoni, 1910: 184
fiorii Leoni, 1910: 183
hispanica Erichson, 1847: 605
immaculata Heller, 1900: 56
intermedia Leoni, 1910: 184
lucidula Fieber, 1831: 472
meridionalis Mulsant, 1842: 564
nigra Gautier des Cottes, 1863: 490
olivicolor Reitter, 1896f: 244
pseudonigra Leoni, 1910: 182
semicyanea Reitter, 1899a: 62
sordida Heller, 1900: 56
thoracalis Heller, 1900: 56
tingens Reitter, 1896f: 246
valesiaca Heer, 1842: 552
violacea Fieber, 1831: 472
viriditarsis Heller, 1900: 55

aurata pokornyi Rataj, 2000: 11 A: WS (Altai)
aurata sicula Aliquò, 1983: 35 E: IT (Sicilia)
aeneicolor Leoni, 1910: 183

aurata viridiventris Reitter, 1896f: 245 E: CT A: ES KI KZ UZ WS

370
viridana Nedelkow, 1909: 25

aurataeformis Curti, 1913: 141 E: PT SP
castiliana Báguena, 1955b: 295
dichroa Báguena, 1955b: 295
esmeraldina Seabra, 1905: 12
pyrenaica Báguena, 1955b: 295
viridis Seabra, 1905: 22

carthami Gory & Percheron, 1833: 243 E: FR (Corse) IT (Sardegna, Sicilia)
bella Reitter, 1891d: 56
italica Reitter, 1891d: 56

delagrangei cypriaca Alexis, 1994: 469 A: CY
delagrangei delagrangei Boucard, 1893: 40 E: AR N: EG A: IS LE SY TR
chrysoprasina Reitter, 1896f: 243
chrysosoma Reitter, 1896a: 4
fulgida Reitter, 1896f: 243
lumenifera Reitter, 1896f: 242
praelustris Reitter, 1896f: 243
syriaca Boucard, 1893: 40

funeraria Gory & Percheron, 1833: 243 N: AG MO TU
gerardi Kraatz, 1879b: 246

kemali Delpont, 1995: 340 A: TR
sexguttata Olsoufieff, 1916: 161 E: AB A: IN
viridescens Reitter, 1891d: 53 A: IN SY
subgenus Cetonia, nomen dubium
delfilsi Le Comte, 1904: 80 E: FR
subgenus Eucetonia Schoch, 1894: 211 type species Cetonia magnifica Ballion, 1871
angulicollis Kraatz, 1889b: 671 A: GAN
chinensis Schürhoff, 1942: 292 (Eucetonia) A: GAN HUB JIX SHA
filchnerae Kolbe, 1908a: 86 A: GAN
kolbei Curti, 1914: 125 A: GAN SHA SHN
magnifica Ballion, 1871a: 348 A: ES FE HEB HEI LIA NC SHN
cupreola Kraatz, 1879b: 243

pakistanica Krajčík & Jákl, 2004: 3 A: PA
pilifera izuensis Nomura, 1969: 93 A: JA (Izu-shotō)
pilifera pilifera Motschulsky, 1861a: 15 (Glycyphana) A: FE JA SC NE
pililineata Schürhoff, 1942: 292 (Eucetonia) A: SCH
prasinata Bourgoin, 1915d: 174 A: TAI
pygidionotis Schürhoff, 1942: 292 (Eucetonia) A: YUN
roelofsi gotoana Nakane, 1964b: 48 A: JA (Gotô)
roelofsi iijimai K. Sakai, 1996: 11 A: JA (Nishino-shima)
roelofsi roelofsi Harold, 1880e: v A: FE (Iturup, Kunashir) JA SC

SCARABAEIDAE

SCARABAEIDAE

371

bodemeyeri Curti, 1915: 20
nitidula Reitter, 1899a: 55
villifrons Reitter, 1899a: 55

roelofsi tane Nakane, 1993a: 2 A: JA (Tanega-shima)
sichuana Krajčík, 2002a: 13 A: SCH
viridiopaca Motschulsky, 1858a: 18 (Glycyphana) A: BEI ES FE HEB HEI NC NMO SC SHA
amouriensis J. Thomson, 1878a: 24 (Euryomia)

subgenus Indocetonia Mikšić, 1965b: 14 type species Protaetia bensoni Westwood, 1849
bensoni Westwood, 1849b: 145 (Protaetia) A: KA PA UP ORR
rhododendri Gestro, 1891: 847 (Cetonia) A: SD UP ORR
assamica Schoch, 1898b: 181 (Eucetonia)
purpurascens Kraatz, 1897b: 405 (Cetonia)

rutilans Janson, 1881: 608 (Glycyphana) A: NP SD
sehnali Krajčík, 2012b: 2 A: SCH
wrzecionkoi Krajčík, 2008a: 2 A: AP XIZ
genus Gametis Burmeister, 1842: 356 type species Cetonia versicolor Fabricius, 1775
Oxycetonia Arrow, 1910a: 163 type species Cetonia versicolor Fabricius, 1775

andrewesi Janson, 1901: 182 A: SD ORR
indecora Mikšić, 1982: 197

bealiae Gory & Percheron, 1833: 282 (Cetonia) A: “Indes orientales” FUJ GUA HEB ZHE ORR
forticula forticula Janson, 1881: 608 (Glycyphana) A: JA
forticula formosana Nomura, 1959b: 53 (Oxycetonia) A: TAI
forticula ishigakiana Nomura, 1959b: 52 (Oxycetonia) A: JA (Ishigaki-jima)
forticula kotoensis Nomura, 1959b: 52 (Oxycetonia) A: TAI (Lanyu)
forticula lutaoensis Kobayashi, 1990e: 86 A: TAI (Lutao)
forticula miyakoana Nomura, 1959b: 52 (Oxycetonia) A: JA (Miyako-jima)
forticula yonakuniana Nomura, 1959b: 52 (Oxycetonia) A: JA (Yonaguni-jima)
incongrua Janson, 1878a: 388 (Glycyphana) A: TAI
costigera Bourgoin, 1915c: 168 (Oxycetonia)

histrio A.G. Olivier, 1789: no. 6: 45 (Cetonia) A: AP HP NP UP “Thibet” ORR
albopunctata Fabricius, 1798: 129 (Cetonia)
atra Mikšić, 1982: 201
bivittata Burmeister, 1842: 363

jucunda Faldermann, 1835: 386 (Cetonia) A: BEI CHQ FE FUJ GAN GUI GUX HAI HEB HEI HKG HUB JA JIA
JIX LIA NC NMO NP PA SC SCH SD SHG SHN SHX YUN ZHE ORR
albosetosa Motschulsky, 1862a: 9 (Glycyphana)
argyrosticta Burmeister, 1842: 360
californica LeConte, 1863: 80 (Euphoria)
dolens Kraatz, 1879a: 236 (Glycyphana)
fallax Mikšić, 1982: 191

372
ferruginosa Reitter, 1899a: 53
goryi Guérin-Méneville, 1840a: 81 (Cetonia)
hainanensis Mikšić, 1982: 195
kuperi Schaum, 1848a: 69 (Glycyphana)
lateriguttata Fairmaire, 1887b: 117 (Glycyphana)
nigra Kraatz, 1893b: 74
obscura Gory & Percheron, 1833: 285 Cetonia)
obscurina Gory & Percheron, 1833: 396 (Cetonia)
prasina Hope, 1831: 25 (Cetonia)
pretiosa Mikšić, 1982: 194
quadrimaculata Mikšić, 1982: 191
rubra Kraatz, 1893b: 74
sanguinalis Hope, 1831: 25 (Cetonia)
subfasciata Reitter, 1896c: 70 (Glycyphana)
variolosa Motschulsky, 1860e: 135 (Glycyphana)
viridiobscura Gory & Percheron, 1833: 406 (Cetonia)
vitticollis Reitter, 1899a: 54

versicolor Fabricius, 1775: 51 (Cetonia) A: AF BT NP PA UP AFR ORR
albella Gmelin, 1790: 1577 (Cetonia)
cruenta Pallas, 1781: 21 (Scarabaeus)
detrita Gory & Percheron, 1833: 281 (Cetonia)
glabra Mikšić, 1982: 205
luctuosa Gory & Percheron, 1833: 283 (Cetonia)
sanguinolenta Gmelin, 1790: 1583 (Scarabaeus)
taprobanica Mikšić, 1982: 203
thebana Herbst, 1779: 324 (Scarabaeus)
variegata Fabricius, 1775: 51 (Cetonia)

genus Glycosia Schoch, 1896a: 86 type species Cetonia tricolor A.G. Olivier, 1789
Sternoplidius Moser, 1908a: 89 type species Sternoplidius alboguttatus Moser, 1908

luctifera doubleti Pouillaude, 1914c: 186 A: SHX YUN
luctifera dureli Pouillaude, 1914c: 187 A: BT SD
luctifera louisae Fairmaire, 1888d: xxxv (Glycyphana) A: YUN XIZ
luctifera luctifera Fairmaire, 1878a: 107 (Glycyphana) A: BT GAN SCH SD SHX TAI YUN
nigra Kometani, 1940a: 155

mawenzhena Krajčík, 2011c: 34 [RN] A: XIZ
nigra Ma, 1988: 274 [HN]

tricolor A.G. Olivier, 1789: no 6: 88 (Cetonia) A: SD ORR
nagpurensis Arrow, 1910a: 130
plagiata Schoch, 1896a: 86

SCARABAEIDAE

SCARABAEIDAE

373

genus Chiloloba Burmeister, 1842: 501 type species Cetonia acuta Wiedemann, 1823
acuta Wiedemann, 1823: 87 (Cetonia) A: HP NP PA SD UP ORR
perplexa Gory & Percheron, 1833: 284 (Cetonia)

genus Glycyphana Burmeister, 1842: 345 type species Cetonia horsfieldii Hope, 1831
subgenus Caloglycyphana Mikšić, 1968a: 139 type species Cetonia binotata Gory & Percheron, 1833
catena catena Arrow, 1910a: 122 A: BT SD ORR
reischigi Krajčík, 2009a: 2 A: FUJ
subgenus Glycyphana Burmeister, 1842: 345 type species Cetonia horsfieldii Hope, 1831
Glycetonia Reitter, 1891d: 52 type species Glycyphana fulvistemma Motschulsky, 1858

aurocincta Arrow, 1910a: 122 A: BT ORR
fulvistemma Motschulsky, 1858a: 18 A: BEI ES FE FUJ GUX HEI HUB JA LIA NC SCH SHG TAI
YUN ZHE
sieboldi Snellen van Vollenhoven, 1864: 158 (Euryomia)
nagoyana Takagi, 1936: 24

horsfieldii Hope, 1831: 25 (Cetonia) A: BT GUI JIX NP SCH SD TAI YUN ORR
aurulenta Arrow, 1910a: 121
biargentata J. Thomson, 1878a: 26 (Euryomia)
chinensis Mikšić, 1967: 22
cingalensis Schein, 1956b: 22
marginicollis Gory & Percheron, 1833: 251 (Cetonia)

hybrida Mikšić, 1970a: 106 A: TAI
subgenus Glycyphaniola Mikšić, 1968a: 140 type species Cetonia modesta Fabricius, 1792
gracilis gracilis Sawada, 1942c: 240 A: TAI
gracilis viridis Sawada, 1942c: 241 A: JA
laotica Mikšić, 1968c: 48 A: HKG ORR
nepalensis Kraatz, 1894f: 294 A: FUJ HAI NP SD YUN ORR
fujianensis Krajčík, 2012b: 7
nigropicea Kraatz, 1894f: 294
roquesi Bourgoin, 1916a: 111

genus Glycyphana, species incertae sedis
minima Bates, 1891a: 21 A: HP
nasalis Boheman, 1858: 58 A: HKG
genus Heterocnemis Albers, 1853: 46 type species Cetonia graeca Brullé, 1832
subgenus Celidotella Reitter, 1909b: 83 type species Celidotella matthiesseni Reitter, 1909
matthiesseni Reitter, 1909b: 83 (Celidotella) A: IN
subgenus Heterocnemis Albers, 1853: 46 type species Cetonia graeca Brullé, 1832
graeca Brullé, 1832: 185 (Cetonia) E: GR A: TR
quadrata Gory& Percheron, 1833: 294 (Cetonia)

374

SCARABAEIDAE

genus Pachnoda Burmeister, 1842: 511 type species Cetonia marginella Fabricius, 1775
Eupachnoda H.J. Kolbe, 1892: 253 type species Cetonia inscripta Gory & Percheron, 1833
Pachnodula Schoch, 1898a: 62 type species Cetonia interrupta A.G. Olivier, 1789

cervenkai Krajčík, 2002b: 16 A: OM
fasciata Fabricius, 1775: 46 (Cetonia) N: EG A: YE SA
alexandrina Gmelin, 1790: 1576 (Cetonia)
cincta A.G. Olivier, 1789: 6: 87 (Cetonia)

histrio Fabricius, 1775: 51 (Cetonia) N: EG A: “Arabie”
histrioides Pouillaude, 1914a: 23 A: YE
interrupta A.G. Olivier, 1789: no. 6: 38 (Cetonia) N: EG A: YE “Arabia” AFR
bipunctata A.G. Olivier, 1789: no 6: 48 (Cetonia)
soudanica Ancey, 1886: 224 (Cetonia)
spreta C.É. Blanchard, 1842: 2

leclercqi Rigout, 1985: 4 A: SA YE
nigricans Rigout, 1985: 4

marginata marginata Drury, 1773: 59 (Scarabaeus) N: EG A: “Arabia” AFR
monacha Gory & Percheron, 1833: 184 (Cetonia)

marginata mirei Ruter, 1963: 1141 N: MO AFR
marginata tunisiensis Rigout, 1984: 65 N: TU
polita C.É. Blanchard, 1842: 2 A: SA
savignyi Gory & Percheron, 1833: 56 (Cetonia) N: AG EG LB MO AFR
aurentia Gory & Percheron, 1833: 185 (Cetonia)
consentanea Schaum, 1844: 386
fasciata Burmeister, 1842: 518
marginata Gory & Percheron, 1833: 183 (Cetonia)

thoracica Fabricius, 1775: 47 (Cetonia) N: EG A: SA YE
arabica C.É. Blanchard, 1842: 3

genus Protaetia Burmeister, 1842: 472 type species Cetonia mandarina Weber, 1801 (=Cetonia fusca
Herbst, 1790)
subgenus Calopotosia Reitter, 1899a: 66 type species Cetonia submarmorea Burmeister, 1842
formosana Moser, 1910c: 356 (Liocola) A: TAI
inquinata Arrow, 1913c: 406 [RN] A: TAI
formosana Moser, 1910c: 357 (Calopotosia) [HN]

lewisi leachi Kurosawa, 1959: 337 A: JA (Amami-Ôshima)
aeneonitens Ruter, 1978: 128

lewisi lewisi Janson, 1888b: 194 (Cetonia) A: JA (Okinawa-shotō)
descarpentriesi Ruter, 1978: 127

orientalis orientalis Gory & Percheron, 1833: 193 (Cetonia) A: CHQ FE FUJ GUI HKG HUB HUN JIX NC SC SCH
ZHE ORR
aerata Erichson, 1834: 240 (Cetonia)

SCARABAEIDAE

375

confuciusana J. Thomson, 1878a: 28 (Cetonia)
ignea Kraatz, 1889a: 380 (Cetonia)

orientalis sakaii Kobayashi, 1994: 148 A: JA (Okinawa-shotō: Kitadaitojima, Minamidaitojima) TAI
orientalis submarmorea Burmeister, 1842: 460 (Cetonia) A: JA
orientalis tokarana Nomura, 1964a: 59 A: JA (Tokara: Naka-no-shima)
subgenus Cetonischema Reitter, 1899a: 68 type species Scarabaeus aeruginosus Linnaeus, 1767 sensu Drury,
1773 (=Potosia aeruginosa Medvedev, 1964)
speciosissima Scopoli, 1786: 48 (Scarabaeus) E: AL AU BH BU BY CR CT CZ FR GE GR HU IT KO MC MD ME
PL RO SB SK SL SP ST SZ TR UK A: TR
aeruginosa Medvedev, 1964: 197 (Potosia)
aureocuprea Mulsant, 1842: 547 (Cetonia)
fastuosa Fabricius, 1792a: 127 (Cetonia)
frischi Schrank, 1798: 416 (Cetonia)
ignea Reitter, 1899a: 69 (Potosia)
immaculata Scriba, 1790: 68 (Cetonia)
miribella Reitter, 1899a: 69 (Potosia)
nudiventris Germar, 1824: 132 (Cetonia)
posnaniensis Szulczewski, 1922: 241 (Potosia)
smaragda Brahm, 1790: 153 (Scarabaeus)
superba Villers, 1789: 36 (Scarabaeus)
viridis Fuessly, 1786: 93 (Scarabaeus)

speciosa cyanochlora Schauer, 1941: 180 (Potosia) A: IN
farsensis Petrovitz, 1980: 637 (Cetonischema)

speciosa jousselini Gory & Percheron, 1833: 222 (Cetonia) A: IN IQ IS LE SY TR
akbesiana Balthasar, 1929a: 116 (Potosia)

speciosa marchei Alexis & Delpont, 1999: 389 (Cetonischema) A: CY
speciosa nematiani Montreuil, 2006b: 428 A: IN
speciosa speciosa Adams, 1817: 302 (Cetonia) E: AB AR GG ST UK (Krym) A: IN LE SY TM TR
caucasica Reitter, 1899a: 69 (Potosia)
ignea Nonfried, 1905b: 331 (Cetonischema)
impressicollis Nonfried, 1905b: 331 (Cetonischema)
moseri Nonfried, 1905b: 331 (Cetonischema)
nobilitata Nonfried, 1905b: 331 (Cetonischema)
obscuripennis Nonfried, 1905b: 332 (Cetonischema)
psittacina Ménétriés, 1832: 189 (Cetonia)

speciosa venusta Ménétriés, 1836a: 150 (Cetonia) E: TR A: TR
subgenus Chrysoliocola Alexis & Delpont, 1998b: 167 type species Potosia nodieri Bourgoin, 1919
elegans Kometani, 1938: 443 (Potosia) A: TAI
subgenus Chrysopotosia Mikšić, 1966a: 7 type species Calopotosia mandschuriensis Schürhoff, 1933
flutschi Legrand, 2005a: 32 A: HAI

376

SCARABAEIDAE

mandschuriensis drumonti Alexis, 1995: 350 (Chrysopotosia) A: ANH GAN GUI HEB HUB HUN QIN SCH SHA
SHN XIZ YUN
minshanensis Alexis & Delpont, 1996a: 283 (Chrysopotosia)

mandschuriensis hainanensis Devecis & Flutsch, 2015: 20 (Chrysopotosia) A: HAI
mandschuriensis mandschuriensis Schürhoff, 1933a: 29 (Calopotosia) A: BEI FE HEI LIA NC NMO SC
subgenus Dicranobia Reitter, 1900a: 11 [RN] type species Cetonia potanini Kraatz, 1889
Dicranophorus Reitter, 1899a: 63 [HN] type species Cetonia potanini Kraatz, 1889

neonata Smetana, 2006: 290 [RN] A: CHQ SCH
potanini S.I. Medvedev, 1964: 248 (Potosia) [HN]

potanini legrandi Krajčík, 2011c: 40 [=2014: 1] [RN] A: HAI
hainanensis Legrand, 2005a: 36 [HN]

potanini potanini Kraatz, 1889b: 669 (Cetonia) A: GAN SCH
rochei Bourgoin, 1916a: 111

sauteri Moser, 1915d: 592 (Potosia) A: TAI
subgenus Endrodia Mikšić, 1958a: 8 type species Cetonia szechenyi J. Frivaldszky, 1890
latimarginalis Ma, 1992a: 198 A: YUN
ligularis Ma, 1992a: 197 A: YUN
sutchuenica Mikšić, 1984: 303 A: SCH
szechenyi J. Frivaldszky, 1890: 202 (Cetonia) A: SCH YUN
subgenus Eumimimetica Kraatz, 1881: 264 type species Cetonia terrosa Gory & Percheron, 1833
Pseudanthracophora Kraatz, 1898b: 407 type species Pseudanthracophora striatipennis Kraatz, 1898 (=Cetonia terrosa
Gory & Percheron, 1833)

terrosa Gory & Percheron, 1833: 264 (Cetonia) A: SD ORR
irrorata Wallace, 1867: xcvii (Cetonia)
striatipennis Kraatz, 1898b: 407 (Pseudanthracophora)

subgenus Eupotosia Mikšić, 1954a: 5 type species Cetonia affinis Andersch, 1797
affinis affinis Andersch, 1797: 154 (Cetonia) E: AB AL AR AU BE BH BU BY CR CT CZ FR GE GG GR HU IT KO
MD ME RO SB SK SL SP ST SZ TR UK A: IN IS LE SY TM TR
aenea Illiger, 1806: 235 (Cetonia)
albonotata Le Comte, 1906: 249 (Potosia)
pyrochroa Reitter, 1899a: 74 (Potosia)
quercus Bonelli, 1812: 59 (Cetonia)

affinis falsafii Montreuil, 2006b: 429 A: IN
cyaniventris Tauzin, 2008: 139

affinis pseudospeciosa S.I. Medvedev, 1964: 210 (Potosia) E: AB AR GG A: IN TM
affinis pyrodera Reitter, 1891d: 60 (Potosia) A: IS LE SY TR
affinis tyrrenica S.I. Medvedev, 1964: 209 (Potosia) E: FR (Corse) IT (Sardegna, Sicilia)
lamberti A. Paulian, 1986: 193 (Eupotosia)
pauliani Moretto & Baraud, 1982: 147 (Eupotosia)

mirifica koenigi Reitter, 1894d: 127 (Potosia) A: IS LE SY TR
mirifica mirifica Mulsant, 1842: 549 (Cetonia) E: AL BH BU FR GR IT SP SZ A: TR

SCARABAEIDAE

377

amethystina Benderitter, 1896: 40 (Cetonia)
balcanica Mikšić, 1965a: 223 (Potosia)

subgenus Fairmairiana Alexis & Delpont, 1998c: 535 type species Cetonia delavayi Fairmaire, 1888
acantha Ma, 1993a: 175 [DA] A: BEI
bousqueti Alexis & Delpont, 1998c: 539 (Fairmairiana) A: YUN
davidiana Fairmaire, 1889a: 31 (Cetonia) A: GUI SCH
delavayi Fairmaire, 1888a: 21 (Cetonia) A: YUN
subgenus Foveopotosia Mikšić, 1959a: 43 type species Cetonia judith Reiche, 1871
judith Reiche, 1871: 87 (Cetonia) E: GR (Ródos) N: EG A: CY IS LE SY TR
foveicauda Reitter, 1899a: 82 (Potosia)

subgenus Indoprotaetia Mikšić, 1968b: 4 type species Cetonia alboguttata Vigors, 1825
alboguttata Vigors, 1825b: 238 (Cetonia) A: NP SD UP ORR
saundersi Bainbridge, 1842: 219 (Cetonia)

wiebesi Mikšić, 1966b: 299 A: SD ORR
subgenus Liocola C.G. Thomson, 1859: 77 type species Cetonia marmorata Fabricius, 1792
Pachnotosia Reitter, 1891d: 56 type species Cetonia marmorata Fabricius, 1792

auripes Hope, 1831: 24 (Cetonia) A: NP SD ORR
ignipes Burmeister, 1842: 467 (Cetonia)

brevitarsis Lewis, 1879b: 463 (Cetonia) A: ANH BEI CHQ FE FUJ GAN GUA GUI GUX HEB HEI HEN HUB HUN
JA JIA JIL JIX LIA NC NIN NMO QIN SCH SHA SHG SC SHN SHX XIN XIZ YUN ZHE
bipunctata Kraatz, 1889a: 380 (Cetonia)
confluens Kraatz, 1889a: 380 (Cetonia)
crassa Harold, 1880e: vi (Cetonia)
cyaniventris Kraatz, 1883a: 11 (Cetonia)
fairmairei Kraatz, 1889a: 379 (Cetonia)
impunctata Kraatz, 1889a: 380 (Cetonia)
marginicollis Kraatz, 1889a: 380 (Cetonia)
mimuloides Reitter, 1899a: 66 (Pachnotosia)
nigrocyanea Kraatz, 1879a: 238 (Cetonia)
seulensis H.J. Kolbe, 1886: 194 (Cetonia)
sexpunctata Kraatz, 1889a: 380 (Cetonia)
thibetana Kraatz, 1889a: 380 (Cetonia)
vaclavi Mikšić, 1958b: 212 (Potosia)
viridis Kraatz, 1889a: 380 (Cetonia)
viridiventris Kraatz, 1889a: 380 (Cetonia)

cataphracta Arrow, 1913c: 405 A: JA TIA
cathaica Bates, 1890: 245 (Cetonia) A: GAN GUI JIA SCH
chicheryi gillesflutschi Krajčík, 2014: 1 [RN] A: GUX YUN
yunnana Flutsch & Jingke, 2013: 27 [HN]

formosana formosana Moser, 1910c: 356 (Liocola) A: TAI
formosana nozatoi Miyake, 1986: 5 A: JA (Saki-shima: Yonaguni)

378

SCARABAEIDAE

hainanensis Devecis, 2004a: 23 (Sinoliocola) A: HAI
himalayana Mikšić, 1987a: 511 A: NP “Himalaya” ORR
insperata Lewis, 1879b: 463 (Cetonia) A: FE (Kunashir, Sakhalin) JA
marmoratoides Reitter, 1899a: 65 (Pachnotosia)

marmorata kalinka Kemal & Kocak, 2009: 2 [RN] A: ES FE HEB HEI NC
orientalis S.I. Medvedev, 1964: 185 (Potosia) [HN]

marmorata marmorata Fabricius, 1792b: 127 (Cetonia) E: AL AU BE BH BU BY CR CT CZ DE EN FI FR GG GR
HU IT LA LT LU MC MD NL NR NT PL RO SB SK SL ST SV SZ UK A: ES KZ MG WS
aenea Scriba, 1790: 69 (Cetonia) [HN]
aeruginea Herbst, 1790: 216 (Cetonia) [HN]
lugubris Herbst, 1786: 157 (Cetonia) [HN]
punctatoundulata L. Petagna, 1819: 35 (Cetonia)
quercus Schrank, 1798: 416 (Cetonia)
zoufali Roubal, 1929b: 101 (Liocola)

miharai Alexis & Delpont, 1998b: 170 (Liocola) A: YUN ORR
miyakoensis Niijima & Kinoshita, 1923: 233 (Liocola) A: JA (Ryukyus: Miyako-jima)
olivacea Ma, 1993a: 171 A: XIZ
pulverisata Devecis, 2004b: 65 (Sinoliocola) A: QIN SHA
rudis Ma, 1993a: 171 A: YUN
speculifera Swartz, 1817: 53 (Cetonia) A: HEN HUN JIX ORR
garreta Bourgoin, 1916a: 110 (Cetonia)
philippensis Gory & Percheron, 1833: 194 (Cetonia)
thyrsis Reitter, 1899a: 68 (Potosia)

tropida Ma, 1993a: 170 A: HAI
subgenus Macroliocola Alexis & Delpont, 1998b: 167 type species Cetonia montana Nonfried, 1892
montana Nonfried, 1892e: 371 (Cetonia) A: SD ORR
nigropurpurea Yawata, 1941: 122 A: TAI
subgenus Macroprotaetia Mikšić, 1965d: 135 type species Cetonia inanis Wallace, 1867
gestroi Mikšić,1970b: 56 A: SD ORR
cuprea Gestro, 1891: 851 (Cetonia) [HN]

subgenus Miksicoprotaetia Legrand & Chew Kea Foo, 2010: 29 type species Protaetia acutissima Mohnike,
1871
gillesi Legrand, 2013: 3 A: YUN
subgenus Netocia A. Costa, 1852: 14 type species Cetonia morio Fabricius, 1781
Melanosa Mulsant & Rey, 1871a: 687 [=1871c: 431] type species Cetonia morio Fabricius, 1781

absidata Ma, 1993a: 173 A: YUN
aethiessina Reitter, 1891d: 74 (Brachytricha) E: AR A: IN TR
vinokurovi Zaitzev, 1918b: 128 (Potosia)

agglomerata Solsky, 1874a: 19 (Cetonia) A: KI KZ TM UZ
alexandra Reitter, 1899a: 94 (Potosia)
belemia Reitter, 1899a: 94 (Potosia)

SCARABAEIDAE
confluens Kraatz, 1884a: 222 (Cetonia)
conglomerata Kraatz, 1886a: 106 (Cetonia)
immarginata Kraatz, 1884a: 223 (Cetonia)
nigroaenea Kraatz, 1883e: 343 (Cetonia)

allardi Rataj, 1986: 16 (Potosia) A: UZ
annae Reitter, 1891d: 71 (Melanosa) A: KI TD UZ
gemina Reitter, 1899a: 95 (Potosia)

araratica Reitter, 1891d: 69 (Melanosa) E: AB AR GG A: AF IN KI KZ TM TR UZ
excavata Faldermann, 1835a: 299 (Cetonia) [HN]
lora Reitter, 1891d: 68 (Melanosa)

asiatica Faldermann, 1835a: 300 (Cetonia) E: AB AR GG A: IN
kaznakovi Olsoufieff, 1916: 167 (Potosia)

aterrima Nonfried, 1895: 293 (Melanosa) A: “Tschi-fu W. China”
bogdanoffi Solsky, 1876b: 277 (Cetonia) E: AF KI KZ TD UZ
cyanea cyanea Kraatz, 1886a: 109 (Cetonia) A: TD TR UZ
cyanea nuristanica Petrovitz, 1959a: 110 (Potosia) [RN] A: AF
afghana Schein, 1956a: 18 (Potosia) [HN]

golestanica Alexis & Delpont, 2001c: 62 (Netocia) A: IN
interruptecostata Ballion, 1871a: 347 (Cetonia) A: AF KI KZ TD TM UZ XIN
kessleri Solsky, 1876b: 276 (Cetonia)
obtusecostata Reitter, 1891d: 69 (Melanosa)

jakli Montreuil & Legrand, 2008: 158 (Netocia) A: IN
karbalayei Alexis & Delpont, 2002: 81 (Netocia) A: IN
karelini Zoubkov, 1829: 159 (Cetonia) E: CT ST A: KI KZ TD UZ XIN
aemula Ballion, 1871a: 347 (Cetonia)
conspersa Ballion, 1871a: 347 (Cetonia)
conspersula Reitter, 1899a: 93 (Potosia)
herminae Reitter, 1891d: 70 (Melanosa)
kardini Gory & Percheron, 1833: 217 (Cetonia)
obesa Gebler, 1830: 111 (Cetonia)
vermicularis Reitter, 1899a: 93 (Potosia)

khorasanica Montreuil & Legrand, 2008: 157 (Netocia) A: IN
loudai Rataj, 1998: 140 (Potosia) A: FE
lugubrides Schürhoff, 1932: 89 (Potosia) A: SCH XIZ
morio heyrovskyi Balthasar, 1935b: 84 (Potosia) N: AG MO TU
morio morio Fabricius, 1781: 51 (Cetonia) E: AN FR IT PT SP SZ
albopunctata Mulsant, 1842: 566 (Cetonia)
codinai Báguena, 1955b: 295 (Netocia)
fuliginosa Scopoli, 1786: 51 (Scarabaeus)
funesta Fabricius, 1781: 497 (Cetonia)
octopunctata Fabricius, 1787a: 27 (Cetonia)

379

380

SCARABAEIDAE

quadripunctata Fabricius, 1781: 52 (Cetonia)

oblonga oblonga Gory & Percheron, 1833: 227 (Cetonia) E: AN FR IT SP
luctifera Mulsant, 1842: 568 (Cetonia)

oblonga raffrayi Desbrochers des Loges, 1869: 46 (Cetonia) E: SP
petrovi Gusakov, 2007: 3 (Netocia) A: AF TD
proctotricha Fischer von Waldheim, 1842a: 13 (Cetonia) A: KI KZ UZ
asteria Reitter, 1899a: 89 (Netocia)
fasciata Fischer von Waldheim, 1844: 52 (Cetonia)
gagarinei Champenois, 1907: 207 (Cetonia)
nadari Champenois, 1907: 206 (Cetonia)
turcomanica Reitter, 1891d: 68 (Melanosa)

renei Rataj, 1986: 13 (Potosia) A: UZ
sardea Gory & Percheron, 1833: 216 (Cetonia) E: FR (Corse) IT (Sardegna, Sicilia)
tancae Krausse, 1913: 64 (Potosia)

schelkovnikovi Zaitzev, 1918b: 127 (Netocia) A: IN
squamosa crassicollis Burmeister, 1842: 443 (Cetonia) E: IT
squamosa squamosa Lefebvre, 1827: 103 (Cetonia) E: IT (Sicilia)
tincta Germar, 1838: 20 (Cetonia)

subpilosa dorchini Legrand & Drumont, 2006: 7 (Netocia) A: IS LE
subpilosa iranica Montreuil & Legrand, 2010b: 22 (Netocia) A: IN
subpilosa subpilosa Desbrochers des Loges, 1869: 46 (Cetonia) E: BU GR A: JO LE SY TR
athalia Reiche, 1871: 86 (Cetonia)

trojana ephesia Tauzin, 1993: 149 (Netocia) A: TR
kohouseki Rataj, 1998: 135 (Potosia)

trojana galathaea Reitter, 1899a: 92 (Melanosa) A: IS JO LE SY
trojana godeti Gory & Percheron, 1833: 229 (Cetonia) E: AB AR GG ST A: IN IQ TR
albilatera Faldermann, 1835a: 291 (Cetonia)
circumdata Faldermann, 1835a: 292 (Cetonia)
kurdistana Nonfried, 1892b: 97 (Melanosa)
magica Harold, 1880a: 160 (Cetonia)
setosula Reitter, 1891d: 69 (Melanosa)

trojana maura Bedel, 1889: 89 (Melanosa) N: AG LB MO TU
trojana syriaca Reitter, 1891d: 69 (Melanosa) A: LE SY TR
trojana trojana Gory & Percheron, 1833: 221 (Cetonia) E: BU GR
turkestanica Kraatz, 1886a: 108 (Cetonia) A: TD TM UZ
vidua Gory & Percheron, 1833: 216 (Cetonia) E: AB AR BU GG GR MC RO TR N: EG A: IN IS JO LE SY TR
adspersa Waltl, 1838: 460 (Cetonia)
aegyptiaca Gory & Percheron, 1833: 214 (Cetonia)
exclamationis Burmeister, 1842: 452 (Cetonia)
ithae Reitter, 1891d: 66 (Melanosa)

SCARABAEIDAE
subcarinata Reitter, 1896g: 266 (Netocia)

subgenus Netocia, nomen dubium
excavata Gory & Percheron, 1833: 225 (Cetonia) E: GR (Pelopónnisos)
subgenus Niponoprotaetia Mikšić, 1966a: 16 type species Cetonia lenzi Harold, 1876
exasperata akitai Masumoto & K. Sakai, 1988a: 4 A: JA (Okinawa-shotō: Kume-jima)
exasperata erabuana Nomura, 1964a: 58 A: JA (Kuchinoerabu-jima)
exasperata exasperata Fairmaire, 1898b: 386 (Cetonia) A: JA (Liou-kiou: Ôshima)
exasperata iheyana K. Sakai, 1989: 18 A: JA (Iheya-shima)
exasperata nakana Nakane, 1956a: 61 A: JA (Tokara: Naka-no-shima)
exasperata nomurai K. Sakai, 1998: 11 A: JA (Ryukyus: Tokuno-shima)
exasperata satoi Nakane, 1956a: 61 A: JA (Okinawa-shotō)
exasperata suwanoseana Nomura, 1964b: 69 A: JA (Tokara: Suwanose-jima)
exasperata uenoi Nomura, 1960b: 75 A: JA (Okinoerabu-jima)
lenzi Harold, 1878b: 128 (Cetonia) A: JA
hondana Arrow, 1913c: 404
nitidiscutellata Niijima & Kinoshita, 1923: 234 (Potosia)

subgenus Oreopotosia S.I. Medvedev, 1964: 204 type species Cetonia thibetana Kraatz, 1889
adelpha Ruter, 1978: 129 A: “Tibet: Vriatong”
dayaoensis Ma, 1993a: 172 A: YUN
rufescens Ma, 1987: 79 A: XIZ
thibetana Kraatz, 1889a: 380 (Cetonia) A: GAN SCH SHX
subgenus Pachyprotaetia Mikšić, 1965d: 83 type species Cetonia mixta Weber, 1801
confusa Gory & Percheron, 1833: 266 (Cetonia) A: UP ORR
piperina Westwood, 1849b: 144

subgenus Philhelena J. Thomson, 1880b: 278 type species Cetonia afflicta Gory & Percheron, 1833
afflicta afflicta Gory & Percheron, 1833: 215 (Cetonia) E: GR TR N: EG A: CY IQ IS JO LE SY TR
atrocoerulea Waltl, 1838: 460 (Cetonia)
leucogramma Gory & Percheron, 1833: 218 (Cetonia)
libanii Gory & Percheron, 1833: 215 (Cetonia)
osmanlis Gory & Percheron, 1833: 220 (Cetonia)
titea Reitter, 1899a: 88 (Potosia)

afflicta servillii Burmeister, 1842: 447 (Cetonia) A: SI YE
ignisternum ignisternum Reitter, 1891d: 68 (Melanosa) [DA] E: AB A: IN IQ TM TR
lucina Reitter, 1899a: 90 (Potosia)
paulina Reitter, 1899a: 90 (Potosia)

ignisternum zagrosica Alexis & Delpont, 2001d: 69 (Netocia) [DA] A: IN
minima Krajčík, 2011b: 2 [DA] A: NP
shamil Olsoufieff, 1916: 177 (Netocia) [DA] E: ST (Gunib) A: TM
margiana Zaitzev, 1918b: 126 (Netocia)

ungarica armeniaca Ménétriés, 1832: 190 (Cetonia) [DA] E: AB AR GG ST A: IN SY TM TR

381

382

SCARABAEIDAE

cirsii Motschulsky, 1845a: 58 (Cetonia)

ungarica auliensis Reitter, 1901a: 76 (Potosia) [DA] E: AB TR A: IN KZ TM TR
anatolica S.I. Medvedev, 1947: 128 (Potosia)
ignithorax Reitter, 1891d: 68 (Melanosa)
tristicula Reitter, 1893b: 73 (Cetonia)

ungarica inderiensis Krynicki, 1832: 129 (Cetonia) [DA] E: ST A: KI KZ TM WS
melancholica Zoubkov, 1833: 325 (Cetonia)
purpurea Burmeister, 1842: 447 (Cetonia)
zubkovii Faldermann, 1836: 371 (Cetonia)

ungarica persica Kraatz, 1886a: 108 (Cetonia) [DA] A: IN IQ TM TR
ungarica pseudoviridana S.I. Medvedev, 1964: 280 (Potosia) [DA] A: IN TM
ungarica sibirica Gebler, 1830: 112 (Cetonia) [DA] A: ES HEI MG NMO WS
hylata Reitter, 1899a: 90 (Potosia)
mongolica Reitter, 1899a: 89 (Potosia)

ungarica ungarica Herbst, 1790: 220 (Cetonia) [DA] E: AU BH BU CR CT CZ HU IT MC MD RO SB SK SL ST TR
UK A: TR
quadriguttata Motschulsky, 1845a: 59 (Cetonia)
simplex Schilsky, 1888a: 186 (Cetonia)
viridis Fabricius, 1792b: 128 (Cetonia)

ungarica viridana Brullé, 1832: 184 (Cetonia) [DA] E: GR
subgenus Potosia Mulsant & Rey, 1871a: 669 [=1871c: 413] type species Cetonia floricola Herbst, 1790 (=Cetonia
metallica Herbst, 1782)
afghana Petrovitz, 1955: 286 (Potosia) A: AF
angustata angustata Germar, 1817: 213 (Cetonia) E: AL BH BU CR GR HU IT MC ME RO SK SL SZ TR
atrorubens I. Frivaldszky, 1835: 262 (Cetonia)
coerulescens Schilsky, 1888a: 186 (Cetonia)
diocletiana Reitter, 1891d: 63 (Potosia)
muelleri Reitter, 1897a: 47 (Potosia)
nasuta Germar, 1824: 131 (Cetonia)
nigra Duftschmid, 1805: 170 (Cetonia)
purpurascens Reitter, 1891d: 63 (Potosia)
vicina Schönherr, 1817: 120 (Cetonia)

angustata angustula Reitter, 1898: 82 (Potosia) A: SY TR
eibesensis Reitter, 1902b: 139 (Potosia)
orientalis Alexis & Delpont, 1996b: 532 (Potosia) [HN]

besucheti Alexis & Delpont, 1996b: 525 (Potosia) E: GR (Ródos, Sámos) A: TR (southwest)
caucasica Kolenati, 1846: 30 (Cetonia) E: AB AR A: IN TR
alboflagellata Olsoufieff, 1916: 176 (Potosia)
araxicola Reitter, 1891d: 62 (Potosia)
azurea König, 1901: 10 (Potosia)

SCARABAEIDAE

383

karsiana Olsoufieff, 1916: 176 (Potosia)
smaragdina Olsoufieff, 1916: 176 (Potosia)

cuprea alainilerestifi Montreuil & Legrand, 2010a: 9 A: IN
cuprea bourgini Ruter, 1967: 36 (Potosia) E: BE FR GE LU
cuprea brancoi Baraud, 1992: 810 (Potosia) E: PT SP N: CI
cuprea cuprea Fabricius, 1775: 48 (Cetonia) E: FR (Corse) IT SZ
florentina Herbst, 1790: 210 (Cetonia)
incerta A. Costa, 1852: 10 (Cetonia)
rubrocuprea Mulsant, 1842: 554 (Cetonia)
nigrifulgens Mancini, 1924: 46 (Potosia)
obscuriventris J. Müller, 1900: 22 (Potosia)

cuprea daurica Motschulsky, 1860e: 135 (Cetonia) A: ES FE HEB HEI
lederi Reitter, 1899a: 76 (Potosia)
purpureipes Matsumura, 1911: 119 (Cetonia)

cuprea hesperica Motschulsky, 1849: 115 (Cetonia) E: SP
cuprea ikonomovi Mikšić, 1958b: 192 (Potosia) A: CY
cuprea ignicollis Gory & Percheron, 1833: 192 (Cetonia) N: EG A: IQ IS LE SAi SI SY TR
camilla Reitter, 1899a: 79 (Potosia)
edda Reitter, 1899a: 78 (Potosia)
fidia Reitter, 1899a: 78 (Potosia)
nerine Reitter, 1899a: 78 (Potosia)
phoebe Reitter, 1899a: 78 (Potosia)
prasiniuscula Reitter, 1899a: 79
sibylla Reitter, 1899a: 79 (Potosia)
xyphia Reitter, 1899a: 78 (Potosia)

cuprea levantina Schatzmayr, 1938a: 22 (Potosia) E: GR (Aegean Islands)
cuprea mandli Balthasar, 1930c: 64 (Potosia) A: ES
cuprea mehrabii Montreuil & Legrand, 2008: 161 A: IN
cuprea metallica Herbst, 1782: 314 (Cetonia) E: AU BE BY CR CZ DE EN FI FR GB GE HU IT LA LT LU NL NR NT
PL RO SB SK SL ST SV SZ UK A: ES WS
excisa Fieber, 1831: 471 (Cetonia)
floricola Herbst, 1790: 218 (Cetonia)
ottomana C.É. Blanchard, 1842: 5 (Cetonia)
sexundulata L. Petagna, 1819: 34 (Cetonia)
sternohirta Seidlitz, 1888b: 154 (Potosia)

cuprea obscura Andersch, 1797: 161 (Cetonia) E: AL AU BH BU CR CZ GR HU IT KO MC ME RO SB SK SL SZ
aenea Nedelkow, 1905: 5 (Potosia)
fabriciana Reitter, 1899a: 81 (Potosia)
subcuprea Reitter, 1899a: 78 (Potosia)

cuprea olivacea Mulsant, 1842: 554 (Cetonia) E: FR

384

SCARABAEIDAE

cuprea Mulsant, 1842: 554 (Cetonia) [HN]
mulsanti Reitter, 1899a: 75 (Potosia)
rubrocuprea Mulsant, 1842: 554 (Potosia)

cuprea viridiaurata Fuente, 1897: 130 E: SP
cuprea volhyniensis Gory & Percheron, 1833: 198 (Cetonia) E: CT ST UK A: KZ
steppensis Motschulsky, 1861: 426 (Potosia)

cuprina Motschulsky, 1849: 117 (Cetonia) E: AB AR BU GR MD RO ST TR UK A: TR
bessarabica Panin, 1942: 186 (Potosia)
ino Reitter, 1899a: 75 (Potosia)
pseudacuminata Reitter, 1899a: 75 (Potosia)
pseudocuprea Reitter, 1899a: 78 (Potosia)
purpurina Reitter, 1899a: 75 (Potosia)
rhodensis Rataj, 1998: 97 (Potosia)
subalboguttata Schaufuss, 1882c: clxxxi (Cetonia)
transfuga Schaufuss, 1882c: clxxxi (Cetonia)

famelica Janson, 1878b: 539 (Cetonia) A: BEI CHQ ES FE HEB HEI HUB JIX LIA NC NMO SC SCH SHA SHN
YUN
aeneiventris Kraatz, 1879a: 238 (Cetonia)
amurensis Heyden, 1886d: 285 (Cetonia)
mimula Harold, 1880e: vi (Cetonia)
scheini Mikšić, 1959a: 29 (Potosia)

fausti Kraatz, 1891a: 75 (Cetonia) E: AR GG A: TR
fieberi boldyrevi Jakobson, 1909: 128 (Potosia) E: CT MD ST
fieberi borysthenica S.I. Medvedev, 1964: 240 (Potosia) E: UK
fieberi fieberi Kraatz, 1880a: 57 (Cetonia) E: AL AU BH CR CT CZ FR GE GR HU IT PL RO SB SK SL ST SZ UK
aenea Fieber, 1831: 469 (Cetonia)

flutschiana Montreuil & Legrand, 2008: 159 A: IN
funebris aethiessoides Reitter, 1898c: 343 E: AR A: SY TR
funebris funebris Gory & Percheron, 1833: 229 (Cetonia) E: AB AR GG A: AF IQ SY TM TR
exclamationis Faldermann, 1835a: 297 (Cetonia)

funebris funesta Ménétriés, 1836a: 150 (Cetonia) A: IN IS LE SY TR
amandana Reitter, 1899a: 74 (Potosia)
angustata Paliatsea, 1937: 58
indigoaenea Nonfried, 1890a: 18 (Potosia)
kindermanni Schoch, 1895: 95 (Melanosa)
notativentris Reitter, 1899a: 74 (Potosia)
tenebrionis Ménétriés, 1839: 31 (Cetonia)

funebris lambillionea Alexis & Delpont, 2000: 279 A: TR
hieroglyphica depressiuscula Reitter, 1891d: 62 (Potosia) A: IN TM
densesculpta Reitter, 1899a: 75 (Potosia)

hieroglyphica hieroglyphica Ménétriés, 1832: 189 (Cetonia) E: AB GG ST A: IN TM

SCARABAEIDAE
ganglbaueri Reitter, 1891d: 57 (Pachnotosia)
pulverulenta Motschulsky, 1845a: 59 (Cetonia)

hypocrita Ragusa, 1905: 291 (Potosia) E: IT (Sicilia) MA
cuprea Gory & Percheron, 1833: 192 (Cetonia) [HN]

impavida Janson, 1878b: 538 (Cetonia) A: AF HP KA NP PA XIZ
dohrni Harold, 1880e: vi (Cetonia)
klapperichi Schein, 1956a: 19

indica Mikšić, 1965c: 26 (Potosia) A: AF HP KA PA
intricata W.W. Saunders, 1852: 31 A: FUJ ZHE
jelineki Petrovitz, 1981: 318 A: TR
keithi Montreuil & Legrand, 2008: 160 A: IN
marginicollis Ballion, 1871a: 346 (Cetonia) A: AF KI KZ TD UZ XIN
adippe Reitter, 1899a: 77 (Potosia)
laeviuscula Kraatz, 1886a: 107 (Cetonia)
maculicollis Kraatz, 1886a: 107 (Cetonia)

mayeti Le Comte, 1906: 247 (Cetonia) E: MA N: LB
multifoveolata Reitter, 1899a: 76 (Potosia) A: GAN HUN SCH XIZ
degrevei Alexis & Delpont, 1998a: 98 (Potosia)

neglecta Hope, 1831: 24 (Cetonia) A: AF AP HP KA NP PA SD UP XIZ ORR
dalmani Gory & Percheron, 1833: 195 (Cetonia)
puncticollis Burmeister, 1842: 470 (Cetonia)

nitididorsis Fairmaire, 1889a: 31 (Cetonia) A: CHQ BEI FE GUI GUX HEI
esquiroli Pouillaude, 1913e: 426 (Cetonia)

opaca cretica Kraatz, 1880b: 104 (Potosia) E: GR (Kríti)
opaca opaca Fabricius, 1787a: 27 (Cetonia) E: FR IT MA PT SP N: AG CI LB MO TU
cardui Gyllenhal, 1817: 47 (Cetonia)
corvina Motschulsky, 1849: 117 (Cetonia)
nudiventris Germar, 1837: 578 (Cetonia)
paulianiana P. Antoine, 1989: 6

splendidula diademata Reitter, 1891d: 62 (Potosia) A: IN
splendidula kermanensis Montreuil & Legrand, 2008: 161 A: IN
splendidula splendidula Faldermann, 1835a: 296 (Cetonia) E: AB AR A: IQ SY TR
chrysis Reitter, 1899a: 79 (Potosia)
erivana Reitter, 1899a: 77 (Potosia)
persplendens Reitter, 1891d: 62 (Potosia)

yunnana Mikšić, 1965c: 28 (Potosia) A: YUN
atomaria Fairmaire, 1887b: 116 (Cetonia) [HN]

subgenus Potosia, nomina dubia
aenea Andersch, 1797: 157 (Cetonia) E: SV
albiguttata Andersch, 1797: 158 (Cetonia) E: GB
algerica Motschulsky, 1849: 117 (Cetonia) N: AG

385

386

SCARABAEIDAE

difficilis Mulsant, 1842: 557 (Cetonia) E: FR
metallica Paykull, 1799: 203 (Cetonia) [HN] E: SV
nobilissima Reitter, 1891d: 62 (Potosia) N: MO
subgenus Protaetia Burmeister, 1842: 472 type species Cetonia mandarina Weber, 1801 (=Cetonia fusca Herbst,
1790)
Heteroprotaetia Mikšić, 1963: 355 type species Cetonia fusca Herbst, 1790

cariana cariana Gestro, 1891: 850 (Cetonia) [DA] A: AP NP SD YUN ORR
coenosa Westwood, 1849b: 146 (Anoplocheila) [DA] A: HP PA
brunneoaenea Westwood, 1849b: 146 (Anoplocheila)

fusca Herbst, 1790: 257 (Cetonia) A: FUJ GUA GUI HAI HKG HUB JA JIX SD SHG TAI ZHE AFR AUR ORR
atomaria Fabricius, 1801b: 153 (Cetonia)
bourgoini R. Paulian, 1960: 76
fictilis Newman, 1838a: 169 (Cetonia)
mandarina Weber, 1801: 68 (Cetonia)
taiwana Niijima & Kinoshita, 1923: 176

kurosawai Kobayashi, 1994: 149 [DA] A: TAI
rana Arrow, 1910a: 153 [DA] A: SD ORR
fucata Bourgoin, 1919b: 324 (Glycyphana)

subgenus Protaetiola Mikšić, 1963: 360 type species Cetonia multiguttulata Mohnike, 1873
bellula Ma, 1993a: 174 A: YUN
caudata Arrow, 1910a: 147 A: BT SD ORR
chalcites Bourgoin, 1917a: 69 (Cetonia)

subgenus Pseudocalopotosia Mikšić, 1966a: 6 type species Cetonia ishigakia Fairmaire, 1898
ishigakia ishigakia Fairmaire, 1898a: 14 (Cetonia) A: JA (Saki-shima: Ishigaki-jima)
ishigakia iriomoteana Nomura, 1964a: 59 A: JA (Saki-shima: Iriomote-shima)
ishigakia kikaiana Nakane, 1960: 6 A: JA (Amami: Kikaiga-shima)
ishigakia miyakona Kurosawa, 1959: 339 A: JA (Ryukyus: Miyako-jima)
ishigakia okinawana Kurosawa, 1959: 339 A: JA (Okinawa-shotō)
ishigakia yonakuniana Nomura, 1964: 59 A: JA (Yonaguni-jima)
subgenus Pseudonetocia S.I. Medvedev, 1964: 300 type species Cetonia cyanescens Kraatz, 1883
cyanescens cyanescens Kraatz, 1883e: 344 (Cetonia) A: KI KZ TD XIN
cupriflua Reitter, 1909b: 84 (Potosia)
imitatoria Reitter, 1909b: 84 (Potosia)
viridescens Heyden, 1892: 106 (Cetonia)

cyanescens jacobsoni Kiseritzky, 1910: 281 (Potosia) A: ES KZ
kulabensis Reitter, 1893e: 223 (Potosia) A: AF HP TD UZ
bucharica Curti, 1915: 22 (Potosia)

subgenus Pyropotosia Reitter, 1899a: 63 type species Cetonia pryeri Janson, 1888
pryeri akusekiana Nomura, 1964b: 68 A: JA (Tokara: Akuseki-jima)
pryeri esakii Nakane, 1956a: 113 A: JA (Kyūshū)
pryeri nitidicosta Yawata, 1941: 122 A: JA (Miyako-jima, Oghami-shima)

SCARABAEIDAE

387

pryeri oschimana Nonfried, 1895: 293 (Potosia) A: JA (Amami-Ôshima)
obenbergeri Balthasar, 1929a: 114 (Pyropotosia)

pryeri pryeri Janson, 1888b: 195 (Cetonia) A: JA (Okinawa-shotō)
okinavana Mikšić, 1971: 210

pryeri tsutsuii Nakane, 1956a: 112 A: JA (Tokara: Naka-no-shima)
subgenus Tomentoprotaetia Mikšić, 1987a: 444 type species Cetonia culta C.O. Waterhouse, 1879
bokonjici Mikšić, 1966b: 302 A: SCH YUN
culta culta C.O. Waterhouse, 1879: 247 (Cetonia) A: TAI
grisea Niijima & Kinoshita, 1923: 234 (Potosia)

culta multimaculata Kurosawa, 1959: 337 A: TAI (Botel Tobago)
culta yaeyamana Nomura, 1965: 9 A: JA (Yunaguni)
subgenus Vietnamoprotaetia Mikšić, 1971: 209 type species Glycyphana aeneipes Bourgoin, 1920
sericophora Seillière, 1910b: 327 (Glycyphana) A: GUI ORR
genus Protaetia, species incertae sedis
flavoguttata Kollar & L. Redtenbacher, 1844: 530 A: “Kashmir”
genus Rhabdotis Burmeister, 1842: 526 type species Cetonia sobrina Gory & Percheron, 1833
picta Fabricius, 1775: 51 (Cetonia) A: SA YE
albosculpta C.É. Blanchard, 1842: 3 (Cetonia)
arabica V. Allard, 1992: 55

genus Stalagmosoma Burmeister, 1842: 808 type species Scarabaeus albellus Pallas, 1781
Stalagmopygus Kraatz, 1882d: 66 type species Scarabaeus albellus Pallas, 1781

albellum Pallas, 1781: 17 (Scarabaeus) E: AB AR A: AE AF IN IQ IS JO KI KZ OM PA SI SY TD TM TR UZ
alternum Gory & Percheron, 1833: 211 (Cetonia)
korini Faldermann, 1835a: 302 (Cetonia)
lepidum Faldermann, 1836: 373 (Cetonia)

cynanki Gory & Percheron, 1833: 295 (Cetonia) N: AG EG LB A: AE AFR
genus Thyreogonia Reitter, 1899a: 46 type species Oxythyrea costata P.H. Lucas, 1858
costata P.H. Lucas, 1858: clxxviii (Oxythyrea) N: AG MO
albomaculata Thérond & Hollande, 1965: 873
fatima Fairmaire, 1860c: 446 (Tropinota)
niveopicta Fairmaire, 1864: 639 (Oxythyrea)

genus Tropinota Mulsant, 1842: 575 type species type species Tropinota reyi Mulsant, 1842 (=Scarabaeus
squalidus Scopoli, 1783)
subgenus Epicometis Burmeister, 1842: 434 type species Scarabaeus hirtellus Linnaeus, 1767 (=Scarabaeus
hirtus Poda von Neuhaus, 1761)
annabrunae Crovetti, 1973: 330 (Epicometis) N: LB
hirta crispa Petrovitz, 1971e: 589 (Epicometis) A: TR (Izmir: Menemen)

388

SCARABAEIDAE

hirta hirta Poda von Neuhaus, 1761: 21 (Scarabaeus) E: AL AN AU BE BH BU BY CR CT CZ EN FR GE GG GR
HU IT KO LA LS LT LU MC MD ME PL PT RO SB SK SL SP ST SZ TR UK N: MO A: KZ TR
hirtella Linnaeus, 1767: 556 (Scarabaeus)
nigrina Mulsant, 1842: 578 (Tropinota)
subfasciata Mulsant, 1842: 578 (Tropinota)
tonsa Burmeister, 1842: 437 (Epicometis)
vestita Say, 1825: 201 (Cetonia)

hirta suturalis Reitter, 1913d: 225 E: AB AL AR BU CR GG GR MC SB A: AF CY IN IQ IS JO LE SY TR
hirtiformis Reitter, 1913d: 227 E: GG A: IN JO KZ TM TR SY UZ WS
iec Ruiz, 2015: 3 N: MO
ilariae Dutto, 2007: 87 A: IS JO SA SY
paulae Leo, 2010: 2 E: IT (Sardegna)
senicula Ménétriés, 1832: 191 (Cetonia) E: AB AR GG A: IN TR
spinifrons jakesi Balthasar, 1967a: 116 (Epicometis) A: AF
spinifrons spinifrons Reitter, 1889b: 107 E: AB AR A: AF IN SY TD TM
turanica Reitter, 1889b: 107 A: AF IN KI TD TM UZ XIN
villiersi Baraud, 1984a: 61 A: IN
subgenus Hemiopta Bedel, 1896: 385 type species Epicometis bleusei Bedel, 1896
bleusei Bedel, 1896: 384 (Epicometis) N: AG
subgenus Tropinota Mulsant, 1842: 575 type species Tropinota reyi Mulsant, 1842 (=Scarabaeus squalidus
Scopoli, 1763)
squalida canariensis Lindberg, 1950: 12 N: CI MR
squalida pilosa Brullé, 1832: 186 (Cetonia) N: AG EG LB MO TU A: IN IQ IS JO KU LE SA SI SY TR
lethierryi Reiche, 1871: 85

squalida squalida Scopoli, 1763: 6 (Scarabaeus) E: AL BH BU CR FR GR IT MA MC ME PT SB SL SP SZ TR
crinita T. Charpentier, 1825: 213 (Cetonia)
heyrovskyi Obenberger, 1917a: 43 (Tropinota)
luctuosa Mulsant, 1842: 575
reyi Mulsant, 1842: 575
sabulicola Motschulsky, 1849: 121
spoliata A. Villa & G.B. Villa, 1833: 34 (Cetonia)
submaculata Mulsant, 1842: 575

vittula Reiche & Saulcy, 1856: 374 A: IS JO LE SY

subtribe Leucocelina Kraatz, 1882

genus Enoplotarsus P.H. Lucas, 1859e: xcviii type species Oxythyrea deserticola P.H. Lucas, 1857
deserticola P.H. Lucas, 1857a: lvi (Oxythyrea) N: AG MO
bedeli Bleuse, 1912a: 127

genus Homothyrea H.J. Kolbe, 1895a: 287 type species Oxythyrea helenae Schaum, 1848
helenae Schaum, 1848a: 71 (Oxythyrea) A: “Arabien” AFR

SCARABAEIDAE

389

collaris Harold, 1878c: 214 (Oxythyrea)

inornatipennis Gahan, 1903: 269 A: YE (Suqutra)
thoracica Schaum, 1841a: 47 (Oxythyrea) A: YE “Arabien” AFR
ruficollis C.É. Blanchard, 1842: 7 (Oxythyrea)

genus Oxythyrea Mulsant, 1842: 572 type species Scarabaeus sticticus Linnaeus, 1767 (=Scarabaeus funestus
Poda von Neuhaus, 1761)
Leucocelidia Olsoufieff, 1916: 169 type species Leucocelidia natalia Olsoufieff, 1916

abigail Reiche & Saulcy, 1856: 372 N: EG A: CY IS LE SI SY
albopicta Motschulsky, 1845a: 59 (Cetonia) E: AB AR GG MC ST A: IN KZ LE SY TD TR UZ
leucosticta Reiche, 1859b: ccix

cinctella Schaum, 1841a: 38 (Cetonia) E: AB AL AR BU GG GR MC ME SB ST TR UK (Krym) A: AF IN IQ IS JO
KI KZ LE PA SI ST SY TD TM TR UZ XIZ
albella Illiger, 1802a: 166 (Cetonia)
cinctelloides Reitter, 1898c: 343 (Leucocelis)
confluens Petrovitz, 1970c: 45
longula Desbrochers des Loges, 1872: 358 (Cetonia)
natalia Olsoufieff, 1916: 169 (Leucocelidia)
orientalis Iablokoff-Khnzorian, 1967: 206
taftanensis Montreuil & Legrand, 2008: 157
variegata Gory & Percheron, 1833: 294 (Cetonia) [HN]

dulcis Reitter, 1899a: 48 E: GR ME A: TR
abigailoides Mikšić, 1978: 70
borchmanni Schein, 1958: 93
goetzi Muche, 1962a: 33

funesta Poda von Neuhaus, 1761: 20 (Scarabaeus) E: AB AL AN AR AU BE BH BU BY CR CT CZ EN FR GE GG
GR HU IT LA LS LT LU KO MA MC MD ME NL NT PL PT RO SB SK SL SP ST SZ TR UK N: AG CI LB MO TU
A: CY IN KZ TR WS
albopunctata De Geer, 1774: 301 (Scarabaeus)
atlantis Escalera, 1914a: 189
consobrina A. Villa & G.B. Villa, 1833: 35 (Cetonia)
deleta Mulsant, 1842: 573
funeraria Goeffroy, 1785: 8 (Scarabaeus)
greeni Donovan, 1807: 418 (Scarabaeus)
nigrominuta Seabra, 1905: 19 (Leucocelis)
stictica Linnaeus, 1767: 552 (Scarabaeus)
viridana Seabra, 1905: 19 (Leucocelis)

gronbechi Petrovitz, 1955: 284 A: AF KA PA
muelleri Schein, 1956a: 15

noemi Reiche & Saulcy, 1856: 371 A: CY IS JO LE SI SY TR
pantherina Gory & Percheron, 1833: 293 (Cetonia) N: AG LB MO TU

390

SCARABAEIDAE

amina Fairmaire & Coquerel, 1860: 448

subcalva Marseul, 1878: 72 N: AG LB MO TU
biskrensis Fairmaire, 1884f: cxliii

tripolitana Reitter, 1891b: 25 (Leucocelis) N: LB TU
genus Paleira Reiche, 1871: 83 type species Cetonia femorata Illiger, 1803
femorata Illiger, 1803: 231 (Cetonia) E: PT SP N: CI MO
hispanica Gory & Percheron, 1833: 280 (Cetonia)

genus Pseudooxythyrea Baraud, 1985a: 587 type species Leucocelis bordesi Peyerimhoff, 1929
bordesi Peyerimhoff, 1929a: 123 (Leucocelis) N: AG LB MO

tribe Cremastocheilini Burmeister & Schaum, 1841
subtribe Coenochilina Burmeister, 1842

genus Coenochilus Schaum, 1841b: 268 type species Cetonia maura Fabricius, 1801
Anatochilus Péringuey, 1907: 492, 517 type species Coenochilus glabratus Boheman, 1857
Xenogenius H.J. Kolbe, 1892: 71 type species Coenochilus conradti H.J. Kolbe, 1892

arrowi Schein, 1954a: 322 A: UP
curtipes Westwood, 1873: 47 A: AP ORR
hervillardi Estienne, 2011: 61 A: PA
solidus Arrow, 1910a: 209 A: BT ORR
striatus Westwood, 1873: 46 A: FUJ GUA GUX HKG JA TAI ORR
uncinatipes Moser, 1915d: 596 A: FUJ

subtribe Cremastocheilina Burmeister & Schaum, 1841

genus Clinterocera Motschulsky, 1858b: 112 type species Cremastochila scabrosa Motschulsky, 1854
Callynomes Westwood, 1873: 26 type species Callynomes vollenhovii Westwood, 1873
Cholerastoma Mohnike, 1871: 315 type species Cholerostoma spondylidea Mohnike, 1871

cervenkai Krajčík, 2012b: 4 A: GUI
davidis davidis Fairmaire, 1878a: 107 (Callynomes) A: SCH YUN ORR
donckieri Bourgoin, 1924a: 138 (Callynomes)
humeralis Moser, 1902: 529

davidis nigra Kano, 1931: 131 (Callynomes) A: TAI
discipennis Fairmaire, 1889a: 32 A: ANH GUA HAI
ishikawai Kurosawa, 1973: 111 (Callynomes) A: JA (Iriomote-jima)
opaca Fairmaire, 1887b: 118 (Callynomes) A: YUN
pusilla Arrow, 1910a: 216 (Callynomes) A: SD ORR
rubra Ma, 1992b: 453 A: HUN
scabrosa Motschulsky, 1854c: 47 (Cremastochila) A: BEI FE GAN HEB NC SC SCH SHN
helleri Fairmaire, 1897f: 248 (Callynomes)
mandarina Westwood, 1873: 27 (Callynomes)

391

SCARABAEIDAE
obsoleta Fairmaire, 1878a: 108 (Callynomes)

trimaculata Ma, 1993c: 284 A: GUI ORR
tuberculata Krajčík, 2011c: 73 A: SCH
yunnana Moser, 1911b: 134 (Callynomes) A: YUN
genus Platysodes Westwood, 1873: 23 type species Platysodes verloreni Westwood, 1873
formosanus Kobayashi, 1990d: 78 A: TAI
madoni Bourgoin, 1923a: 220 A: GUA GUI GUX ORR

subtribe Cymophorina Krikken, 1984

genus Cymophorus Kirby, 1827: 153 type species Cymophorus undatus Kirby, 1827
Ptychophorus Schaum, 1841b: 271 [RN] type species Cymophorus undatus Kirby, 1827

subgenus Pseudocymophorus Schein, 1954b: 361 type species Ptychophorus hilaris Boheman, 1857
pulchellus Arrow, 1910a: 203 A: “S China” ORR
tonkinensis Schein, 1954b: 383

subtribe Goliathopsidina Krikken, 1984

genus Goliathopsis Janson, 1881: 609 type species Pilinurgus despectus Westwood, 1873
camptotropus C.-J. Yang, 1988: 8 A: GUI
esquiroli Pouillaude, 1913e: 421 A: GUI GUX
lameyi Fairmaire, 1893f: vii A: SE ORR
maolanus C.-J. Yang, 1988: 10 A: GUI
polystrictus C.-J. Yang, 1988: 8 A: GUI
velutinus Pouillaude, 1913e: 423 A: YUN ORR

subtribe Macromina Burmeister & Schaum, 1840

genus Campsiura Hope, 1831: 25 type species Campsiura xanthorhina Hope, 1831
subgenus Campsiura Hope, 1831: 25 type species Campsiura xanthorhina Hope, 1831
Estenomenus Faldermann, 1835b: 384 Estenomenus mirabilis Faldermann, 1835

melanopus Schaum, 1848b: 60 (Macroma) A: SD ORR
nigripennis Hope, 1841b: 65 [HN]

mirabilis Faldermann, 1835b: 385 (Estenomenus) A: CHQ GUI HEB HUB SCH SHA YUN
oberthueri Janson, 1888a: 112 (Macroma) A: BT SCH XIZ ORR
ochreipennis Fairmaire, 1887b: 117 (Macroma) A: GUI SCH YUN
omisiena Heller, 1923: 77 A: CHQ GUI GUX SCH
xanthorhina Hope, 1831: 25 A: NP SD XIZ ORR
bicolor Gory & Percheron, 1833: 149

subgenus Eucampsiura Mikšić, 1987b: 137 type species Macroma javanica Gory & Percheron, 1833
lijingkei Legrand & Flutsch, 2007: 96 A: HAI
nigripennis nigripennis Schaum, 1841b: 279 (Macroma) A: HKG SCH SD YUN ORR
genus Campsiura, species incertae sedis

392

SCARABAEIDAE

feistmanteli Nonfried, 1892a: 90 (Macroma) A: GUA

subtribe Pilinurgina Krikken, 1984

genus Parapilinurgus Arrow, 1910a: 204 type species Parapilinurgus variegatus Arrow, 1910
chinensis Krajčík, 2010a: 2 A: HUB
inexpectatus Krajčík, 2010a: 3 A: JIA
masumotoi Nomura, 1977a: 83 A: TAI

tribe Diplognathini Burmeister, 1842

genus Anthracophora Burmeister, 1842: 623 type species Anthracophora rusticola Burmeister, 1842
Poecilophilides Kraatz, 1898b: 406 type species Anthracophora rusticola Burmeister, 1842

crucifera A.G. Olivier, 1789: no 6: 39 (Cetonia) A: BT SD UP ORR
atromaculata Fabricius, 1792b: 141 (Cetonia)
ceylonensis Kraatz, 1895c: 110

dalmanni Hope, 1831: 24 (Cetonia) A: NP SD UP ORR
bohemanni Westwood, 1849b: 149

eddai eddai Tesař, 1959: 6 (Poecilophilides) A: TAI
eddai rongi Alexis & Delpont, 1997: 503 (Poecilophilides) A: GUA GUI SCH YUN
rusticola Burmeister, 1842: 624 A: CHQ FE GAN HEI HUB HUN JA JIX NC SC SCH SHG SHN YUN ZHE
rama Bainbridge, 1842: 217 (Diplognatha)
sinensis W.W. Saunders, 1852: 32 (Porphyronota)

tribe Goliathini Latreille, 1829
subtribe Coryphocerina Burmeister, 1842

genus Bietia Fairmaire, 1898b: 385 type species Bietia rudicollis Fairmaire, 1898
Atropinota Heller, 1923: 78 type species Atropinota funkei Heller, 1923

barclayi Krajčík, 2012b: 10 A: SCH
minima Krajčík, 2011c: 10 A: SCH
naxiorum Jákl, Král & Kubáň, 2010: 520 A: YUN
rudicollis Fairmaire, 1898b: 386 A: SCH
funkei Heller, 1923: 78 (Atropinota)

simillima Fairmaire, 1898b: 386 A: SCH
genus Chloresthia Fairmaire, 1905: 91 type species Chloresthia clypeata Fairmaire, 1905
clypeata Fairmaire, 1905: 91 A: SHN
genus Cosmiomorpha W.W. Saunders, 1852: 28 type species Cosmiomorpha modesta W.W. Saunders, 1852
subgenus Cosmiomorpha W.W. Saunders, 1852: 28 type species Cosmiomorpha modesta W.W. Saunders, 1852
cheni Qiu & Xu, 2013: 424 A: YUN
decliva Janson, 1890: 127 A: CHQ FUJ GAN GUA GUI GUX HEB HEN HUB HUN JIX SCH SHA SHG SHX YUN
ZHE ORR

SCARABAEIDAE

393

angulosa Fairmaire, 1898b: 385
baryi Bourgoin, 1916a: 109
squamulosa Schürhoff, 1933b: 101

fortis Qiu & Xu, 2013: 418 A: GUI GUX HUN
maolanensis Qiu & Xu, 2013: 422 A: GUI
modesta W.W. Saunders, 1852: 29 A: ANH BEI CHQ FUJ GUI GUX HEN HUB JIA JIX SHA SHN ZHE
nigripedis Qiu & Xu, 2013: 420 A: GUI
sauteri Bourgoin, 1931: 45 A: TAI
subgenus Microcosmiomorpha Mikšić, 1974c: 770 type species Cosmiomorpha setulosa Westwood,
1854
horni Bourgoin, 1931: 46 A: TAI
schneideri Jákl, 2014: 454 A: SCH XIZ
setulosa cribellata Fairmaire, 1893e: 314 A: GUI GUX YUN ORR
setulosa intermedia Mikšić, 1974: 236 A: FUJ HKG HUB JIX YUN ZHE
setulosa setulosa Westwood, 1854: 70 A: CHQ GAN GUX HEN JIA SCH SHA
similis myiakoana Nomura, 1964b: 70 A: JA (Myiako-jima)
similis nigra Niijima & Kinoshita, 1927: 69 A: JA
similis similis Fairmaire, 1900b: 621 A: TAI
similis yonakuniana Nomura, 1964a: 57 A: JA (Yonaguni-jima)
taiwanomontana Masumoto & K. Sakai, 1988c: 117 A: TAI
genus Dicheros Gory & Percheron, 1833: 300 type species Cetonia bicornis Latreille, 1816
subgenus Coryphe W.S. Macleay, 1838: 28 type species Cetonia macleayi Kirby, 1819
Coryphocera Burmeister, 1842: 220 [RN] type species Cetonia macleayi Kirby, 1819

bimacula gracilis Janson, 1909: 225 A: BT ORR
subgenus Hemicoryphocera Mikšić, 1974c: 753 type species Heterorhina childrenii Westwood, 1842
childrenii Westwood, 1842e: 139 (Heterorhina) A: “North India” ORR
nigrotestaceus Wallace, 1867: xciii (Heterorhina) A: CH ORR
genus Diphyllomorpha Hope, 1843c: 107 type species Diphyllomorpha mearesi Hope, 1843 (=Anomalocera
parryi Westwood, 1842)
Anomalocera Westwood, 1842c: 120 [HN] type species Anomalocera parryi Westwood, 1842
Meretricula Özdikmen, 2008: 251 [RN] type species Anomalocera parryi Westwood, 1842

glaberrima Westwood, 1842e: 136 (Anomalocera) A: HP KA NP PA SD UP XIZ
hirtiventris Kollar & L. Redtenbacher, 1844: 528 (Coryphocera)

olivacea insularis Moser, 1910a: 293 (Rhomborhina) A: TAI
olivacea olivacea Janson, 1883: 63 (Rhomborhina) A: FUJ GUA SCH
nigroolivacea S.I. Medvedev, 1964: 44 (Rhomborhina)

olivacea subopaca Arrow, 1907a: 348 (Anomalocera) A: NP SD ORR
pseudovitalisi Krajčík, 2014: 3 A: YUN
paotao Masumoto & K. Sakai, 1987a: 51 (Anomalocera) A: TAI

394

SCARABAEIDAE

parryi Westwood, 1842c: 120 (Anomalocera) A: NP SD ORR
mearesi Hope, 1843c: 107

genus Euchloropus Arrow, 1907a: 350 type species Cetonia laeta Fabricius, 1801
laetus Fabricius, 1801b: 150 (Cetonia) A: SD ORR
sylheticus J. Thomson, 1860: 30 (Heterorhina)

genus Gnathocera Kirby, 1825: 571 type species Gnathocera immaculata Kirby, 1825 (=Cetonia afzelii
Swartz, 1817)
Amphistoros Gory & Percheron, 1833: 34 type species Scarabaeus trivittatus Swederus, 1787
Gnathocerida Péringuey, 1907: 422 type species Gnathocera hirta Burmeister, 1842

trivittata aegyptiaca Kraatz, 1886c: 441 N: EG
genus Hemiheterorrhina Mikšić, 1974c: 762 type species Cetonia mutabilis Hope, 1831
dispar Arrow, 1907a: 347 (Heterorhina) A: NP SD
mutabilis Hope, 1831: 24 (Cetonia) A: BT HP KA NP UP ORR
affinis Kollar & L. Redtenbacher, 1844: 530 (Coryphocera)
bengalensis Hope, 1831: 24 (Cetonia)
dorsalis Gory & Percheron, 1833: 143 (Gnathocera)
hopei Gory & Percheron, 1833: 134 (Gnathocera)
melanaria Gory & Percheron, 1833: 143 (Gnathocera)
melanura Gory & Percheron, 1833: 153 (Gnathocera)

genus Herculaisia Seillière, 1910a: 239 type species Neophaedimus melaleucus Fairmaire, 1899
melaleuca Fairmaire, 1889: 103 (Neophaedimus) A: YUN ORR
genus Heterorhina Westwood, 1842e: 132 type species Cetonia nigritarsis Hope, 1831
subgenus Heterorhina Westwood, 1842e: 132 type species Cetonia nigritarsis Hope, 1831
chayuensis chayuensis Devecis, 2008: 105 A: XIZ
coxalis C.É. Blanchard, 1850: 26 (Coryphocera) A: NP UP ORR
flutschi yunnana Devecis, 2008: 104 A: YUN
jingkelii Devecis, 2008: 109 A: GUX
micans Guérin-Méneville, 1840a: 80 (Gnathocera) A: NP ORR
olivacea Guérin-Méneville, 1840a: 80 (Gnathocera)

nigritarsis magna Devecis, 2008: 100 A: NP
nigritarsis nigritarsis Hope, 1831: 24 (Cetonia) A: HP NP UP XIZ ORR
obesa obesa Janson, 1884: 104 A: HP NP SD XIZ ORR
punctatissima Westwood, 1842e: 135 A: AP BT NP SD YUN ORR
jucunda Hope, 1841b: 64 (Coryphe)
purpurascens Schoch, 1896b: 328 (Coryphocera)

tibialis Westwood, 1842e: 136 A: BT NP SD XIZ ORR
hookeri White, 1856: 14 (Trigonophorus)

SCARABAEIDAE

395

subgenus Heterorrhiniolla Mikšić, 1974c: 764 type species Cetonia amoena Hope, 1841
amoena Hope, 1841b: 64 (Cetonia) A: BT NP SD UP ORR
barmanica Gestro, 1888b: 99 A: XIZ ORR
porphyretica Westwood, 1849b: 144 A: HP
genus Ingrisma Fairmaire, 1893d: 292 type species Ingrisma rasuta Fairmaire, 1893
Cosmiorrhina Bourgoin, 1917e: 365 type species Cosmiorrhina bilobiceps Bourgoin, 1917

hainanica Krikken, 1984: 41 A: HAI
paralleliceps Bourgoin, 1914d: 446 A: TAI
insulana Moser, 1916b: 141

whiteheadi C.O. Waterhouse, 1900b: 310 A: HAI
genus Jumnos W.W. Saunders, 1839: 176 type species Jumnos ruckeri W.W. Saunders, 1839
roylei Westwood, 1839b: liv (Cetonia) A: HP KA
ruckeri ruckeri W.W. Saunders, 1839: 176 A: HP SD ORR
ruckeri nosei Nagai, 1992: 4 A: HAI
genus Mawenzhena Alexis & Delpont, 2001b: 26 type species Moseriana nitida Ma, 1990
nitida Ma, 1990a: 344 (Moseriana) A: GUX
genus Moseriana Ruter, 1965: 205 type species Cosmiomorpha bimaculata Moser, 1907
bimaculata Moser, 1907: 142 (Cosmiomorpha) A: YUN ORR
brevipilosa Ma, 1990a: 346 A: GUX
laevicostata Fairmaire, 1889a: 30 (Cetonia) A: GUI
longipilosa Ma, 1990a: 345 A: GUI GUX
mai Alexis & Delpont, 2001a: 19 A: YUN
genus Neophaedimus P.H. Lucas, 1870: lxxx type species Neophaedimus auzouxii P.H. Lucas, 1870
auzouxii P.H. Lucas, 1870: lxxxi A: CHQ GAN SCH SHA
maculicrus Fairmaire, 1897f: 248 (Rhomborhina)

castaneus Ma, 1989: 226 A: JIA ZHE
genus Periphanesthes Kraatz, 1880d: 213 type species Macroma aurora Motschulsky, 1858
Bonsiella Ruter, 1965: 206 type species Coryphocera blanda Jordan, 1895a (=Macroma aurora Motschulsky,
1858)

aurora Motschulsky, 1858d: 57 (Macroma) A: ANH FUJ GUA HAI HUN JIA JIX ZHE ORR
blanda Jordan, 1895a: 266 (Coryphocera)

genus Petrovitzia Mikšić, 1965e: 303 type species Cetonia guillotii Fairmaire, 1891
guillotii Fairmaire, 1891d: xii (Cetonia) A: CHQ GAN GUI GUX HUB SCH YUN
rugulosa Ma, 1990a: 344 (Moseriana)

pilosa Krajčík, 2012b: 9 A: SCH

396

SCARABAEIDAE

genus Pseudodiceros Mikšić, 1974c: 751 type species Diceros nigrocyaneus Bourgoin, 1917
nigrocyaneus Bourgoin, 1917c: 278 (Diceros) A: HUB SCH YUN ORR
genus Pseudotorynorrhina Mikšić, 1967: 309 type species Rhomborhina japonica Hope, 1841
Cosmiomorphomima Mikšić, 1976b: 6 type species Cosmiomorpha arrowi Paulian, 1960

fortunei fortunei W.W. Saunders, 1852: 30 (Rhomborhina) A: BEI FUJ GUI HAI HUB HUN JIA SHG ZHE
japonica Hope, 1841b: 64 (Rhomborhina) A: FUJ CHQ GAN HEI HEN HUB HUN JA JIX SC SCH SHG SHN
TAI ZHE
coreana Ruter, 1965: 196 (Rhomborhina)
clypeata Burmeister, 1842: 199 (Rhomborhina)
cupripes Nonfried, 1889: 533 (Rhomborhina)
glauca J. Thomson, 1878a: 9 (Rhomborhina)
ignita Nonfried, 1890b: 90 (Rhomborrhina)
kuytchuensis Ruter, 1965: 196 (Rhomborrhina)
nickerlii Nonfried, 1889: 533 (Rhomborhina)
nigra W.W. Saunders, 1852: 29 (Rhomborhina)
occidentalis Ruter, 1965: 196 (Rhomborhina)
reitteri Nonfried, 1890b: 90 (Rhomborhina)
squamuligera J. Thomson, 1878a: 9 (Rhomborhina)

tonkiniana Ruter, 1965: 165 (Rhomborhina) A: YUN ORR
genus Rhomborhina Hope, 1837: 120 type species Goliathus heros Gory & Percheron, 1833
subgenus Pseudorhomborrhina Mikšić, 1956: 216 type species Rhomborhina yunnana Moser, 1907
fuscipes Fairmaire, 1893e: 314 A: CHQ GUI HUB SCH SHA YUN ORR
yunnana Moser, 1907: 142 A: FUJ YUN ORR
subgenus Rhomborhina Hope, 1837: 120 type species Goliathus heros Gory & Percheron, 1833
aokii K. Sakai, 1993: 5 A: TAI
castanea K. Sakai, 1997: 57 A: YUN ORR
bousqueti Alexis & Delpont, 1998b: 177
vietnamica Alexis & Delpont, 1998b: 178

formosana Moser, 1909: 313 A: TAI
hamai Nomura, 1964a: 57 A: JA (Ishigaki-jima)
hiekei Ruter, 1965: 201 A: ZHE
jeanneli Ruter, 1965: 198 A: YUN ORR
aterrima Mikšić, 1967: 292

kurosawai Masumoto & K. Sakai, 1987b: 104 A: TAI
mellyi diffusa Fairmaire, 1897f: 247 A: GUI HUB SCH ORR
rufitibiis Bates, 1891b: 79
setchuenensis Ruter, 1965: 199
tonkinensis Ruter, 1965: 200

mellyi huangcaoshanensis J. Li, 2015c: 44 A: SCH
mellyi knirschi Schürhoff, 1933b: 101 A: YUN

SCARABAEIDAE

397

mellyi mellyi Gory& Percheron, 1833: 156 (Goliathus) A: NP SD XIZ ORR
dives Westwood, 1845a: 90

microcephala Westwood, 1842c: 119 A: PA SD ORR
mearesii Burmeister, 1842: 781 (Anomalocera)
staudingeri Nonfried, 1890a: 17

polita C.O. Waterhouse, 1875a: 113 A: JA
resplendens heros Gory & Percheron, 1833: 155 (Goliathus) A: NP SD ORR
resplendens resplendens Swartz, 1817: 51 (Cetonia) A: FUJ GUA HAI HKG JIX SCH ZHE
hainanensis Schürhoff, 1942: 284

splendida Moser, 1913b: 156 A: TAI
taiwana Sawada, 1949: 70 A: TAI
unicolor fukueana K. Sakai, 2000: 25 A: JA (Fukue-jima)
unicolor unicolor Motschulsky, 1862a: 8 A: JA
unicolor vernicata Fairmaire, 1897f: 246 A: CHQ GAN SCH SHA XIZ
continentalis Ruter, 1965: 200

violacea Schürhoff, 1942: 284 A: HUN JIX ZHE
genus Torynorrhina Arrow, 1907b: 433 type species Rhomborhina distincta Hope, 1841
apicalis Westwood, 1842c: 118 (Rhomborhina) A: NP SD ORR
distincta Hope, 1841b: 63 (Rhomborhina) A: AP BT NP SD YUN ORR
nigriventris Bourgoin, 1917a: 69 (Rhomborhina)
pilipes Burmeister, 1842: 779 (Rhomborhina)
ultramarinea Nonfried, 1906: 222 (Rhomborhina)

flammea flammea Gestro, 1888b: 115 (Rhomborhina) A: YUN ORR
cariana Gestro, 1891: 837 (Rhomborhina)

fulvopilosa Moser, 1911a: 120 (Rhomborhina) A: CHQ FUJ GUA GUI JIA SCH YUN ORR
hyacinthina Hope, 1841b: 62 (Rhomborhina) A: BT HP NP SD ORR
nigrina Kraatz, 1901: 415 (Rhomborhina)

opalina Hope, 1831: 24 (Cetonia) A: AP BT HP NP PA SD UP XIZ
pilifera Moser, 1914c: 573 (Rhomborhina) [RN] A: TAI
aurora Bourgoin, 1914d: 447 (Rhomborhina)
formosana Moser, 1909: 313 (Rhomborhina) [HN]

thiemei Moser, 1902: 526 (Rhomborhina) A: SE ORR
genus Trigonophorinus Pouillaude, 1913a: 43 type species Trigonophorinus leemei Pouillaude, 1913
(=Trigonophorus riaultii Fairmaire, 1897)
riaultii Fairmaire, 1897f: 245 (Trigonophorus) A: GUI YUN ORR
lemeei Pouillaude, 1913a: 45

genus Trigonophorus Hope, 1831: 24 type species Trigonophorus nepalensis Hope, 1831
subgenus Trigonophorus Hope, 1831 type species Trigonophorus nepalensis Hope, 1831
bonnardi Delpont, 2009: 61 A: GUX

398

SCARABAEIDAE

dechambrei Masumoto & K. Sakai, 1988b: 63 A: SCH
dilutus Bourgoin, 1914c: 438 A: TAI
gracilipes Westwood, 1845a: 88 A: BT NP SD ORR
hookeri White, 1856: 14 A: NP “Himalaya” ORR
parvus Kraatz, 1894g: 295

ligularis Ma, 1995: 226 A: GUI
nepalensis Hope, 1831: 24 (Cetonia) A: AP BT NP SD XIZ ORR
cantori Hope, 1841b: 62 (Rhomborhina)
hardwickei Hope, 1831: 24

pucholti Krajčík, 2007a: 6 A: SCH
rothschildii rothschildii Fairmaire, 1891f: ccvi A: FUJ GUI HUB SCH SHA XIZ ZHE
politus S.I. Medvedev, 1964: 49

rothschildii varians Bourgoin, 1914c: 437 A: TAI
murasaki Kometani, 1940b: 223

saundersii Westwood, 1842c: 122 A: NP SD ORR
scintillans Arrow, 1910a: 105 A: SD
yunnanus Schürhoff, 1935: 24 A: YUN
xisana Ma, 1985: 117 A: YUN
xizangensis Ma & Y.-W. Zhang, 1981: 390 A: XIZ

subtribe Dicronocephalina Krikken, 1984

genus Dicronocephalus Hope, 1831: 24 type species Dicronocephalus wallichii Hope, 1831
Acindrocephalus Keith & Delpont, 2004: 363 [RN] type species Dicranocephalus wallichii Hope, 1831
Dicranocephalus Hope, 1831: 24 [original spelling]
Dicranoceps S.I. Medvedev, 1972: 112 [RN] type species Dicranocephalus wallichii Hope, 1831

adamsii Pascoe, 1863: 25 (Dicranocephalus) A: CHQ GAN GUI HEI HUB JIL LIA NC QIN SC SCH SHA
XIZ ORR
drumonti Legrand, 2005b: 4

bieti Pouillaude, 1914d: 273 (Dicranocephalus) A: SCH YUN
bourgoini Pouillaude, 1914d: 293 (Dicranocephalus) A: TAI
bowringi Pascoe, 1863: 25 (Dicranocephalus) A: FE FUJ GUA GUI GUX HAI HKG HUN JIA JIX SCH SHN XIZ
YUN ZHE ORR
diminuata Young, 2012: 203

dabryi P.H. Lucas, 1872: 284 (Dicranocephalus) A: CHQ GAN HUB SCH SHA XIZ YUN
shimomurai Kurosawa, 1986: 31 (Dicranocephalus) A: TAI
uenoi katoi Kurosawa, 1968a: 226 (Dicranocephalus) A: TAI (central)
uenoi uenoi Kurosawa, 1968a: 226 (Dicranocephalus) A: TAI (southern)
wallichii Hope, 1831: 24 (Dicranocephalus) A: BT NP SD ORR
yui cheni Kurosawa, 1986: 33 (Dicranocephalus) A: TAI (southern)
yui yui Kurosawa, 1968a: 228 (Dicranocephalus) A: TAI (central)

399

SCARABAEIDAE
tribe Gymnetini Kirby, 1827
subtribe Gymnetina Kirby, 1827

genus Clinteria Burmeister, 1842: 299 type species Cetonia guttifera Burmeister, 1842 (=Gymnetis auronotata
C.É. Blanchard, 1842)
Tinclirea J. Thomson, 1880a: 268 type species Clinteria hilaris Burmeister, 1842 (=Cetonia klugi Hope, 1831)
Triclirea Schoch, 1895: 61 type species Cetonia pumila Swartz, 1819

aeneofusca Bourgoin, 1915c: 167 A: TAI
arunachala Chatterjee & Saha, 1981: 160 A: AP
confinis confinis Hope, 1831: 25 (Cetonia) A: BT ORR
confinis pseudoconfinis Schürhoff, 1942: 289 A: HP KA PA SD UP
sikkimensis Schürhoff, 1942: 289

ducalis davidis Fairmaire, 1887b: 116 A: FUJ HKG ORR
ducalis ducalis White, 1856: 15 A: SD ORR
oberthueri Arrow, 1910a: 180

hoffmeisteri White, 1847: 341 (Cetonia) A: NP ORR
klugi klugi Hope, 1831: 25 (Cetonia) A: NP PA ORR
duodecimguttata C.É. Blanchard, 1842: 16 (Gymnetis)
flavopicta C.É. Blanchard, 1850: 37
hilaris Burmeister, 1842: 303

nepalensis Krajčík, 2009b: 2 A: NP
spilota Hope, 1831: 25 (Cetonia) A: HP KA NP SD UP
tetraspilota Schoch, 1895: 77 (Triclirea)
viridipes Gory & Percheron, 1833: 364 (Gymnetis)

yunnana Krajčík, 2011a: 2 A: YUN
genus Clinteria, nomen dubium
spuria Burmeister, 1847: 555 A: KA

tribe Osmodermatini Schenkling, 1922

genus Osmoderma LePeletier & Audinet-Serville, 1828: 702 type species Scarabaeus eremita Scopoli, 1763
Gymnodus Kirby, 1837: 139 type species Scarabaeus eremita Scopoli, 1763

barnabita Motschulsky, 1845a: 58 E: AL AU BH BU BY CR CT CZ EN FI GE HU KO LA LT MC MD ME NT PL RO
SB SK SL ST UK
septentrionale Tauzin, 1994: 223

brevipenne Pic, 1904c: 74 A: TR
caeleste Gusakov, 2002: 39 (Gymnodus) A: FE
sikhotense Boucher, 2002: 426

cristinae Sparacio, 1994: 306 E: IT (Sicilia)
meridionale Tauzin, 1994: 228

davidis Fairmaire, 1887d: 321 A: BEI ES FE HEB HEI NC SCH
eremita Scopoli, 1763: 7 (Scarabaeus) E: AU BE DE FR GE IT LU NL NR SL SP SV SZ

400

SCARABAEIDAE

coriarium De Geer, 1774: 300 (Scarabaeus)
eremiticum Knoch, 1801: 107 (Cetonia)
sociale Horn, 1871b: 338

italicum Sparacio, 2001: 232 E: IT (Basilicata, Calabria, Campania)
lassallei Baraud & Tauzin, 1991: 159 E: GR TR
opicum coreanum Tauzin, 2013a: 175 A: FE NC
opicum opicum Lewis, 1887a: 49 A: JA
richteri S.I. Medvedev, 1954: 297 E: GG
genus Osmoderma, nomen dubium
dallieri Pic, 1945: 10 [distribution unknown]

tribe Phaedimini Schoch, 1894

genus Philistina W.S. Macleay, 1838: 25 type species “Macronata rhynophylla of Gory” (=Goliathus rhinophyllus Wiedemann, 1823)
Mycteristes Laporte, 1840: 162 type species Goliathus rhinophyllus Wiedemann, 1823

subgenus Cephalocosmus Kraatz, 1895b: 107 type species Cephalocosmus moewisii Kraatz, 1895 (=Mycteristes
microphyllus Wood-Mason, 1881)
Cosmocephalus Schoch, 1896: 363 type species Cephalocosmus moewisii Kraatz, 1895 (=Mycteristes microphyllus WoodMason, 1881)

benesi Drumont, 1998: 296 (Mycteristes) A: SCH
microphylla Wood-Mason, 1881: 411 (Mycteristes) A: BT SD YUN ORR
moewisii Kraatz, 1895b: 107 (Cephalocosmus)

tonkinensis Moser, 1904: 315 (Mycteristes) A: FUJ ORR
yunnana Krajčík, 2012b: 8 A: YUN
subgenus Hemiprigenia Mikšić, 1974b: 188 type species Prigenia khasiana Jordan, 1894
khasiana Jordan, 1894b: 691 (Prigenia) A: XIZ ORR
tibetana Janson, 1917: 5 (Mycteristes) A: “S. E. Tibet Vrianatang” ORR

tribe Schizorhinini Burmeister, 1842
subtribe Lomapterina Burmeister, 1842

genus Agestrata Eschscholtz, 1829: 13 type species Agestrata luconica Eschscholtz, 1829
Tetragonus Gory & Percheron, 1833: 302 type species Agestrata dehaani Gory & Percheron, 1833

orichalca orichalca Linnaeus, 1769: 504 (Scarabaeus) A: AP GUA GUX HAI HKG SD ORR
chinensis Fabricius, 1775: 42 (Cetonia)
gagates Hope, 1841a: 33
oblonga P. Brown, 1776: 122 (Scarabaeus)
samson Sharp, 1874a: 35
sinensis Gmelin, 1790: 1850 (Cetonia)
viridicyanea Schürhoff, 1934: 54
withilli Hope, 1841a: 33

401

SCARABAEIDAE
genus Thaumastopeus Kraatz, 1885b: 350 type species Lomaptera mohnikii J. Thomson, 1877
nigritus Frölich, 1792: 110 (Cetonia) A: BT HKG HP SD ORR
anthracinus Wiedemann, 1823: 83 (Macronota)
ebenus Burmeister, 1842: 315 (Lomaptera)
pullus Billberg, 1817: 46 (Cetonia)
simmillimus Schoch, 1898b: 157

shangaicus van de Pool, 1886: 181 (Lomaptera) A: HAI SHG YUN ORR
nigrovirens Bourgoin, 1916d: 298

tribe Taenioderini Mikšić, 1976a
subtribe Chalcotheina Mikšić, 1976a

genus Clerota Burmeister, 1842: 317 type species Macronota budda Gory & Percheron, 1833
bodhisattva Künckel d’Herculais, 1912: 403 A: YUN ORR
bourgoini Valck Lucassen, 1931: 163 A: SD
jeanvoinei Valck Lucassen, 1931: 165 A: “SE China” ORR
shiva Valck Lucassen, 1931: 158 A: BT SD ORR

subtribe Taenioderina Mikšić, 1976a

genus Bombodes Westwood, 1848a: 36 type species Bombodes ursus Westwood, 1848
dejeani Pouillaude, 1914e: 341 A: SCH
klapperichi Schein, 1953: 118 A: FUJ HUN JIX
latimaculata Ma, 1993b: 179 (Pleuronota) A: FUJ
mangshanensis Ma, 1992b: 438 (Pleuronota) A: HUN
nigellus Bourgoin, 1916c: 254 A: YUN ORR
tangi H. Huang & C.-C. Chen, 2015b: 3 A: XIZ
unimaculata Ma, 1993b: 180 (Pleuronota) A: YUN
ursus Westwood, 1848a: 36 A: GUX NP ORR
affinis Bourgoin, 1917b: 231

vitalisi Bourgoin, 1914b: 352 A: GUX HUN YUN ORR
vitalisiana R. Paulian, 1960: 30 (Coilodera) [RN]

westwoodii J. Thomson, 1857b: 284 A: BT SD XIZ
genus Coilodera Hope, 1831: 25 type species Coilodera penicillata Hope, 1831
mearesii Westwood, 1842b: 104 (Macronota) A: NP SD ORR
penicillata formosana Moser, 1910a: 297 A: TAI
hopponum Niijima & Kinoshita, 1923: 232 (Macronota)

penicillata nigroscutellaris Moser, 1902: 527 A: FUJ HAI ORR
penicillata penicillata Hope, 1831: 25 A: NP SD ORR
dives Gory & Percheron, 1833: 314 (Macronota)

402

SCARABAEIDAE

genus Euselates J. Thomson, 1880b: 277 type species Euselates magnus J. Thomson, 1880
subgenus Euselates J. Thomson, 1880b: 277 type species Euselates magnus J. Thomson, 1880
antennatus Wallace, 1867: xvi (Macronota) A: BT SD ORR
gracilis Arrow, 1907a: 350 (Macronota)

furcatus Kraatz, 1893a: 73 A: HAI
haeckelsehnali Jákl & Krajčík, 2008: 7 A: GUX
jansoni Arrow, 1910a: 64 (Macronota) A: SD ORR
kudrnaorum Jákl & Krajčík, 2008: 8 A: NP
laoticus Mikšić, 1974a: 80 A: YUN ORR
magnus J. Thomson, 1880b: 277 A: HAI HKG HUB ORR
schoenfeldti Kraatz, 1893a: 72
galei C.O. Waterhouse, 1900b: 312 (Taeniodera)

moupinensis Fairmaire, 1891d: xii (Taeniodera) A: FUJ GAN HEN HUB HUN JIX SCH SHA YUN ZHE
bipunctatus Schürhoff, 1935: 25 (Macronota)
donckieri Bourgoin, 1926: 71 (Macronota)
reitteri Schürhoff, 1934: 57 (Macronota)

ornatus W.W. Saunders, 1852: 31 (Taeniodera) A: FUJ GUX HAI SHG ZHE ORR
perraudierii taivanicus Mikšić, 1974a: 97 A: TAI
formosanus Niijima & Kinoshita, 1923: 230 (Macronota) [HN]

proximus Bourgoin, 1915b: 156 (Macronota) A: TAI
kagiensis Niijima & Kinoshita, 1923: 231 (Macronota)

quadrilineatus Hope, 1831: 24 (Macronota) A: AP BT NP SD XIZ ORR
aubaili Bourgoin, 1917d: 323 (Macronota)

tonkinensis formosanus Moser, 1910a: 298 A: TAI
anachoralis Kriesche, 1921d: 124
confluens Kriesche, 1921d: 124
fuscomaculatus Niijima & Kinoshita, 1923: 229 (Macronota)
procerus Bourgoin, 1915b: 160 (Macronota)

tonkinensis trivittatus Kriesche, 1921d: 123 A: FUJ GUA GUX
virgata Janson, 1892: 59 (Taeniodera) A: AP SD ORR
quadristrigata Kraatz, 1892c: 316 (Taeniodera)

whiteheadi C.O. Waterhouse, 1900b: 311 (Taeniodera) A: HAI
wittmeri Sabatinelli, 1984a: 159 A: NP
subgenus Pseudoeuselates Mikšić, 1974a: 61 type species Macronota setipes Westwood, 1854
setipes Westwood, 1854: 73 (Macronota) A: FUJ “China septentrionalis”
genus Euselates, nomen dubium
sanguinosus Motschulsky, 1854d: 64 (Macronota) A: SHG
genus Ixorida J. Thomson, 1880b: 277 type species Macronota mouhotii Wallace, 1867
mouhotii Wallace, 1867: xcvi (Macronota) A: YUN ORR

SCARABAEIDAE

403

genus Macronotops Krikken, 1977b: 200 type species Pleuronota sexmaculata Kraatz, 1894
curvimarginatus Ma, 1993b: 181 (Pleuronota) A: XIZ
fulvoguttatus Fairmaire, 1891d: xi (Macronota) A: HUB SCH YUN
fulvopilosus Fairmaire, 1894a: 219 (Taeniodera) A: SCH
hefengensis Ma, 1992b: 439 (Pleuronota) A: HUB
nigropubescens Mikšić, 1971: 208 (Pleuronota) A: TAI
sexmaculatus Kraatz, 1894a: 141 (Pleuronota) A: BT SD ORR
subsexmaculatus Ma, 1992b: 438 (Pleuronota) A: ZHE
vuilleti olivaceofuscus Bourgoin, 1916b: 136 (Macronota) A: FUJ HAI SCH ZHE
genus Meroloba J. Thomson, 1880b: 278 type species Macronota antiqua Gory & Percheron, 1833
vittidorsis Bourgoin, 1919a: 218 (Macronota) A: CH ORR
genus Stenonota Fairmaire, 1889a: 29 type species Stenonota semirugata Fairmaire, 1889
semirugata Fairmaire, 1889a: 30 A: GUI
genus Taeniodera Burmeister, 1842: 325 type species Macronota monacha Gory & Percheron, 1833
Ataenia Schoch, 1894: 203 type species Macronota biplagiata Gory & Percheron, 1833
Carolina J. Thomson, 1880b: 277 type species Macronota annae Wallace, 1867 (=Macronota malabariensis Gory &
Percheron, 1833)
Melinospila Kraatz, 1890: 277 type species Macronota flavomaculata Gory & Percheron, 1833
Polydomia J. Thomson, 1880b: 278 type species Macronota cervina Wallace, 1867

batillifera Bourgoin, 1914a: 292 (Macronota) A: BT ORR
coomani fencli Krajčík, 2007a: 5 A: HAI
crucicollis van Lansberge, 1887: 165 A: SD ORR
flavofasciata flavofasciata Moser, 1902: 527 (Carolina) A: FUJ HAI ORR
lurida Moser, 1902: 528 (Carolina)

flavofasciata formosana Moser, 1910a: 298 (Carolina) A: TAI
lata Bourgoin, 1915b: 158 (Macronota)
monaldaoi Kano, 1931: 132 (Macronota)
polischana Kriesche, 1921d: 123 (Carolina)

garnieri Bourgoin, 1917a: 68 (Macronota) A: HAI ORR
soror Bourgoin, 1924b: 153 (Macronota)

idolica Janson, 1909: 225 A: CH ORR
indica Janson, 1909: 226 A: SD ORR
luteovaria Bourgoin, 1917d: 324 (Macronota) A: YUN
malabariensis simillima Bourgoin, 1924b: 153 (Macronota) A: CH ORR
miksiciana Krajčík, 2010b: 4 A: HAI ORR
oberthueri van Lansberge, 1887: 167 A: SD ORR
humilis van Lansberge, 1887: 166

rutilans Ma, 1988: 272 A: XIZ YUN

404

SCARABAEIDAE

viridula Niijima & Kinoshita, 1923: 232 (Macronota) A: TAI
nigricolor Kano, 1931: 133 (Macronota)

zebraea Fairmaire, 1893d: 294 A: FUJ GUX ORR
bruyanti Bourgoin, 1916d: 297 (Macronota)

tribe Trichiini Fleming, 1821
subtribe Trichiina Fleming, 1821

genus Agnorimus Miyake & Iwase, 1991: 187 type species Gnorimus tibialis Chûjô, 1938 (=Gnorimus pictus
Moser, 1902)
pictus Moser, 1902: 531 (Gnorimus) A: FUJ HAI TAI YUN ORR
tibialis Chûjô, 1938: 444 (Gnorimus)
yunnanus Moser, 1908b: 257 (Gnorimus)

genus Epitrichius Tagawa, 1941: 18 type species Trichius elegans Kano, 1931
bowringii J. Thomson, 1857a: 118 (Trichius) A: BEI GUA GUX HAI HEB JIX SHG ZHE ORR
mandarinus L. Redtenbacher, 1868: 82 (Trichius)
miyashitai Krajčík, 2006: 25 (Trichius)

cupreipes Bourgoin, 1915e: 175 (Trichius) A: TAI
formosanus Niijima & Kinoshita, 1923: 189 (Gnorimus)
uraiensis Kano, 1931: 130 (Trichius)

elegans Kano, 1931: 127 (Trichius) A: TAI
taiheisanus Kano, 1931: 128 (Trichius)

lagopus Fairmaire, 1897d: 166 (Trichius) A: JA (Amami-Ôshima)
ferriei Pouillaude, 1913b: 125 (Trichius)

magnificus Krajčík & Chou, 2008: 4 A: TAI
shinshuingensis C.-L. Li & P.-S. Yang, 2008: 17

genus Gnorimotrichius Krajčík, 2009a: 4 type species Gnorimotrichius dolezali Krajčík, 2009
dolezali Krajčík, 2009a: 4 A: SCH
genus Gnorimus Le Peletier & Audinet-Serville, 1828: 702 type species Scarabaeus nobilis Linnaeus, 1758
Aleurostictus Stephens, 1839: 170 type species Scarabaeus nobilis Linnaeus, 1758

armeniacus Reitter, 1887d: 528 A: TR
baborensis Bedel, 1919: 248 N: AG
diversus Pic, 1925: 14
internenotatus Pic, 1925: 14

bartelsi Faldermann, 1835a: 289 E: GG ST A: TR
decempunctatus Helfer, 1833: 495 E: IT (Sicilia)
rufipennis Bedel, 1911a: 150
velutinus Ragusa, 1882a: 250

nobilis bolshakovi Gusakov, 2002: 7 (Aleurostictus) E: CT
nobilis macedonicus Baraud, 1992: 771 E: MC

SCARABAEIDAE

405

nobilis nobilis Linnaeus, 1758: 353 (Scarabaeus) E: AL AU BE BH BU BY CR CZ DE EN FR GB GE GR HU IT LA
LS MC ME NL NR LU MD PL PT RO SB SK SL SP SV SZ UK A: TR
auratus Schrank, 1781: 9 (Scarabaeus)
basipennis Pic, 1915a: 2
cupreicollis Mulsant, 1842: 533
cuspidatus Fabricius, 1787a: 27 (Scarabaeus)
cyaneoviolaceus Mancini, 1924: 46
immaculatus Mulsant, 1842: 533
rubrocupreus Mulsant, 1842: 533
subparallelus Pic, 1945: 11
superbissimus Roubal, 1933a: 128
tatari Ješátko, 1938: 40
thoracicus Hanuš, 1925: 96
viridulus De Geer, 1774: 297 (Scarabaeus)

signatus Chûjô, 1940b: 426 (Trichius) A: TAI
sumizome Kobayashi & C. K. Yu, 1993a: 78

subopacus Motschulsky, 1860e: 134 A: ES FE HEI JA JIL JIX LIA NC
subcostatus Ménétriés, 1832: 188 (Trichius) E: AB A: IN
cupreus Kraatz, 1895f: 415

variabilis Linnaeus, 1758: 352 (Scarabaeus) E: AU BE BU BY CR CT CZ DE EN FR GB GE GR HU IT LA LT MD
NL PL PT RO SB SK SL SP ST SV SZ UK A: TR
albopunctatus De Geer, 1774: 301 (Scarabaeus)
ambiguus Mulsant, 1842: 530
angularis Mulsant, 1842: 530
cordatus Fabricius, 1787a: 27 (Cetonia)
decempunctatus Schrank, 1798: 414 (Trichius)
flavopilosus Devecis, 1992: 2
heydeni Beckers, 1888: 24
hyperalbus Devecis, 1992: 2
juvencus Mulsant, 1842: 530
nigricollis Mulsant, 1842: 530
octopunctatus Fabricius, 1775: 44 (Cetonia)
seminiger Pic, 1915a: 2

viridiopacus Lewis, 1887a: 49 (Trichius) A: FE (Kunashir) HEB JA
genus Indognorimus Krikken, 2009a: 4 type species Gnorimus costipennis Janson, 1890
costipennis Janson, 1890: 128 (Gnorimus) A: XIZ ORR
viridis Jordan, 1894a: 486 (Gnorimus)

genus Indotrichius Krikken, 2009a: 7 type species Trichius ornatus Jordan, 1895b
gorodinskii Tauzin, 2013b: 190 (Paratrichius) A: XIZ
sikkimensis Krajčík, 2008b: 11 (Paratrichius) A: SD

406

SCARABAEIDAE

genus Lasiotrichius Reitter, 1899a: 101 type species Scarabaeus succinctus Pallas, 1781
sichuanicus Krajčík, 2001: 67 A: SCH
succinctus hananoi Sawada, 1943b: 6 (Trichius) A: CHQ GAN SCH SHA
succinctus shirozui Sawada, 1949: 72 (Trichius) [RN] A: TAI
formosanus Sawada, 1943b: 6 (Trichius) [HN]

succinctus succinctus Pallas, 1781: 18 (Scarabaeus) A: BEI ES FE HEB HEI JA JIL MG NC
amurensis Pic, 1923d: 6
dahuricus Gebler, 1823: 129 (Trichius)

succinctus tokushimus Krajčík, 2007b: 14 A: JA
turnai Krajčík, 2011c: 77 A: FUJ HUB ZHE
genus Paratrichius Janson, 1881: 610 type species Paratrichius longicornis Janson, 1881 (=Gnorimus doenitzi
Harold, 1879)
akane Myiake & Yamaya, 1999: 113 A: SCH
alboguttatus Moser, 1905a: 215 (Trichius) A: NP ORR
alexisi Krajčík, 2007b: 16 A: YUN ORR
castanus Ma, 1992c: 531 A: YUN
circularis Ma, 1990b: 457 A: SCH
diversicolor Bourgoin, 1915e: 177 (Trichius) A: TAI
doenitzi Harold, 1879c: 366 (Gnorimus) A: JA
longicornis Janson, 1881: 610
otsukae Niijima & Kinoshita, 1923: 190 (Gnorimus)

duplicatus duplicatus Lewis, 1895a: 406 A: JA (Amami)
osimanus Chûjô, 1941a: 35

duplicatus okinawanus Nomura, 1959b: 53 A: JA (Okinawa-shotō)
geibiensis Miyake, 1997: 7 A: JA
guttatus Iwase, 1993: 61 A: TAI
itoi Tagawa, 1938: 15 A: JA
kucerai Krajčík, 2007b: 17 A: SHA
kyushuensis Miyake, 1990: 28 A: JA (Kyūshū)
loi Kobayashi & Tanikado, 2003: 67 A: TAI
marmoreus Moser, 1902: 535 (Trichius) A: YUN ORR
nicoudi Bourgoin, 1920: 285 (Trichius) A: YUN ORR
nomurai Tesař, 1941a: 69 A: TAI
pilosonotus Yawata, 1943: 6
takasagonus Yawata, 1943: 8

oberthuri Moser, 1903: 286 (Trichius) A: SCH YUN
papilionaceus Ma, 1990b: 458 A: HAI
pauliani Tesař, 1941b: 130 A: SCH
riekoae Iwase, 1996: 82 A: GAN HEN SCH SHA
rotundatus Ma, 1990b: 459 A: YUN ORR
lianae Ricchiardi, 1993: 5

SCARABAEIDAE

407

rubrodecoratus Tesař, 1952: 59 (Trichius) A: FUJ SD
rufescens Ma, 1990b: 457 A: FUJ HAI
septemdecimguttatus Snellen van Vollenhoven, 1864: 159 (Trichius) A: HUB JA
taiwanus Iwase, 1993: 63 A: TAI
thibetanus Pouillaude, 1913b: 127 (Trichius) A: SCH
tigris Iwase, 1996: 79 A: HEN SHA
turnai Krajčík, 2007b: 18 A: HEN
versicolor Moser, 1902: 537 (Trichius) A: FUJ HAI YUN ORR
vitalisi Bourgoin, 1915a: 87 A: FUJ ORR
vittatus Sawada, 1939a: 44 A: TAI
zideki Krajčík, 2011: 4 A: SCH
genus Pseudagenius Heller, 1923: 78 type species Pseudagenius testaceipennis Heller, 1923
testaceipennis Heller, 1923: 79 A: SCH
viridicatus Ma, 1992c: 528 A: SCH YUN
genus Tibiotrichius Miyake, 1994a: 47 type species Tibiotrichius fujiokai Miyake, 1994
anoguttatus Fairmaire, 1891d: xiii (Gnorimus) A: HUB
dubernardi Pouillaude, 1913c: 157 (Trichius) A: YUN
flavitarsis Fairmaire, 1887b: 118 (Gnorimus) A: YUN
guerryi Moser, 1905a: 215 (Gnorimus)

klapperichi Tesař, 1942: 216 (Trichius) A: SCH
kuatunensis Tesař, 1952: 59 (Trichius) A: FUJ
miwai Chûjô, 1941b: 37 (Trichius) A: TAI
sinensis Pouillaude, 1913c: 160 (Trichius) A: SCH XIZ
genus Trichius Fabricius, 1775: 25 type species Scarabaeus fasciatus Linnaeus, 1758
Polytrichius Heer, 1834: 82 type species Scarabaeus fasciatus Linnaeus, 1758

abdominalis Ménétriés, 1832: 189 E: AB A: IN
heydeni G. Rossi, 1891: 44

albomaculatus Moser, 1903: 285 (Gnorimus) A: SCH
fasciatus Linnaeus, 1758: 352 (Scarabaeus) E: AB AL AU BE BH BU BY CR CT CZ DE EN FI FR GB HU IT KZ LA
LS LT LU MC NL NR NT PL PT RO SB SK SL SP ST SV SZ UK A: ES FE HEB JA KI KZ MG NO UZ WS XIN
abdominalis Mulsant, 1842: 540
abbreviatus Mulsant, 1842: 537
abruptus G. Rossi, 1888: 209
andersoni Bogdanov-Katjkov, 1914: 471
beckersi Geilenkeuser, 1892: 207
bimaculatus Gebler, 1830: 111
bipunctatus Kraatz, 1891d: 198
bisbinotatus Pic, 1923d: 6
borealis Schulze, 1910: 3

408
commutatus G. Rossi, 1882: 209
confluens G. Rossi, 1882: 209
divisus Mulsant, 1842: 537
dubius Mulsant, 1842: 536
erichsoni G. Rossi, 1882: 209
fabricii G. Rossi, 1882: 209
interruptus Mulsant, 1842: 537
kuenii Friese, 1896: 225
lineatocollis Kraatz, 1891d: 198
linnei G. Rossi, 1882: 209
noui Pellet, 1871a: 472 [=1873b: 13]
obliquus Mulsant, 1842: 537
prolongatus Mulsant, 1842: 537
pseudosibiricus Schulze, 1910: 8
pulchellus G. Rossi, 1882: 209
quadrimaculatus Kraatz, 1891d: 198
rangnowi Schulze, 1910: 2
reitteri Kraatz, 1891d: 198
sahlbergi Schulze, 1910: 5
scopolii Depoli, 1924: 107
scutellaris Kraatz, 1891d: 198
sibiricus Reitter, 1890e: 143
vulgaris G. Rossi, 1882: 209

gallicus gallicus Dejean, 1821: 61 E: BE CZ DE FR GB GE IT NL PL PT SK SL SP SZ UK
conjunctus Kraatz, 1891b: 195
connexus Kraatz, 1891b: 195
corsicus Kraatz, 1891b: 195
externenotatus Pic, 1923d: 6
fasciolatus Gené, 1836: 192
nudiventris Kraatz, 1891b: 194
suturalis Kraatz, 1891b: 195
vodozi Schneider, 1902: 48

gallicus zonatus Germar, 1831: 3 N: AG MO TU
edoughensis Pic, 1923d: 6
fortunatorum C.É. Blanchard, 1842: 21
multinotatus Pic, 1923d: 6
tangerianus Pic, 1947a: 6
theresae Pic, 1947a: 6
vaucheri Pic, 1947a: 6

japonicus Janson, 1885: 152 A: FE (Iturup, Kunashir) JA
orientalis Reitter, 1894a: 6 E: UK (Krym) A: SY TR

SCARABAEIDAE

409

SCARABAEIDAE
akbesianus Pic, 1923d: 6
brevefasciatus Pic, 1923d: 6
delagrangei Pic, 1947a: 7
macedonicus Pic, 1947a: 7
multidivisus Pic, 1947a: 7
notatipennis Pic, 1923d: 6

sexualis Bedel, 1906: 92 [NP] E: AL AU BE BH BU CR CZ FR GE GR HU IT KO MC ME PL PT RO SB SK SL SP
SZ UK
apicalis Mulsant, 1842: 540 [NO]
bipartitus Heyden, 1889b: 385 [NO]
bipunctatus Kraatz, 1891c: 196 [NO]
bivittatus Mulsant, 1842: 540 [NO]
bivittulatus Kraatz, 1891c: 196 [NO]
dentatus Mulsant, 1842: 540 [NO]
dorsalis Mulsant, 1842: 540 [NO]
intermedius Mulsant, 1842: 540 [NO]

trilineatus Ma, 1992c: 533 A: SCH YUN
tronqueti P. Antoine, 2002: 134 A: GUX
genus Trichius, nomen dubium
quadrilineatus Hope, 1831: 24 A: NP

tribe Valgini Mulsant, 1842

genus Charitovalgus H.J. Kolbe, 1904a: 21 type species Valgus pulcher Kraatz, 1883
banzai Sawada, 1941a: 8 A: TAI
formosanus Sawada, 1939a: 42 A: TAI
genus Chromovalgus H.J. Kolbe, 1897: 215 type species Valgus peyroni Mulsant & Wachanru, 1852
klapperichi Endrődi, 1956: 185 A: AF
peyroni Mulsant & Wachanru, 1852a: 7 [=1852b: 167] (Valgus) A: IN LE SY TR
genus Dasyvalgoides Endrődi, 1952b: 64 type species Dasyvalgoides denticulatus Endrődi, 1952
denticulatus Endrődi, 1952b: 65 A: FUJ ORR
genus Dasyvalgus H.J. Kolbe, 1904a: 34 type species Valgus vethi Ritsema, 1879
Anepsiovalgus H.J. Kolbe, 1904a: 30 type species Anepsiovalgus mimus H.J. Kolbe, 1904
Nannovalgus H.J. Kolbe, 1904a: 24, 26 type species Nannovalgus pusio H.J. Kolbe, 1904
Plocovalgus H.J. Kolbe, 1904a: 45 type species Plocovalgus waterstradti H.J. Kolbe, 1904
Syngonovalgus H.J. Kolbe, 1904a: 28 type species Syngonovalgus subnitidus H.J. Kolbe, 1904
Trichovalgus H.J. Kolbe, 1904a: 44 type species Valgus niger Kraatz, 1883

becvariensis Ricchiardi, 2015: 119 A: SCH YUN
benesi Ricchiardi, 2015: 120 A: GAN HUB SCH SHA

410

SCARABAEIDAE

carbonarius Arrow, 1910a: 239 A: HAI NP SD ORR
formosanus Moser, 1915d: 598 A: TAI
fulvicauda Arrow, 1910a: 242 A: BT ORR
grandis Pic, 1928e: 264 A: SD NP ORR
ichangicus Moser, 1915d: 599 A: HUB
inouei Sawada, 1939b: 88 A: FUJ GUX JA TAI
sebastiani Endrődi, 1952b: 69

jordansi Endrődi, 1952b: 65 A: FUJ
laligantii Fairmaire, 1889b: 343 (Valgus) A: FUJ GUX HKG ORR
lightbrowni Ricchiardi, 2015:124 A: YUN
makiharai Miyake, 1985: 11 A: TAI
minutus Ricchiardi, 2015:125 A: XIZ
montivagus Moser, 1915d: 601 A: GUX ORR
motuoensis Ricchiardi, 2015:125 A: XIZ
paramentatus Ricchiardi, 2015:126 A: YUN
penicillatus C.É. Blanchard, 1850: 45 (Valgus) A: BT HP NP
proximus Ricchiardi, 1994: 168 A: SD
pubicollis Pic, 1928b: 4 A: “Thibet”
rufipes Ricchiardi, 2015:128 A: GUX HAI ORR
rugosus Ricchiardi, 2013b: 136 A: YUN
sabatinellii Ricchiardi, 1994: 169 A: NP SD
sauteri Ricchiardi, 1998: 245 A: TAI
shimomurai Kobayashi, 1994: 147 A: TAI
sommershofi Endrődi, 1952b: 67 A: FUJ
submontanus Ricchiardi, 1994: 172 A: BT NP
taiwanus Miyake, 1991: 30 [RN] A: TAI
similis Miyake, 1985: 8 [HN]

tomentatus Ricchiardi, 2015:129 A: SCH YUN
tuberculatus Lewis, 1887b: 201 (Valgus) A: JA
varius Ricchiardi, 2015:130 A: SCH
wadai Miyake, 1985: 9 A: TAI
genus Excisivalgus Endrődi, 1952b: 62 type species Excisivalgus klapperichi Endrődi, 1952
csikii Endrődi, 1952b: 63 A: FUJ
klapperichi Endrődi, 1952b: 63 A: FUJ
genus Hybovalgus H.J. Kolbe, 1904a: 24 type species Hybovalgus bioculatus H.J. Kolbe, 1904
kamiyai Sawada, 1941a: 11 (Dasyvalgus) A: TAI
laticollis Arrow, 1944b: 238 A: SHN
major Sawada, 1939b: 84 A: JA TAI
matsudai Miyake, 1985: 6 A: TAI

SCARABAEIDAE
obsoletsquamosus Miyake, 1985: 6 A: TAI
takasagoensis Sawada, 1941a: 12 (Dasyvalgus) A: TAI
thibetanus Nonfried, 1892e: 372 (Valgus) A: GAN SCH SHA YUN XIZ
thoracicus Moser, 1911a: 128 A: TAI
tonkinensis Moser, 1905b: 272 A: FUJ ORR
versicolor Moser, 1908: 259 A: YUN
yunnanus Moser, 1906: 403 A: YUN
genus Indovalgus Ricchiardi, 2013a: 325 type species Valgus podicalis C.É. Blanchard, 1850
podicalis C.É. Blanchard, 1850: 44 (Valgus) A: BT NP UP ORR
propygidialis Moser, 1905b: 267 (Spilovalgus)

genus Neovalgus Miyake, 1985: 2 type species Valgus fumosus Lewis, 1887
assamensis Krajčík, 2011d: 2 A: AP
chinensis Krajčík, 2011d: 3 A: SHA
formosanus Miyake, 1985: 2 A: TAI
fumosus Lewis, 1887b: 201 (Valgus) A: JA
shikokuensis Sawada, 1941a: 7 (Valgus)

laetus laetus Arrow, 1913c: 408 (Chromovalgus) A: JA (Ryukyus: Ôshima)
laetus ootsuboi Kobayashi, 2011: 11 A: JA (Tokunoshima)
laterufus Pic, 1928d: 247 (Valgus) A: BT NP “Thibet”
pictus Hope, 1831: 24 (Acanthurus) A: NP ORR
taiwanus Sawada, 1941a: 2 (Valgus) A: TAI
genus Nipponovalgus Sawada, 1941a: 9 type species Valgus angusticollis C.O. Waterhouse, 1875
angusticollis abgusticollis Nakane, 1956a: 62 A: JA (Tokara-rettō)
angusticollis angusticollis C.O. Waterhouse, 1875a: 115 (Valgus) A: JA
yonakuniensis Sawada, 1941a: 10 A: JA (Yonaguni-jima)
genus Oreoderus Burmeister, 1842: 726 type species Valgus argillaceus Hope, 1842
arrowi Ricchiardi, 2001: 521 A: YUN
bhutanus Arrow, 1910a: 225 A: BT SD
bidentatus Ricchiardi, 2001: 526 A: YUN ORR
coomani Paulian, 1961: 31 A: YUN ORR
crassipes Arrow, 1944b: 243 A: UP
maculipennis Gestro, 1891: 869 A: FUJ YUN ORR
sikkimensis Ricchiardi, 2001: 552 A: SD
genus Valgus Scriba, 1790: 66 type species Scarabaeus hemipterus Linnaeus, 1758
Acanthurus Kirby, 1827: 155 type species Scarabaeus hemipterus Linnaeus, 1758
Homovalgus H.J. Kolbe, 1897: 214 type species Trichius seticollis Palisot de Beauvois, 1805

411

412

SCARABAEIDAE

hemipterus hemipterus Linnaeus, 1758: 351 (Scarabaeus) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR
GB GE GG GR HU IT KO KZ LA LT LU MC MD ME NL NR PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO
TU A: ES FE IN KI KZ TD TR UZ NARi
rubi Báguena, 1955: 293
squamulatus O.F. Müller, 1776: 55 (Scarabaeus)
variegatus Scopoli, 1763: 12 (Scarabaeus)

hemipterus meridionalis Rössner, 2014: 204 A: CY IN IS LE SY TR
heydeni Semenov, 1891: 331 A: SCH
koreanus Sawada, 1944: 12 A: FE SC
okajimai Kobayashi, 1994: 145 A: TAI
parvicollis Fairmaire, 1891f: ccvi A: HUB
savioi Pic, 1928c: 7 A: CH
genus Yanovalgus Nomura, 1952b: 29 type species Yanovalgus planiusculus Nomura, 1952
planiusculus Nomura, 1952b: 30 A: JA (Yaku-shima)
family Scarabaeidae, nomen dubium
Scarabaeus pellucidus Roemer, 1789: 2 E: SZ

series SCIRTIFORMIA Fleming, 1821
superfamily SCIRTOIDEA Fleming, 1821

family SCIRTIDAE Fleming, 1821
subfamily Nipponocyphoninae Lawrence & Yoshitomi, 2007

genus Nipponocyphon Lawrence & Yoshitomi, 2007: 509 type species Nipponocyphon nakanei Lawrence &
Yoshitomi, 2007
nakanei Lawrence & Yoshitomi, 2007: 514 A: JA

subfamily Scirtinae Fleming, 1821

genus Calvarium Pic, 1918: 14 type species Calvarium maxi Pic, 1918
hashimotorum Yoshitomi, 1998: 155 (Cyphon) A: JA (Ryukyus)

genus Contacyphon Gozis, 1886: 23 type species Cantharis variabilis Thunberg, 1785
abeillei Bourgeois, 1884b: xlii (Cyphon) E: FR A: IS TR
albanicus Klausnitzer, 1990f: 401 (Cyphon) E: AL
algenib Klausnitzer, 2003a: 94 [RN] (Cyphon) A: IN
simulans Klausnitzer, 1990d: 160 (Cyphon) [HN]

andreaspuetzi Klausnitzer, 2012d: 244 (Cyphon) A: SCH
arietizans Nyholm, 1970: 30 (Cyphon) E: AR GG A: TR
atripennis Pic, 1918a: 5 (Cyphon) A: TAI
australis Klausnitzer, 1991: 163 (Cyphon) N: AG TU

SCIRTIDAE

413

baoshanensis Yoshitomi, 2009b: 123 (Cyphon) A: YUN
brevirostris Yoshitomi, 2009b: 114 (Cyphon) A: NP
buceros Nyholm, 1949: 2 (Cyphon) E: AB AR GG ST A: TR
caspicus Klausnitzer, 1975a: 66 (Cyphon) E: AB
coarctatus Paykull, 1799: 120 (Cyphon) E: AU BH BY CR CZ CT DE EN FI FR GB GE HU IR IT LA LS LT LU NL
NR NT PL RO SK ST SV SZ UK A: KZ TM WS
alpinus Bourgeois, 1884a: 36 (Cyphon) [DA]
angulosus Marsham, 1802: 228 (Crioceris) [DA]
barnevillei Tournier, 1868a: 71 (Cyphon) [DA]
cartusiensis Gozis, 1882: 197 (Cyphon) [DA]
concolor Marsham, 1802: 226 (Crioceris) [DA]
convexus Rey, 1891d: 102 (Cyphon) [DA]
grandis Tournier, 1868a: 62 (Cyphon)
griseus Fabricius, 1801a: 502 (Cyphon)
immunis Stephens, 1833a: 43 (Cyphon) [RN]
kuenckeli Mulsant & Rey, 1865: 73 (Cyphon) [DA]
meridianus Mulsant & Rey, 1865: 72 (Cyphon) [DA]
nigricans Marsham, 1802: 226 (Crioceris) [DA]
nitidulus C.G. Thomson, 1855: 319 (Cyphon)
pallidiventris C.G. Thomson, 1855: 320 (Cyphon)
paykullii Guérin-Méneville, 1843: 10 (Elodes)

consobrinus Nyholm, 1949: 3 (Cyphon) A: FE JA NC SC
coreanicus Klausnitzer, 1975b: 308 (Cyphon) A: FE NC
corsicus Nyholm, 1964: 54 (Cyphon) E: FR (Corse)
derelictum Peyerimhoff, 1931b: 60 (Cyphon) E: SP N: AG MO
baeticus Nyholm, 1964: 50 (Cyphon)

designandus Nyholm, 1957: 78 (Cyphon) E: AL GR MC
friesei Klausnitzer, 1969: 840 (Cyphon)
jelineki Klausnitzer, 1969: 841 (Cyphon)

difficilis Klausnitzer, 1976b: 257 (Cyphon) E: GR (Kríti)
dinolabis Nyholm, 1970: 40 (Cyphon) A: TR
echinatus Klausnitzer, 1982: 282 (Cyphon) A: FE (Kunashir)
euoplus Nyholm, 1970: 38 (Cyphon) E: ST A: TR
euryceros Nyholm, 1970: 34 (Cyphon) A: TR
fasciatus Pic, 1931: 4 [DA] (Cyphon) A: HP
formosanus Pic, 1918a: 5 (Cyphon) A: JA TAI
furcillatus Nyholm, 1948: 3 (Cyphon) E: BU GR
fuscomarginatus Nakane, 1963c: 139 (Cyphon) A: FE (Kunashir, Iturup) JA
aberratus Klausnitzer, 1982: 279 (Cyphon)

414

SCIRTIDAE

geiseri Klausnitzer, 2013a: 45 (Cyphon) A: YUN
gracilicornis Wollaston, 1864: 214 (Cyphon) N: CI
granulosus Sasagawa, 1985: 43 (Cyphon) A: JA
haplous Klausnitzer, 1976a: 216 (Cyphon) A: BT NP
helmutsteueri Klausnitzer, 2011b: 103 (Cyphon) A: YUN
hilaris Nyholm, 1944: 21 (Cyphon) E: BY DE FI FR GB GE IR LA LT NL NR PL SP SV
iberus Nyholm, 1976: 287 (Cyphon) E: FR PT SP
jaechi Yoshitomi, 2002: 41 (Cyphon) A: HKG
jaegeri Klausnitzer, 2005a: 69 (Cyphon) A: NP
kongsbergensis Munster, 1923: 292 (Cyphon) E: AU BY CT CZ DE EN FI GB GE IT LA LT NR NT PL RU ST SV SZ
UK A: ES FE MG WS NAR
laevipennis Tournier, 1868a: 64 (Cyphon) E: AB AL AR AU BU BY CT CZ DE EN FI FR GB GE GR HU IT KZ LA
LT LU MC MD ME NL NR NT PL RO SK SP ST SV SZ UK N: EG LB MO TU A: AF ES IN IQ IS KZ MG NC SA
TD TM TR
kaszabi Klausnitzer, 1967: 275 (Cyphon)
phragmiteticola Nyholm, 1955: 259 (Cyphon)

lepidulus Nyholm, 1968: 250 (Cyphon) E: AB GR A: CY IN IQ SY TR
lindbergi Nyholm, 1948: 6 (Cyphon) E: PT SP (Baleares) N: AG MO
lithophilus Hernando, Aguilera & Ribera, 2003: 275 (Cyphon) N: MO (Anti-Atlas)
litigiosus Klausnitzer, 1980e: 210 (Cyphon) A: SD NP
loebli Klausnitzer, 2006b: 442 (Cyphon) A: YUN
longispinus Nyholm, 1968: 250 (Cyphon) A: TR
longiusculus Nyholm, 1970: 26 (Cyphon) E: AB GG GR (Ródos) TR A: IN IS TR
macedonicus Nyholm, 1957: 77 (Cyphon) E: GR
graecus Klausnitzer, 1976b: 257 (Cyphon)

magicus Klausnitzer, 1973c: 233 (Cyphon) A: JA (Ryukyus) TAI
manaslu Klausnitzer, 2009e: 249 (Cyphon) A: NP
masahiroi Yoshitomi, 2009b: 116 (Cyphon) A: NP
mizoro Nakane, 1963c: 140 (Cyphon) A: JA
nepalensis Yoshitomi, 2009b: 112 (Cyphon) A: NP
nipponicus Yoshitomi, 2005: 161 (Cyphon) A: JA
ochraceus nyholmi Klausnitzer, 1991: 164 (Cyphon) N: AG TU
ochraceus ochraceus Stephens, 1830: 287 (Cyphon) E: AU BY CR CT CZ DE FI FR GB GE GR HU IR IT LA LT LU
NL NR NT PL SK SP ST SV SZ TR UK A: TR
pallidulus Boheman, 1850: 212 (Cyphon)
suturalis Tournier, 1868a: 75 (Cyphon)

osellai Klausnitzer, 1989: 112 (Cyphon) A: TR
ozensis Satô, 1982: 389 (Cyphon) A: JA
padi Linnaeus, 1758: 369 (Chrysomela) E: AL AU BU BY CT CZ DE EN FI FR GB GE HU IR IT LA LT LU MD ME
NL NR NT PL RO SB SK ST SV SZ UK YU N: AG A: ES FE IS JA JO KZ NC TR WS
discolor Panzer, 1805: 99: 8 (Cyphon)

SCIRTIDAE

415

graciosus Kolenati, 1846: 40 (Cyphon)
pusillus Guérin-Méneville, 1843a: 8 (Elodes) [DA]
simplex Schilsky, 1888a: 188 (Cyphon) [DA]

palmi Nyholm, 1948: 5 (Cyphon) E: FR IT N: MO
palustris C.G. Thomson, 1855: 320 (Cyphon) E: AR AU BU BY CR CT CZ DE FI FR GB GE GR HU IR IT LA
LS LT LU MC ME NL NR NT PL RO RU SK SL SP ST SV SZ UK N: AG A: CY ES FE IN IS KZ LE NC SY TD
TR WS
elongatus Tournier, 1868a: 55 (Cyphon)
fuscicornis C.G. Thomson, 1855: 320 (Cyphon)
intermedius Tournier, 1868a: 59 (Cyphon)
puncticollis Tournier, 1868a: 56 (Cyphon)

pandellei Bourgeois, 1884a: 34 (Cyphon) E: FR PT SP
pareuoplus Klausnitzer, 2012c: 366 (Cyphon) A: IN
perses Nyholm, 1976: 283 (Cyphon) A: IN
poggii Klausnitzer, 1990b: 111 (Cyphon) E: IT (Sardegna)
ponticus Nyholm, 1976: 285 (Cyphon) A: TR
pubescens Fabricius, 1792b: 47 (Cistela) E: AL AU BU BY CR CT CZ DE EN FI FR GB GE GR HU IT LA LT LU ME
NL NR NT PL RO SK ST SV SZ TR UK A: ES FE KZ TR WS NAR
carectorum Nyholm, 1945: 63 (Cyphon)

puncticeps hisamatsui Sasagawa, 1985: 45 (Cyphon) A: JA (Ryukyus)
puncticeps puncticeps Kiesenwetter, 1874: 245 (Cyphon) A: JA (Honshu, Kyushu)
puncticeps shikokensis Sasagawa, 1985: 45 (Cyphon) A: JA (Shikoku)
thunbergi Nakane, 1963c: 139 (Cyphon) A: JA

punctipennis Sharp, 1872a: 155 (Cyphon) E: AU BY CT CZ DE FI GB GE IR LA LT NR NT SV A: ES FE WS
putonii Brisout de Barneville, 1863: 83 (Cyphon) E: AU FR GE PT SP SZ AURi
depressus Mulsant & Rey, 1865: 89 (Cyphon)
macer Kiesenwetter, 1863: 720 (Cyphon)

putzi Yoshitomi, 2009a: 87 (Cyphon) A: YUN
quadrum Klausnitzer, 1980c: 125 (Cyphon) E: AB AR
reitteri Klausnitzer, 1976b: 256 (Cyphon) E: AB
rotundosternalis Klausnitzer, 2012b: 342 (Cyphon) A: NP
rotundulus Klausnitzer, 1980b: 211 (Cyphon) A: SD
ruficeps Tournier, 1868a: 54 (Cyphon) E: AU BU BY CT CZ FR GE GG GR HU IT LA LU MD NL PL RO SK SL ST
SZ A: TR
schawalleri Klausnitzer, 2006b: 445 (Cyphon) A: YUN
schmidti Klausnitzer, 2006b: 446 (Cyphon) A: NP
scurrula Nyholm, 1970: 39 (Cyphon) E: BU SB TR YU A: TR
siculus Tournier, 1868a: 74 (Cyphon) E: FR IT (Sardegna, Sicilia) SP N: AG MO
impressus Kiesenwetter, 1871a: 76 (Cyphon)

simplex Klausnitzer, 1981: 63 (Cyphon) A: IN
solarii Nyholm, 1957: 75 (Cyphon) E IT

416

SCIRTIDAE

sulcicollis Mulsant & Rey, 1865: 87 (Cyphon) E: FR SP N: MO TU
marocanus Pic, 1934a: 17 (Cyphon)

suspiciosus Klausnitzer, 1980e: 209 (Cyphon) A: SD
tabidus Kiesenwetter, 1860: 412 [DA] (Cyphon) E: GR
tertius Klausnitzer, 1976a: 218 (Cyphon) A: BT
tsushimanus Yoshitomi, 2005: 164 (Cyphon) A: JA (Tsushima)
unguiculatus Nyholm, 1949: 2 (Cyphon) E: IT
unidentatus Klausnitzer, 2006a: 116 (Cyphon) A: GUX
ussuricus Nyholm, 1948: 4 (Cyphon) A: FE JA
sakaii Yoshitomi, 2002: 228 (Cyphon)

variabilis Thunberg, 1787: 54 (Cantharis) E: AU BY CT CZ DE EN FI FR GB GE HU IR IT LA LS LT LU MD NL
NR NT PL RO SK SP ST SV SZ UK A: ES FE JA KZ MG UZ WS AURi NAR NTR
dubius Stephens, 1830: 288 (Cyphon)
fusciceps Kirby, 1837: 245 (Cyphon)
modestus J. LeConte, 1854b: 355 (Helodes)
nebulosus J. LeConte, 1854b: 355 (Helodes)
nigriceps Kiesenwetter, 1860: 413 (Cyphon)
nigricornis Schilsky, 1888: 188 (Cyphon)
ovalis Say, 1825: 161 (Cyphon)
piceus J. LeConte, 1854: 354 (Helodes)
punctatus J. LeConte, 1854: 354 (Helodes)
rufipectus Rey, 1891d: 101 (Cyphon)

weigeli Klausnitzer, 2005c: 300 (Cyphon) A: NP
wittmeri Nyholm, 1970: 32 (Cyphon) A: TR
wuorentausi Nyholm, 1949: 4 (Cyphon) A: FE
yakushimanus Yoshitomi, 2005: 166 (Cyphon) A: JA
genus Dermestocyphon Pic, 1918d: 18 type species Cyphon drianti Pic, 1918
subgenus Dermestocyphon Pic, 1918d: 18 type species Cyphon drianti Pic, 1918
drianti Pic, 1918d: 18 (Cyphon) A: YUN
subgenus Himacyphon Ruta, Yoshitomi & Klausnitzer, 2013: 278 type species Cyphon optatus Klausnitzer, 1980
optatus Klausnitzer, 1980e: 203 (Cyphon) A: NP UP ORR
subgenus Oreocyphon Klausnitzer, 2008a: 81 type species Cyphon honorus Klausnitzer, 1980
anticetestaceus Klausnitzer, 1976a: 219 (Prionocyphon) A: BT
honorus Klausnitzer, 1980e: 205 (Cyphon) A: NP ORR
umbratilis Klausnitzer, 1976a: 218 (Prionocyphon) A: BT NP
genus Elodes Latreille, 1797: 44 type species Cistela pallida Fabricius, 1775 [as Cyphon pallidus Fabricius]
(=Lampyris minuta Linnaeus, 1767)
Cyphon Paykull, 1799: 117 type species Cistela pallida Fabricius, 1775 (=Lampyris minuta Linnaeus, 1767)
Helodes Agassiz, 1846: 393, 506 [HN] [unjustified emendation]

SCIRTIDAE

417

abeillei Klausnitzer, 1990e: 238 (Helodes) E: FR N: AG
akbesianus Pic, 1898b: 26 (Helodes) A: LE TR
algirinus Pic, 1898b: 27 (Helodes) N: AG
amicula Klausnitzer, 1980d: 123 (Helodes) E: GR
angelinii Klausnitzer, 1987: 249 (Helodes) E: IT
arcanus Klausnitzer, 1972a: 69 (Helodes) E: CR ME
australis Klausnitzer, 1990f: 248 (Helodes) E: GR
bertiae Klausnitzer, 1988: 36 (Helodes) E: FR (Corse)
burmensis Klausnitzer, 1974: 19 (Helodes) A: AP NP SD ORR
calabriae Klausnitzer, 1987: 247 (Helodes) E: IT
caucasicus Klausnitzer, 1970a: 181 (Helodes) E: AB
chobauti Abeille de Perrin, 1894c: 92 (Helodes) N: AG
chrysocomes Abeille de Perrin, 1872: xlii (Helodes) [DA] E: FR (Alpes-de-Haute-Province)
cognatus Klausnitzer, 1980b: 85 (Flavohelodes) A: KA
corsicus Pic, 1898d: 137 (Helodes) E: FR (Corse)
creticus Klausnitzer, 1973a: 37 (Helodes) E: GR (Kríti)
denticulatus Klausnitzer, 1973a: 37 (Helodes) E: IT
dubius Klausnitzer, 1972a: 71 (Helodes) N: AG TU
eberti Klausnitzer, 1970a: 181 (Helodes) E: AB UK A: IN TR
elegans Yoshitomi, 1997: 372 A: JA
elongatus Tournier, 1868a: 34 (Helodes) E: AU BE CZ DE FR GB GE GR HU IR IT LU NL PL RO SK SP SZ UK
koelleri Klausnitzer, 1970a: 180 (Helodes)
subterraneus Mulsant & Rey, 1876: 189 (Helodes)

flavoangularis Klausnitzer, 1990a: 22 (Helodes) A: TR
genei Guérin-Méneville, 1843a: 12 E: FR (Corse) IT (Sardegna, Sicilia)
gerdmuelleri Klausnitzer, 2009b: 723 A: KZ
globulus Klausnitzer, 1990e: 240 (Helodes) E: FR
holgeri Klausnitzer, 2003b: 199 E: GR
improvisus Klausnitzer, 1990c: 125 (Helodes) A: TR
indubius Klausnitzer, 1980e: 196 (Flavohelodes) A: SD
jelineki Klausnitzer, 2009b: 715 A: UZ
johni Klausnitzer, 1975c: 329 (Helodes) E: BE CZ GE HU RO SK
joosti Klausnitzer, 2004c: 90 A: SCH
kiesenwetteri Tournier, 1868a: 34 (Helodes) [DA] E: GE SL
kojimai Nakane, 1963c: 140 A: FE (Kunashir) JA
nigratus Klausnitzer, 1982: 275 (Flavohelodes)
ohbayashii Satô, 1985: 55 (Helodes)

lohsei Klausnitzer, 2000: 24 E: ST
longiusculus Klausnitzer, 1980e: 197 (Flavohelodes) A: SD
longulus Klausnitzer, 1990e: 247 (Helodes) E: BU
luteatus Klausnitzer, 1979b: 335 (Helodes) E: TR

418

SCIRTIDAE

major Pic, 1954a: 63 [DA] A: FUJ
malickyi Klausnitzer, 1976c: 699 (Helodes) A: CY
megalopus Klausnitzer, 2002a: 129 A: TR
miaoershanensis Yoshitomi & Satô, 1997: 75 [RN] A: GUX
bicolor Yoshitomi & Satô, 1996a: 245 [HN]

microps Klausnitzer, 2002a: 130 A: TR
minutus Linnaeus, 1767: 645 (Lampyris) E: AU BE BU BY CZ DE EN FI FR GB GE HU IT LA LS LT LU NL NR NT
PL RU SK SP SV SZ TR A: TR
caudatus Scopoli, 1763: 40 (Cantharis)
discicollis Pic, 1941: 1 (Helodes) [DA]
laetus Panzer, 1793c: 8: 8 (Cistela)
melanurus Fabricius, 1792b: 22 (Galleruca)
nebrodensis Ragusa, 1884: 335 (Helodes) [DA]
nigripennis Tournier, 1868a: 35 (Helodes)
pallidus Fabricius, 1775: 117 (Cistela)
testaceus Schilsky, 1888a: 188 (Helodes)

nocturnus Klausnitzer, 1979a: 17 (Helodes) E: GR
novaki J. Müller, 1916: 90 (Helodes) E: CR
novacreticus Klausnitzer, 1990e: 250 (Helodes) E: GR (Kríti)
orientalis Iablokoff-Khnzorian, 1973: 112 A: IN TD
pechlaneri Klausnitzer, 2003c: 279 A: YUN
pendens Klausnitzer, 1990c: 246 (Helodes) E: BH CR
persicus Klausnitzer, 1975a: 331 A: IN
persimilis Klausnitzer, 2002a: 125 A: TR
pollux Klausnitzer, 2008c: 221 E: FR (Corse)
pseudominutus Klausnitzer, 1971: 6 (Helodes) E: AU BE BY CT CZ DE GB GE HU IT LA LU NL NT PL RO SP ST
SZ A: TR
pseudoscutellaris Klausnitzer, 1973a: 36 (Helodes) E: AR GG
rufotestaceus Klausnitzer, 1976a: 215 (Helodes) A: BT SD ORR
scutellaris Tournier, 1868a: 33 (Helodes) E: SP
secundocreticus Klausnitzer, 1980d: 125 (Helodes) E: GR (Kríti)
sericeus Kiesenwetter, 1859a: 22 (Helodes) E: GR
sieberi Klausnitzer, 1973b: 113 (Helodes) E: GR (Aegean Islands) TR A: TR
sororum Pic, 1918d: 17 [DA] A: SD
spinidens Klausnitzer, 2001: 123 N: MO
takahashii Yoshitomi, 2005: 32 A: JA (Ryukyus)
tegminis Klausnitzer, 2012a: 191 A: PA
tournieri Kiesenwetter, 1871b: 88 E: FR (Corse) IT (Sardegna) N: TU A: TR
africanus Klausnitzer, 1972a: 70 (Helodes)
pallidithorax Pic, 1898d: 137 (Helodes)

tricuspis Nyholm, 1985: 438 E: AU BE CT CZ DE FI FR GB GE HU NL NR NT PL SV SZ UK

SCIRTIDAE

419

unicolor Klausnitzer, 2012c: 368 A: IN
venetae Klausnitzer, 1987: 247 (Helodes) E: IT
venustulus Klausnitzer, 2002a: 122 A: TR
genus Exochomoscirtes Pic, 1916e: 6 type species Exochomoscirtes binotatus Pic, 1916
arunachalensis Ruta & Yoshitomi, 2010: 74 A: AP
bezdeki Ruta & Yoshitomi, 2010 A: AP ORR
herthae Klausnitzer, 2010d: 1365 A: AP ORR
holgerdathei Klausnitzer, 2010c: 285 A: AP
maculatus Waterhouse, 1880: 568 (Scirtes) A: NP ORR
nipponicus Ruta & Yoshitomi, 2010: 70 A: JA (Yaeyama)
genus Herthania Klausnitzer, 2006c: 71 type species Cyphon japonicola Nakane, 1963
certa Klausnitzer, 2006c: 72 A: YUN
japonicola Nakane, 1963c: 139 (Cyphon) A: JA
obscurata Klausnitzer, 1982: 278 (Cyphon) A: FE (Kunashir; Sachalin) JA YUN
paludosa Sasagawa, 1985: 39 (Cyphon)

satoi Sasagawa, 1985: 38 (Cyphon) A: JA
intermedia Nakane, 1963b: 139 (Cyphon) [HN]
ishiharai Sasagawa, 1985: 40 (Cyphon)
sasagawai Yoshitomi & Klausnitzer, 2003: 93 (Cyphon) [RN]

genus Hydrocyphon L. Redtenbacher, 1858: 519 type species Cyphon deflexicollis P.W.J. Müller, 1821
alticola Klausnitzer, 1976a: 217 (Cyphon) A: BT SD
amaurus Klausnitzer, 1980e: 213 (Cyphon) A: AP NP SD
aritai Yoshitomi, 2001: 103 A: TAI
australis Linder, 1864: xiv E: BH FR IT PT SP TR
bhutanensis Klausnitzer, 1976a: 215 A: BT NP
bicornis Yoshitomi & Klausnitzer, 2003: 527 A: YUN
bifidus Yoshitomi & Satô, 2005a: 156 A: GUI
celatus Klausnitzer, 1980e: 199 A: HP NP
championi Reitter, 1903b: 177 E: PT SP
consolatorius Klausnitzer, 1990d: 159 A: IN
deflexicollis P.W.J. Müller, 1821: 223 (Cyphon) E: AU BE BH BU CR CZ FR GB GE GR HU IR IT LU MC NL PL RU
SK SL SZ TR A: TR
ater Stephens, 1830: 287 (Cyphon)
pini Curtis, 1836: 602 (Elodes)

dubius Klausnitzer, 1980e: 208 (Cyphon) A: SD NP
dudgeoni Yoshitomi & Klausnitzer, 2003: 522 A: HGK
finitimus Nyholm, 1977: 97 E: AR A: TR
forficulatus Nyholm, 1981: 261 A: NP ORR

420
fulvescens Nyholm, 1977: 104 E: SP
gereckei Hernando, Aguilera & Ribera, 2004: 389 N: MO
graseri Klausnitzer, 2006b: 448 A: NP
guangxiensis Yoshitomi & Klausnitzer, 2003: 521 A: GUX ORR
hainanensis Yoshitomi & Klausnitzer, 2003: 522 A: HAI
hamiota Nyholm, 1972: 36 E: SP
hydrocyphonoides Tournier, 1868a: 66 (Cyphon) E: IT N: AG TU
illiesi Klausnitzer, 1991: 162 N: AG
interrogationis Klausnitzer, 1980c: 201 A: PA
iriomotensis Yoshitomi, 2001: 98 A: JA (Ryukyus)
jaechi Yoshitomi & Klausnitzer, 2003: 526 A: JIA
komareki Yoshitomi, 2008b: 78 A: JIX
kopetdaghensis Ruta, 2007: 59 A: TM
laeticolor Nyholm, 1967: 45 E: SP
lii Yoshitomi & Klausnitzer, 2003: 524 A: ZHE
masatakai Yoshitomi, 2008b: 77 A: HUB
minous Nyholm, 1967: 28 E: GR (Kríti)
mirabilis Yoshitomi & Satô, 2005a: 159 A: GUI NP ORR
nakanei Yoshitomi, 2001: 101 A: JA
nepalensis Yoshitomi & Satô, 2005a: 166 A: NP
novaki Nyholm, 1967: 37 E: AL BH CR GR IT
fuscatus Klausnitzer, 1970b: 206

nuristanicus Klausnitzer, 2004b: 129 A: AF
nyholmi Yoshitomi & Satô, 2005a: 160 A: NP
oblongulus Nyholm, 1967: 27 A: CY
ovatus Nyholm, 1967: 35 E: IT
pallidicollis Raffray, 1873: 374 E: FR (Corse) IT (Sardegna) N: AG MO
pernigrans Nyholm, 1967: 39 E: SP
proximus Nyholm, 1967: 34 E: IT
pulchellus Klausnitzer, 1980e: 200 A: NP
rectangulus Klausnitzer, 1991: 162 N: AG
rivulorum Nyholm, 1977: 102 A: TR
satoi Yoshitomi, 2001: 90 A: JA NC TAI
schoenmanni Yoshitomi & Klausnitzer, 2003: 523 A: YUN
segrex Nyholm, 1972: 34 E: AB GR (Chios, Sámos) TR A: IN TR
sieberi Klausnitzer, 2010c: 123 (Cyphon) A: SD
sinicus Pic, 1934b: 23 A: SCH
steueri Klausnitzer, 2006b: 447 A: NP
subinterrogationis Klausnitzer, 2012b: 329 A: SD
taiwanus Yoshitomi, 2001: 99 A: TAI
uenoi Yoshitomi & Klausnitzer, 2003: 529 A: YUN

SCIRTIDAE

SCIRTIDAE

421

verrucosus Klausnitzer, 2012b: 333 A: NP
vicinans Nyholm, 1972: 33 A: IS JO TR
wangi Yoshitomi, 2008b: 80 A: JIX
yoshitomii Klausnitzer, 2002b: 259 A: NP PA
zwicki Klausnitzer, 2011c: 263 A: PA
genus Mescirtes Motschulsky, 1863: 485 type species Mescirtes gagatinus Motschulsky, 1863
ruthsteuerae Klausnitzer, 2009a: 295 (Prionocyphon) A: NP
squamatilis Klausnitzer, 2013b: 127 A: YUN
weigeli Klausnitzer, 2009a: 296 (Prionocyphon) A: NP
genus Microcara C.G. Thomson, 1859: 107 type species Galleruca livida Fabricius, 1792 (=Silpha testacea
Linnaeus, 1767)
dispar Seidlitz, 1874: 329 E: PT SP
luteicornis Reitter, 1888b: 207 E: AB GG ST TR A: IN TR
caspica Reitter, 1889i: 372

omissa Klausnitzer, 1972b: 150 E: BH CR RO
pilosula Reitter, 1911a: 244 E: IT
testacea Linnaeus, 1767: 574 (Silpha) E: AU BU BY CR CT CZ DE EN FI FR GB GE HU IR IT KZ LA LS LT LU NL
NR NT PL RO RU SK SP ST SV SZ UK
assimilis Stephens, 1830: 284 (Cyphon)
bohemani Mannerheim, 1844: 196 (Cyphon)
livida Fabricius, 1792b: 22 (Galleruca)
mollis Marsham, 1802: 225 (Crioceris)
obscura Stephens, 1830: 285 (Cyphon)
tenella Olivier, 1795: nr. 54: 9 (Cistela)

genus Nyholmia Klausnitzer, 2013c: 255 type species Cyphon collaris Guérin-Méneville, 1843
ainu Nakane, 1963a: 31 (Cyphon) A: FE (Kunashir, Sachalin) JA (Hokkaidō)
kerzhneri Klausnitzer, 1982: 283 (Cyphon)

aomoriana Yoshitomi, 2005: 110 (Cyphon) A: JA
hasegawai Nakane, 1963c: 139 (Cyphon) A: JA
kyushuana Yoshitomi, 2005: 125 (Cyphon) A: JA
occidens Yoshitomi, 2005: 121 (Cyphon) A: JA
ohbayashii Yoshitomi, 2005: 122 (Cyphon) A: JA
patiens Klausnitzer, 1982: 280 (Cyphon) A: FE (Kunashir, Sachalin) JA
propria Yoshitomi, 2005: 115 (Cyphon) A: JA
sanno Nakane, 1963a: 139 (Cyphon) A: JA
sannoides Yoshitomi, 1996: 97 (Cyphon) A: JA
seryu Nakane, 1963c: 139 (Cyphon) A: JA
tohokuana Yoshitomi, 2005: 110 (Cyphon) A: JA

422

SCIRTIDAE

uenoi Yoshitomi, 2005: 119 (Cyphon) A: JA
vulgaris Yoshitomi, 2005: 117 (Cyphon) A: JA
genus Odeles Klausnitzer, 2004a: 80 type species Cistela marginata Fabricius, 1798
alienata Klausnitzer, 1976a: 214 (Helodes) A: BT
apenninicola Klausnitzer, 2008b: 187 E: IT
armilabris Nyholm, 1974: 48 (Helodes) E: ST A: TR
asturica Klausnitzer, 2008b: 193 E: SP
auriensis Klausnitzer, 2008b: 192 E: SP
bulgharensis Klausnitzer, 1980a: 59 (Helodes) E: BU MC
calpensis Klausnitzer, 2008b: 188 E: PT, SP
cantabrica Klausnitzer, 2008b: 193 E: SP
granosicollis Klausnitzer, 2008b: 191 E: SP
gredleri Kiesenwetter, 1863: 719 (Helodes) E: AU BH CR CZ FR IT RO SL SZ
signaticornis Baudi di Selve, 1873: 226 (Helodes)

hausmanni Gredler, 1857: 65 (Cyphon) E: AU CZ FR GE HU IT SK SZ
bonvouloirii Tournier, 1868a: 39 (Helodes) [DA]

horstaspoecki Klausnitzer, 2004b: 262 E: FR, FR (Corse) IT
inornata Lewis, 1895b: 107 (Helodes) A: FE (Kunashir) JA “Korea”
piceata Klausnitzer, 1982: 277 (Helodes)

marginata Fabricius, 1798: 103 (Cistela) [NP] E: AU BE CZ DE FR GB GE HU IR IT LS LU NL NR PL RO SK SV
SZ UK
circumfusa Marsham, 1802: 227 (Crioceris)
delagrangei Pic, 1898b: 26 (Helodes) [DA]
fuliginosa Bourgeois, 1884a: 20 (Helodes) [DA]
limbata Laporte, 1840: 259 (Cyphon)
nigricans Schilsky, 1888a: 188 (Helodes) [HN]
nimbata Panzer, 1794b: 24: 15 (Cistela)
schilskyi Pic, 1914a: 37 (Cyphon) [RN]
xanthopoda Bourgeois, 1884a: 20 (Helodes) [DA]

marocana Pic, 1939a: 21 (Helodes) [DA] N: MO
morimotoi Yoshitomi & Satô, 2003: 195 A: NP
necopinata Klausnitzer, 2011a: 400 A: SHX
nepalensis Yoshitomi & Satô, 2003: 191 A: NP
nigrovittata Bourgeois, 1896: 119 (Helodes) A: HP
peninsularis Pic, 1898b: 27 (Helodes) [DA] E: SP
scapularis Lewis, 1895: 107 (Helodes) A: JA NC
styriaca Klausnitzer, 2008d: 55 E: AU SL
tenebrica Klausnitzer, 2008b: 196 E: SP
trilineata Chevrolat, 1865: 352 (Helodes) E: FR SP
pyrenaea Pic, 1898b: 26 (Helodes)

SCIRTIDAE

423

wilsoni Pic, 1918d: 17 A: FE (Kunashir) JA
monticola Nakane, 1963c: 140 (Sarabandus)

wittmeri Klausnitzer, 1976a: 213 (Helodes) A: BT HP KA SD
genus Ora Clark, 1865: 385 type species Ora grayii Clark, 1865
mawatarii Nakane, 1958: 90 (Scirtes) A: JA
okinawana Nakane, 1963c: 139 (Scirtes) A: JA
yayeyamana Satô & Chûjô, 1972: 21 (Scirtes) A: JA (Ryukyus)
genus Prionocyphon L. Redtenbacher, 1858: 517 type species Cyphon serricornis P.W.J. Müller, 1821
babai Yoshitomi, 2010: 529 A: TAI
costipennis Ruta, 2010a: 53 A: TAI
fuscipennis Kiesenwetter, 1874: 244 A: JA
macrodascilloides Ruta, 2010a: 54 A: TAI
majusculus Nyholm, 1971: 41 N: MO
numidicus Nyholm, 1971: 40 N: AG
ornatus Abeille de Perrin, 1881: 103 E: AB BU IT FR GR A: LE SY TR
ovalis Kiesenwetter, 1874: 243 A: HEN JA
serricornis P.W.J. Müller, 1821: 221 (Cyphon) E: AL AU BE BH BU CR CT CZ DE FR GB GE GR HU IR IT LA LT LU
MD NL NR PL RO RU SK SP ST SV SZ
chrysomeloides Stephens, 1830: 283 (Cyphon)
serraticornis Zetterstedt, 1824: 154 (Cyphon) [HN]

sexmaculatus Lewis, 1895: 105 A: JA
genus Sacodes LeConte, 1854b: 356 type species Elodes thoracica Guérin-Méneville, 1843
Flavohelodes Klausnitzer, 1980a: 61 type species Helodes flavicollis Kiesenwetter, 1859

dux Lewis, 1895b: 106 (Helodes) A: JA
flavicollis Kiesenwetter, 1859a: 22 (Helodes) E: AL AU BH BU CR GR HU IT PL RO SK SB SL TR UK
carolinae Tschapeck, 1859: 425 (Elodes)

humeralis Yoshitomi & Satô, 1996b: 306 (Flavohelodes) A: TAI
kaszabi Klausnitzer, 1973b: 113 (Helodes) A: NC
leei Yoshitomi & Satô, 2004: 211 A: TAI
mamaevi Klausnitzer, 1977: 167 (Helodes) A: FE
minima Klausnitzer, 1973b: 107 (Helodes) A: FE (Kunashir) JA
nakanei Klausnitzer, 1973b: 107 (Helodes) A: JA
okinawana Yoshitomi, 2005: 36 A: JA (Ryukyus)
protecta Harold, 1880a: 169 A: FE JA
shibatai Yoshitomi, 2008a: 235 A: JA (Yaku-shima)
taiwanensis Yoshitomi & Satô, 1996b: 303 (Flavohelodes) A: TAI
testaceoapicalis Pic, 1954a: 64 (Elodes) A: JA (Ryukyus) TAI
amamiensis Satô, 1966a: 13 (Elodes)

424

SCIRTIDAE

tsushimensis Yoshitomi, 1997: 399 A: JA (Tsushima)
genus Scirtes Illiger, 1807: 301 type species Chrysomela hemisphaerica Linnaeus, 1758
biinterruptus Pic, 1918e: 16 A: SD
borneensis Pic, 1918e: 16 [DA] A: CH ORR
elongatus Waterhouse, 1880: 571 (Scirtes) A: HKG NC SC
sinensis Pic, 1918b: 16

hamifer Klausnitzer, 2005b: 42 A: NP ORR
hartmanni Klausnitzer, 2005b: 40 A: NP
hemisphaericus Linnaeus, 1758: 373 (Chrysomela) E: AU BE BU BY CR CT CZ DE FI FR GB GE GR HU IR IT KZ
LA LS LT LU MC MD NL NR NT PL RO RU SB SK SL SP ST SV SZ UK YU A: RU TR UZ
attenuatus Baudi di Selve, 1872: 92 (Scirtes)
fuscus De Geer, 1775: 348 (Chrysomela)
latiusculus O.F. Müller, 1776: 84 (Altica)

japonicus Kiesenwetter, 1874: 244 (Scyrtes) A: FE JA NC TAI YUN NAR ORR
nigricans Waterhouse, 1880: 567 (Scirtes) A: CH
orbicularis Panzer, 1793: 6 (Altica) E: AB AU BE BY CT CZ DE FR GB GE GG HU IR IT LA LU NL PL SK ST SV
SZ UK A: IN TR
fulviculus Reitter, 1898b: 121 (Scyrtes)
pallescens Stephens, 1830: 282 (Scirtes) [DA]

ovatulus Lewis, 1895b: 105 [DA] A: FE JA
rufonotatus Pic, 1915b: 6 A: TAI
sobrinus Lewis, 1895b: 105 A: FE JA NC
sakishimanus Satô & Chûjô, 1972: 20 (Scirtes)

tsumaguro Satô & Chûjô, 1972: 21 A: JA
unicolor Pic, 1914b: 5 A: YUN ORR
ussuriensis Nyholm, 2002: 57 A: FE
genus Yoshitomia Klausnitzer, 2013c: 260 type species Cyphon beattyi Pic, 1918
beattyi Pic, 1918d: 18 (Cyphon) A: JA
genus Ypsiloncyphon Klausnitzer, 2009d: 248 type species Cyphon chlorizans Klausnitzer, 1973
amami Yoshitomi, 2005: 134 (Cyphon) A: JA (Ryukyus)
africumsinis Klausnitzer, 2015a: 162 A: YUN
honshuanus Yoshitomi, 2005: 131 (Cyphon) A: JA
jiangxiensis Yoshitomi, 2002: 43 (Cyphon) A: JIX
mendosus Klausnitzer, 1980e: 206 (Cyphon) A: NP
minutulus Klausnitzer, 1980e: 206 (Cyphon) A: SD
nepalensis Yoshitomi, 2015b: 481 A: NP
okinawanus Yoshitomi, 2005: 134 (Cyphon) A: JA (Ryukyus)
pannussustentans Klausnitzer, 2015a: 158 A: YUN
serratus Klausnitzer, 2015b: 474 A: AP
sinuosus Sasagawa, 1985: 36 (Cyphon) A: JA (Ryukyus)

425

EUCINETIDAE
spinifer Yoshitomi, 2005: 136 (Cyphon) A: JA (Ryukyus)
vegrandis Klausnitzer, 2015a: 160 A: YUN
yayeyamanus Yoshitomi, 2005: 138 (Cyphon) A: JA (Ryukyus)
zimmermanni Klausnitzer, 2015a: 165 A: AP
family Scirtidae, nomen dubium
Cyphon rugosus Olivier, 1795: 54: 10 (Cistela) [DA] distribution unknown

family DECLINIIDAE Nikitsky, Lawrence, Kirejtshuk & Gratshev, 1994

genus Declinia Nikitsky, Lawrence, Kirejtshuk & Gratshev, 1994: 4 type species Declinia relicta Nikitsky,
Lawrence, Kirejtshuk & Gratshev, 1994
relicta Nikitsky, Lawrence, Kirejtshuk & Gratshev, 1994: 5 A: FE SC
versicolor M. Sakai & Satô, 1996: 104 A: JA

family EUCINETIDAE Lacordaire, 1857

genus Bisaya Reitter, 1884a: 3 type species Bisaya nossidiiformis Reitter, 1884
nossidiiformis Reitter, 1884a: 5 E: AB A: IN
genus Eucilodes Vít, 1985: 429 type species Eucinetus caucasicus Reitter, 1880
caucasicus Reitter, 1880a: 475 (Eucinetus) E: FI GG ST
nikkon Vít, 1977: 928 (Euscaphurus) A: JA
ussurensis Nikitsky, 1989: 438 A: FE
genus Eucinetella Nikitsky, 1996: 1366 type species Eucinetella kurbatovi Nikitsky, 1996
kurbatovi Nikitsky, 1996: 1366 A: SCH
genus Eucinetus Germar, 1818: 255 type species Scaphidium haemorrhoidale Germar, 1818
Apeosina Broun, 1881: 692 type species Apeosina stewardi Broun, 1881
Cryptomera Broun, 1893: 1358 type species Cryptomera nigra Broun, 1893
Hamaxobium Duftschmid, 1825: 79 type species Hamaxobium haemorrhoum Duftschmid, 1825

haemorrhoidalis Germar, 1818: 255 (Scaphidium) E: AU BU CT CZ DE EN FI FR GB GE GR HU IT (Sardegna)
LT MA NL NT PL RO SK ST SV SZ UK N: AG MO TU A: ES GG JA MG NC SC NARi
haemorrhoum Duftschmid, 1825: 80 (Hamaxobium)
haemorhous Laporte, 1841: 260 (Nycteus)
reitteri Portevin, 1908b: 264 [RN]
strigosus Reitter, 1888d: 423 [HN]

rufus Portevin, 1908b: 264 A: JA
genus Nycteus Latreille, 1829: 462 type species Nycteus meridionalis Laporte, 1838
bicolor Reitter, 1887d: 514 (Eucinetus) E: GG ST TR
hopffgarteni hopffgarteni Reitter, 1885a: 71 (Eucinetus) E: BH BU GR HU IT PL RO SK YU
hopffgarteni prospector Vít, 1985: 451 (Eucinetus) E: IT (Calabria)

426

CLAMBIDAE

meridionalis Laporte, 1838: 25 E: FR (Corse) GB GE IT (Sardegna, Sicilia) NL SP N: AG MO MR TU
ovum Wollaston, 1854: 242

oertzeni Reitter, 1887d: 514 (Eucinetus) E: GR
rugosus Portevin, 1908b: 264 (Eucinetus) A: JA
subrutilus Pic, 1906b: 33 (Eucinetus) E: FR N: TU
wollastoni Vít, 1999: 392 (Eucinetus) N: CI (Tenerife)
genus Proeuzkus Vít, 2000: 124 type species Proeuzkus pachys Vít, 2000
pachys Vít, 2000: 125 A: NP
genus Tohlezkus Vít, 1977: 920 type species Tohlezkus ponticus Vít, 1977
Subulistomella M. Sakai, 1980: 83 type species Subulistomella rufa M. Sakai, 1980

orientalis Vít, 1981: 769 A: TAI
ponticus Vít, 1977: 922 E: ST A: TR
rufus M. Sakai, 1980: 85 (Subulistomella) A: JA

family CLAMBIDAE Fischer von Waldheim, 1821
subfamily Calyptomerinae Crowson, 1955

genus Calyptomerus L. Redtenbacher, 1847: 159 type species Calyptomerus alpestris L. Redtenbacher,
1847
Comazus Fairmaire & Laboulbène, 1854: 328 type species Clambus enshamensis Stephens, 1829

alpestris L. Redtenbacher, 1847: 159 E: AU CT CZ FR GE IT RO SK SL SZ UK “Caucasus”
caucasicus Reitter, 1876: 289 E: GG ST A: TR
dubius Marsham, 1802: 234 (Scaphidium) E: AU BE BH CR CZ DE FR GB GE GR IR IT LT MR NL NR SK SP SV
SZ UK N: AG CI MO A: IS TR
cephalotes Dejean, 1837: 455 (Cyrtocephalus)
enshamensis Stephens, 1829a: 184 (Clambus)
troglodytes Fauvel, 1861: 576

subfamily Clambinae Fischer von Waldheim, 1821

genus Clambus Fischer von Waldheim, 1821b: 52 type species Dermestes armadillo De Geer, 1774
Sternuchus LeConte, 1850: 222 type species Sternuchus gibbulus LeConte, 1850

armadillo De Geer, 1774: 220 (Dermestes) E: AU BE BH CZ DE EN FI FR GB GE HU IR IT LA LS LT NL NR NT
PL SK SV SZ UK NAR
atomarius Sturm, 1807: 65 (Agathidium)
convexus Marsham, 1802: 73 (Dermestes)
nanus Stephens, 1829a: 183 (Agathidium)

bengalensis Endrödy-Younga, 1986: 111 A: NP SD
besucheti Endrödy-Younga, 1974: 541 E: TR A: TR
bipartitus Endrödy-Younga, 1990: 307 A: NP
caucasus Endrödy-Younga, 1960: 300 E: AB IT “Caucasus” A: TR

CLAMBIDAE

427

cilicicus J.R. Sahlberg, 1913: 80 E: GR A: LE TR
crinitus Endrödy-Younga, 1978: 72 A: UP
croceus Endrödy-Younga, 1986: 105 A: UP
dux dux Endrödy-Younga, 1960: 281 E: AL AU BH FR GR HU IT RO SK SZ UK A: TR
dux lanceolatus Endrödy-Younga, 1960: 284 E: SP
dux proximus Endrödy-Younga, 1960: 284 E: AR A: AF TR
dux xerxes Endrödy-Younga, 1960: 285 E: “Caucasus” A: IN IS PA
evae Endrödy-Younga, 1960: 301 E: AU BH FR GB GE HU IR IT PL SZ
felix Endrödy-Younga, 1960: 280 E: SP N: AG EG MO
filii Endrödy-Younga, 1960: 285 E: FR (Corsica)
formosanus formosanus Endrödy-Younga, 1959: 103 A: TAI ORR
formosanus indicus Endrödy-Younga, 1978: 83 A: PA UP
formosanus japonicus Endrödy-Younga, 1960: 274 A: JA NP
gibbulus LeConte, 1850: 222 (Sternuchus) E: BE CZ FI FR GB GE IT LT NR NT PL RO SK SL SP ST SV SZ UK NAR
radula Endrödy-Younga, 1960: 271

gibbus Endrödy-Younga, 1986: 98 A: TAI
hayekae Endrödy-Younga, 1960: 295 E: FR IT SP N: AG MO
hisamatsui Endrödy-Younga, 1986: 95 A: JA
kaszabi Endrödy-Younga, 1960: 276 N: MO
klapperichi Endrödy-Younga, 1986: 95 A: TAI
kumaonis Endrödy-Younga, 1986: 108 A: UP
loebli Endrödy-Younga, 1986: 93 A: JA
loeblianus Endrödy-Younga, 1990: 309 A: NP
lohsei Meybohm, 2004: 14 E: FI GE PL
minutus complicans Wollaston, 1864: 101 E: BE FR GR IT NL SP SZ N: AG CI MO TU A: IS LE
ruficapillus J.R. Sahlberg, 1903a: 9

minutus minutus Sturm, 1807: 64 (Agathidium) E: AU BE BY DE FR GE GR HU IT LS NL PL UK A: LE
mus Endrödy-Younga, 1986: 102 A: SD
nigrellus Reitter, 1914a: 38 E: AU BE BH CR CZ DE FI FR GB GE HU IT LS NL NR NT PL RO SV SZ UK
aegrepilosus Endrödy-Younga, 1960: 289

nigriclavis Stephens, 1835: 404 E: AU BE BH BU CR CZ FR GB GE IT LS NL PL RO SK SZ
nipponicus Endrödy-Younga, 1960: 275 A: JA
pallidulus pallidulus Reitter, 1911c: 278 E: BU CR DE FR GB GE GR HU IR IT NL PL SV SZ N: AG A: IS
bicolor A. Fiori, 1915: 72

pallidulus shinto Endrödy-Younga, 1986: 92 A: JA
pilosellus Reitter, 1876: 289 E: AB AU BU CR GG HU IT RO SK ST UK A: NP TAI TR
pubescens L. Redtenbacher, 1847: 158 E: AU BE BU CT CZ DE EN FI FR GB GE HU IR IT LA LT NL NR NT PL RO
SK SP ST SV SZ UK A: FE MG
punctillatus Rey, 1889a: 5

punctulum L. Beck, 1817: 8 (Agathidium) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE HU IR IT LA LS LT
NL NR NT PL RO SK SL SP ST SV SZ A: FE TR

428

DASCILLIDAE

borealis A. Strand, 1946: 168
nitidus Stephens, 1835: 405

ruber Endrödy-Younga, 1986: 103 A: NP SD UP ORR
simsoni Blackburn, 1902: 288 Ei: AU BE DE FR GB GE NL SV AFR AUR
tuberculatus Endrödy-Younga, 1986: 97 A: TAI
villosus Endrödy-Younga, 1978: 78 A: NP ORR
vulneratus LeConte, 1879: 512 A: JA NAR
wittmeri Endrödy-Younga, 1986: 107 A: NP SD UP
genus Loricaster Mulsant & Rey, 1861a: 134 [=1862: 118] type species Loricaster testaceus Mulsant & Rey, 1861
cribripennis Reitter, 1904b: 147 E: FR
glaber Portevin, 1908a: 26 A: JA
testaceus atomus Reitter, 1884c: 119 E: GR A: IS SY TR
testaceus caspicus Reitter, 1884c: 119 E: AB ST
testaceus pumilus Reitter, 1884c: 120 E: FR IT SP N: AG MO
testaceus testaceus Mulsant & Rey, 1861a: 135 [=1862: 119] E: FR GE IT SZ
testaceus viertli Reitter, 1884c: 119 E: RO

series ELATERIFORMIA Crowson, 1960
superfamily DASCILLOIDEA Guérin-Méneville, 1843 (1834)
family DASCILLIDAE Guérin-Méneville, 1843 (1834)
subfamily Dascillinae Guérin-Méneville, 1843 (1834)
tribe Cinnabariini Pic, 1914

genus Coptocera A. Murray, 1868: 93 type species Coptocera gallerrucoides A. Murray, 1868
Cinnabarium Fairmaire, 1895d: ccvi type species Cinnabarium seminigrum Faimaire, 1895

nigronotata Pic, 1910: 28 (Cinnabarium) A: YUN
obliterata Pic, 1910: 29 (Cinnabarium)

sasajii Satô & Osawa, 2001: 305 (Metallidascillus) A: TAI
yangi Satô, Li & Lee, 2004: 367 (Metallidascillus) A: TAI

tribe Dascillini Guérin-Méneville, 1843 (1834)

genus Dascillus Latreille, 1797: 43 type species Chrysomela cervina Linnaeus, 1758
Atopa Paykull, 1799: 116 type species Chrysomela cervina Linnaeus, 1758

acutus Jin, Ślipiński & Pang, 2013: 562 A: GAN HUB SCH SHX
atripes Pic, 1915c: 4 A: CH
cavaleriei Pic, 1930b: 272 A: GUI HUB
cervinus Linnaeus, 1758: 376 (Chrysomela) E: AU BY CR CZ DE FI FR GB GE GR HU IR IT LA LS LT NL NR NT
PL PT SK SL SP ST SV SZ UK
cinereus Fabricius, 1781: 146 (Cistela)
semiobscurus Pic, 1908b: 41
testaceovillosus De Geer, 1774: 235 (Ptinus)

chifengi Jin, Ślipiński & Pang, 2013: 564 A: TAI

DASCILLIDAE
compressus Jin, Ślipiński & Pang, 2013: 564 A: NP SD
congruus Pascoe, 1860: 44 A: ANH FUJ GUA HUN JIX TAI ZHE
fortunei Pic, 1913c: 166
klapperichi Pic, 1955: 25
perroudi Pic, 1939a: 4
taiwanus Nakane, 1995: 24

costatus Pic, 1927a: 1 A: BT NP SD UP ORR
costulatus Fairmaire, 1886: 333 A: YUN
elongatus Faldermann, 1835a: 183 (Atopa) E: AR GG ST A: IN TR
obscuricollis Pic, 1908b: 41
ruficollis Pic, 1908b: 41

formosanus Jin, Ślipiński & Pang, 2013: 568 A: TAI
jaspideus Fairmaire, 1878: 115 (Therius) A: GAN HEN HUB NIN SHX SCH YUN
lanceus Jin, Ślipiński & Pang, 2013: 570 A: TAI
largus Jin, Ślipiński & Pang, 2013: 570 A: HUB SCH
latipennis Jin, Ślipiński & Pang, 2013: 571 A: SCH ORR
longicornis Westwood, 1849a: 6 A: NP
maculosus Fairmaire, 1889a: 35 A: GUI HEN HUB HUN SCH YUN ZHE
mongolicus Heyden, 1889c: 675 A: GAN HEN NIN SHX SCH YUN
montanus Jin, Ślipiński & Pang, 2013: 573 A: HUB SCH
nigronotatus Pic, 1914: 2 (Pseudolichas) A: GUI YUN ORR
fulvovittatus Pic, 1914: 3

nivipictus Fairmaire, 1904: 86 A: GUI JIX YUN ORR
cambodjensis Pic, 1914: 2
obliteratus Pic, 1908a: 55
ochraceus Pic, 1913d: 172
ruficornis Pic, 1914: 2
suturellus Fairmaire, 1904: 87

pallidofemoratus Pic, 1911b: 142 A: NP SD UP YUN ORR
planus Jin, Ślipiński & Pang, 2013: 576 A: SCH YUN
praestans Fairmaire, 1891d: xiv A: HUB
russus Jin, Ślipiński & Pang, 2013: 577 A: AP
sicanus Fairmaire, 1861: 582 E: IT PT ST
aprutianus Depoli, 1936: 137
parallelus Dorn, 1872: 482

sublineatus Pic, 1915c: 4 A: HUB SHX SCH XIZ YUN
superbus Pic, 1907a: 142 (Pseudolichas) A: YUN
reductus Pic, 1909d: 146

thibetanus Pic, 1915c: 3 A: “Thibet”
tibetensis Jin, Ślipiński & Pang, 2013: 580 A: XIZ
transversus Jin, Ślipiński & Pang, 2013: 580 A: YUN
vittatus Pic, 1914: 7 (Cladotoma) A: BT NP SD

429

430

DASCILLIDAE

uniformis Pic, 1908a: 54 (Pseudolychas) A: YUN
nigriceps Pic, 1909a: 19 (Pseudolychas)

genus Petalon Schönherr, 1833: 111 type species Bruchus fulvulus Wiedemann, 1819
Pseudolichas Fairmaire, 1878a: 111 type species Pseudolichas sulcifrons Fairmaire, 1878

acerbus Jin, Ślipiński & Pang, 2013: 586 A: YUN ORR
bengalensis Pic, 1911b: 142 (Dascillus) A: NP SD UP
calvescens Bourgeois, 1892: 234 (Dascillus) A: YUN ORR
brevesulcatus Pic, 1933: 23 (Dascillus)
holzi Pic, 1911b: 142 (Dascillus)
rubropubens Pic, 1934c: 562 (Dascillus)
rufocinctus Pic, 1914c: 500 (Dascillus)

digitatus Jin, Ślipiński & Pang, 2013: 589 A: YUN ORR
fulvulus Wiedemann, 1819: 173 (Bruchus) A: BT UP ORR
corporaali Pic, 1923f: 86 (Dascillus)
obscuricolor Pic, 1933: 23 (Dascillus)
striatus Pic, 1911b: 142 (Dascillus)

indicus Guérin-Méneville, 1861: 541 (Dascillus) A: “Inde boréale” ORR
iviei Jin, Ślipiński & Pang, 2013: 592 A: YUN ORR
nigripennis Guérin-Méneville, 1861: 540 (Dascillus) A: “Bengale” ORR
renardi Bourgeois, 1891: cxxxvii (Dascillus) A: NP ORR
rufovillosus Bourgeois, 1892: 234 (Dascillus)

sulcifrons Fairmaire, 1878a: 112 (Pseudolichas) A: GUX YUN ORR
genus Sinocaulus Fairmaire, 1878a: 117 type species Sinocaulus rubrovelutinus Fairmaire, 1878
clypeatus Jin, Ślipiński & Pang, 2013: 596 A: GUI
laticollis Fairmaire, 1891d: xv A: HUB
omiensis Jin, Ślipiński & Pang, 2013: 597 A: SCH
rubrovelutinus Fairmaire, 1878a: 117 A: CHQ GUX GUI SCH
atriceps Pic, 1910: 45

truncatus Jin, Ślipiński & Wang, 2013: 583 A: GUI

subfamily Karumiinae Escalera, 1913
tribe Emmitini Escalera, 1914

genus Emmita Escalera, 1914b: 349 [RN] type species Emma serricornis Escalera, 1913
Emma Escalera, 1913: 319 [HN] type species Emma serricornis Escalera, 1913

serricornis Escalera, 1913: 320 (Emma) N: MO

tribe Genecerini Pic, 1914

genus Genecerus Walker, 1871: 13 type species Genecerus cervinus Walker, 1871
cervinus Walker, 1871: 14 A: YE AFR

431

RHIPICERIDAE
tribe Karumiini Escalera, 1913

genus Karumia Escalera, 1913b: 321 type species Karumia estafilinoides Escalera, 1913
Drilocephalus Pic, 1918b: 3 type species Drilocephalus pallidipennis Pic, 1918
Escalerina Bolívar y Pieltain, 1926: 199 type species Karumia microcephala Escalera, 1913
Zarudniola Semenov & Martynov, 1925: 74 type species Zarudniola staphylinus Semenov & Martynov, 1925

eberti Mandl, 1967b: 450 A: AF
estafilinoides Escalera, 1913b: 321 A: IN
loeffleri kermanensis Wittmer, 1983: 270 (Zarudniola) A: IN
loeffleri loeffleri Mandl, 1967b: 450 A: IN
microcephala Escalera, 1913b: 222 A: IN
spinigera Mandl, 1967: 448 A: AF
staphylinus Semenov & Martynov, 1925: 74 (Zarudniola) A: IN
vartianorum Mandl, 1967b: 451 A: IN

family RHIPICERIDAE Latreille, 1834
subfamily Sandalinae Jakobson, 1913

genus Arrhaphipterus Schaum, 1862a: 61 type species Arrhaphus olivetorum Kraatz, 1859
blanchei Chevrolat, 1854b: 432 (Callirrhipis) A: LE SY
dioni Chobaut, 1896c: 423 N: AG
larclausei Reitter, 1894c: 107 N: AG EG MO
olivetorum Kraatz, 1859: 54 (Arrhaphus) E: GR A: TR
phlomidis K. Daniel, 1900: 139 A: TR
schelkownikoffi Reitter, 1893c: 111 E: AB AR A: TM TR
areschanus Pic, 1916a: 1

vanderwolfi Wurst, 1999: 74 E: GR
zoroastres Wurst, 1997: 105 A: IN
genus Chamoerrhipes Latreille, 1834: 169 type species Chamoerrhipes ophthalmicus Latreille, 1834
Eurhipis Laporte, 1834: 228 type species Eurhipis senegalensis Laporte, 1834 (=Chamoerrhipes ophthalmicus
Latreille, 1834)

bifoveolatus Taschenberg, 1883: 177 (Chamaerhipis) A: YE (Suqutra)
genus Sandalus Knoch, 1801: 131 type species Sandalus petrophya Knoch, 1801
Megarhipis Laporte, 1834: 266 type species Ptiocerus brunneus Laporte, 1834
Microrhipis Guérin-Méneville, 1831a: 1 type species Microrhipis dumerilii Guérin-Méneville, 1831
Ptyocerus C. Thunberg, 1806a: 3 type species Ptyocerus mystacinus C. Thunberg, 1806

bourgeoisii Théry, 1900: 33 A: CH
chinensis Fairmaire, 1900b: 623 A: SCH
kani M. Sakai & A. Sakai, 1981: 151 A: JA (Amami-Ôshima)
nigripennis Pic, 1916b: 8 A: YUN
sauteri Emden, 1924a: 28 A: JA TAI

432

BUPRESTIDAE

takizawai Nakane, 1985: 35

segnis Lewis, 1887c: 316 A: JA TAI
semitestaceus Pic, 1906a: 1

superfamily Dascilloidea, incertae sedis
Spermodenops mollipennis Abeille de Perrin, 1881: 87 A: “Palestina”

superfamily BUPRESTOIDEA Leach, 1815
family BUPRESTIDAE Leach, 1815
subfamily Julodinae Lacordaire, 1857

genus Aaata Semenov, 1906: 152 type species Julodis finchi C.O. Waterhouse, 1884
finchi C.O. Waterhouse, 1884a: 429 (Julodis) A: IN PA
genus Julodella Semenov, 1893a: 311 type species Buprestis globithorax Steven, 1830
abeillei Théry, 1893: cclviii (Julodis) A: IN IQ TM TR SY
steppensis Obenberger, 1940k: 114 (Julodis)

brevilata Semenov, 1893a: 317 (Julodis) A: IN TM
cymbiformis Bílý, 1990b: 32 A: SA
dilaticollis Semenov, 1893a: 316 (Julodis) E: AB GG A: IN TR
fairmairei Théry, 1895b: clvii (Julodis)
veselyi Obenberger, 1940k: 114

globithorax Steven, 1830: 159 (Buprestis) E: AB AR A: IN TR
propinqua Fischer von Waldheim, 1830: 184 (Buprestis)
rotundicollis Laporte & Gory, 1835: 23 (Julodis)

haarlovi Descarpentries, 1965: 260 A: AF
impluviata Semenov, 1893a: 316 (Julodis) A: AF IN
impressithorax Bílý, 1983a: 37 A: IN
iranica Bílý, 1983a: 33 A: IN TR
kaufmanni Ballion, 1871a: 348 (Julodis) A: TD TM UZ
cyphodera Fairmaire, 1892d: clii (Julodis)
futura Semenov, 1893a: 315 (Julodis)
kauffmanni Ballion, 1871a: 348 [unused original spelling]

mesopotamica Holdhaus, 1920: 52 (Julodis) A: IQ SY
parvula Bílý, 1983a: 36 A: IN
pfeufferi Mühle, 2013: 188 A: PA
plasoni Marseul, 1889: 241 (Julodis) A: IN
schochi Théry, 1896: 107 (Julodis) A: IN
shestoperovi Stepanov, 1959a: 47 A: TM
testaceipes Obenberger, 1928c: 66 A: IN
zarudniana Semenov, 1903d: 163 A: IN

BUPRESTIDAE
genus Julodis Eschscholtz, 1829: 9 type species Buprestis fascicularis Linnaeus, 1758
Jalodis Eschscholtz, 1829: 9 [unused original spelling]
Phyllis Gistel, 1834: 10 type species Buprestis fascicularis Linnaeus, 1758
Saccosoma Motschulsky, 1859b: 304 type species Buprestis variolaris Pallas, 1773

aeneipes E. Saunders, 1869: 2 A: AF IN
afghanica Obenberger, 1924b: 7

aequinoctialis aequinoctialis A.G. Olivier, 1790a: no 32: 56 (Buprestis) N: MO AFR
aequinoctialis coelatocollis Reiche, 1877: clx N: AG
caelatocollis Reiche, 1877: clx [unused original spelling]

aequinoctialis cyrenaea Obenberger, 1940k: 114 N: LB
aequinoctialis deserticola Fairmaire, 1859c: ccxxxvi N: AG MO AFR
vermiculata Chevrolat, 1861: 147

aequinoctialis floccosa Klug, 1829: [no 5] (Buprestis) N: EG
aequinoctialis tubu Descarpentries & Bruneau de Miré, 1963: 165 N: LB AFR
algirica Laporte, 1835: 162 N: AG MO
albopilosa Chevrolat, 1838b: 50
algerica Laporte, 1835: 162 [unused original spelling]

andreae andreae A.G. Olivier, 1790a: no 32: 53 (Buprestis) E: AB AR A: IN IQ TR SY
ampliata Marseul, 1865: 66
costifera Obenberger, 1919b: 142
derasa Abeille de Perrin, 1904a: 208
hampei J. Thomson, 1878b: 11
lineata Kerremans, 1886: xxxvii
lineigera Marseul, 1865: 62
setosa Steven, 1830: 156 (Buprestis) [HN]
sovitzii Steven, 1830: 158 (Buprestis)
stevensii E. Saunders, 1871: 5 [RN]
subviolacea Abeille de Perrin, 1904a: 208
tenuelineata Abeille de Perrin, 1904a: 208

andreae obesa Obenberger, 1923b: 23 A: TR SY
andreae obsoletesulcata Obenberger, 1917e: 47 A: TR
mandli Pochon, 1969: 50

andreae sulcata L. Redtenbacher, 1843: 982 A: IS LE SY TR
carinulosa Obenberger, 1917e: 46
luteogramma Marseul, 1865: 67
media Abeille de Perrin, 1904a: 208

aristidis P.H. Lucas, 1860: xxvii N: AG LB MO TU
angustior Fairmaire, 1884a: 142

armeniaca armeniaca Marseul, 1865: 80 A: IN IQ SY TR
cyrus Obenberger, 1924b: 18

433

434
orientalis Kerremans, 1899: 272

armeniaca cypria Obenberger, 1917e: 48 A: CY
armeniaca oertzeni Ganglbauer, 1886e: 309 A: SY TR
atkinsoni Kerremans, 1896: 48 A: PA
audouinii Laporte & Gory, 1835: 16 A: IN IQ TR
auduinii Laporte & Gory, 1835: 16 [unused original spelling]
theryi Abeille de Perrin, 1900a: 3

balucha Obenberger, 1923b: 17 A: AF IN PA
semenovi Alexeev, 1990: 63

bleusei Abeille de Perrin, 1896a: 262 N: AG MO
cailliaudi cailliaudi Latreille, 1827: 277 (Buprestis) N: AG EG AFR
aethiopica Laporte, 1835: 162
obscura Bílý, 1971: 174
cyanura Kerremans, 1914b: 244

cailliaudi mniszechii Reiche, 1860a: 24 N: EG AFR
berberae Abeille de Perrin, 1900a: 4

cailliaudi spectabilis Gory, 1840: 23 A: SA YE
candida Hołyński, 1996: 135 A: AE SA YE
chrysesthes chrysesthes Chevrolat, 1860a: 208 N: AG TU
chrysesthes nivea Alfieri, 1976: 93 N: EG LB A: SI
clouei Buquet, 1843: 22 A: YE (Suqutra) AFR
consobrina consobrina Kerremans, 1914b: 247 A: IN
bushirensis Obenberger, 1928c: 66
hauseri Obenberger, 1928c: 65

consobrina gotwendensis Obenberger, 1924b: 6 A: IN
cylindrica Théry, 1925a: 82 A: IN
ehrenbergii ehrenbergii Laporte, 1835: 162 E: AL BU GR MC N: EG A: CY IN IQ SI SY TR
bohemanni Mannerheim, 1837: 19

ehrenbergii olivieri Laporte, 1835: 161 E: GR (Chíos, Sýros) A: CY
escalerae Abeille de Perrin, 1904a: 206 A: IN
bouvieri Kerremans, 1905: 227

euphratica aegyptiorum Marseul, 1865: 42 N: EG A: SA SI
euphratica beludjistana Obenberger, 1924b: 6 A: PA
euphratica euphratica Laporte & Gory, 1835: 18 A: AE IN IQ OM SA
euphratica karelinii Mannerheim, 1837: 123 A: AF IN TD TM UZ
mrazi Obenberger, 1924b: 6

euphratica scenica Kerremans, 1905: 231 A: IN PA
klapaleki Obenberger, 1924b: 9
nusskina Obenberger, 1924b: 6
ormarensis Obenberger, 1924b: 10

faldermanni Mannerheim, 1837: 14 E: AB AR GG A: IN TR

BUPRESTIDAE

BUPRESTIDAE
fidelissima Rosenhauer, 1856: 133 E: PT SP
pilosipennis Abeille de Perrin, 1904a: 208

fimbriata fimbriata Klug, 1829: [no 2] (Buprestis) N: AG EG LB A: SA YE AFR
arabica Gory, 1840: 15
chevalieri Kerremans, 1914b: 245
kustai Nonfried, 1892d: 335 (Sternocera)

fimbriata lacunosa Fairmaire, 1882a: 49 A: AE OM AFR
argodi Abeille de Perrin, 1900a: 3

interpunctata interpunctata J. Thomson, 1878b: 10 A: PA ORR
interpunctata kabakovi Alexeev, 1990: 66 A: AF PA
intricata L. Redtenbacher, 1843: 982 A: IN IQ IS SY TR
cupreacaelata Marseul, 1865: 53
ruginota Marseul, 1865: 54
rugosa J. Thomson, 1878b: 9 [HN]
thomsoni Kerremans, 1892a: 23 [RN]

iris Laporte & Gory, 1835: 12 N: EG A: IQ IS JO SI
ethema Gory, 1840: 13

kerimi Fairmaire, 1875a: 510 N: LB TU
tunetana Obenberger, 1927f: 70

klapperichi Cobos, 1966d: 307 A: AF
laevicostata Gory, 1840: 16 A: IN IQ TM
longicollis Abeille de Perrin, 1904a: 207 [distribution unknown]
lucasi E. Saunders, 1871: 4 [RN] N: AG LB TU
cicatricosa P.H. Lucas, 1859f: clxxxiii [HN]

manipularis Fabricius, 1798: 134 (Buprestis) N: MO
marmottani Escalera, 1913a: 32 N: MO
matthiesseni Reitter, 1905b: 204 A: IN
nemethi Théry, 1932c: 141 N: MO
onopordi chalcostigma Chevrolat, 1860a: 209 N: MO
kaenigii Gory, 1840: 19 [HN]

onopordi longiseta Abeille de Perrin, 1904a: 207 N: EG A: SI
gassneri Obenberger, 1917e: 47

onopordi onopordi Fabricius, 1787a: 181 (Buprestis) N: AG LB MO TU
jaminii P.H. Lucas, 1859f: clxxxiii
koenigii Mannerheim, 1837: 22
leucostica Fairmaire, 1859: ccxxxvi [unused original spelling]
leucosticta Fairmaire, 1859: ccxxxvi
puberula Reiche, 1877: clx
reboudi Fairmaire, 1876b: 49
setifensis P.H. Lucas, 1844a: 49

onopordi sommeri Marseul, 1865: 56 E: FR SP

435

436
kuesteri Schaefer, 1935: 46

onopordi splichali Obenberger, 1917e: 47 E: IT (Lampedusa) N: TU
lampedusanus Tassi, 1966: 17

partha Obenberger, 1923b: 23 A: IN IQ
amrishi Makhan, 2012: 1
straussi Pochon, 1967: 205

peregrina Chevrolat, 1838b: 48 N: EG A: IS JO SI
caiffensis Abeille de Perrin, 1904a: 206
iridescens Reiche, 1877: clx
noe Abeille de Perrin, 1896a: 263

pietzchmanni Kerremans, 1914b: 249 A: IN IQ
pietzehmanni Kerremans, 1914b: 249 [unused original spelling]

pilosa marseulii E. Saunders, 1871: 5 N: MO
pilosa pilosa Fabricius, 1798: 135 (Buprestis) N: MO
tingitana Gory, 1840: 21

proxima Gory, 1840: 12 A: IN IQ
beesoni Obenberger, 1934o: 186

pubescens pubescens A.G. Olivier, 1790a: no 32: 55 (Buprestis) E: GR
brullei Laporte & Gory, 1835: 25
helladica Obenberger, 1919b: 142

pubescens yvenii Mannerheim, 1837: 21 E: GR (Kríti)
ivenii Mannerheim, 1837: 21 [unused original spelling]

punctatocostata punctatocostata Gory, 1840: 17 A: IN
quadricostata Marseul, 1865: 71

punctatocostata ramifera Marseul, 1865: 69 A: IN
rothii Sturm, 1843: 328 A: IS JO SA SY
corrosa Reiche, 1877: clix
mucescens Dohrn, 1874: 256
zoufali Obenberger, 1914e: 112

speculifer dicksonae Théry, 1936c: 118 A: JO KU SA
speculifer speculifer Laporte, 1835: 160 A: IN IQ
iranica Obenberger, 1923b: 17
monstrosa Abeille de Perrin, 1904a: 206

syriaca anatolica Kerremans, 1903: 11 A: TR SY
syriaca distincta Gory, 1840: 11 A: IQ JO SY
balthasari Obenberger, 1934o: 187
loeffleri Mandl & Pochon, 1957: 323
palmyrensis Obenberger, 1923b: 16

syriaca syriaca A.G. Olivier, 1790a: no 32: 87 (Buprestis) N: EG A: IS JO LE SI SY
philistina Obenberger, 1934o: 187

variolaris badakschanica Cobos, 1966d: 305 A: AF

BUPRESTIDAE

BUPRESTIDAE
variolaris brendelli Hołyński, 1984a: 524 A: IN
variolaris bucharica Semenov, 1893c: 496 A: TD
acuminata Abeille de Perrin, 1904a: 211

variolaris freygessneri Meyer-Darcis, 1883: xxxix A: AF IN TM UZ
undulata Heyden, 1883a: 107

variolaris indica J. Thomson, 1878b: 10 A: AF IN PA “India or.”
aberrans Obenberger, 1924b: 7
eoa Obenberger, 1923b: 15

variolaris variolaris Pallas, 1771: 464 (Buprestis) E: AB KZ ST A: AF GAN IN KI KZ TM UZ XIN
amurensis Abeille de Perrin, 1904a: 211
kirghisica Marseul, 1865: 47
kokandensis Obenberger, 1924b: 6
oberthuri Kerremans, 1898a: 134
seminata Abeille de Perrin, 1904a: 211
zablodskii Motschulsky, 1845a: 33

whitehillii wazira Hołyński, 2014b: 25 A: PA
whitehillii whitehillii G. R. Gray, 1832b: 358 (Buprestis) A: PA ORR
genus Sternocera Eschscholtz, 1829: 18 [NP] type species Buprestis chrysis Fabricius, 1775
subgenus Sternocera Eschscholtz, 1829: 18 type species Buprestis chrysis Fabricius, 1775
Charytonia Gistel, 1834: 10 type species Buprestis chrysis Fabricius, 1775
Sternoxus Billberg, 1820a: 23 [NO] type species Buprestis chrysis Fabricius, 1775

aequisignata E. Saunders, 1866: 298 A: YUN ORR
bannaiensis Wu & Hou, 1996: 4
menghaiensis Wu & Hou, 1996: 5
punctatofoveata E. Saunders, 1866: 298

aurosignata J. Thomson, 1878b: 8 A: “Himalaya” NP SD ORR
bengalensis Obenberger, 1939: 35
birmanica Théry, 1922b: 193
flammea Obenberger, 1923f: 110
glabriventris Obenberger, 1932a: 206
pseudopunctatofoveata Obenberger, 1932a: 207
rufomaculata Obenberger, 1932a: 207

chrysis Fabricius, 1775: 218 (Buprestis) A: AF PA SD UP ORR
bramina Chevrolat, 1838b: 46
chrysidioides Laporte & Gory, 1835: 7
chrysites Pallas, 1781: 62 (Buprestis)
hybrida Théry, 1898: 367
marseuli Obenberger, 1923f: 109
nitens Kerremans, 1886: xxxv
nitidicollis Laporte & Gory, 1835: 8

437

438

BUPRESTIDAE

simulatrix Obenberger, 1923f: 109
singularis Kerremans, 1890a: xix
splendida Goeze, 1777: 595 (Buprestis)

dasypleuros Kollar, 1844: 504 A: KA NP “Himalaya” “India bor.”
bicolor Kerremans, 1886: xxxv
mniszechii J. Thomson, 1879a: 5

subgenus Afrocera Hołyński, 2014: 39 type species Sternocera orissa Buquet, 1837
castanea castanea A.G. Olivier, 1790a: no 32: 25 (Buprestis) N: AG EG LB A: “Arabia mer.” AFR
basinigra J. Müller, 1941: 327
druryi C.O. Waterhouse, 1904a: 247
embrikiana Obenberger, 1940a: 28
fischeri Gerstäcker, 1884: 51 [HN]
gerstaeckeri Kerremans, 1888a: 78 [RN]
gerstaekeri Kerremans, 1888a: 78 [RN] [unused original spelling]
irregularis Latreille, 1827: 276 (Buprestis)
nigerica Obenberger, 1940a: 26

subfamily Polycestinae Lacordaire, 1857
tribe Acmaeoderini Kerremans, 1893

genus Acmaeodera Eschscholtz, 1829: 9 type species Buprestis cylindrica Fabricius, 1775
subgenus Acmaeodera Eschscholtz, 1829: 9 type species Buprestis cylindrica Fabricius, 1775
Andromeda Gistel, 1834: 10 type species Buprestis ornata Fabricius, 1775

affinis P.H. Lucas, 1846a: pl. 14 N: AG LB MO TU
babatauensis Obenberger, 1935r: 207 A: KI TD TM UZ
gussakovskii Stepanov, 1958a: 114

bellamyola Volkovitsh, 2014: 37 A: SCH YUN
bilyi Volkovitsh, 1988: 37 A: KI UZ
brevipes abeilleana Obenberger, 1940k: 115 [RN] N: AG
abeillei Obenberger, 1914h: 251 [HN]

brevipes brevipes Kiesenwetter, 1858: 242 E: AL BU GR MC ME TR A: TR
praecox Marseul, 1865: 284

brevipes saducaea Obenberger, 1946: 12 A: IN IQ IS JO LE SY TR
brunneipennis Kerremans, 1906: 212 N: EG TU
elater Théry, 1930a: 101

bushirensis Obenberger, 1940k: 126 A: IN
cecropia Kiesenwetter, 1858: 242 E: GR A: CY IS TR
rhodica Obenberger, 1935q: 202

cerasina Marseul, 1865: 293 E: GR A: TR
chalcithorax Volkovitsh, 1986a: 19 E: AB AR A: IN IQ TM
cylindrica Fabricius, 1775: 220 (Buprestis) E: FR IT PT SP N: AG LB MO TU

BUPRESTIDAE

439

acuminipennis Laporte & Gory, 1835: 25
convolvuli Waltl, 1835: 59

damasensis Pic, 1936a: 25 A: IS SY
distigma Volkovitsh, 1983: 559 A: SY TR
eckweileri Volkovitsh, 1983: 555 A: IN
flavolineata cypricola Volkovitsh, 1983: 561 A: CY
flavolineata destructovittata Volkovitsh, 1988: 39 E: GR (Southern Sporades)
flavolineata flavolineata Laporte & Gory, 1835: 20 E: AB AR GG GR TR A: IN IQ IS JO LE SY TR
dorsalis Spinola, 1838: 392
krueperi Obenberger, 1927b: 81
suturalis Gory, 1840: 44

flavolineata piloselloides Volkovitsh, 1983: 562 E: GR (Kríti)
ghilarovi Volkovitsh, 1988: 34 E: AR A: TM
guichardi Levey & Volkovitsh, 1996: 142 A: AE OM
holzschuhi Volkovitsh & Bílý, 1979: 330 A: TR
inquirenda Volkovitsh, 1977a: 42 A: TD
kabakovi Volkovitsh, 1990: 74 A: AF PA
lata Heyden, 1885a: 287 A: KI KZ TD UZ
lopatini Volkovitsh, 2005: 358 A: KI
lucida Volkovitsh, 1983: 562 A: IN
medvedevi Volkovitsh, 1976a: 198 A: MG SHA SHX
moralesi Español & Mateu, 1947: 223 N: MO
pici Obenberger, 1924e: 49 A: “Arabia”
pilosellae persica Mannerheim, 1837: 124 E: AB AR GG A: IN TM TR
araxicola Reitter, 1890o: 339
polyxantha Semenov, 1895c: 245

pilosellae pilosellae Bonelli, 1812: 177 (Buprestis) E: AL BH BU CR GR FR IT MC ME PT RO SL SP TR N: AG
A: TR
dayremi Théry, 1930a: 77
discoidea A.G. Olivier, 1790a: no 32: 82 (Buprestis) [HN]

pilosellae syriaca Abeille de Perrin, 1900a: 10 A: IQ IS JO SY TR
planidorsis Semenov, 1896c: 321 E: AB AR
pulchra Fabricius, 1792b: 201 (Buprestis) E: PT SP N: AG MO TU A: IS TR
postverta Buquet, 1840b: 394

revelierii Mulsant & Rey, 1859a: 234 [=1859b: 170] E: FR (Corse) IT (Sardegna)
revelierii Mulsant & Rey, 1861b: 167 [HN]

sycophanta afghanica Volkovitsh, 1990: 73 A: AF PA
sycophanta sycophanta Abeille de Perrin, 1900a: 10 E: BU GR A: IN SY TR
edmundi Obenberger, 1935h: 107 [RN]
irrorata Reitter, 1890o: 338 [HN]

440

BUPRESTIDAE

rufocincta Baudi di Selve, 1870: 82 A: CY
semenovi Obenberger, 1935s: 211 A: SCH XIZ YUN
transcaucasica Semenov, 1895c: 246 E: AB AR GG A: IN TR
tunisiana Krajčík, 2012a: 1 N: TU
wethloi Obenberger, 1940k: 125 A: IN TR
xanthelytra Obenberger, 1940k: 124 A: TR
yunnana Fairmaire, 1888a: 23 A: YUN
nigrita Kerremans, 1895: 213

subgenus Acmaeotethya Volkovitsh, 1979b: 343 type species Elater degener Scopoli, 1763
acaciae Mayet, 1887b: xcv N: AG EG LB MO TU A: SI
bartoni Obenberger, 1940k: 127 E: GR (Kríti)
batelkai Volkovitsh, 2011b: 180 A: AE OM
bernardi Descarpentries, 1953: 251 N: AG LB A: IS JO OM AFR
biseriata Reitter, 1890o: 340 A: IN IS LE SY TR
bodoani Kerremans, 1911: 633 A: IN TR
boreoafricana Bellamy, 1998: 91 [RN] N: AG LB
stellata Abeille de Perrin, 1891: 273 [HN]

cisti cisti Wollaston, 1862: 439 N: CI
palmensis Lindberg, 1953: 8

cisti kudrnaorum Krajčík, 2012a: 4 N: CI (Gomera)
coluber Abeille de Perrin, 1904a: 219 A: IN
crinita abigail Obenberger, 1946: 11 N: EG A: IS JO LE SY
crinita crinita Gory, 1840: 41 E: AL BU CR GR HU IT MC ME SL A: SY TR
reichei Boieldieu, 1865: 6

crinita melanosoma P.H. Lucas, 1844a: 88 E: SP PT N: EG AG MO TU
crinita perinella Obenberger, 1934o: 218 [RN] A: CY
reichei Abeille de Perrin, 1891: 274 [HN]

degener degener Scopoli, 1763: 92 (Elater) E: AU BH BU CR CZ FR GG GR HU IT PT RO SK SL SP ST SZ TR UK
A: IN SY TR
degener mlokossewiczi Semenov, 1896c: 324 E: GG
guayarmina Volkovitsh & Liberto, 2002: 10 N: CI
hadiboe Volkovitsh, 2012: 202 A: YE (Suqutra)
hoberlandti Bílý, 1983a: 40 A: IN
holmi Levey & Volkovitsh, 1996: 144 A: YE (Suqutra)
instabilis Cobos, 1966d: 309 A: AF IN TM
kabateki Volkovitsh, 2012: 199 A: YE (Suqutra)
kachetica Semenov, 1896c: 325 E: AB GG
mesmini Obenberger, 1934o: 201

kerremansi Stebbing, 1914: 195 A: PA
maghrebica Volkovitsh, 2006b: 56 N: AG MO TU
maroccana Obenberger, 1916d: 237 E: SP N: AG MO
mirabilis Volkovitsh, 1990: 77 A: AF

BUPRESTIDAE

441

morio Gory, 1840: 42 N: AG
octodecimguttata octodecimguttata Piller & Mitterpacher, 1783: 68 (Buprestis) E: AB AR AU BH BU CR CZ FR
GE GR HU IT MC MD PL RO SB SK SL ST TR UK A: IQ SY TR
friwaldskyi Marseul, 1865: 278
sedecimpunctata Schrank, 1789: 83 (Buprestis)

octodecimguttata quattuordecimpunctata Villers, 1789: 338 (Buprestis) E: FR IT PT SP N: AG MO TU
feisthameli Gory, 1840: 37
insolita Théry, 1908: 68
multipunctata P.H. Lucas, 1844a: 87
quattuordecimguttata Fabricius, 1798: 136 (Buprestis)

ottomana confluens Baudi di Selve, 1870: 81 A: CY
ottomana menradi Mühle, 1997: 33 N: LB
ottomana ottomana I. Frivaldszky, 1837: 175 (Buprestis) E: AB AL BU GR MC TR A: IN IS JO LE SY TR
incognita Kerremans, 1898b: 309

pallidepicta Reitter, 1895d: 152 A: IN TM
patricki Brechtel, 1998: 96 A: OM YE
prepsli Volkovitsh, 2013: 78 A: IN
quadrifasciata octodecimpunctata Gory, 1840: 36 E: FR (Corse) IT (Sardegna)
quadrifasciata prunneri Spinola, 1838: 375 E: FR IT (Sardegna)
quadrifasciata quadrifasciata P. Rossi, 1790a: 187 (Buprestis) E: FR IT SP N: AG LB MO A: TR
graptelytra Obenberger, 1914h: 250
kabyliana Obenberger, 1914h: 250
mutabilis Spinola, 1838: 372

quadristriata Volkovitsh, 2013: 75 A: PA
quadrizonata Abeille de Perrin, 1891: 269 E: BU GR MC A: CY IN IS JO LE SY TR
rotundicollis Volkovitsh, 2013: 76 A: IN
rufoguttata Reitter, 1890o: 340 E: AB A: IN
necatrix Abeille de Perrin, 1893: 133
truculenta Abeille de Perrin, 1900a: 10

saxicola bijuga Marseul, 1865: 282 A: CY
cisti Baudi di Selve, 1870: 82 [HN]
cisticola Obenberger, 1940k: 131 [RN]

saxicola minoa Obenberger, 1940k: 119 E: GR (Kríti)
saxicola saxicola Spinola, 1838: 372 E: AB GR A: IN IQ IS JO LE SY TR
guttifera Marseul, 1865: 281 [HN]
spilophora Marseul, 1869a: 122 [RN]

sedecimmactata Abeille de Perrin, 1891: 272 A: TR
simulans albipilis Abeille de Perrin, 1904a: 219 A: IN
simulans simulans Abeille de Perrin, 1891: 272 A: IS JO LE
deprivata Volkovitsh, 1983: 564

socotraensis Volkovitsh, 2012: 204 A: YE (Suqutra)

442

BUPRESTIDAE

tassii Schaefer, 1965: 36 E: IT
truquii Abeille de Perrin, 1891: 275 E: GR (Kríti, Southern Sporades) A: CY IS TR
fusa Abeille de Perrin, 1900a: 10

undulata Abeille de Perrin, 1891: 274 A: IN IS LE SY
uvarovi Obenberger, 1928a: 119 A: AF HP PA
vanharteni Volkovitsh, 2011b: 178 A: AE OM
virgo soudana Marseul, 1867a: 49 N: EG A: YE AFR
berberae Abeille de Perrin, 1900a: 11

subgenus Cobosiella Volkovitsh, 1979b: 345 type species Acmaeodera chotanica Semenov, 1890
chotanica Semenov, 1891: 338 A: PA TD TM UZ XIN
holynskii Volkovitsh, 2011b: 181 A: AE IN PA QA UP ORR
luzonica luzonica Nonfried, 1895: 302 A: JA (Ryukyus) ORR
luzonica volkovitshi Hołyński, 1993c: 6 A: TAI
stictipennis Laporte & Gory, 1835: 26 A: UP ORR
subgenus Lisposcelis Volkovitsh, 1979b: 344 type species Acmaeodera jakobsoni Obenberger, 1928
jakobsoni Obenberger, 1928a: 119 A: NP “West Himalaya”
eberti Cobos, 1966c: 247

zaitzevi Volkovitsh & Niehuis, 2012: 241 A: IS
subgenus Loepotethya Volkovitsh, 1979b: 342 type species Acmaeodera ocellata Abeille de Perrin, 1891
ocellata Abeille de Perrin, 1891: 275 A: IN IQ SY TR
hauseri Théry, 1936c: 118
hetita Obenberger, 1925c: 55
kheili Obenberger, 1924b: 13

tibetica Volkovitsh, 2006a: 36 A: WP
subgenus Palaeotethya Volkovitsh, 1979b: 343 type species Buprestis bipunctata A.G. Olivier, 1790
algirica Kerremans, 1907b: 315 N: AG MO
anatolica Volkovitsh & Bílý, 1979: 332 A: TR
bipunctata bipunctata A.G. Olivier, 1790a: no 32: 52 (Buprestis) E: AL BH BU CR FR GR IT PT SP TR A: IS JO LE
SY TR
pittneri Rosenhauer, 1847: 96
senex Abeille de Perrin, 1891: 279
sexpustulata Laporte & Gory, 1835: 12
vaillant Spinola, 1838: 370 [RN]
vaillanti Marseul, 1865: 297 [unjustified emendation]
bipunctata flavopunctata P.H. Lucas, 1844a: 88 N: AG LB TU
riffensis Cobos, 1955b: 201

bipunctata guillebeaui Abeille de Perrin, 1891: 277 A: CY
bipunctata kureimatica Escalera, 1914a: 195 N: AG MO
bipunctata obenbergeriana Schaefer, 1936b: 117 [RN] E: FR (Corse)
obenbergeri Schaefer, 1936a: 5 [HN]

bipunctata plagiata Wollaston, 1864: 206 N: CI

BUPRESTIDAE
bipunctata romanoi Sparacio, 1992: 124 E: IT (Pantelleria, Sicilia)
ceylonensis Obenberger, 1932a: 209 A: CY
flavonotata P.H. Lucas, 1846a: pl. 14 N: AG MO
montilloti Raffray, 1873: 372

leonhardi leonhardi Obenberger, 1914h: 252 N: AG LB
leonhardi punica Obenberger, 1934o: 272 N: TU
lugubris fracta Wollaston, 1864: 205 N: CI
wollastoniana Obenberger, 1921a: 90

lugubris grosseseriata Cobos, 1960: 286 N: MO
lugubris lugubris Spinola, 1838: 370 N: AG EG LB MO TU A: IS JO
flavoangulata Kerremans, 1907b: 332
oranensis Obenberger, 1923b: 24
pasqualinii Baudon, 1958b: 148
ramosa Chevrolat, 1860b: 454
rubromaculata P.H. Lucas, 1844a: 88
semiopaca Abeille de Perrin, 1893: 134
variivestis Abeille de Perrin, 1904a: 220

lugubris malacensis Cobos, 1954b: 283 E: PT SP
lugubris oertzeni Ganglbauer, 1889: 55 E: GR (Kríti, Kiklades, Southern Sporades)
marania Obenberger, 1940k: 132

lugubris segurensis Escalera, 1904: 225 E: SP
macchabaea Abeille de Perrin, 1891: 276 A: IS JO SY
houskai Obenberger, 1946: 14
pharisaea Obenberger, 1946: 13

nigellata Abeille de Perrin, 1904a: 222 E: PT SP N: AG MO
lusitanica Obenberger, 1914h: 251
satanula Reitter, 1907b: 205

quadrifaria Baudi di Selve, 1870: 83 E: GR A: CY SY TR
sahariensis Volkovitsh, 1987a: 50 N: MO
santschii Théry, 1930a: 93 N: TU
tagoror Arnáiz, García & Bercedo, 2009: 84 N: CI (Lanzarote)
subgenus Paracmaeodera Théry, 1946: 28, 31 type species Acmaeodera fascigera Harold, 1869
aurifera aurifera Laporte & Gory, 1835: 31 A: NP PA UP ORR
hoscheki Obenberger, 1916d: 242
scintillans Kerremans, 1892a: 179 [RN]
varicolor Kerremans, 1888b: lxvi [HN]

aurifera sindhia Hołyński, 1993a: 2 A: PA ORR
aurifera siwalika Hołyński, 1993a: 2 A: UP
elevata Klug, 1829: [no 6] (Buprestis) N: AG EG LB A: IS SA SI YE AFR
costulipennis Obenberger, 1916c: 98
gibba Laporte & Gory, 1835: 29

443

444
nubica Obenberger, 1936k: 185
ovis Gory, 1840: 49 [HN]
sculptilis Harold, 1878c: 217
semicostata Kerremans, 1907b: 505

luculenta Boheman, 1860: 19 N: EG AFR
bella Nonfried, 1892c: 126
dives Nonfried, 1892c: 136
fossicollis Harold, 1878c: 216
keniae Obenberger, 1924f: 99
politiformis Obenberger, 1916b: 17
viridis Kerremans, 1898b: 308
sincera Obenberger, 1928a: 108

subgenus Ptychomus Marseul, 1865: 322 type species Buprestis polita Klug, 1829
aenea Kerremans, 1899: 269 A: OM SA AFR
inornata Théry, 1931b: 104

arabica Gory, 1840: 50 A: AE IS OM SA YE AFR
omana Krajčík, 2015: 5

foudrasii foudrasii Solier, 1833: 276 N: AG MO AFR
bourgoini Obenberger, 1928a: 117

grandicollis Abeille de Perrin, 1900: 13 N: AG MO AFR
nocturna Holm, 1978: 119 A: YE “Arabia”
polita chudeaui Descarpentries, 1958b: 971 N: AG MO
polita hedjazensis Théry, 1931: 103 A: SA
polita permicans Abeille de Perrin, 1900a: 12 A: AE IS OM SA SI YE AFR
polita polita Klug, 1829: [no 7] (Buprestis) N: AG EG LB MO AFR
klugii E. Saunders, 1871: 85
monodi Théry, 1930b: 64
viridipennis Théry, 1931c: 104

pseudopolita Descarpentries & Mateu, 1965: 1029 N: LB
sancta Théry, 1926b: 50 N: EG A: IQ SA YE AFR
yerburyi C.O. Waterhouse, 1904a: 265 A: IS JO OM SA YE AFR
curtus Théry, 1931c: 105

subgenus Rugacmaeodera Holm, 1978: 50 type species Buprestis ruficaudis De Geer 1778
mirei Descarpentries & Mateu, 1965: 1025 N: LB A: OM SA SI YE AFR
dispersemaculata Alfieri, 1976: 96
minima Holm, 1978: 59

mombassica Nonfried, 1895: 303 N: EG AFR
denudata Obenberger, 1916d: 237
discoidalis Nonfried, 1895: 304
fascipennis Kraatz, 1896b: 82

BUPRESTIDAE

BUPRESTIDAE

445

fulvipennis Kerremans, 1898b: 310
orientalis Kerremans, 1898b: 310
zanzibarica Kerremans, 1898b: 309

genus Acmaeoderella Cobos, 1955a: 5 type species Buprestis discoida Fabricius, 1787
subgenus Acmaeoderella Cobos, 1955a: 5 type species Buprestis discoida Fabricius, 1787
angorana Obenberger, 1940k: 138 (Acmaeodera) A: TR
carolusi Volkovitsh, 1989a: 57 A: TR
caspica caspica Ganglbauer, 1888: 195 (Acmaeodera) A: IN TM
caspica suturifera Reitter, 1904d: 256 (Acmaeodera) A: AF IN KZ SY TM UZ
caspica turkestanica Obenberger, 1934o: 249 (Acmaeodera) A: AF KZ TD TM UZ
cinerea Volkovitsh, 1982: 30 A: IN KI KZ TD TM UZ
circassica Reitter, 1890o: 342 (Acmaeodera) E: AB AR BU GR IT MC RO ST UK A: KZ TR
faldermanni Obenberger, 1934o: 270 (Acmaeodera)
florilega Obenberger, 1934o: 266 (Acmaeodera)
hellenica Obenberger, 1914h: 252 (Acmaeodera)
kubanica Obenberger, 1934o: 263 (Acmaeodera)
macedonica Obenberger, 1934o: 267 (Acmaeodera)
normanna Sparacio, 2006: 36

coarctata coarctata P.H. Lucas, 1846b: pl. 15 (Acmaeodera) E: FR PT SP N: AG MO TU
maurorum Obenberger, 1934o: 263 (Acmaeodera)
navasi Obenberger, 1934o: 262 (Acmaeodera)

coarctata seminata Abeille de Perrin, 1895c: cxx (Acmaeodera) E: BU GR MC TR A: TR
abeillei Pic, 1899: 256 (Acmaeodera)

cypriota cypriota Obenberger, 1940k: 138 (Acmaeodera) A: CY
cypriota gerae Volkovitsh, 1989a: 52 E: GR A: TR
discoida Fabricius, 1787a: 184 (Buprestis) E: FR IT MA PT SP N: AG EG LB MO TU A: IS JO SY
albosetosa Rosenhauer, 1856: 133 (Acmaeodera)
barbara Gory, 1840: 45 (Acmaeodera)
chevrolati Levrat, 1858: 262 (Acmaeodera) [HN]
ferghanensis Obenberger, 1934o: 245 (Acmaeodera)
laesicollis Chevrolat, 1859: 382 (Acmaeodera)
levrati Marseul, 1865: 306 (Acmaeodera) [RN]
parandelli Théry, 1895b: clix (Acmaeodera)
praeoccupata Lotte, 1938b: 150 (Acmaeodera) [RN]
pygmaea Obenberger, 1916d: 240 (Acmaeodera)
theryi Pic, 1936b: 213 (Acmaeodera) [HN]

dlabolai Bílý, 1983a: 49 A: IN
elbursi Obenberger, 1924b: 13 (Acmaeodera) E: AR GR (Kríti, Southern Sporades) TR A: CY IN IQ IS JO LE
SY TR

446

BUPRESTIDAE

aramaea Obenberger, 1934o: 254, 256 (Acmaeodera)
damasica Obenberger, 1940k: 136 (Acmaeodera)

fulvinaeva Reitter, 1890o: 345 (Acmaeodera) E: AB AR A: IS LE TR
iphigenia Obenberger, 1934o: 250 (Acmaeodera)

jonica Volkovitsh, 1989a: 58 E: AL GR A: IS LE TR
levantina Obenberger, 1934o: 255 (Acmaeodera) E: BU GR MC
moroderi Reitter, 1906d: 375 (Acmaeodera) E: PT SP
ardoisi Obenberger, 1934o: 265 (Acmaeodera)
cercedillana Obenberger, 1934o: 264 (Acmaeodera)
oliveirae Obenberger, 1934o: 264 (Acmaeodera)

muehlei Volkovitsh, 1989a: 54 A: SY TR
oresitropha Obenberger, 1924b: 11 (Acmaeodera) E: AB AR GG A: KI TD TM UZ
plavilscikovi Obenberger, 1936d: 46 (Acmaeodera) E: AB AR KZ ST A: KI KZ TM UZ
samosicola Volkovitsh, 1989a: 55 E: GR (Sámos)
serricornis Abeille de Perrin, 1900a: 11 (Acmaeodera) E: AB AR A: IN IS LE SA SY TR
beduina Cobos, 1963: 361
erinaceiformis Obenberger, 1922a: 19 (Acmaeodera)
staneki Obenberger, 1934o: 266 (Acmaeodera)

stepaneki Obenberger, 1940k: 139 (Acmaeodera) E: GR (Kríti)
stricta Abeille de Perrin, 1895c: cxix (Acmaeodera) E: AB AR A: CY IN IQ IS JO LE SY TR
bodenheimeri Théry, 1932d: 230 (Acmaeodera)
strumosa Obenberger, 1934o: 259 (Acmaeodera)

trinacriae Obenberger, 1923b: 25 (Acmaeodera) E: IT (Sicilia)
turanica Reitter, 1890o: 345 (Acmaeodera) A: AF KZ TD TM UZ
deminuta Semenov, 1895d: 265 (Acmaeodera)
sogdiana Semenov, 1895d: 264 (Acmaeodera)
warentzoffi Théry, 1895b: clviii (Acmaeodera)

volkovitshi Bílý, 1985: 161 A: SA
zeravshanica Volkovitsh, 1987b: 139 A: UZ
subgenus Carininota Volkovitsh, 1979b: 352 type species Buprestis flavofasciata Piller & Mitterpacher, 1783
christophi Obenberger, 1935h: 106 (Acmaeodera) A: IN TM
crucifera Abeille de Perrin, 1904a: 221 (Acmaeodera) A: IN
dsungarica Obenberger, 1918b: 19 (Acmaeodera) A: KZ MG XIN
zaisanicola Obenberger, 1935h: 107 (Acmaeodera)

farinosa Reiche, 1856b: 410 (Acmaeodera) E: GR A: CY IS LE SY TR
dermestoides I. Frivaldszky, 1845: 180 (Buprestis) [HN]
lisa Bellamy, 1998: 91 [RN]

flavofasciata aequistriata Abeille de Perrin, 1904a: 221 (Acmaeodera) A: IN IQ IS TR
flavofasciata albifrons Abeille de Perrin, 1891: 278 (Acmaeodera) E: AB AR A: TR
flavofasciata chorasanica Volkovitsh & Alexeev, 1994: 425 A: IN TM

BUPRESTIDAE

447

flavofasciata ciliciana Volkovitsh, 2006b: 56 A: SY TR
flavofasciata elaeagni Volkovitsh, 1982: 29 A: TD
flavofasciata flavofasciata Piller & Mitterpacher, 1783: 38, 84 (Buprestis) E: AB AL AR AU BH BU CR CT CZ FR
GE GG GR HU IT KO KZ MA MC MD ME PT RO SB SK SL SP ST SZ TR UK A: IS KZ MG NW TR WS
deorum Obenberger, 1940k: 135 (Acmaeodera)
hirta Villers, 1789: 338 (Buprestis)
taeniata Fabricius, 1787a: 180 (Buprestis)
veselyi Obenberger, 1934o: 229 (Acmaeodera)

flavofasciata hirsutula Gory, 1840: 47 (Acmaeodera) E: SP N: AG MO TU
tristis P.H. Lucas, 1844a: 87 (Acmaeodera)

flavofasciata ornatipennis Cobos, 1958: 239 N: MO
flavofasciata pilivestis Abeille de Perrin, 1904a: 222 (Acmaeodera) E: SP PT
flavofasciata placida Baudi di Selve, 1870: 84 (Acmaeodera) A: CY
flavofasciata tschitscherini Semenov, 1895a: 134 (Acmaeodera) A: KI KZ TD TM UZ
sterbai Obenberger, 1934c: 38 (Acmaeodera)

glasunovi Semenov, 1895d: 265 (Acmaeodera) A: KI KZ TD TM UZ
judinae Stepanov, 1954b: 1307 (Acmaeodera)
varsobica Stepanov, 1958a: 112 (Acmaeodera)

impunctata Abeille de Perrin, 1891: 279 (Acmaeodera) A: IN IQ TR
medvedevi Volkovitsh, 2006a: 38 A: SHX
mimonti decorata Marseul, 1865: 299 (Acmaeodera) E: AB AR A: TR
mimonti mimonti Boieldieu, 1865: 5 (Acmaeodera) E: AB AL AR BU CT GG GR HU IT KZ MC RO ST UK A: IN
KZ TM TR WS
arenicola Csiki, 1905: 578 (Acmaeodera)
latisquammis Obenberger, 1914h: 254 (Acmaeodera)

mongolorum Volkovitsh, 1989b: 302 A: MG
repetekensis Obenberger, 1934o: 234 (Acmaeodera) A: KZ TM UZ
troniceki Obenberger, 1935h: 105 (Acmaeodera) A: KZ TD TM UZ
zarudniana Volkovitsh, 1977a: 45 A: IN TM
subgenus Euacmaeoderella Volkovitsh, 1979b: 352 type species Buprestis gibbulosa Ménétriés, 1832
adamantina Reitter, 1890o: 343 (Acmaeodera) A: AF SY TD TM UZ
alepidota Volkovitsh, 1977a: 53 A: TM
alfierii Théry, 1929b: 115 (Acmaeodera) N: EG
andresi Théry, 1929b: 118 (Acmaeodera) N: EG
antoinei Baudon, 1956a: 129 (Acmaeodera) N: MO
ballioni Ganglbauer, 1888: 196 (Acmaeodera) A: AE AF IN KZ OM TD TM UZ
xerxes Obenberger, 1916d: 236 (Acmaeodera)

brandli Volkovitsh, 1981: 847 A: IN SY TR
candens Volkovitsh, 1977a: 61 A: TD TM
canescens Semenov, 1895d: 260 (Acmaeodera) A: IN KZ TM UZ

448

BUPRESTIDAE

tamerlani Obenberger, 1934j: 112 (Acmaeodera)

chusistanica Obenberger, 1934o: 288 (Acmaeodera) A: IN
coelestina Volkovitsh, 1977a: 48 A: TM
cyanipennis cyanipennis P.H. Lucas, 1846d: 144 (Acmaeodera) E: FR SP N: AG MO TU
coelicolor Abeille de Perrin, 1895e: cclxxiv (Acmaeodera) [RN]
virgo Abeille de Perrin, 1895c: cxxi (Acmaeodera) [HN]

cyanipennis hispana Abeille de Perrin, 1900a: 12 (Acmaeodera) E: PT SP
dilatatisquamis Obenberger, 1928b: 18 (Acmaeodera) A: TM UZ
dubia Ballion, 1871a: 350 (Acmaeodera) A: AF IN KI KZ TD TM UZ
chalcopracta Obenberger, 1940k: 142 (Acmaeodera)
chloe Obenberger, 1940k: 143 (Acmaeodera)
krali Obenberger, 1923b: 26 (Acmaeodera)
pamirigena Obenberger, 1940k: 141 (Acmaeodera)
steppicola Obenberger, 1940k: 144 (Acmaeodera)

gibbulosa Ménétriés, 1832: 153 (Buprestis) E: AB AR BU GR MC ST TR UK A: CY IN IQ IS JO KZ LE SY TM TR
araxigena Obenberger, 1940k: 141 (Acmaeodera)
corsica Obenberger, 1922a: 19 (Acmaeodera)
cuprinula Reitter, 1890o: 341 (Acmaeodera)
lederi Obenberger, 1934o: 276 (Acmaeodera)
lugens Gory, 1840: 45 (Acmaeodera)
pseudocuprinula Obenberger, 1940k: 142 (Acmaeodera)
strumosa Abeille de Perrin, 1895c: cxviii (Acmaeodera)
tenuifrons Obenberger, 1924b: 12 (Acmaeodera)

hamadanica Volkovitsh, 1983: 567 A: IN TR
henoni Abeille de Perrin, 1894g: ccclv [=1894a: 48] (Acmaeodera) N: AG MO
holocyanea Volkovitsh, 2006a: 40 N: MO
insueta Volkovitsh, 1977a: 55 A: IN TD TM UZ
iranica Obenberger, 1934o: 223 (Acmaeodera) A: AF IN KZ TM
jezeki Bílý, 1983a: 52 A: IN
judeorum Obenberger, 1914h: 253 (Acmaeodera) A: IS JO SY
kubani Volkovitsh, 1987a: 59 A: TD TM UZ
lanuginosa baudoni Cobos, 1958: 242 N: MO
lanuginosa lanuginosa Gyllenhal, 1817: 125 (Buprestis) E: CR FR GR IT MA PT SP N: AG LB MO TU A: IQ IS JO
SI SY
moltonii Schatzmayr, 1949: 27 (Acmaeodera)
ovis Chevrolat, 1838b: 51 (Acmaeodera)

lanuginosa mogadorica Cobos, 1958: 241 N: MO
lanuginosa reducta Schaefer, 1938a: 80 (Acmaeodera) E: FR (Corse) IT (Sardegna)
leucotricha Obenberger, 1940k: 144 (Acmaeodera) A: TM
mauritanica P.H. Lucas, 1844a: 87 (Acmaeodera) N: AG MO
nigrentis Volkovitsh, 1979a: 75 A: UZ
nivetecta Volkovitsh, 1976b: 637 A: AF IN TD TM UZ

BUPRESTIDAE
nivifera cheopis Obenberger, 1934o: 222 (Acmaeodera) N: EG A: IS SI
mochii Lotte, 1943: 152 (Acmaeodera)

nivifera nivifera Abeille de Perrin, 1894e: lxv (Acmaeodera) N: AG TU
oblonga Volkovitsh, 1977a: 60 A: TD or UZ (Fergana Valley)
obscura Reitter, 1889f: 281 (Acmaeodera) E: AB AR A: IN IQ IS LE SA SY TM TR
cyaniventris Reitter, 1890o: 344 (Acmaeodera)
kalalae Obenberger, 1935h: 108 (Acmaeodera)
nigroopaca Cobos, 1963: 357

olivacea Abeille de Perrin, 1904a: 223 (Acmaeodera) A: IN
opacicollis Abeille de Perrin, 1900a: 12 (Acmaeodera) A: KZ UZ
pellitula Reitter, 1890o: 342 (Acmaeodera) E: AB AR
perroti perroti Schaefer, 1949b: 493 (Acmaeodera) E: FR
perroti franzinii Curletti & Magnani, 1987: 173 E: IT (Sardegna)
personata Semenov, 1896e: 378 (Acmaeodera) A: KZ MG TM UZ
pseudonivetecta Volkovitsh, 2011b: 189 A: AE IN OM
richteri Volkovitsh, 1976b: 639 A: TD
safavii Volkovitsh, 1981: 844 A: IN TR
semiviolacea cyaneomixta Semenov, 1895d: 264 (Acmaeodera) A: KI TD TM UZ
semiviolacea semiviolacea Semenov, 1895d: 263 (Acmaeodera) A: KZ UZ
solskyi Obenberger, 1934o: 223 (Acmaeodera) A: TM UZ
squamosa Théry, 1914: 131 (Acmaeodera) N: EG TU A: IQ IS
isis Obenberger, 1934o: 222 (Acmaeodera)
squamidorsis Obenberger, 1926a: 70 (Acmaeodera) [RN]

staudingeri Abeille de Perrin, 1900a: 10 (Acmaeodera) A: AF KI KZ TD TM UZ
ganglbaueri Obenberger, 1940k: 140 (Acmaeodera)

strandi Obenberger, 1918b: 20 (Acmaeodera) A: IN TM
subcyanea Reitter, 1890o: 344 (Acmaeodera) E: AB AR BU GG GR A: AF IN TM TR
cyrta Iablokoff-Khnzorian, 1966: 309 (Acmaeodera)
laticornis Abeille de Perrin, 1900a: 11 (Acmaeodera)

syrdarjensis Obenberger, 1934o: 285 (Acmaeodera) A: KI KZ UZ
theryana Abeille de Perrin, 1900a: 11 (Acmaeodera) A: IN
tragacanthae kopetdagica Volkovitsh, 1987a: 56 A: TM
tragacanthae tragacanthae Volkovitsh, 1987a: 52 E: AB AR
valentinae Volkovitsh, 1977a: 56 A: KI KZ TD TM UZ
vaulogeri muscula Obenberger, 1934o: 277 (Acmaeodera) N: AG
vaulogeri vaulogeri Abeille de Perrin, 1893: 133 (Acmaeodera) E: SP N: AG MO TU
vetusta Ménétriés, 1832: 152 (Buprestis) E: AB AR BU GR A: CY IN IQ IS JO LE SY TM TR
cuprifera Laporte & Gory, 1835: 25 (Acmaeodera)
producta Laporte & Gory, 1835: 26 (Acmaeodera)

villosula Steven, 1830: 161 (Buprestis) E: AB AR GG GR MC N: EG A: AF CY IN IQ IS JO LE SY TD TM TR
antitauri Obenberger, 1934o: 285 (Acmaeodera)
boryi Brullé, 1832: 135 (Buprestis)

449

450

BUPRESTIDAE

cyanescens Gory, 1840: 46 (Acmaeodera)
hyacinthina I. Frivaldszky, 1845: 180 (Buprestis)
jakowlewi Semenov, 1895c: 243 (Acmaeodera)
steveni Faldermann, 1838: 68 (Acmaeodera) [RN]
tiberiadis Abeille de Perrin, 1891: 280 (Acmaeodera)

subgenus Kocheridia Cobos, 1958: 238 type species Acmaeodera trifoveolata Lucas, 1846
trifoveolata Lucas, 1846b: pl. 15 (Acmaeodera) N: AG LB MO TU A: IS
subgenus Liogastria Volkovitsh, 1979b: 349 type species Buprestis virgulata Illiger, 1803
bolivari Obenberger, 1934o: 254 (Acmaeodera) N: AG MO
cerastes Abeille de Perrin, 1900a: 12 (Acmaeodera) N: AG MO
chobauti Théry, 1930a: 113 (Acmaeodera) N: AG
chrysanthemi Chevrolat, 1854a: 394 (Acmaeodera) E: AB AR BU GR TR A: CY IN IQ IS JO LE SA SY TR
klapaleki Obenberger, 1924b: 12 (Acmaeodera)
noemi Obenberger, 1940k: 137 (Acmaeodera)
rudepilosa Obenberger, 1940c: 20 [=1940k: 136] (Acmaeodera)
syriaca Obenberger, 1924b: 13 (Acmaeodera) [HN]
syriae Obenberger, 1926a: 70 (Acmaeodera) [RN]
tonstrix Reitter, 1895d: 154 (Acmaeodera)

confusissima Cobos, 1966a: 67 N: AG MO
elegans elegans Harold, 1869d: 124 (Acmaeodera) [RN] N: CI
ornata Wollaston, 1864: 207 (Acmaeodera) [HN]

elegans kairouanensis Lotte, 1938b: 150 (Acmaeodera) [RN] N: AG LB MO TU
rufipes Reitter, 1895d: 153 (Acmaeodera)
simillima Théry, 1930a: 114 (Acmaeodera) [HN]

elegans miribella Obenberger, 1946: 16 (Acmaeodera) A: IS JO SY
eucera Obenberger, 1946: 14 (Acmaeodera)

fossulicollis Escalera, 1914a: 197 (Acmaeodera) N: MO
maculipennis maculipennis Pic, 1897b: 203 (Acmaeodera) N: AG EG MO TU A: IS
cairensis Obenberger, 1923b: 27 (Acmaeodera)

maculipennis susica Escalera, 1913a: 33 (Acmaeodera) N: MO
paradoxa Escalera, 1914a: 198 (Acmaeodera) N: AG MO
piciella Obenberger, 1926a: 57 (Acmaeodera) [RN] N: AG
suturalis Pic, 1895a: 12 (Acmaeodera) [HN]

pseudovirgulata Volkovitsh & Bílý, 1979: 335 A: CY
rubroornata Escalera, 1914a: 194 (Acmaeodera) N: MO
rufomarginata Lucas, 1846b: pl. 15 (Acmaeodera) E: PT SP N: AG MO TU
lateralis Reitter, 1890o: 341 (Acmaeodera) [HN]
ordinata Abeille de Perrin, 1894f: cxxxi (Acmaeodera) [RN]
regularis Abeille de Perrin, 1894g: cccliv [=1894a: 48] (Acmaeodera) [HN]
reitteri Kerremans, 1892a: 179 (Acmaeodera) [RN]

sefrensis Pic, 1895a: 12 (Acmaeodera) N: AG EG MO TU A: IS
stramineipennis Chobaut, 1922: 25 (Acmaeodera)

BUPRESTIDAE

451

vazquezi Cobos, 1966a: 68 N: MO
virgulata Illiger, 1803: 235 (Buprestis) E: FR IT PT SP N: AG EG LB MO TU
alesi Obenberger, 1936e: 74 (Acmaeodera)
flavovittata Lucas, 1844a: 89 (Acmaeodera)
inedita Lotte, 1938b: 150 (Acmaeodera)
lineola Marseul, 1865: 308 (Acmaeodera)
nigra Ragusa, 1893c: 297 (Acmaeodera)
scabiosae Chevrolat, 1859: 381 (Acmaeodera)

subgenus Omphalothorax Volkovitsh, 1979b: 349 type species Buprestis adspersula Illiger, 1803
adspersula adspersula Illiger, 1803: 237 (Buprestis) E: AL BH BU CR FR GR IT MC ME PT SP N: AG EG LB MO
TU A: CY IQ IS SY TR
dermestoides Solier, 1833: 275 (Acmaeodera)

adspersula squamiplumis Peyerimhoff, 1931b: 70 (Acmaeodera) N: AG MO A: IS SI
arabica Cobos, 1963: 360 A: IN IS OM SA
sheherzad Pochon, 1971: 241 (Acmaeodera)

argentea Volkovitsh, 2011b: 188 A: AE IN OM
densisquammis Abeille de Perrin, 1904a: 218 (Acmaeodera) A: IN IS JO SY TR
despecta Baudi di Selve, 1870: 85 (Acmaeodera) N: EG A: CY IN IS JO LE SY TR
polygonalis Obenberger, 1916d: 241 (Acmaeodera)
reflexangula Reitter, 1890o: 346 (Acmaeodera)

domenicoi Volkovitsh, 2013: 80 A: IN
filiformis Reitter, 1904d: 257 (Acmaeodera) A: GAN XIN
gianassoi Curletti & Magnani, 1988: 137 N: AG MO
longissima cypraea Krajčík, 2012a: 5 A: CY
longissima longissima Abeille de Perrin, 1904a: 218 (Acmaeodera) E: GR A: AE IN IQ IS JO TR
akbesiana Escalera, 1904: 224 (Acmaeodera)
amanicola Pic, 1905a: 97 (Acmaeodera)
fatima Pochon, 1971: 242 (Acmaeodera)

marcaisi Descarpentries & Mateu, 1967: 647 (Acmaeodera) N: AG A: IS YE AFR
pectinata Bílý, 1973a: 12

nannorrhopsicola Volkovitsh & Bílý, 1979: 333 A: IN PA
pharao Obenberger, 1923e: 84 (Acmaeodera) N: EG LB TU A: SI
prosopiphaga Volkovitsh, 2013: 81 A: IN PA ORR
samai Magnani, 1995b: 101 N: AG
vayssieresi Cobos, 1984: 70 N: AG
zygophylli Curletti & Magnani, 1988: 139 N: AG
genus Acmaeoderella, specie incertae sedis
lopatini Volkovitsh, 2013: 79 A: IN
genus Microacmaeodera Cobos, 1966d: 310 type species Acmaeodera longicornis Cobos, 1966
subgenus Microacmaeodera Cobos, 1966d: 310 type species Acmaeodera longicornis Cobos, 1966
kucerai Volkovitsh, 2007: 73 A: YUN

452

BUPRESTIDAE

longicornis Cobos, 1966d: 310 (Acmaeodera) A: AF PA
wittmeri Volkovitsh, 1986b: 133 A: KA
genus Xantheremia Volkovitsh, 1979b: 345 type species Acmaeodera koenigi Ganglbauer, 1888
subgenus Paratethya Bílý, 1983a: 43 type species Acmaeodera mazandaranica Bílý, 1983
convoluta Klug, 1829: [no 8] (Buprestis) N: AG EG A: JO OM AFR
fasciata Roth, 1851: 119 (Acmaeodera) N: EG A: YE AFR
cadabae Roth, 1851: 19 (Acmaeodera)
minuta Kerremans, 1898b: 311 (Acmaeodera)

mazandaranica Bílý, 1983a: 43 A: IN PA
subgenus Xantheremia Volkovitsh, 1979b: 345 type species Acmaeodera koenigi Ganglbauer, 1888
brancsiki Obenberger, 1935n: 178 (Acmaeodera) A: KZ TD TM UZ
chivensis Volkovitsh, 1978: 39 (Acmaeodera) A: KZ TD UZ
flavipennis Klug, 1829: [no 9] (Buprestis) N: AG EG LB MO A: JO SI AFR
brunneicostata Alfieri, 1976: 96 (Acmaeodera)

freidbergi Volkovitsh, 2004: 119 A: IS
jelineki Bílý, 1983a: 46 A: IN
kaplini Volkovitsh, 1984: 1576 A: TM
koenigi Ganglbauer, 1888: 193 (Acmaeodera) A: AF IN KZ TD TM UZ
chalcoxantha Brancsik, 1899: 99 (Acmaeodera)
subscalaris Reitter, 1897h: 214 (Acmaeodera)

niehuisi Volkovitsh, 2011a: 51 A: IN TR
pantherina Bílý, 1979b: 216 (Acmaeodera) A: AE IQ IS QA SA
philistina Marseul, 1865: 298 (Acmaeodera) A: IS JO SA SI SY
immaculata Andres, 1920: 72 (Acmaeodera)

prepsli Volkovitsh, 2011b: 185 A: AE IN OM PA SA
steinbergi Volkovitsh, 1978: 36 (Acmaeodera) A: TD TM UZ
volkovitshi Bílý, 1983a: 45 A: IN
tribe Acmaeoderini, species incertae sedis
Acmaeodera beesoni Obenberger, 1928a: 117 A: PA (Punjab)
Acmaeodera gardneri Obenberger, 1928a: 118 A: PA (Punjab)
Acmaeodera inamoena Faldermann, 1835a: 138 E: “Caucasus”
Acmaeodera malvina Gistel, 1857a: 45 E: FR or SP (Pyrenees)
Acmaeodera refleximargo Reitter, 1890o: 346 E: UK (Krym)

tribe Haplostethini LeConte, 1861

genus Ankareus Kerremans, 1894a: 349 type species Ankareus cyanicollis Kerremans, 1894
Obenbergerietta E. Strand, 1942: 392 [RN] type species Sicardia tristis Théry, 1909
Pseudianthe Fairmaire, 1901a: 167 type species Pseudianthe tenebrosa Fairmaire, 1901
Sicardia Théry, 1909b: 447 [HN] type species Sicardia tristis Théry, 1909

453

BUPRESTIDAE
luciphilus Bílý, Curletti & van Harten, 2003: 78 A: YE
genus Mastogenius Solier, 1851: 507 type species Mastogenius parallelus Solier, 1851
Haplostethus LeConte, 1860: 253 type species Haplostethus subcyaneus LeConte, 1860

insperatus Kurosawa, 1972: 617 A: JA TAI
relictus Bílý, 1979b: 216 A: SA YE
taoi Tôyama, 1983: 58 (Haplostethus) A: YUN ORR

tribe Paratrachyini Cobos, 1980

genus Paratrachys E. Saunders, 1873: 523 type species Paratrachys hederae E. Saunders, 1873
subgenus Paratrachys E. Saunders, 1873: 523 type species Paratrachys hederae E. Saunders, 1873
chinensis Obenberger, 1958a: 227 A: YUN ORR
hederae formosana Kurosawa, 1985b: 169 A: TAI
hederae hederae E. Saunders, 1873: 523 A: FUJ GUA GUX HUN JA YUN ORR
hederoides Cobos, 1980: 51 A: YUN
hypocrita cobosi Hołyński, 1992: 132 A: “China”
hypocrita hypocrita Fairmaire, 1889a: 349 (Trachys) A: HKG YUN ORR
mixtipubescens Kurosawa, 1985b: 168 A: JA (Ryukyus)
nigricans nigricans Kerremans, 1890c: ccix (Trachys) A: NP ORR
taboureli Cobos, 1980: 52, 55

princeps chujoi Kurosawa, 1976a: 53 A: JA (Ryukyus)
princeps kasaharai Kurosawa, 1985b: 169 A: JA (Ryukyus)
princeps princeps Kurosawa, 1976a: 52 A: JA (Ryukyus)
sinicola Obenberger, 1958a: 227 A: FUJ YUN ZHE ORR
genus Sponsor Gory & Laporte, 1839: 1 type species Sponsor convexus Gory & Laporte, 1839
subgenus Sponsor Gory & Laporte, 1839: 1 type species Sponsor convexus Gory & Laporte, 1839
Neoptosima Théry, 1905: 35 type species Neoptosima spinosa Théry, 1905

gianassoi G. Novak, 2002a: 39 A: OM YE

tribe Polycestini Lacordaire, 1857
subtribe Polycestina Lacordaire, 1857

genus Polycesta Dejean, 1833: 78 type species Buprestis porcata Fabricius, 1775
subgenus Polycesta Dejean, 1833: 78 type species Buprestis porcata Fabricius, 1775
Lycaste Gistel, 1834: 10 type species Buprestis porcata Fabricius, 1775

afghanica Volkovitsh, 1990: 70 A: AF
iranica Obenberger, 1923b: 24 (Pseudocastalia) A: IN PA ORR
tonkinea Fairmaire, 1888a: 345 A: YUN ORR
genus Pseudocastalia Kraatz, 1896b: 84 type species Pseudocastalia benningseni Kraatz, 1896
Svatacesta Zábranský, 2004: 119 type species Strigoptera (Svatacesta) socotra Zábranský, 2004

454

BUPRESTIDAE

arabica arabica Gestro, 1877: 354 (Polycesta) A: AE SA YE AFR
socotra Zábranský, 2004: 119 (Strigoptera) A: YE (Suqutra)
genus Strigopteroides Cobos, 1981: 36 type species Buprestis aegyptiaca Gmelin, 1790 (=Buprestis depressa
Fabricius, 1775)
cypria Mühle & Kakouris, 2013: A: CY
depressa depressa Fabricius, 1775: 219 (Buprestis) N: AG EG A: IN IQ SA SY TR
aegyptiaca Gmelin, 1790: 1928 (Buprestis)
castelnaudi E. Saunders, 1871: 59 (Polycesta) [RN]

depressa turcica G. Novak, 1995: 4 A: TR
hellenica Cobos, 1990: 277 E: GR
margotanae G. Novak, 1995: 1 E: GR (Kríti)
semenowi Obenberger, 1934o: 188 (Pseudocastalia) A: AF IN TD TM UZ
genus Strigoptera Dejean, 1833: 78 type species Buprestis bimaculata Linnaeus, 1758
Castalia Gory & Laporte, 1839: 1 [HN] type species Buprestis bimaculata Linnaeus, 1758

bimaculata Linnaeus, 1758: 662 (Buprestis) A: GUX HAI AUS ORR
annamica Nonfried, 1895: 299 (Castalia)
bipustulata Boisduval, 1835: 83 (Buprestis)
impustulata Gory, 1841: 189 (Castalia)
obscura Gestro, 1877: 354
pacifica Boisduval, 1835: 83 (Buprestis)

genus Thurntaxisia Schatzmayr, 1929: 16 type species Xenopseina alexandri Schatzmayr, 1929
Acassidiocesta Mandl, 1972: 27 type species Polycesta cottyi Fairmaire, 1866

alexandri Schatzmayr, 1929: 17 (Xenopseina) E: GR A: CY IQ
madoni Théry, 1931b: 75 (Polycesta)

cottyi Fairmaire, 1866a: 25 (Polycesta) N: AG MO A: TR

subtribe Xenopseina Volkovitsh, 2008

genus Theryola Nelson, 1997: 318 type species Polycesta touzalini Théry, 1922
touzalini Théry, 1922b: 193 (Polycesta) A: YUN

tribe Polyctesini Cobos, 1955

genus Bellamyina Bílý, 1994: 39 type species Bellamyina cylindrica Bílý, 1994 (=Polyctesis hunanensis
Peng, 1992)
hunanensis Peng, 1992: 387 (Polyctesis) A: SCH SHA YUN
cylindrica Bílý, 1994c: 41

genus Chrysophana LeConte, 1860: 219 type species Ancylocheira placida LeConte, 1854
holzschuhi Bílý, 1984a: 5 A: PA

455

BUPRESTIDAE
genus Polyctesis Marseul, 1865: 264 type species Polyctesis rhois Marseul, 1865
foveicollis Fairmaire, 1889b: 344 A: “China” ORR
hauseri Obenberger, 1934o: 189 A: YUN ORR
johanidesi Bílý, 1997: 15 A: IN TR
rhois Marseul, 1865: 265 A: CY IS LE SY TR
strandi Obenberger, 1934o: 189 A: GUI HEN SCH YUN ZHE

genus Schoutedeniastes Burgeon, 1941: 185 [RN] type species Schoutedenia kerremansi Obenberger, 1924
(=Ptosima magnifica C.O. Waterhouse, 1875)
Schoutedenia Obenberger, 1924c: 15 [HN]

amabilis amabilis Laporte & Gory, 1835: 3 (Ptosima) A: NP ORR
apicata C.O. Waterhouse, 1882: 172 (Ptosima) A: NP ORR
igorrota Heller, 1891a: 160 (Polyctesis) A: TAI ORR

tribe Ptosimini Kerremans, 1903

genus Ptosima Dejean, 1833: 79 type species Buprestis novemmaculata Fabricius, 1775 (=Buprestis undecimmaculata Herbst, 1784)
bowringi C.O. Waterhouse, 1882: 172 A: GUA JIX ZHE
chinensis Marseul, 1867a: 54 A: BEI FUJ GAN GUA GUI GUX HEB HEN HUB HUN JA JIA JIX SHA SHG SHX TAI
YUN “Korea” ORR
bilyi Hołyński, 1993d: 153
elegans Nonfried, 1895: 301
sennae Nonfried, 1895: 301
staudingeri Obenberger, 1914g: 133

undecimmaculata metallescens Bílý, 1982: 186 E: GR (Kríti)
undecimmaculata undecimmaculata Herbst, 1784: 120 (Buprestis) E: AL AU BH BU CR CZ FR GE GG GR HU IT
KO MA MC MD ME PL PT RO SB SK SL SP ST SZ TR UK N: AG EG MO TU A: CY IN IQ IS JO LE SY TR NTRi
confusa A. Villa & G.B. Villa, 1833a: 33 (Buprestis)
cyclops Marseul, 1865: 263
flavoguttata Illiger, 1803: 238 (Buprestis)
intermedia Demaison, 1904: 285
sexmaculata Herbst, 1784: 121 (Buprestis)
sexpunctata Villers, 1789: 338 (Buprestis)
tredecemmaculata Baudi di Selve, 1870: 80
volucris Gistel, 1857a: 70

subfamily Galbellinae Reitter, 1911

genus Galbella Westwood, 1848a: 83 type species Galbella violacea Westwood, 1848
subgenus Galbella Westwood, 1848a: 83 type species Galbella violacea Westwood, 1848
Ianthe Marseul, 1865: 27 [unused original spelling]
Janthe Marseul, 1865: 27 type species Janthe felix Marseul, 1866

456

BUPRESTIDAE

Oncomoea E. Saunders, 1866: 321 type species Oncomoea coerulea E. Saunders, 1866
Sponsorinus Fairmaire, 1900a: 241 type species Sponsorinus lata Fairmaire, 1900

acaciae Descarpentries & Mateu, 1965: 1053 N: AG AFR
atricolor Abeille de Perrin, 1907a: l A: YE AFR
ennediana Descarpentries & Mateu, 1965: 1051 A: OM YE AFR
felix Marseul, 1866: 504 (Janthe) E: GR (Ródos) A: CY IS JO LE SY TR
felicissima Abeille de Perrin, 1897c: 36

harti Janson, 1891: 184 A: IS SA SI
holzschuhi Volkovitsh, 2008: 40 A: IN

subfamily Chrysochroinae Laporte, 1835
tribe Chrysochroini Laporte, 1835

genus Agelia Laporte & Gory, 1835: 5 type species Chrysochroa pectinicornis Laporte & Gory, 1835
subgenus Agelia Laporte & Gory, 1835: 5 type species Chrysochroa pectinicornis Laporte & Gory,
1835
pectinicornis Laporte & Gory, 1835: 5 (Chrysochroa) A: SD UP ORR
cardoni Théry, 1911: 178
shiva Gory, 1840: 52 (Chrysochroa)

subgenus Afrelia Hołyński, 2009: 124 type species Chrysochroa petelii Gory, 1840
lordi Walker, 1871: 13 (Chrysochroa) N: EG A: “Arabia” AFR
aegyptiaca Obenberger, 1924b: 14
analis Kraatz, 1896b: 84
humeralis Théry, 1910b: 374
montana Obenberger, 1924b: 14
morosa Fairmaire, 1887a: 141
nickerleana Obenberger, 1928d: 98
nickerli Obenberger, 1924c: 37
placida Gerstäcker, 1884: 51
ragazzii Gestro, 1895: 343
tricolor Fairmaire, 1884d: lxxiii

genus Catoxantha Dejean, 1833: 75 type species Buprestis bicolor Fabricius, 1778
Epacmene Gistel, 1848b: xi [RN]
Megaloxantha Kerremans, 1903: 47 type species Buprestis bicolor Fabricius, 1778

brunnea hainana Kurosawa, 1991: 207 (Megaloxantha) A: HAI
alia Bellamy, 1998: 92 (Megaloxantha) [RN]
hainana C. Yang & Xie, 1993: 277 (Megaloxantha) [HN]

gigantea Schaller, 1783a: 304 (Buprestis) A: BT NP SD ORR
assamensis J. Thomson, 1879d: lxx

luodiana C. Yang & Xie, 1993: 276 A: GUI (Megaloxantha)

BUPRESTIDAE

457

genus Chrysochroa Dejean, 1833: 75 type species Buprestis fulminans Fabricius, 1787
subgenus Chrooxantha Hołyński, 2009: 171 type species Chrysochroa edwardsi Hope, 1843
edwardsii Hope, 1843c: 109 A: SD ORR
subgenus Chrysochroa Dejean, 1833: 75 type species Buprestis fulminans Fabricius, 1787
Euchloris Billberg, 1820a: 23 type species Buprestis ignita Linnaeus, 1758

alternans Waterhouse, 1888b: 264 A: JA (Ryukyus)
coreana Han & Park, 2012: 92 A: SC
deyrollei E. Saunders, 1867a: 430 [justified emendation] A: PA ORR
fulgidissima adachii Akiyama & Ohmomo, 1998: 4 A: JA (Danjo)
fulgidissima fulgidissima Schönherr, 1817: 229 (Buprestis) [RN] A: JA FUJ GUA GUX HUN JIX TAI “Korea” ORR
coeruleocephala Motschulsky, 1862a: 6
fulgida A.G. Olivier, 1790a: no 32: 32 (Buprestis) [HN]

holstii Waterhouse & Gahan, 1890: 169 A: JA (Ogasawara) ORR
ocellata ocellata Fabricius, 1775: 218 (Buprestis) A: NP ORR
pseudofulgidissima Han & Park, 2012: 95 A: HAI
rajah assamensis Guérin-Méneville, 1847a: 6 A: BT SD ORR
rajah rajah Gory, 1840: 53 A: GUI SCH YUN ORR
chinensis Laporte & Gory, 1835: 7 [HN]

tonkinensis Descarpentries, 1948b: 89 A: HAI ORR
unidentata Fabricius, 1775: 216 (Buprestis) A: AP BT SD ORR
didyma Gory, 1840: 62
levior Kerremans, 1893b: 328
marginata Gory, 1840: 63
mutabilis A.G. Olivier, 1790a: no 32: 7 (Buprestis)
sublineata White, 1843: 343

subgenus Chrysoxantha Hołyński, 2009: 66, 129 type species Buprestis buqueti Gory, 1833
buqueti Gory, 1833b: 61 (Buprestis) A: FUI GUA GUX NP SD YUN ORR
fruhstorferi C.O. Waterhouse, 1904a: 266
mirabilis J. Thomson, 1878b: 14
obliqua Kerremans, 1908: 36
rugicollis E. Saunders, 1866: 300
suturalis Kerremans, 1893c: 504

subgenus Pyranthe Gistel, 1834: 12 type species Buprestis vittata Fabricius, 1775
Charytonia Gistel, 1848a: 128 [RN] [HN]

vittata Fabricius, 1775: 216 (Buprestis) A: SCH YUN ORR
aurea De Geer, 1778: 634 (Buprestis)

genus Demochroa White, 1859b: 6615 type species Catoxantha carinata White, 1859 (=Catoxantha lacordairei
J. Thomson, 1859)
subgenus Demoxantha Hołyński, 2009: 86 type species Demochroa gratiosa Deyrolle, 1864
gratiosa indica Csiki, 1900: 402 A: SD “Himalaya“ ORR
bowringii C.O. Waterhouse, 1904a: 265

458

BUPRESTIDAE

genus Iridotaenia Deyrolle, 1864a: 25 type species Chrysodema sumptuosa Laporte & Gory, 1835
subgenus Euiridotaenia Hołyński, 1997: 182 type species Buprestis cyaniceps Fabricius, 1801
vitalisi Bourgoin, 1922: 20 A: YUN ORR
tonkinea Théry, 1922: 213

subgenus Iridomroczkowskia Hołyński, 1997: 182 type species Iridotaenia sulcifera E. Saunders, 1874
kotoensis Miwa & Chûjô, 1940: 66 A: TAI
subgenus Iridotaenia Deyrolle, 1864a: 25 type species Chrysodema sumptuosa Laporte & Gory, 1835
Iridotoenia Deyrolle, 1864a: 25 [unused original spelling]

hainanensis Kurosawa, 1982: 199 A: HAI
genus Philocteanus Deyrolle, 1864a: 10 type species Chrysochroa leucophthalma Laporte & Gory, 1835
buphthalmus J. Thomson, 1878b: 13 A: SD ORR
leucophthalmus Laporte & Gory, 1835: 18 (Chrysochroa) A: GUX ORR
moricii Fairmaire, 1878b: 270 A: YUN ORR
plutus Laporte & Gory, 1835: 18 (Chrysochroa) A: AP ORR
rubroaureus De Geer, 1778: 63 (Buprestis) A: SD ORR
genus Steraspis Dejean, 1833: 85 type species Buprestis scabra Fabricius, 1775
subgenus Steraspis Dejean, 1833: 85 type species Buprestis scabra Fabricius, 1775
Glyphana Gistel, 1848a: xi [SN]
Phalantha Gistel, 1834: 10 type species Buprestis scabra Fabricius, 1775

ceardi Théry, 1930a: 545 N: AG MO
speciosa arabica C.O. Waterhouse, 1904b: 348 A: AE OM SA YE
speciosa speciosa Klug, 1829: [no 11] (Buprestis) N: AG EG LB A: SI AFR
fulgens J. Thomson, 1879c: 296
intermedia Kerremans, 1891b: clvii
obscura J. Thomson, 1879c: 295

squamosa Klug, 1829: [no 10] (Buprestis) N: AG EG LB MO A: CY IS JO LE SI SY AFR
boyeri Solier, 1833: 269
kindermanni Marseul, 1865: 89
tamariscicola J. Thomson, 1878b: 13

genus Xanthocata Kubáň, nom. nov. [see new Acts] type species Buprestis opulenta Gory, 1832
Catoxantha Solier, 1833: 266 [HN] type species Buprestis opulenta Gory, 1832

bonvouloirii Deyrolle, 1861: 395 (Catoxantha) A: SD ORR
regina Schaufuss, 1863c: 168 (Catoxantha)

tribe Chalcophorini Lacordaire, 1857

genus Chalcophora Dejean, 1833: 77 type species Buprestis mariana Linnaeus, 1758
Anaglyptes Gistel, 1848a: 128 type species Anaglyptes merianus Gistel, 1848 (=Buprestis mariana Linnaeus, 1758)

alternans Abeille de Perrin, 1904a: 212 (Buprestis) A: SY TR

BUPRESTIDAE

459

anachoreta Szallies, 1991: 447 A: TR
detrita detrita Klug, 1829: [no 19] (Buprestis) E: IT GR (Aegean Islands) A: IS LE SY TR
detrita marani Obenberger, 1935a: 8 E: AL BU GR TR
detrita margotana G. Novak, 2002b: 45 A: CY
intermedia fagniezi Schaefer, 1936c: 239 E: FR SP
intermedia intermedia Rey, 1890b: 172 (Buprestis) E: AL BH CR FR (Corse) GR IT ME ST UK (Krym) A: TR
proscheki Obenberger, 1935a: 7

japonica bourgoini Obenberger, 1935a: 6 A: YUN ORR
japonica chinensis Schaufuss, 1879a: 480 A: ANH FUJ GUA GUX HEN HKG HUB HUN JIA JIX SCH ZHE
sinica Jakobson, 1913: 780 (Buprestis)

japonica japonica Gory, 1840: 81 (Buprestis) A: HEI JA NC SC SHN
japonica kumejimana Kurosawa, 1974b: 173 A: JA (Ryukyus)
japonica miwai Kurosawa, 1974b: 173 A: JA (Ryukyus) GUA HAI TAI
japonica oshimana Schönfeldt, 1890: 172 A: JA (Ryukyus)
japonica takarajimana Kurosawa, 1974b: 172 A: JA (Ryukyus)
mariana Linnaeus, 1758: 409 (Buprestis) E: AL AU BE BH BU BY CR CT CZ DE EN FI FR GE GG GR HU IT LA
LS LT LU MC MD ME NL NR NT PL RO SB SK SL ST SV SZ UK A: ES IN/IQ border (Khosrovi) TR WS
bischofi Pochon, 1966: 66
hiulca Pallas, 1781: 409 (Buprestis)
meriana Gistel, 1848a: 128 (Anaglyptes)
zoufali Obenberger, 1924e: 49

massiliensis Villers, 1789: 339 (Buprestis) E: FR IT PT SP N: AG CI MO TU
florentina Kiesenwetter, 1857: 65

maura Español, 1933: 123 N: AG MO TU
yunnana abnormalis Miwa & Chûjô, 1935: 270 A: JA (Ryukyus)
yunnana formosana Kurosawa, 1974b: 177 A: TAI
yunnana insularis Théry, 1908: 78 A: JA (Ryukyus)
yunnana nakanei Kurosawa, 1974b: 178 A: JA (Ryukyus)
yunnana nonfriedi Obenberger, 1935a: 7 A: FUJ GUA GUI GUX HEN HUB HUN JIX SCH SHA “Tibet”
yunnana satzumae Lewis, 1896: 334 A: JA
yunnana yunnana Fairmaire, 1888a: 24 A: YUN ORR
genus Chalcophorella Kerremans, 1903: 79 type species Buprestis stigmatica Dalman, 1817
subgenus Chalcophorella Kerremans, 1903: 79 type species Buprestis stigmatica Dalman, 1817
morgani Théry, 1925b: 171 (Chalcophora) A: IN IQ TR
stigmatica Dalman, 1817: 119 (Buprestis) E: AL BH BU GR MC TR N: EG A: CY IN IQ IS JO LE SY TR
balcanica Obenberger, 1936j: 175
biimpressa Pic, 1916g: 9
lefebvrei Laporte & Gory, 1836: 17 (Buprestis)
marseuli Garbiglietti, 1867: lxvii (Buprestis)
quadrimaculata L. Redtenbacher, 1850: 47 (Chalcophora)

460

BUPRESTIDAE

quadrinotata Klug, 1829: [no 20] (Buprestis)

subgenus Rossiella Obenberger, 1942: 9 type species Buprestis fabricii P. Rossi, 1794
fabricii P. Rossi, 1794: 100 (Buprestis) E: AL BH BU CR GR IT MC TR A: IN IQ LE SY TR
semiviridis Pic, 1916g: 9

subgenus Stigmatophorella Tôyama, 1986: 192 type species Chalcophora quadrioculata L. Redtenbacher,
1843
bagdadensis bagdadensis Laporte & Gory, 1836: 125 (Buprestis) E: AR A: IN IQ SY TR
amarensis Obenberger, 1942: 8
aureoscripta Mandl & Pochon, 1957: 325
berhauti Mandl & Pochon, 1957: 323

bagdadensis freyi Obenberger, 1942: 8 A: IN
escalerae Abeille de Perrin, 1904a: 212 (Buprestis) A: IN
orientalis Obenberger, 1924b: 14 A: AF IN PA
quadrioculata L. Redtenbacher, 1843: 982 (Chalcophora) E: GG A: IN IQ SY TR
akbesiana Cobos, 1957: 200

genus Chlorophorella Descarpentries, 1973: 250 type species Chlorophorella gerlingi Descarpentries, 1973
gerlingi Descarpentries, 1973: 251 A: IN
genus Chrysodema Laporte & Gory, 1835: 2 type species Chrysodema sonnerati Laporte & Gory, 1835
subgenus Chrysodema Laporte & Gory, 1835 type species Chrysodema sonnerati Laporte & Gory, 1835
lewisii E. Saunders, 1873: 511 A: JA TAI
oschimana Nonfried, 1895: 297

subgenus Cyalithoides Fisher, 1922: 4 type species Cyalithoides fulgida Fisher, 1922 (=Chrysodema robusta
Deyrolle, 1864)
Mitshekia Hołyński, 1994: 71 type species Buprestis smaragdula A.G. Olivier, 1790

aurostriata E. Saunders, 1866: 302 A: YUN ORR
tonkinea Kerremans, 1909b: 540

eximia berliozi Descarpentries, 1948a: 63 A: TAI (Lanyu)
subgenus Marcsikiella Hołyński, 2014a: 374 type species Buprestis radians Guérin-Ménéville, 1830
dalmanni Mannerheim, 1837: 40 (Evides) A: JA (Ryukyus) TAI ORR
granulosa Kerremans, 1895: 197
manilarum J. Thomson, 1879a: 9
mira Obenberger, 1928a: 147

yasumatsui Kurosawa, 1954a: 29 A: TAI (Lanyu)
genus Nipponobuprestis Tôyama, 1986: 188 type species Chalcophora amabilis Snellen van Vollenhoven,
1864
subgenus Nipponobuprestis Tôyama, 1986: 188 type species Chalcophora amabilis Snellen van Vollenhoven,
1864
amabilis Snellen van Vollenhoven, 1864: 163 (Chalcophora) A: HUN JA “Korea”

461

BUPRESTIDAE
rubrocinctus Peng, 1995: 97 A: HUB JIX
subgenus Nipponobuprestisia Peng, 1995: 99 type species Chalcophora querceti E. Saunders, 1873
bilyi Peng, 1995: 101 A: JIX
guangxiensis Peng, 1995: 99 A: GUX ORR
orientalis Peng, 1995: 100 A: FUJ
querceti E. Saunders, 1873: 510 (Chalcophora) A: CE JA SE
genus Sapaia Bílý, 1994a: 8 type species Sapaia brodskyi Bílý, 1994
brodskyi Bílý, 1994a: 10 A: YUN ORR
genus Tamamushia Miwa & Chûjô, 1935: 274 type species Tamamushia virida Miwa & Chûjô, 1935
virida fujitai Ohmomo & Karube, 2004: 95 A: JA (Ogasawara)
virida virida Miwa & Chûjô, 1935: 275 A: JA (Ogasawara)

tribe Dicercini Gistel, 1848

genus Capnodis Eschscholtz, 1829: 9 type species Buprestis tenebricosa A.G. Olivier, 1790
Caloptera Gistel, 1834: 10 type species Buprestis tenebrionis Linnaeus, 1760

anthracina Fischer von Waldheim, 1830: 184 (Buprestis) A: IN IQ TD TM
anthracina Faldermann, 1835a: 140 [HN]
bactriana Semenov, 1891: 334
cribricollis Abeille de Perrin, 1896b: 270

carbonaria Klug, 1829: [no 16] (Buprestis) E: AB AR BH BU GG GR MC ST UK (Krym) N: EG A: CY IS JO LE
SY TR
abeillei Obenberger, 1926b: 202 [RN]
cribricollis Abeille de Perrin, 1904a: 212 [HN]
lefebvrei Laporte & Gory, 1836: 4

cariosa cariosa Pallas, 1776: 708 (Buprestis) E: AB AL AR BH BU CR GG GR IT MC ME RO ST TR UK N: EG A:
CY IQ IS JO LE SY TR
bruttia V. Petagna, 1787: 22 (Buprestis)

cariosa hauseri Obenberger, 1928c: 67 A: IN IQ TR
iranica Bogatchev, 1947: 17

excisa alfierii Théry, 1929b: 120 N: EG A: IS OM SA SI
aericolor Blair, 1931: 271

excisa excisa Ménétriés, 1848: 28 E: AB AR A: AE IN IQ KU KZ PA SA TD TM TR UZ
henningii Faldermann, 1835a: 141 E: AB AR A: IN IQ TM TR SY
indica J. Thomson, 1879b: 176 A: HP NP PA UP “Central Asia” “Kashmir”
fuksai Obenberger, 1924b: 15
mysteriosa Obenberger, 1917e: 48
orientalis Abeille de Perrin, 1896b: 270
vermiculata Fairmaire, 1891e: cxxv

jacobsoni Richter, 1952: 159 A: IN IQ TM

462

BUPRESTIDAE

marquardti Reitter, 1913a: 66 A: TR
miliaris metallica Ballion, 1871a: 349 A: AF GAN KA KZ PA TD UZ XIN
afghanica Obenberger, 1934k: 123
melattica Ballion, 1871a: 349 [incorrect original spelling]

miliaris miliaris Klug, 1829: [no 15] (Buprestis) E: AB AR GG GR IT ST A: CY IN IQ IS JO KI KZ LE SY TM TR UZ
daedalea Steven, 1830: 170 (Buprestis)

parumstriata Ballion, 1871a: 349 A: AF IN KI PA TD TM UZ “India”
costulata Fairmaire, 1902a: 39

porosa persica Théry, 1936e: 221 A: IN
porosa porosa Klug, 1829: [no 17] (Buprestis) E: AB AL AR BH BU CR GG GR MC ME ST TR A: CY IN IQ IS JO
LE SY TR
hypocrita Géhin, 1855: 55
mannerheimii Faldermann, 1835b: 142

semisuturalis Marseul, 1865: 127 A: IS LE SY TR
perrini Théry, 1936e: 222

sexmaculata Ballion, 1871a: 349 A: AF IN KI KZ PA TD TM UZ “Kashmir”
kashmirensis Fairmaire, 1891e: cxxv

tenebricosa alia Bellamy, 2004: 259 [RN] A: IN
bushirensis Kubáň, 2006: 40 [RN]
iranica Bellamy, 2003: 155 [RN] [HN]
persica Obenberger, 1945: 16 [HN]

tenebricosa bucharica Obenberger, 1945: 18 A: AF IN KA KI KZ TD TM UZ
tenebricosa tenebricosa A.G. Olivier, 1790a: no 32: 62 (Buprestis) E: AB AL AR BH BU CR FR GG GR IT KZ MA
MC MD ME PT RO SP ST TR UK N: AG MO TU A: CY IN IQ IS JO KZ LE SI SY TM TR
alternata Obenberger, 1945: 16
anomala Fairmaire, 1895c: cx
aurosparsa Abeille de Perrin, 1891: 258
lugens Küster, 1846: 52
occidentalis Obenberger, 1945: 18
simulatrix Obenberger, 1916d: 259

tenebrionis Linnaeus, 1760: 213 (Buprestis) E: AB AL AR AU BH BU CR CT CZ FR GE GG GR HU IT KZ MA MC
MD ME PL PT RO SB SK SL SP ST SV TR UK N: AG MO TU A: CY IN IQ IS JO LE SY TM TR
aerea Laporte & Gory, 1836: 9
naevia Gmelin, 1790: 1939 (Buprestis)
tenebrioides Pallas, 1781: 67 (Buprestis)
variegata Goeze, 1777: 596 (Buprestis)

genus Cardiaspis E. Saunders, 1866: 307 type species Cardiaspis mouhotii E. Saunders, 1866
Bellamyola Hołyński, 1993b: 13 [SN]

mouhotii E. Saunders, 1866: 307 A: BT SD ORR

BUPRESTIDAE

463

genus Cyphosoma Mannerheim, 1837: 91 [conserved name] type species Buprestis tatarica Pallas, 1771
Coeculus Gory & Laporte, 1839: 1 type species Buprestis sibirica Fabricius, 1781 (=Buprestis tataricum Pallas, 1771)
[Cyphonota Dejean, 1833: 89, suppressed]

euphraticum euphraticum Laporte & Gory, 1839: 4 (Coeculus) A: IN IQ IS SY
escalerai Obenberger, 1913a: 23, 27 [justified emendation]

euphraticum insulare Kiesenwetter, 1858: 244 (Buprestis) E: AL BU GR IT ME ST TR UK (Krym) A: SY TR
curtulum Abeille de Perrin, 1900a: 4
lanciferum Abeille de Perrin, 1891: 258

lawsoniae kalalae Obenberger, 1929e: 59 A: IQ JO
lawsoniae lawsoniae Chevrolat, 1838a: 56 (Buprestis) E: FR (Corse) IT SP PT N: AG EG LB MO TU
buqueti Laporte & Gory, 1839: 3 (Coeculus)
gravidum Laporte & Gory, 1839: 3 (Coeculus)
lausoniae Chevrolat, 1838a: 55 (Buprestis) [unused original spelling]

lawsoniae orientalis Bílý, 1983a: 55 A: IN ORR
paganettii Obenberger, 1914d: 115 E: GR A: SY TR
tataricum Pallas, 1771: 464 (Buprestis) E: AB AR KZ ST A: GAN IN KI KZ TD TM UZ WS XIN
repandum Ménétriés, 1832: 150 (Buprestis)
sibiricum Fabricius, 1781: 279 (Buprestis)

turcomanicum Kraatz, 1883f: 358 (Coeculus) A: AF IN PA TD TM UZ
fulvovittis Reitter, 1887d: 511 (Coeculus)

veselyi Obenberger, 1925c: 55 E: AR A: IN TR
luristanicum Richter, 1952: 173

genus Dicerca Eschscholtz, 1829: 9 type species Buprestis aenea Linnaeus, 1760
Argante Gistel, 1834: 10 type species Buprestis moesta Fabricius, 1792
Hemidicerca Richter, 1952: 132 type species Dicerca fritillum Ménétriés, 1832
Neoargante Nelson, 2006: 272 [RN] type species Buprestis moesta Fabricius, 1792
Stenuris Kirby, 1837: 154 type species Buprestis lurida Fabricius, 1775

aenea aenea Linnaeus, 1760: 213 (Buprestis) E: AL AU BH BU BY CR CT CZ EN FI FR GE GR HU IT KZ LA LT
MC MD ME NR NT PL PT RO SB SK SL SP ST SV UK N: AG MO A: ES KZ WS
carniolica Fabricius, 1801b: 189 (Buprestis)
oxyptera Pallas, 1781: 70 (Buprestis)
reticulata Fabricius, 1794: 451 (Buprestis)
scabrosa Mannerheim, 1837: 54
subrugosa Paykull, 1799: 218 (Buprestis)

aenea bella Abeille de Perrin, 1891: 259 A: SY TR
aenea chinensis Obenberger, 1929d: 12 A: ES FE HEB SHG TAI
aenea validiuscula Semenov, 1896c: 319 E: AB AR GG A: IN KI KZ TD TM UZ
zicharevi Obenberger, 1928b: 17

aino Lewis, 1893: 328 A: BEI ES FE HEB HEI HUB JA (Hokkaido) JIL KZ LIA MG NMO WS YUN
alni Fischer von Waldheim, 1 824: 197 (Buprestis) E: AL AU BH BU BY CR CT CZ EN FI FR GE GR HU IT LA LT
MC MD ME NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG TU A: KZ TR
fagi Laporte & Gory, 1837: 103 (Buprestis)

464

BUPRESTIDAE

sahlbergi Gistel, 1857a: 12

amphibia Marseul, 1865: 145 E: BH BY IT RO ST UK A: ES FE KZ TR WS
miranda Reitter, 1904a: 24

berolinensis Herbst, 1779: 321 (Buprestis) E: AL AU BH BU BY CR CZ FR GE GR HU IT MC MD ME NL PL RO SB
SK SL SP SZ TR UK A: TR
calcarata Schaller, 1783a: 310 (Buprestis)
calcarata Fabricius, 1801b: 188 (Buprestis) [HN]

chlorostigma Mannerheim, 1837: 55 E: AB AR GG ST UK A: TR
corrugata corrugata Fairmaire, 1902b: 268 A: GUI SCH YUN ORR
corrugata thibetana Hołyński, 2006: 22 A: XIZ YUN
corrugata vitalisi Descarpentries et Villiers, 1963e: 263 A: GUI ORR
fritillum Ménétriés, 1832: 149 E: AB A: IN TM
furcata Thunberg, 1787: 52 (Buprestis) E: AU BY CT CZ EN FI GE HU IT LA LT NR NT PL SK ST SV UK
acuminata Pallas, 1781: 69 (Buprestis) [HN]

herbstii Kiesenwetter, 1857: 40 (Buprestis) E: AU BU CR GG GR IT PL RO
konstantinoui Brandl & Makris, 2011: 54 A: CY
kurosawai Hattori & Akiyama, 1999: 423 A: TAI
moesta Fabricius, 1792b: 206 (Buprestis) E: AL AU BH BU BY CR CT CZ DE EN FI GE GR HU IT LA LT NR NT
PL RO SK SL SP ST SV SZ UK N: CI A: SHN TR WS
quadrilineata Herbst, 1801: 104 (Buprestis)

obtusa Kraatz, 1882f: 112 A: KI KZ TD UZ
scabida Marseul, 1865: 140 E: AB A: IN
tibialis Lewis, 1893: 328 A: BEI FUI JIX JA
latouchei Fairmaire, 1900b: 622

unokichii Hattori, 1991: 57 A: TAI
genus Lampetis Dejean, 1833: 76 type species Buprestis bioculata A.G. Olivier, 1790
subgenus Lampetis Dejean, 1833: 76 type species Buprestis bioculata A.G. Olivier, 1790
bioculata bioculata A.G. Olivier, 1790a: no 32: 65 (Buprestis) N: MO AFR
bioculata oculifera Laporte & Gory, 1836: 45 (Buprestis) N: EG LB AFR
bardii Obenberger, 1927f: 71 (Psiloptera)
farinosa Kerremans, 1903: 96 (Psiloptera)
macalisteri C.O. Waterhouse, 1900c: 219 (Psiloptera)

subgenus Spinthoptera Casey, 1909: 71 type species Psiloptera valens LeConte, 1858 (=Buprestis drummondi
Laporte & Gory, 1836)
Buprestodes Carter, 1924: 23 type species Buprestodes corruscans Catrer, 1924

affinis E. Saunders, 1866: 305 A: SD ORR
arabica Gahan, 1895: 287 A: OM SA YE
sancta Théry, 1922b: 249

argentata Mannerheim, 1837: 61 (Buprestis) E: AB AR A: AF IN KZ TD TM UZ XIN
catenulata catenulata Klug, 1829: [no 24] (Buprestis) N: EG AFR

BUPRESTIDAE

465

catenulata chopardi Obenberger, 1950: 193 (Psiloptera) N: LB AFR
fastuosa Fabricius, 1775: 216 (Buprestis) A: KA SD ORR
caerulea A.G. Olivier, 1790a: no 32: 21 (Buprestis) [HN]
coerulescens Herbst, 1801: 192 (Buprestis) [RN]
japanensis Obenberger, 1914e: 113 (Psiloptera)
variegata Carter, 1924: 24 (Buprestodes)

orientalis Laporte & Gory, 1836: 86 (Buprestis) A: SD ORR
persica Kerremans, 1910: 170 A: IN
mimosae Klug, 1829: [no 23] (Buprestis) E: GR (Ródos) N: AG EG LB MO TU A: AE IN IS JO LE OM PA SA SI SY
TR YE AFR
catenulata Laporte & Gory, 1837: 82 (Buprestis) [HN]
composita Marseul, 1865: 104 (Psiloptera) [RN]
cyanea Alfieri, 1976: 99 (Psiloptera) [HN] N: EG
interrupta Laporte & Gory, 1837: 81 [ICZN 2002: 211] (Buprestis)
kubani Bellamy, 2007: 82 [RN]
rugosa Palisot de Beauvois, 1807: 44 (Buprestis) [HN]

svobodai Krajčík, 2009: 7 A: AE OM SA YE
viridicuprea E. Saunders, 1866: 304 A: AP SCH YUN ORR
cambodiensis J. Thomson, 1879a: 13

genus Latipalpis Solier, 1833: 11 type species Buprestis plana A.G. Olivier, 1790
subgenus Latipalpis Solier, 1833: 11 type species Buprestis plana A.G. Olivier, 1790
cypria Niehuis, 2005a: 9 A: CY
margotana G. Novak, 1990: 82 E: GR (Kríti)
plana berythensis G. Novak, 1990: 81 A: IN LE SY TR
plana plana A.G. Olivier, 1790a: no 32: 93 (Buprestis) E: IT FR SP
gallica Abeille de Perrin, 1891: 257 (Psiloptera)
pisana P. Rossi, 1794: 101 (Buprestis)

subgenus Palpilatis Bílý, 1980a: 50 type species Dicerca plasoni Reitter, 1888
johanidesi Niehuis, 2002b: 106 A: IN IQ TR
persica Bílý, 1980a: 52 A: IN
plasoni Reitter, 1888a: 70 (Dicerca) A: TR SY
tauricola Pic, 1916g: 9 (Dicerca)

stellio Kiesenwetter, 1857: 31 (Buprestis) E: BU CR GR A: TR SY
latipennis Obenberger, 1914g: 130

genus Perotis Dejean, 1833: 77 type species Buprestis lugubris Fabricius, 1777
Aurigena Laporte & Gory, 1836: 1 type species Buprestis lugubris Fabricius, 1777

chlorana Laporte & Gory, 1836: 3 (Aurigena) A: IS JO LE SY
viriditarsis Schaufuss, 1879a: 480 (Psiloptera)

cuprata Klug, 1829: [no 18] (Buprestis) E: AB AR A: IN IQ JO LE SY TR

466

BUPRESTIDAE

capnodiformis Reitter, 1883b: 114 (Aurigena)
cuprea Hampe, 1852: 311
orientalis Laporte & Gory, 1836: 4 (Aurigena)

lugubris aereiventris Reiche, 1861b: 4 (Aurigena) A: IS LE SY TR
lugubris longicollis Kraatz, 1880c: 142 E: AB AR GG A: IN IQ TM TR
subcostata Reitter, 1889f: 281 (Aurigena)

lugubris lugubris Fabricius, 1777: 236 (Buprestis) E: AL AU BH BU BY CR CZ GG GR HU IT MC MD ME RO SB
SK SL ST TR UK A: TR (western part)
europaea Abeille de Perrin, 1896b: 273 (Aurigena)
laportea Brullé, 1832: 133 (Buprestis)
prolongata Obenberger, 1918b: 21
punctulata Abeille de Perrin, 1896b: 273 (Aurigena)

lugubris meridionalis Izzillo & Sparacio, 2011: 154 E: IT
lugubris mutabilis Abeille de Perrin, 1896b: 273 (Aurigena) A: TR (Cilician Taurus mts)
lugubris transcaspica Semenov, 1891: 332 (Psiloptera) A: IN TM
guttipennis Abeille de Perrin, 1904a: 213

margotana G. Novak, 1983b: 112 (Aurigena) E: GR (Kríti)
susannae G. Novak, 1983b: 113 (Aurigena) A: CY
unicolor A.G. Olivier, 1790a: no 32: 63 (Buprestis) E: IT (Marettimo) FR PT SP N: AG MO TU
igniventris Escalera, 1914a: 199
lyonii Vigors, 1826: 514 (Buprestis)
planidorsis Liskenne, 1994: 322 (Aurigena)
striata Spinola, 1837: 111
tarsata Fabricius, 1792b: 208 (Buprestis)
tersata Fabricius, 1792b: 208 (Buprestis) [incorrect original spelling]

xerxes malachitica Abeille de Perrin, 1896b: 273 (Aurigena) A: IS LE SY TR
xerxes xerxes Marseul, 1865: 109 (Psiloptera) E: GR (Karpathos, Ródos) A: IN TR
xerces Marseul, 1865: 109 (Psiloptera) [incorrect original spelling]

genus Phelix Marseul, 1865: 134 type species Coeculus teter Laporte & Gory, 1839
teter Laporte & Gory, 1839: 5 (Coeculus) N: “Nubia”
genus Tokaranodicerca Hattori, 2004: 140 type species Dicerca nishidai Toyama, 1986
nishidai Tôyama, 1986: 19 (Dicerca) A: JA (Ryukyus)
shimonoi Hattori, 2005: 444 A: JA (Ryukyus)
genus Touzalinia Théry, 1922b: 256 type species Touzalinia psilopteroides Théry, 1922
psilopteroides psilopteroides Théry, 1922b: 256 A: YUN
genus Zoolrecordia Hołyński, 2006: 7 [RN] type species Dicercomorpha cupreomaculata E. Saunders, 1866
Tristria Hołyński, 2001a: 132 [HN] type species Dicercomorpha cupreomaculata E. Saunders, 1866

467

BUPRESTIDAE
cupreomaculata E. Saunders, 1866: 306 (Dicercomorpha) A: YUN ORR

tribe Evidini Tôyama, 1987

genus Evides Dejean, 1833: 77 type species Buprestis elegans Fabricius, 1781
elegans Fabricius, 1781: 277 (Buprestis) A: NP ORR

tribe Paratassini Bílý & Volkovitsh, 1996

genus Paratassa Marseul, 1882: 167 type species Sphenoptera caroli Marseul, 1882 (=Sphenoptera ­coraebiformis
Fairmaire, 1875)
acuminata Bílý & Volkovitsh, 1996: 344 N: IS SI
aegyptiaca Bílý & Volkovitsh, 1996: 344 N: EG
agadeziaca Bílý, 2004: 11 N: MO
arabica Bílý & Volkovitsh, 1996: 345 A: SA
aurulenta Bílý & Volkovitsh, 1996: 343 N: AG
coraebiformis Fairmaire, 1875a: 511 (Sphenoptera) N: AG LB TU
caroli Marseul, 1882: 168 (Sphenoptera)

medioatlassica Bílý & Volkovitsh, 1996: 340 N: MO
meridionalis Bílý & Volkovitsh, 1996: 340 N: AG
occidentalis Bílý & Volkovitsh, 1996: 339 N: MO
orientalis Bílý & Volkovitsh, 1996: 345 A: AE IN OM
ringenbachi Liberto & Gigli, 2005: 58 N: LB
tunesiaca Bílý & Volkovitsh, 1996: 343 N: TU

tribe Poecilonotini Jakobson, 1913

genus Lamprodila Motschulsky, 1860a: 11 [RN] type species Buprestis rutilans Fabricius, 1777
subgenus Lamprodila Motschulsky, 1860a: 11 [RN] type species Buprestis rutilans Fabricius, 1777
Castalia Laporte & Gory, 1836: 114 [HN] type species Buprestis rutilans Fabricius, 1777
[Dendrochariessa Gistel, 1848: ix, suppressed name]
Lampra Dejean, 1833: 78 [HN] type species Buprestis rutilans Fabricius, 1777
Scintillatrix Obenberger, 1956a: 41 [RN]

amurensis Obenberger, 1924b: 19 (Lampra) A: FE NE NC
rambouseki Obenberger, 1929c: 122 (Lampra)

decipiens achaica Brandl, 1986: 2 (Scintillatrix) E: GR
decipiens decipiens Gebler, 1847: 407 (Poecilonota) E: AB AR AU BU BY CR CT CZ EN FR GE GR HU IT LA LS
LT MD NT PL RO SK SL ST SZ UK N: EG A: ES IN KI KZ TM TR UZ WS
dives Guillebeau, 1889: 7 (Poecilonota)
modesta Guillebeau, 1889: 5 (Poecilonota)

decipiens kamikochiana Obenberger, 1940k: 163 (Lampra) A: FE (Kunashir, Sakhalin) JA
gloriosa Marseul, 1865: 160 (Poecilonota) E: BU CR MC GR TR A: CY IN IQ SY TR
hoschecki Obenberger, 1917e: 51 (Poecilonota) A: ANH FE SCH SHX “Korea”
hoscheki Obenberger, 1917e: 51 (Poecilonota) [unused original spelling]

468

BUPRESTIDAE

iranica Obenberger, 1952e: 333 (Lampra) A: IN TR
klapaleki kairouzi Magnani, Brandl & Baiocchi, 2001: 202 (Ovalisia) A: LE
klapaleki klapaleki Obenberger, 1924b: 17 (Lampra) E: GG A: SY TR
kucerai Bílý, 1999a: 87 (Palmar) A: YUN
limbata Gebler, 1832: 41 (Buprestis) A: ES FE GAN HEB HEI HEN HUB JIA JIL JIX LIA MG NIN NMO QIN SHA
SHN SHX XIN ZHE
semenoviella Obenberger, 1952e: 368 (Lampra)

lukjanovitshi Richter, 1952: 72 (Lampra) A: ES HEB NMO SHX
mirifica barbarica Hellrigl, 1972: 681 (Lampra) N: AG
mirifica mirifica Mulsant, 1855b: 146 (Lampra) E: AL AU BH BU BY CR CT CZ FR GE GR HU IT KZ MC MD ME
PL PT RO SK SL SP ST TR UK A: KZ TR UZ
mirifica nadezhdae Semenov, 1909: 314 (Lampra) E: AB AR GG A: IN TM
caspica Obenberger, 1930a: 102 (Lampra)
circumciliata Obenberger, 1927b: 82 (Lampra)
montana Obenberger, 1924b: 17 (Lampra)

mirifica vicina Guillebeau, 1889: 3 (Poecilonota) E: AB A: IN TR “Syrie”
caucasica Obenberger, 1934h: 115 (Lampra)

nipponensis Kurosawa, 1953a: 72 (Lampra) A: JA
nobilissima bellula Lewis, 1893: 329 (Poecilonota) A: FE HEI HEN HUN JA JIL LIA SHX “Korea”
charbinensis Obenberger, 1952e: 367 (Lampra)
inexpecta Kurosawa, 1953a: 77 (Lampra) [unused original spelling]
inexspecta Kurosawa, 1953a: 77 (Lampra)
integripennis Obenberger, 1952e: 341 (Lampra)
mandjurica Obenberger, 1934h: 110 (Lampra)
suvorovi Obenberger, 1934h: 109 (Lampra)

nobilissima nobilissima Mannerheim, 1852b: 278 (Lampra) A: ES MG NMO SHA
pretiosa Mannerheim, 1852b: 275 (Lampra) A: BEI FE HEI HUB HUN GUX LIA MG NC NMO SC SCH
SHX YUN
djingischani Obenberger, 1952e: 343 (Lampra)

provosti Fairmaire, 1887d: 321 (Poecilonota) A: BEI FUI HEB JIX NMO TIA SHA ZHE
bourgoini Obenberger, 1930a: 103 (Lampra)
generosa Obenberger, 1952e: 342 (Lampra)
prosternalis Obenberger, 1930a: 104 (Lampra)

rutilans podolica Obenberger, 1952e: 335 (Lampra) E: MD ST UK
rutilans rutilans Fabricius, 1777: 235 (Buprestis) E: AB AL AU BH BU BY CR CT CZ EN FI FR GE GG GR HU IT
LA LS LT MC MD NR NT PL RO SB SK SL SP SV SZ TR UK N: AG
aeruginosa Herbst, 1780: 91 (Buprestis)
fastuosa Well, 1781: 385 (Buprestis)

savioi Pic, 1923c: 14 (Lampra) A: HKG JIA SCH SHG
solieri Laporte & Gory, 1837: 116 (Lampra) E: IT MC SP N: AG MO A: TR
guiraoi Fairmaire, 1855a: 107 (Lampra)

BUPRESTIDAE

469

hieroglyphica Mulsant & Godart, 1855a: 5 (Lampra)

suyfunensis Obenberger, 1934h: 108 (Lampra) A: ES FE HEI JIL MG NC NMO
chinganensis Obenberger, 1940k: 164 (Lampra)

tschitscherini Semenov, 1895a: 130 (Poecilonota) A: FE NE SHA SCH “Korea”
ussuriensis Obenberger, 1924b: 18 (Lampra)

tuerki Ganglbauer, 1882b: 135 (Lampra) E: AB A: IN
subgenus Palmar Schaefer, 1949b: 175 type species Buprestis festiva Linnaeus, 1767
assamensis Stebbing, 1914: 211 (Lampra) A: AP ORR
ardoini Baudon, 1966a: 51 (Ovalisia)
jeanvoinei Descarpentries & Villiers, 1963e: 273 (Ovalisia)

balcanica Kirchsberg, 1876: 29 (Poecilonota) E: AB AL AR BU GG GR MC TR A: IN TR
ottomana Obenberger, 1952e: 322 (Lampra)

beauchenei Fairmaire, 1889a: 346 (Poecilonota) A: FUI YUN ORR
beauchenii Fairmaire, 1889a: 346 (Poecilonota) [incorrect original spelling]
bedeli Obenberger, 1921b: 236 (Lampra)

clermonti Obenberger, 1924b: 15 (Lampra) A: XIZ YUN
cretica Zabransky, 1994: 11 (Palmar) E: GR (Kríti)
cupraria Fairmaire, 1898b: 387 (Poecilonota) A: YUN ORR
cupraria Théry, 1910a: 13 (Lampra) [HN]

cupreosplendens cupreosplendens Kerremans, 1895: 210 (Poecilonota) A: GUI GUX HAI HUB HUN JIX SCH
cupreosplendens kerremansi Obenberger, 1924b: 16 (Lampra) [RN] A: TAI
cupreosplendes Kerremans, 1912c: 203 (Lampra) [HN]

cupreosplendens miribella Obenberger, 1952g: 319 (Lampra) A: FUJ
cupreosplendens obliterata Descarpentries & Villiers, 1963e: 273 (Ovalisia) A: YUN ORR
cupreosplendens rodeti Nonfried, 1895: 298 (Poecilonota) A: ZHE
davidis davidis Fairmaire, 1887b: 120 (Poecilonota) A: ANH FUJ GUI GUX HUB HUN JA JIX SCH YUN TAI ORR
tonkinea Kerremans, 1895: 210 (Poecilonota)

davidis intermedia Kurosawa, 1956: 33 (Lampra) A: JA TAI
tapanensis Hattori, 1996: 90 (Ovalisia)

elongata Kerremans, 1895: 209 (Poecilonota) A: FUI GUA GUI HUN JIX SCH SHA YUN
adonis Obenberger, 1934h: 107 (Lampra)

festiva festiva Linnaeus, 1767: 662 (Buprestis) E: AL AU BH BU CR FR GE GR HU IT MC ME PT RO SL SP SZ TR
N: AG MO TU
bonnairei Fairmaire, 1884c: lxiv (Lampra)
bonnairii Fairmaire, 1884c: lxiv (Lampra) [unused original spelling]
decemmaculata P. Rossi, 1794: 101 (Buprestis)
decempunctata Fabricius, 1798: 136 (Buprestis)

festiva holzschuhi Hellrigl, 1972: 666 (Palmar) A: LE SY TR
igneilimbata Kurosawa, 1946: 4 (Lampra) A: TAI
igneolimdata Kurosawa, 1946: 4 (Lampra) [unused original spelling]

kheili Obenberger, 1925a: 100 (Lampra) A: FUJ HEN JIX YUN

470

BUPRESTIDAE

magnifica Kerremans, 1892b: 171 (Poecilonota) A: AP BT SD ORR
perroti Descarpentries & Villiers, 1963e: 273 (Ovalisia) A: YUN ORR
pseudovirgata Ohmomo, 2005: 227 (Ovalisia) A: TAI
pulchra Obenberger, 1921a: 93 (Lampra) A: YUN ORR
refulgens Obenberger, 1924b: 16 (Lampra) A: GUA GUX HUN JIX ORR
chamensis Descarpentries & Villiers, 1963e: 273 (Ovalisia)

shirozui Ohmomo, 2005: 230 (Ovalisia) A: TAI
subcoerulea Kerremans, 1895: 210 (Poecilonota) A: FUI GUI SD SHA SHX YUN ORR
croesus Obenberger, 1921a: 92 (Lampra)

virgata Motschulsky, 1860h: 490 (Poecilonota) A: ANH FE HEI HUB JA JIX MG NC NMO SC SCH SHA YUN
beata Obenberger, 1952e: 329 (Lampra)
subangulosa Fairmaire, 1889b: 32 (Lampra)

vivata Lewis, 1893: 329 (Poecilonota) A: HEI JA
masudai Kano, 1929: 95 (Poecilonota)

genus Lamprodila, nomen dubium
bella Gory, 1840: 116 (Buprestis) A: TR
genus Poecilonota Eschscholtz, 1829: 9 type species Buprestis variolosa Paykull, 1799
Descarpentriesina Leraut, 1983: 6 [RN]

semenovi Obenberger, 1934n: 148 A: FUJ SCH YUN
variolosa chinensis Théry, 1926c: 155 A: BEI HEB HUB LIA NMO SHA SHN SHX
variolosa cupreomaculata Miwa & Chûjô, 1935: 271 A: JA
variolosa dicercoides Reitter, 1888d: 426 A: FE NC HEI JIL LIA NMO
variolosa populialbae Richard, 1889: no 46: 6 E: AL BH BU CR FR GR IT KZ MC ME PT SP ST TR N: AG MO
TU A: TR
lugdunensis Rey, 1890b: 172 (Lampra)
setulosa Fleischer, 1896: 31
tremulae Abeille de Perrin, 1896b: 275

variolosa yanoi Kurosawa, 1963a: 90 A: JA
variolosa variolosa Paykull, 1799: 219 (Buprestis) E: AB AR AU BH BY CR CT CZ EN FI FR GE GG HU IT LA LT
MD ME NR NT PL RO SB SK SL ST SV SZ UK A: ES KZ MG WS
conspersa Gyllenhal, 1808: 441 (Buprestis)

subfamily Chrysochroinae, nomen dubium
Buprestis plebeja Fabricius, 1777: 236 [may be Perotis or Poecilonota] A: “India”

tribe Sphenopterini Lacordaire, 1857

genus Genestia Théry, 1923a: 94 type species Genestia raffrayi Théry, 1923
subgenus Microgenestia Bellamy, 1986: 319 type species Genestia jakovlevi Obenberger, 1928
mateui Cobos, 1953a: 13 N: AG MO

BUPRESTIDAE
genus Sphenoptera Dejean, 1833: 81 type species Buprestis antiqua Illiger, 1803
subgenus Chiloblemma Obenberger, 1926b: 11 type species Sphenoptera hispidula Reitter, 1890
hispidula Reitter, 1890m: 281 E: AB AR GG
pilosula Obenberger, 1915a: 56 [HN]

subgenus Chilostetha Jakovlev, 1889a: 84 type species Sphenoptera basalis Morawitz, 1861
Chilostethus Nelson, 1982: 446 [unjustified emendation]
Cyphostetha Jakovlev, 1900d: 400 [HN]

acerba Théry, 1929b: 123 N: EG
anchorata Bílý, 1996: 31 [RN] N: AG TU
anchorifera Bílý, 1994c: 327 [HN]

armillata Jakovlev, 1908c: 520 A: TR
basalis basalis Morawitz, 1861a: 168 E: AB BH BU CR CT HU KZ MD RO SB ST UK A: KZ WS
aestivalis Obenberger, 1952d: 90
arcas Obenberger, 1952d: 104
arethusa Obenberger, 1952d: 77
carinivertex Obenberger, 1952d: 93
chankae Obenberger, 1952d: 103
chariessa Obenberger, 1920b: 126
diffusa Obenberger, 1952d: 84
elisa Obenberger, 1952d: 82
hero Obenberger, 1952d: 79
laevigatula Obenberger, 1952d: 82
nogaica Obenberger, 1952d: 105
permixta Obenberger, 1952d: 102
persephone Obenberger, 1952d: 94
phoebia Obenberger, 1952d: 92
phyllis Obenberger, 1952d: 87
pygmaea Jakovlev, 1908c: 518
sareptana Obenberger, 1952d: 83
sublica Obenberger, 1952d: 77
urania Obenberger, 1952d: 90
vernalis Obenberger, 1952d: 86

basalis pirazzolii Obenberger, 1952d: 103 E: FR GR IT SP
astorgae Obenberger, 1952d: 78
schaeferi Obenberger, 1952d: 108

bedeli Abeille de Perrin, 1909: 27 E: SP
berbera Obenberger, 1924g: 115 N: AG MO
pardoi Cobos, 1952: 89

bilyana Niehuis, 2002a: 29 [RN] N: MO
bilyi Niehuis, 2001b: 125 [HN]

boanoi Curletti & Magnani, 1988: 142 N: AG

471

472
brandli Niehuis, 2001b: 128 A: SA
canescens canescens Motschulsky, 1860i: 517 E: KZ A: AF KZ TM UZ
divnogorskii Obenberger, 1952d: 101
sokolovi Jakovlev, 1900d: 434

canescens eximia Jakovlev, 1886: 100 A: AF TM UZ
flora Obenberger, 1952d: 98
karavajevi Obenberger, 1952d: 86

cauta cauta Jakovlev, 1904a: 183 E: AR AU BH BU CR GR HU IT MC SK SP A: IN JO SY TR
damascena Obenberger, 1952d: 58
danubiana Obenberger, 1952d: 58 [RN]
dryadis Obenberger, 1926e: 208
iberica Obenberger, 1952d: 88
maja Jakovlev, 1908c: 516
oeneis Obenberger, 1952d: 88
petriceki Obenberger, 1952d: 57
rambouseki Théry, 1928c: 132 [HN]
wagneri Obenberger, 1926e: 197

cauta palea Obenberger, 1952d: 66 E: KZ ST UK
ussuriensis Obenberger, 1952d: 54
xantho Obenberger, 1952d: 67

chinensis Kerremans, 1898a: 120 A: “China”
chusistanica Obenberger, 1926e: 201 A: IN
cobosi Alexeev, 1989b: 312 [RN] A: MG
tamerlani Cobos, 1968: 379 [HN]

convicta Jakovlev, 1900d: 435 A: IS TR
cribratipennis Escalera, 1914a: 204 E: SP N: AG MO TU A: JO
davidis Théry, 1928c: 131 A: “China”
densesculpta Jakovlev, 1908c: 512 A: ES MG
gengiskani Cobos, 1968: 376
marcopoli Cobos, 1972a: 33

egena Mannerheim, 1852b: 284 A: ES MG WS
carinulata Jakovlev, 1902c: 289

elpha Obenberger, 1926e: 200 A: TR
epistomalis Obenberger, 1927f: 73 E: KZ
eroylandusica Zykov & Alexeev, 1993: 137 A: TM
geghardica Kalashian & Zykov, 1994: 134 E: AB AR
gonyoxys Abeille de Perrin, 1909: 29 N: AG
halperini Niehuis, 2001b: 131 A: IS
hayeki Kalashian & Volkovitsh, 2007: 38 A: IN
impressifrons Fairmaire, 1875a: 511 E: IT (Sardegna) PT SP N: AG EG LB MO TU A: IS AFR
andresi Obenberger, 1924g: 120

BUPRESTIDAE

BUPRESTIDAE
constantinensis Abeille de Perrin, 1900a: 4
dione Jakovlev, 1901a: 176
excisa Jakovlev, 1887a: 68
fernandesi Cobos, 1966b: 123
klugi Obenberger, 1924g: 120
seriola Jakovlev, 1904a: 181

infantula Reitter, 1895a: 37 E: “Araxesthal” [in AB or AR]
insidiosa Mannerheim, 1852b: 282 A: ES MG NC NO
belenois Obenberger, 1952d: 76
erratrix Obenberger, 1920b: 125
forceps Jakovlev, 1908c: 517
improbula Obenberger, 1952d: 75
iphis Obenberger, 1952d: 74
paraegena Cobos, 1972a: 38
placida Cobos, 1972a: 36
pseudoforceps Obenberger, 1952d: 73
shansiana Obenberger, 1952d: 72
subcylindrica Marseul, 1865: 378
zubaci Obenberger, 1952d: 73

jordani Abeille de Perrin, 1909: 26 A: IS
jugoslavica Obenberger, 1926e: 207 E: BH BU CR GR MC TR NARi
verecunda Obenberger, 1952d: 85

kiachtae Obenberger, 1952d: 99 A: ES
krueperi Jakovlev, 1887a: 81 E: GR
laportei E. Saunders, 1871: 88 E: AL BH BU CR FR GR IT MC ME SP A: TR
comita Obenberger, 1952d: 48
sceptrifera Obenberger, 1952d: 55

leventi Kalashian & Volkovitsh, 2007: 39 A: TR
libanica Fairmaire, 1881a: 87 A: LE
liauteyi Obenberger, 1924g: 115 N: AG A: IS
magnanii Curletti & Sparacio, 1990: 121 N: TU
mannerheimii E. Saunders, 1871: 88 [RN] A: MG
heteronyma Obenberger, 1926e: 192 [RN]
laticollis Mannerheim, 1852b: 283 [HN]

marmottani Théry, 1930a: 218 N: AG
lottei Obenberger, 1952d: 68

mingrelica Obenberger, 1926e: 208 E: GG
minutissima Desbrochers des Loges, 1870: 119 E: SP
muehlei Niehuis, 2001b: 135 A: IS SI
muehlheimi Obenberger, 1916d: 254 A: MG NE NO
nana Jakovlev, 1908c: 513 A: NO

473

474

BUPRESTIDAE

notha Abeille de Perrin, 1909: 28 E: SP
paradoxa Abeille de Perrin, 1898: 255 N: AG TU
parvula Fabricius, 1798: 137 (Buprestis) E: SP N: AG MO TU
pilosula Jakovlev, 1887a: 69 E: SP
almeriana Cobos, 1986: 205
castellana Cobos, 1986: 206
gadetinica Verdugo & Coello, 2001: 98
mallorquensis Théry, 1926a: 20
pilosulella Cobos, 1986: 206

plena Obenberger, 1952d: 91 A: MG
popovii Mannerheim, 1852b: 280 A: ES FE MG NO
balthasari Obenberger, 1928c: 68
kaszabiana Cobos, 1968: 373
mongolica Jakovlev, 1887b: 115

puberula Jakovlev, 1887b: 116 E: AB AR A: IN KI KZ TD TM UZ
cataonia Obenberger, 1926e: 209
inderiensis Obenberger, 1927f: 73
rauda Jakovlev, 1908c: 515
seriatosetosa Obenberger, 1926e: 210

puta Marseul, 1865: 388 N: AG MO TU A: IS
robustithoracis Zykov & Alexeev, 1993: 139 A: TM
rotrouiana Cobos, 1965a: 13 N: MO
sikha Obenberger, 1926e: 203 A: “Himalaya”
substriata Krynicki, 1834: 166 E: AU BU CT CZ GR HU KZ MC MD PL RO SB SK ST TR UK
arsilage Obenberger, 1952d: 45
trebinjensis Obenberger, 1916d: 253

syriaca Jakovlev, 1908c: 514 A: IN SY TR
blattnyi Obenberger, 1926e: 202
calosoma Obenberger, 1926e: 203

tezcani Niehuis, 1999b: 43 E: AR A: IN TR
vediensis Kalashian, 1994: 82 E: AR
vestita Jakovlev, 1887b: 117 E: “Caucasus” A: TM
jagdievi Alexeev, 1979a: 73

subgenus Chrysoblemma Jakovlev, 1889a: 84 type species Sphenoptera scovitzii Faldermann, 1835
alexeevi Kalashian & Volkovitsh, 1993: 54 A: KZ
amoena Jakovlev, 1901a: 183 A: IN TM
amplicollis Jakovlev, 1899a: 292 A: KZ TD TM UZ
phryne Jakovlev, 1905: 27
obtusangula Obenberger, 1927g: 23

antoniae Reitter, 1891e: 142 E: AB AR
aradica Niehuis, 2009b: 126 A: IS

BUPRESTIDAE
araxidis Reitter, 1890m: 282 E: “Araxesthal” [in AB or AR]
araxis Fauvel, 1895: 111 [unjustified emendation]

artemisiae Reitter, 1889f: 281 E: AB AR GG A: AF IN IS TM TR
caucasica Jakovlev, 1891b: 138
molitor Reitter, 1890g: 170

batelkai Kalashian, 2014b: 291 A: AE
bellamyi Kalashian, 2014a: 44 A: AR
bifulgida Reitter, 1898a: 12 A: KZ TD TM UZ
chobauti Abeille de Perrin, 1897b: 29 N: AG LB
cornui Théry, 1895b: clviii N: AG
cylindricollis Marseul, 1865: 401 N: AG EG A: IN SA SI
alfredi Reitter, 1900c: 85
henonii Fairmaire & Coquerel, 1866: 26

dumonti Théry, 1922a: 30 N: AG EG TU A: SA
dushakensis Kalashian, 1994: 81 A: TM UZ
eddin eddin Jakovlev, 1903c: 266 A: IN
eddin pakistana Volkovitsh & Kalashian, 2001: 198 A: PA
filistina Obenberger, 1927g: 24 A: SY
formosula Obenberger, 1927d: 27 A: IN
gahani Kerremans, 1913a: 328 A: “Arabia”
glasunovi Jakovlev, 1903c: 273 A: IN
gracilis Jakovlev, 1900d: 443 A: KZ TM
hammadae Kalashian & Volkovitsh, 1997: 360 A: TD
hauseri Reitter, 1895a: 34 A: AF IN KZ TD TM UZ
heroica Jakovlev, 1903c: 270 A: IN
ignita Reitter, 1895a: 33 E: ST KZ A: IN KZ TD TM UZ
aurora Semenov, 1897: 249
prasinula Obenberger, 1937f: 91

israelita Théry, 1930a: 554 A: SI
jakowlewi Reitter, 1895a: 39 E: AB AR
apta Jakovlev, 1903c: 274

kasimi Théry, 1936a: 9 N: EG
kerzhneri Volkovitsh & Kalashian, 2001: 196 A: KZ UZ
khnzoriani Kalashian, 1996: 780 E: AR
limoniastri Théry, 1928a: 9 N: TU
lutulenta Jakovlev, 1905: 30 A: TM
mandarina Théry, 1910a: 4 A: HUB
mauretanica Théry, 1930a: 200 N: TU A: IS SI AFR
medvedevi Kalashian & Volkovitsh, 2006: 133 A: TD
mercedes Obenberger, 1952b: 3 N: MO
mirabilis Kalashian, 2011: 199 A: AE

475

476

BUPRESTIDAE

mutilata Théry, 1929b: 130 N: EG
obenbergeriana Volkovitsh & Kalashian, 2003: 338 [RN] A: “Amu-Darja”
amudarjensis Obenberger, 1952b: 2 [HN]

orichalcea Pallas, 1781: 75 (Buprestis) E: AB AR CT KZ ST A: KI KZ MG NO NW TD TM UZ WS XIN
aciculata Marseul, 1865: 395
astrachanica Reitter, 1890m: 281
australis Gory & Laporte, 1839: 34
chrysis Jakovlev, 1899a: 293
cupraria Mannerheim, 1837: 96
cuprea Ballion, 1878: 291
fossulata Zubkov, 1829: 157 (Buprestis) [HN]
impressicollis Motschulsky, 1860i: 517 [RN]
karelini Faldermann, 1833: 46 (Buprestis)
meyeri Gebler, 1830: 76 (Buprestis)
ostenta Jakovlev, 1908b: 8
phoebas Jakovlev, 1908b: 9
pruinosa Abeille de Perrin, 1891: 267
sinkiangensis Obenberger, 1927f: 74
solskyi Becker, 1867: 108
subaenea Cobos, 1968: 382
wilkinsi Jakovlev, 1887b: 117
zoubkoffii Gory, 1841: 312

ovata Alexeev, 1978: 851 A: TD
parumpunctata parumpunctata Klug, 1829: [no 33] (Buprestis) A: IN IS SA SI AFR
houskai Obenberger, 1946: 27

parumpunctata theryi Pic, 1895b: 78 N: AG EG MO A: SA AFR
pubescens Jakovlev, 1886: 100 A: KZ TD TM UZ
amudarjensis Obenberger, 1929d: 13
anniae Obenberger, 1927g: 26

punctatissima Reitter, 1895a: 41 A: AF KZ TD TM UZ
eos Semenov, 1897: 254
venus Obenberger, 1927g: 25

quezeli Descarpentries & Bruneau de Miré, 1963: 173 N: AG A: IN IS JO SA SI AFR
rotundicollis Gory & Laporte, 1839: 90 N: AG A: IS SI
scabra Gory, 1841: 304

sancta Reitter, 1890m: 280 E: AB AR A: IN TR
quadrisignatula Obenberger, 1937h: 110
roseicollis Obenberger, 1937g: 92
taborskyi Obenberger, 1955a: 15

scovitzii alfierii Obenberger, 1923e: 81 N: AG EG A: AE IS SA SI SY

BUPRESTIDAE
scovitzii scovitzii Faldermann, 1835a: 155 E: AB AR ST A: AF IN IQ IS KZ TD TM TR UZ
angelica Obenberger, 1915a: 55
coelestis Obenberger, 1927g: 21
combinatrix Obenberger, 1920b: 124
latescutata Fairmaire, 1892a: cliii
rhododactyla Semenov, 1897: 251
tedshenigena Obenberger, 1937h: 110
transcaspica Jakovlev, 1893b: 448
vachshenica Stepanov, 1959b: 59

scovitzii splendens Gory & Laporte, 1839: 29 N: MO AFR
staneki Obenberger, 1952b: 3 A: TR
striatipennis Jakovlev, 1885: 132 A: IN KZ MG NW TD TM UZ
cyaneoviridis Cobos, 1968: 385
potanini Jakovlev, 1889a: 86
procera Reitter, 1890m: 277

tamaricis Klug, 1829: [no 30] (Buprestis) A: AF IN IQ IS KI KZ SA SI TD TM UZ
asiatica Gory & Laporte, 1839: 36
dilotti Obenberger, 1929e: 60
filiformis Gory & Laporte, 1839: 39
pseudoignita Alexeev, 1978: 852
walteri Reitter, 1890m: 279

tamarisci beckeri Dohrn, 1866: 249 E: AB AR GG ST UK (Krym) A: AF IN KI KZ MG NW TD TM TR UZ
chrysorrhoea Obenberger, 1927d: 21
potopolskii Stepanov, 1959b: 59
scutellata Obenberger, 1920b: 122
tekkensis Obenberger, 1952b: 4

tamarisci tamarisci Gory & Laporte, 1839: 35 N: EG A: IN “Arabia”
smaragdina Jakovlev, 1902d: 352
sterbai Obenberger, 1927d: 22

theryana Jakovlev, 1903c: 272 N: AG
tigridis Obenberger, 1927d: 22 A: IQ
tomentosa Jakovlev, 1886: 101 A: KZ TD TM UZ
ahngeri Jakovlev, 1900d: 441
scintilla Jakovlev, 1908b: 10

tondui Théry, 1910a: 11 N: AG A: IS
tristicula Reitter, 1895a: 38 E: AB AR A: IN TM TR
elegans Jakovlev, 1900d: 433

tschitscherini Jakovlev, 1900b: 98 E: AB AR A: KZ TM UZ
viridiaurea Kraatz, 1882i: 318 A: AF IN KZ TD TM UZ
cylindracea Reitter, 1897h: 215

477

478
viridiceps Abeille de Perrin, 1891: 267 A: TR
viridula Jakovlev, 1905: 28 A: TM
viridiflua Marseul, 1865: 397 A: TR
zarudniana Volkovitsh & Kalashian, 2003: 332 A: IN
zarudnyi schatinensis Alexeev & Zykov, 1991: 149 E: AR GG A: TR
zarudnyi zarudnyi Jakovlev, 1890b: 99 A: IN
subgenus Cyphostetha Jakovlev, 1893a: 135 type species Sphenoptera coerulea Jakovlev, 1899
Rhaphidochila Jakovlev, 1900d: 445 type species Sphenoptera coerulea Jakovlev, 1899

coerulea Jakovlev, 1899c: 559 E: AR A: AF IN TM TR
nox Jakovlev, 1900d: 445 A: IN
subgenus Deudora Jakovlev, 1899b: 325 type species Buprestis rauca Fabricius, 1787
acuminata Jakovlev, 1899b: 331 A: IN
aeneiventris Jakovlev, 1886: 91 A: TR
aeneomicans Kraatz, 1882i: 317 E: AB AR GG A: AF IN KI KZ LE NW SY TD TM TR UZ
aeneofulgens Obenberger, 1929g: 120
ahriman Obenberger, 1929b: 48
aladaghensis Obenberger, 1920b: 116
amasiae Obenberger, 1929b: 49
baicalensis Obenberger, 1929b: 53
fabichi Obenberger, 1952c: 14
fleischeri Obenberger, 1952c: 15
hoberlandti Obenberger, 1955a: 13
moganensis Obenberger, 1955a: 12
persica Obenberger, 1929g: 121
unidentata Jakovlev, 1891b: 137
ushinskii Obenberger, 1929g: 122
veselyi Obenberger, 1915a: 53

afflicta Jakovlev, 1900d: 406 A: KZ TM UZ
althaeae Obenberger, 1929g: 112 A: IN
ambigua Klug, 1829: [no 28] (Buprestis) A: IN IQ SY
amitina Jakovlev, 1908a: 224 A: TR
antilibanensis Obenberger, 1952c: 7 A: SY TR
nikitaliae Obenberger, 1952c: 16

antoinei Théry, 1930a: 182 N: MO
armena Steven, 1829: 286 (Buprestis) A: IN
astuta Jakovlev, 1899c: 549 A: TR
baumanni Niehuis, 1999c: 51 A: TR
bucharica Jakovlev, 1900d: 422 A: KZ TD TM UZ
bactriana Jakovlev, 1902e: 576

confinis Jakovlev, 1893a: 131 A: SY
conjuncta Jakovlev, 1900d: 416 A: UZ

BUPRESTIDAE

BUPRESTIDAE
costata Jakovlev, 1905: 31 A: TM
curta Jakovlev, 1885: 131 A: IN KZ TD TM UZ
caspica Jakovlev, 1904a: 184
plavilscikovi Obenberger, 1952c: 18

danieli Jakovlev, 1900d: 504 A: TR
finitima Jakovlev, 1908a: 225 A: IN
galobulloni Español & Mateu, 1947: 231 N: MO
gemmata gemmata A.G. Olivier, 1790a: no 32: 90 (Buprestis) E: AB CR FR IT PT SP N: AG EG MO TU
laevis Rey, 1891b: 19
lineata Fabricius, 1775: 217 (Buprestis)
lineola Herbst, 1801: 284 (Buprestis)
raucoides Obenberger, 1920b: 119

gemmata sicelidis Obenberger, 1916d: 251 E: IT (Sicilia)
hamata Jakovlev, 1900b: 101 A: AF KI TD UZ
zarawshanensis Obenberger, 1929b: 51

idonea Jakovlev, 1908a: 223 A: TR
improvisa Jakovlev, 1902e: 575 A: TR
incerta Jakovlev, 1887a: 57 E: AB AR A: IN TR
confusa Kerremans, 1913a: 104
dolens Jakovlev, 1893a: 133
joukli Obenberger, 1929g: 118

inculta Obenberger, 1929b: 42 A: IN
kepelensis Zykov & Alexeev, 1992: 150 A: AF TD TM UZ
khosrovica Kalashian, 1990: 229 E: AR
koenigi Jakovlev, 1891b: 133 A: IN KZ TM
alma Obenberger, 1952c: 11
integripennis Obenberger, 1929g: 116

kryzhanovskii Alexeev, 1991: 859 A: AF
kulzeri Théry, 1942d: 276 A: TR
lederi Jakovlev, 1891b: 134 A: IS
manifesta Jakovlev, 1900d: 428 E: AB AR GG A: IN TR
megarauca Obenberger, 1952c: 21

micans Jakovlev, 1886: 93 E: AR A: IN TR
binominata Kerremans, 1913a: 577
escalerai Obenberger, 1929b: 43
irritatrix Obenberger, 1929b: 37
pseudomicans Obenberger, 1929b: 42
shah Obenberger, 1929b: 37
viridatrix Obenberger, 1929d: 37 [unused original spelling]

misella Jakovlev, 1900d: 415 E: AR GG A: IN TR
callimicra Obenberger, 1929b: 53

479

480

BUPRESTIDAE

liliputana Obenberger, 1929b: 52

mitroshinae Kalashian & Volkovitsh, 2007: 35 A: TD TM
nairae Kalashian & Volkovitsh, 2007: 37 N: AG
orthoglypta M. Antoine, 1953: 208 N: MO
parysatis Obenberger, 1929b: 43 A: IN
isfahanensis Obenberger, 1952c: 8
proserpina Obenberger, 1952c: 10

pierrei Descarpentries, 1958b: 973 N: AG
pseudounidentata Alexeev, 1991: 856 A: AF
radicicola Obenberger, 1929g: 118 A: SY
rauca Fabricius, 1787a: 177 (Buprestis) E: AL BH BU FR GR IT MC ME PT SP TR N: AG EG LB MO TU A: LE
SY TR
africana Jakovlev, 1900d: 402
bassii Gory & Laporte, 1839: 13
bravaisii Gory & Laporte, 1839: 30
carduorum Chevrolat, 1840: 14
celtiberica Gory, 1841: 302
geminata Illiger, 1803: 244 (Buprestis)
heliobia Obenberger, 1929b: 55
metallica Fabricius, 1792b: 210 (Buprestis)
similis Gory & Laporte, 1839: 14
striola Illiger, 1805: 94 (Buprestis)

schimmeli Niehuis, 2005b:143 A: IS TR
schmideggeri Kalashian, 2011: 200 A: AE
sculpticollis Heyden, 1886c: 278 E: AB AR GG A: CY IN IQ IS LE SY TR
akbesiana Obenberger, 1920b: 120
alutaceiceps Obenberger, 1929g: 111
morgani Obenberger, 1929g: 114

segregata Jakovlev, 1900d: 506 E: AB
signata Jakovlev, 1887a: 65 E: AB AR BU GG GR MC A: IN IS SY TM TR
addenda Jakovlev, 1900d: 413
araxicola Obenberger, 1929b: 50
arsinoe Obenberger, 1952c: 22
buresi Obenberger, 1933b: 61
destituta Obenberger, 1929b: 46
detrita Jakovlev, 1902e: 572
gyaurdagensis Obenberger, 1955a: 14
hierosolymitana Obenberger, 1946: 25
jejuna Jakovlev, 1900d: 417
julia Obenberger, 1929b: 50
jureceki Obenberger, 1946: 26

BUPRESTIDAE
obsoleta Jakovlev, 1891b: 136
satelles Jakovlev, 1899b: 331

simplex Jakovlev, 1893a: 130 E: AB AR GR (Thracia, Aegean Islands) A: IN TR
allecta Jakovlev, 1900d: 410
ankarana Obenberger, 1952c: 17
fraterna Jakovlev, 1900d: 408
samotica Obenberger, 1929g: 128

siwaensis Descarpentries, 1954: 165 N: EG
smyrneensis Gory, 1841: 305 E: AB AR A: IN IS LE SY TR
agilis Obenberger, 1952c: 19
anxia Jakovlev, 1900d: 420
captiosa Jakovlev, 1902e: 571
cassia Obenberger, 1929b: 45
congrua Jakovlev, 1900d: 419
delicata Kerremans, 1909a: 279
inconspicua Jakovlev, 1893a: 134
maronita Obenberger, 1929b: 51
migriensis Obenberger, 1916d: 249
nephele Obenberger, 1929b: 43
pseudorauca Obenberger, 1952c: 13
salomonis Obenberger, 1952c: 12
subsulcata Kerremans, 1909a: 282

sobrina Jakovlev, 1886: 97 A: IN TR
sphaerocephala Jakovlev, 1887a: 54 A: SY
splichali Obenberger, 1914c: 34 A: “Himalaya”
subcostata Mulsant, 1851b: 187 A: TR
sulciventris Jakovlev, 1886: 98 A: AF KI KZ TD TM UZ
alaiensis Jakovlev, 1893a: 132
aucta Jakovlev, 1902e: 569
dives Jakovlev, 1902e: 573
dubia Kraatz, 1882i: 315 [HN]
jaxartis Obenberger, 1929b: 36
kraatzi Jakovlev, 1887a: 56 [RN]
kuliabensis Obenberger, 1952c: 9

surobayensis Alexeev, 1991: 860 A: AF
tamerlani Obenberger, 1929g: 127 A: UZ
tenax Jakovlev, 1902e: 577 E: AB “Arax Valley” [in AB or AR]
tibialis Jakovlev, 1886: 88 A: AF “Alai”
subchalybea Kerremans, 1909a: 280

ventrisculpta Obenberger, 1916d: 252 N: AG TU
vittaticollis Lucas, 1844a: 50 N: AG EG LB MO TU A: IS AFR

481

482
aulacophora Jakobson, 1912: 136 [RN]
korfodana Kerremans, 1913a: 213 [RN]
sulcata Marseul, 1865: 367 [HN]

zhelochovtsevi Alexeev, 1978: 850 A: TD
subgenus Hoplistura Jakovlev, 1889a: 84 type species Sphenoptera semenovi Jakovlev, 1889
Hoplandrocneme Semenov, 1899c: 651 type species Sphenoptera flagrans Semenov, 1895
Oplistura Jakovlev, 1889a: 84 [unused original spelling]

alcides Reitter, 1900c: 84 N: IS TU
ardens ardens Klug, 1829: [no 32] (Buprestis) A: IQ SA AFR
ardea Gory & Laporte, 1839: 29
sulcifrons Gory, 1841: 311

arguta Jakovlev, 1900d: 440 A: IN
auricollis Kerremans, 1892b: 191 A: YUN ORR
balassogloi balassogloi Jakovlev, 1885: 131 E: AB AR GG ST A: AF IN IQ KZ TD TM TR UZ
flagrans Semenov, 1896d: 348
morawitzi Semenov, 1897: 248
protracta Jakovlev, 1885: 133
venusta Jakovlev, 1904b: 309

balassogloi seistanica Descarpentries, 1965: 263 A: AF
beesoni Obenberger, 1926b: 22 A: UP
bellatrix Obenberger, 1927g: 185 A: IQ
bergevini Théry, 1922a: 31 N: AG
bogosina Obenberger, 1924g: 119 N: EG AFR
bouvieri Kerremans, 1913a: 445 A: YE
brechteli Niehuis, 2003: 25 N: EG A: IS
brevior Pic, 1896: 140 N: AG MO
crebepunctata Kerremans, 1892b: 189 A: “Himalaya” ORR
cyaniceps Kerremans, 1890c: cciii A: “Himalaya” ORR
decorsei Kerremans, 1913a: 543 N: EG AFR
diabolica Obenberger, 1915a: 55 N: “Egypt”
diffinis Klug, 1835: 30 (Buprestis) N: EG AFR
annexa Jakovlev, 1902d: 322
curvipes J. Thomson, 1878a: 67
gentilis Fåhraeus, 1851: 336
metallescens Gory & Laporte, 1839: 23
obtusa Gory & Laporte, 1839: 25
proxima Gory, 1841: 306

diversa Gory, 1841: 304 N: EG AFR
latona Jakovlev, 1902d: 339

dongolensis Klug, 1829: [no 31] (Buprestis) N: EG AFR

BUPRESTIDAE

BUPRESTIDAE
dumolini Gory, 1841: 308 N: EG AFR
frontalis J. Thomson, 1878a: 71 N: “Sahara” AFR
lesneana Obenberger, 1930b: 116

glyphoderes Abeille de Perrin, 1901a: 8 [RN] N: TU
abeillei Kerremans, 1914b: 145 [RN]
sculpticollis Abeille de Perrin, 1900a: 5 [HN]
tunetana Jakobson, 1912: 136 [RN]

gnezdilovi Kalashian, 2011: 198 A: AE
gordoni Obenberger, 1927g: 171 N: EG AFR
gossypii Kerremans, 1891a: 33 N: EG AFR ORR
gossypicida Obenberger, 1927g: 189 N: EG A: IS SA YE AFR
grata Jakovlev, 1904a: 180 N: EG
heydeni Gredler, 1878: 510 N: EG AFR
heyrovskyi Obenberger, 1924g: 122 N: EG LB AFR
ignescens Escalera, 1914a: 203 N: MO
kerremansi Jakovlev, 1901a: 178 A: “Himalaya” ORR
khartoumensis Obenberger, 1927g: 173 N: EG A: AE YE AFR
krisna Obenberger, 1926b: 20 A: “Himalaya” ORR
laesicollis Abeille de Perrin, 1909: 29 N: AG
lalage Obenberger, 1927g: 179 A: IN
ligulata Jakovlev, 1904a: 178 A: IN
lineifrons Kerremans, 1892a: 190 [RN] A: KA
sulcifrons Fairmaire, 1891e: cxxvi [HN]

marginicollis Hope, 1831: 25 (Buprestis) A: NP
mesopotamica Marseul, 1865: 389 E: AB AR GG UK (Krym) A: AF IN IQ KZ TD TM TR UZ
deserti Obenberger, 1920b: 129
fairmairei Marseul, 1865: 528
fulgurans Obenberger, 1920b: 128
namanganensis Obenberger, 1927g: 176
pelleti Mulsant & Rey, 1866: 87
turkestanica Jakovlev, 1885: 133
sartica Obenberger, 1927g: 171

moises Obenberger, 1924g: 121 A: SI
neglecta neglecta Klug, 1835: 30 (Buprestis) N: EG A: SA AFR
caieana Kerremans, 1907a: 94

nilotica Gory & Laporte, 1839: 31 N: AG AFR
brevis Gory & Laporte, 1839: 32

nitens Kerremans, 1898a: 118 N: EG A: IS OM SA YE
nubiae Obenberger, 1924g: 123 N: “Nubia”
olivieri Obenberger, 1924g: 117 A: “Arabia”

483

484

BUPRESTIDAE

parallelicollis Kerremans, 1913a: 524 N: MO AFR
perrotetii afganica Cobos, 1966d: 312 A: AF
perrotetii perrotetii Guérin-Méneville, 1840c: 328 A: IN ORR
angustata J. Thomson, 1878a: 72

pharia Chevrolat, 1838b: 77 N: EG A: IS SI AFR
aegyptiaca Gory, 1841: 306
angolensis Gory, 1841: 303

pici Théry, 1930a: 234 N: EG A: IS JO SI AFR
punjabensis Obenberger, 1926b: 22 A: “Himalaya”
purpurifera Walker, 1871: 13 A: SI
pyristoma Obenberger, 1916d: 256 N: EG
quadraticollis Gerstäcker, 1871: 53 N: LB AFR
recticollis Harold, 1878c: 217

rotroui Théry, 1930a: 216 N: AG
salamita Kerremans, 1913a: 508 N: EG AFR
semenovi Jakovlev, 1889a: 85 E: AB A: AF IN KZ MG NW TD TM UZ
jedlickai Obenberger, 1927g: 187
lamaica Obenberger, 1920b: 126
prosternalis Reitter, 1890m: 279 [HN]
reitteri Jakovlev, 1891b: 139
sagitta Semenov, 1899c: 651

smaragdifrons Escalera, 1914a: 203 N: MO
trispinosa Klug, 1829: [no 29] (Buprestis) N: EG A: IS SI SY AFR
owariensis Gory & Laporte, 1839: 28
trilineata Palisot de Beauvois, 1807: 44 (Buprestis)

tutankhameni Obenberger, 1924g: 118 N: EG
subgenus Paradeudora Obenberger, 1924g: 114 type species Sphenoptera mossulensis Obenberger, 1920
kermanshahensis Obenberger, 1952a: 4 A: AE IN IQ KU SA SY
mossulensis Obenberger, 1920b: 118 A: IQ SI
saharensis Obenberger, 1924g: 113 N: AG
subgenus Sphenoptera Dejean, 1833: 81 type species Buprestis antiqua Illiger, 1803
Chrysodora Gistel, 1834: 11 type species Buprestis antiqua Illiger, 1803
Depsaria Gistel, 1856: 31 [SN] type species Buprestis antiqua Illiger, 1803

abbreviata Jakovlev, 1900b: 104 E: AR A: IN TR
aerata Jakovlev, 1887a: 62 E: AR A: IN TM
aliena Jakovlev, 1900d: 503 A: TR
altaica Cobos, 1968: 368 A: MG
amasica Obenberger, 1927a: 90 A: TR
anthracina Jakovlev, 1887a: 78 E: AB AR A: IN TM
atramentaria Jakovlev, 1900d: 500

BUPRESTIDAE

485

furva Jakovlev, 1908a: 215
tenebricosa Jakovlev, 1908a: 215

antiqua antiqua Illiger, 1803: 247 (Buprestis) E: AL AR AU BH BU CR CZ FR GG GR HU IT MC MD ME PT RO
SB SK SL SP ST UK N: AG MO A: IN IQ TR
ausonica Obenberger, 1927a: 95
danaica Obenberger, 1927a: 92
dianthi Steven, 1829: 286 (Buprestis)
earo Obenberger, 1929g: 123
georgica Obenberger, 1916d: 248
helladicola Obenberger, 1927a: 92
inaequalis Steven, 1829: 287 (Buprestis)
iridiventris Gory & Laporte, 1839: 7
korbi Obenberger, 1927a: 93
lineigera Obenberger, 1920b: 115
litigiosa Mannerheim, 1837: 94
melitta Obenberger, 1927a: 88
oreophila Obenberger, 1927a: 90
quercii Leoni, 1911b: 5
serena Jakovlev, 1900d: 407
somchetica Kolenati, 1846: 34
variolosa Kerremans, 1898b: 299

antiqua pernitens Obenberger, 1927a: 94 A: TR
antiqua ramosula Obenberger, 1927a: 95 E: AB AR A: IN
arcana Jakovlev, 1908a: 211 A: IN IQ TR
assuricola Obenberger, 1920b: 121
baghdadensis Obenberger, 1929d: 13
heliochaera Obenberger, 1929g: 111
stella Obenberger, 1952c: 5
teheranica Obenberger, 1952c: 6

ardua Gory & Laporte, 1839: 17 E: FR GR
arnoldii Alexeev, 1975: 143 A: ES MG
alexeevi Bellamy, 1998: 93 [HN]

auriceps Jakovlev, 1886: 90 A: TR
banghaasi Obenberger, 1915a: 51 A: TR
barbarica Gmelin, 1790: 1936 (Buprestis) [RN] E: AL AU CR FR GR (Kríti) IT PT RO SP N: AG MO TU A: TR
barbara A.G. Olivier, 1790b: 237 (Buprestis) [RN]
bicolor Fabricius, 1787a: 183 (Buprestis) [HN]
bousaadensis Obenberger, 1924g: 113
conica Gory & Laporte, 1839: 16
cupriventris Gory & Laporte, 1839: 18

486
dilaticollis Gory & Laporte, 1839: 12
gemellata Mannerheim, 1837: 96
laticollis A.G. Olivier, 1790a: no 32: 69 (Buprestis)
oreobata Obenberger, 1927a: 59
osiris Obenberger, 1924g: 113
sicula Jakovlev, 1886: 95
silvestrii Leoni, 1911b: 8

barbata Jakovlev, 1887b: 113 A: “Ala-Taou”
bicarinata Jakovlev, 1887a: 77 A: IN
bifoveolata Marseul, 1865: 382 N: AG
bodemeyeri Jakovlev, 1900d: 501 A: TR
quadrata Kerremans, 1909a: 279

boisduvali Obenberger, 1927a: 77 A: IN
brussae Obenberger, 1927a: 86 A: TR
caesia Jakovlev, 1904a: 175 A: UZ
canaliculata Pallas, 1781: 65 (Buprestis) A: ES MG NO
daurica Marseul, 1865: 361
pallasia Schönherr, 1817: 264 (Buprestis) [RN]
roborowskyi Jakovlev, 1901a: 171
zichyi Csiki, 1901: 109

chalybea anardarensis Alexeev, 1991: 855 A: AF
chalybea chalybea Ménétriés, 1848: 30 A: KZ TM
askhabadensis Obenberger, 1927a: 62
schelli Obenberger, 1927a: 62

chrysostoma Gory & Laporte, 1839: 35 E: AL GR
albanica Obenberger, 1927a: 58
ariadne Obenberger, 1913d: 88
graeca Obenberger, 1927a: 58
minos Obenberger, 1920b: 114

clarescens Kerremans, 1909a: 283 A: IN
commixta Obenberger, 1927a: 52 A: ES
coracina Steven, 1829: 286 (Buprestis) E: AB AR BU GG GR MC ST UK A: IN IQ IS LE SY TR
aethiops Jakovlev, 1891b: 129
babel Marseul, 1865: 340
clauda Marseul, 1865: 341
crassiceps Jakovlev, 1887a: 75
dichroa Jakovlev, 1887b: 112
hartmanni Obenberger, 1927a: 11
korenskyi Obenberger, 1927a: 37, 41
mrazi Obenberger, 1927a: 30
parnassica Jakovlev, 1887a: 74

BUPRESTIDAE

BUPRESTIDAE
poetica Obenberger, 1927a: 35
quadrifoveolata Marseul, 1865: 356
rugata Kerremans, 1903: 112
tatarica Obenberger, 1927a: 38
thoracica Jakovlev, 1887a: 80

cunea Marseul, 1865: 346 A: IN IQ TR
cuprifrons Faldermann, 1835a: 153 E: “Transcaucasia” or A: “Persia”
cuprina agnoscenda Obenberger, 1927a: 75 A: KZ
cuprina cuprina Motschulsky, 1860i: 516 E: AB AR BU CT GR (Kríti) IT (Sicilia) KZ ST UK A: KZ NW
balcanica Jakovlev, 1908a: 219
castelnaudi Kerremans, 1913a: 162
circe Obenberger, 1927a: 85
dilatipes Jakovlev, 1891b: 131
fallaciosa Jakovlev, 1903b: 216
fossithorax Obenberger, 1920b: 116
jermaki Obenberger, 1927a: 63
lutshniki Obenberger, 1927e: 39
nowickii Obenberger, 1927a: 76
uralensis Obenberger, 1927a: 72

cyanea Jakovlev, 1899c: 558 A: AF IN KZ TM
cyaneoniger Kerremans, 1909a: 275 A: IN TR
cyphogastra Jakovlev, 1900d: 426 A: UZ
danilevskyi Kalashian, 2002: 78 A: KZ
demissa Marseul, 1865: 365 E: AR A: AF IN IQ TM TR
agrippina Kerremans, 1911: 633
ajax Kerremans, 1913a: 137
breiti Obenberger, 1920b: 117
ganglbaueri Jakovlev, 1886: 85
kaszabi Obenberger, 1963: 321
komarovi Jakovlev, 1886: 86
obscura Jakovlev, 1891b: 130
profusa Kerremans, 1909a: 277
shahrudensis Obenberger, 1927a: 55

depressiuscula Obenberger, 1927a: 49, 63 A: KI UZ
diluta Jakovlev, 1900c: 202 E: AR
donata Obenberger, 1929g: 125 A: TR
ebenina Jakovlev, 1900d: 498 A: AF TD UZ “Alai”
herzi Jakovlev, 1908a: 220

edmundi Jakovlev, 1900d: 425 E: AB AR A: IN TR
oporina Jakovlev, 1903d: 386

egregia Jakovlev, 1901a: 170 A: TM

487

488

BUPRESTIDAE

enyuseki Obenberger, 1927a: 89 A: TR
eugenii Jakovlev, 1899c: 554 E: AB AR GG A: IN
cyaniventris Obenberger, 1927a: 23
leonhardi Obenberger, 1927a: 25
maga Obenberger, 1927a: 26

euplecta Obenberger, 1927a: 89 A: IN TR
exarata Fischer von Waldheim, 1824: 169 (Buprestis) E: AB AR KZ ST A: AF IN KI KZ NW TD TM UZ
araxana Obenberger, 1915a: 52
dejeanii Zubkov, 1829: 156 (Buprestis)
interrupta Kerremans, 1909a: 276
nomadica Obenberger, 1916d: 247
prosternalis Jakovlev, 1885: 130
spectabilis Kraatz, 1883e: 345
sumakovi Obenberger, 1927a: 29

excavata Faldermann, 1835a: 154 E: “Transcaucasia” A: IN
exigua Jakovlev, 1908a: 222 A: KI
exoleta Jakovlev, 1908a: 217 A: IN
politipennis Obenberger, 1927a: 12, 23

extensocarinata Jakovlev, 1889a: 84 A: ES MG NO
fallatrix Obenberger, 1927a: 87 E: “Caucasus”
hyrcas Obenberger, 1927a: 87

faragi Théry, 1936a: 6 A: SI
fissifrons Marseul, 1865: 350 A: TM
fossa Kerremans, 1909a: 271 A: IN
foveola Gebler, 1825: 46 (Buprestis) E: KZ ST A: KI KZ
strandi Obenberger, 1920b: 113

freyi Théry, 1942d: 274 A: IN
galkae Kalashian & Volkovitsh, 2009: 401 A: PA
hetita Obenberger, 1927a: 13 A: IN TR
hypocrita Mannerheim, 1837: 92 E: AB AR GG ST A: IN JO PA TM TR
dlabolai Obenberger, 1955a: 11
erdshmiadzini Obenberger, 1927a: 20
funebralis Obenberger, 1927a: 19
glabricollis Obenberger, 1929g: 129
ixion Kerremans, 1912a: 74
levantina Obenberger, 1929g: 130
ordubadensis Obenberger, 1916d: 250
sieversi Jakovlev, 1887a: 82
torrida Jakovlev, 1899c: 553
turkmenica Obenberger, 1927a: 44

igniceps Jakovlev, 1886: 89 A: TR

BUPRESTIDAE
inermis Kerremans, 1898a: 119 A: ES MG NO
csikii Obenberger, 1963: 322
dsungaricola Obenberger, 1920b: 111
macra Jakovlev, 1909: 1
planocarpinoi Cobos, 1972a: 28

intaminata Jakovlev, 1908a: 218 E: AB A: TR
irregularis Jakovlev, 1886: 96 A: ES KI KZ MG
sajanensis Obenberger, 1927a: 14
sesquistriata Cobos, 1972a: 26

jacobsonorum Kalashian & Volkovitsh, 2009: 403 A: KA
korshinskii Jakovlev, 1900d: 423 A: UZ
kozlowi Jakovlev, 1890b: 102 A: NW
krali Obenberger, 1927a: 83 A: AF UZ
lapidaria Brullé, 1832: 134 (Buprestis) E: AB AL AR BU CR GG GR IT (Trieste) ST A: IN IQ IS LE SY TR
carcelii Gory & Laporte, 1839: 21
caucasicola Obenberger, 1927a: 56
hellenica Obenberger, 1913d: 87
ogloblini Obenberger, 1927a: 56

lateralis Faldermann, 1836: 366 E: ST A: KI KZ TM UZ
latesulcata Jakovlev, 1886: 99 E: AB AR GG A: TM TR
mannerheimi J. Thomson, 1878a: 63 [HN]

laticeps Jakovlev, 1886: 94 A: KI KZ MG NW TD UZ
karakusensis Obenberger, 1927a: 80
massagetica Obenberger, 1920b: 109
memnonia Obenberger, 1927a: 79
montana Jakovlev, 1887b: 113
montivaga Obenberger, 1927a: 80
strigosa Jakovlev, 1887a: 71
tadjika Obenberger, 1927a: 74

lia Jakovlev, 1901a: 168 A: TM UZ
longipennis Jakovlev, 1904a: 174 A: KI KZ
lopatini Kalashian, 2005: 249 A: XIN
lucidicollis Kraatz, 1882i: 316 A: UZ
luctifica Jakovlev, 1904b: 310 [RN] A: NO
foveithoracica Kerremans, 1913b: 116 [RN]
luctuosa Jakovlev, 1904a: 177 [HN]

magna Gory & Laporte, 1839: 3 A: IN IQ IS JO SA SY TR
alexandri Obenberger, 1927a: 33

manca Obenberger, 1927e: 37 A: UZ
manderstjernae Jakovlev, 1886: 94 A: KI KZ NW UZ
bucharae Obenberger, 1927a: 78

489

490
margellanica Kraatz, 1882f: 112 A: AF UZ
haarlovi Descarpentries, 1965: 262
plana Jakovlev, 1887b: 114

marseuliana Obenberger, 1927a: 82 [RN] A: IQ
marseuli Kerremans, 1913a: 170 [HN]

marseulii E. Saunders, 1871: 87 A: ES
mazuri Hołyński, 1996: 140 A: TR
merkli Kalashian et Volkovitsh, 2008: 205 A: IN
mixta Jakovlev, 1887b: 111 E: AR A: TR
moesta Jakovlev, 1887a: 64 A: IN
uncta Kerremans, 1909a: 277

mora Curletti & Magnani, 1992: 37 N: TU
nausicaa Obenberger, 1927a: 21 A: IN
navicula Jakovlev, 1908a: 212 A: TM
steppensis Obenberger, 1927a: 81

nereis Obenberger, 1927a: 27 A: TR
notata Jakovlev, 1899c: 556 A: AF “Alai”
obruta Kerremans, 1909a: 275 A: AR IN SY
chalcosoma Obenberger, 1927a: 43
hetera Obenberger, 1927a: 21

obscuriventris Motschulsky, 1860i: 517 A: ES KI KZ MG NO WS
olivacea Kraatz, 1882f: 113 A: TM UZ
oresitropha Obenberger, 1927a: 31 A: IN TR
ostentator Kerremans, 1909a: 281 A: SY
ottomana Obenberger, 1927a: 91 A: TR
pharao Gory & Laporte, 1839: 7 N: AG EG LB TU A: IS LE SY
algirica Abeille de Perrin, 1891: 266
breviuscula Abeille de Perrin, 1891: 266
pharaon Gory & Laporte, 1839: 7 [unused original spelling]

pilipes Jakovlev, 1886: 92 A: KI KZ
anchorifera Obenberger, 1952c: 20
romanovi Obenberger, 1927a: 50
winkleri Obenberger, 1920b: 108

pliginskii Obenberger, 1927a: 92 E: UK (Krym)
porrecta Jakovlev, 1908a: 214 A: TR
prava Kerremans, 1909a: 278 A: TR
propinqua Kraatz, 1882i: 315 A: TD UZ
pseudochalybaea Richter, 1945c: 148 A: KZ UZ
purpuriventris Kraatz, 1882i: 314 A: UZ
rangnowi Kerremans, 1909a: 273 E: AB AR A: AF IN TM
strandiana Obenberger, 1927a: 42

BUPRESTIDAE

BUPRESTIDAE
repetekensis Obenberger, 1927a: 80 A: AF IN TM UZ
budshnurdi Obenberger, 1927a: 71
darius Obenberger, 1927a: 81

rugulosa Faldermann, 1835a: 151 E: AB AR A: IN TR
cribrata Faldermann, 1835a: 152

saxosa Kerremans, 1909a: 283 A: IN
serripes Jakovlev, 1900d: 431 A: TM
shindandensis Alexeev, 1991: 853 A: AF
signifera Jakovlev, 1899c: 551 A: KZ UZ
spreta Jakovlev, 1900c: 203 A: KI KZ
stenophthalma Jakovlev, 1900b: 105 A: KZ
subtilis Jakovlev, 1899c: 552 A: IN TM
subtricostata Kraatz, 1882i: 318 A: KZ TD UZ
sulcata eoa Obenberger, 1927a: 67 A: NW
sulcata lukianovici Obenberger, 1927a: 67 E: UK (Krym)
sulcata minussina Obenberger, 1927a: 73 A: ES MG
mongoliana Cobos, 1968: 361

sulcata sulcata Fischer von Waldheim, 1824: 197 (Buprestis) E: AB AR GG KZ ST A: ES IN KZ NW WS
benedicenda Obenberger, 1927a: 72
bergrothi Obenberger, 1927a: 66
fossulata Gebler, 1825: 46 (Buprestis)
gebleri Gory & Laporte, 1839: 12
impressicollis Marseul, 1865: 555 [HN]
mendax Marseul, 1869a: 122 [RN]
tifliensis Obenberger, 1927e: 38
uralicola Obenberger, 1927a: 75

sulcata winkleriana Obenberger, 1920b: 110 A: KZ NW
alatauensis Obenberger, 1927a: 68
borochorensis Obenberger, 1920b: 111
iliensis Obenberger, 1927a: 72
jakovleviana Obenberger, 1927e: 40
komurolensis Obenberger, 1927a: 75
modificata Obenberger, 1920b: 112
sororella Obenberger, 1920b: 112
tarbagataica Obenberger, 1927a: 68

suvorovi Jakovlev, 1908a: 213 A: KI or KZ
tragacanthae Klug, 1829: [no 26] (Buprestis) E: AB AR GG GR UK (Krym) A: AF IN IQ KI LE PA SY TM TR
adusta Jakovlev, 1887b: 111
armenigena Obenberger, 1927a: 12
armenorum Obenberger, 1927a: 37, 39
artaxerxes Obenberger, 1927a: 17

491

492
catenifera Obenberger, 1927a: 16
christophi Jakovlev, 1887a: 73
cilicica Obenberger, 1927a: 8
coeruleolimbata Obenberger, 1927a: 16
corrosa Obenberger, 1927a: 31
corrugata Klug, 1829: [no 27] (Buprestis)
derugata L. Redtenbacher, 1850: 47
elamita Marseul, 1865: 238
encausta Marseul, 1865: 340
exasciata Jakovlev, 1886: 83
gigas Obenberger, 1916d: 246
glabrata Ménétriés, 1832: 151 (Buprestis)
goryi Théry, 1938a: 91
klapaleki Obenberger, 1927a: 16
klickai Obenberger, 1927a: 28
kulpensis Obenberger, 1927a: 6
lobicollis Marseul, 1865: 339
luristana Kerremans, 1909a: 272
maledicta Obenberger, 1920b: 104
maura Jakovlev, 1902d: 317
mniszechi Marseul, 1865: 332
modesta Jakovlev, 1887a: 76
morio Jakovlev, 1886: 84
morosa Jakovlev, 1908a: 216
mrazeki Obenberger, 1927a: 18
nominanda Obenberger, 1920b: 106
obnubila Jakovlev, 1908a: 221
parsa Obenberger, 1920b: 105
prolongata Jakovlev, 1887a: 70
prometheus Obenberger, 1916d: 245
radicigena Obenberger, 1927a: 8
rambouseki Obenberger, 1927a: 10
satrapa Obenberger, 1927a: 32
signaticollis Jakovlev, 1887a: 85
sordida Jakovlev, 1886: 84
speciosa Jakovlev, 1887b: 110
syriae Obenberger, 1927a: 34
tauricola Obenberger, 1927a: 33
vavrai Obenberger, 1927a: 41
zarathustrae Obenberger, 1920b: 107
zobeida Obenberger, 1927a: 36

BUPRESTIDAE

BUPRESTIDAE

493

tristis Jakovlev, 1886: 87 A: “Ferghana”
trisulcata Reiche & Saulcy, 1856: 413 A: IN IQ IS LE SY
ubarchangajensis Cobos, 1968: 365 A: MG
varia Jakovlev, 1887a: 83 A: UZ
samarkanda Obenberger, 1929g: 124

vidua Jakovlev, 1900d: 429 A: NW
violacea Jakovlev, 1899c: 557 A: IN TM
viridicoerulea Kraatz, 1882i: 317 A: TM UZ
subgenus Sphenopterella Volkovitsh & Kalashian, 1994: 99 type species Sphenoptera margaritae Volkovitsh &
Kalashian, 1994
margaritae Volkovitsh & Kalashian, 1994: 100 A: UZ
vlastae Kalashian & Volkovitsh, 2006: 135 A: IN
subgenus Tropeopeltis Jakovlev, 1902d: 283 type species Sphenoptera anthaxoides Reitter, 1895
andamanensis C.O. Waterhouse, 1877a: 6 A: SD SW ORR
cupreotoma J. Thomson, 1878a: 69
impressa Kerremans, 1892b: 182
rebierei Kerremans, 1914b: 111

anthaxoides Reitter, 1895a: 42 E: AB AR A: IN TR
arabica Gory, 1841: 307 N: EG A: AE IS OM SA YE
aterrima Kerremans, 1914d: 204 A: HP “Himalaya”
calligoni Kalashian & Volkovitsh, 1997: 357 A: KZ TM
dadkhani Obenberger, 1955b: 90 A: PA
davatchii Descarpentries, 1960: 175 A: IN
deobani Obenberger, 1926b: 60 A: UP
dichrosoma Obenberger, 1926b: 59 A: “Himalaya”
fossiventris Kerremans, 1914b: 108 N: “Nubia” AFR
fulgens fulgens Gory, 1841: 309 N: AG EG LB A: IN SA AFR
arenosa Obenberger, 1924g: 125
chalcichroa Obenberger, 1914g: 132

fulgens ramses Obenberger, 1916d: 257 N: EG
fusiformis Kerremans, 1913c: 10, 73 A: SA
himalayensis Théry, 1910a: 7 A: “Northwestern Himalaya”
hypsibata Obenberger, 1927g: 197 A: KA
inimica Obenberger, 1924g: 124 N: EG
isis Jakovlev, 1901a: 179 N: EG
kaznakowi bicoloratus Alexeev, 1991: 862 A: AF
kaznakowi kaznakowi Jakovlev, 1899a: 295 A: IN TD TM UZ
klapperichi Cobos, 1966d: 313 A: AF
konbirensis Kerremans, 1892b: 188 A: “Himalaya” ORR
bengalensis Jakobson, 1912: 135 [RN]
cincticollis Kerremans, 1893b: 330

494

BUPRESTIDAE

cinctiventris Kerremans, 1913a: 42
cupriventris Kerremans, 1890c: cciii [HN]
paradoxa Kerremans, 1913c: 6, 49 [HN]

lafertei lafertei J. Thomson, 1878a: 65 A: HP UP
buqueti Obenberger, 1926b: 72

lafertei monticola Cobos, 1966d: 315 A: AF
mediocris Kerremans, 1893b: 330 A: “Himalaya” ORR
nigrescens J. Thomson, 1878a: 64 A: “Himalaya” ORR
oertzeni Jakovlev, 1887b: 115 E: GR N: EG A: CY IN IS SY TR
deltae Obenberger, 1924g: 123

prorsa Kerremans, 1914b: 104 A: TR
punctisternum Obenberger, 1924g: 124 N: EG
schneideri Reitter, 1898a: 11 A: KZ TM UZ
lebedevi Obenberger, 1928b: 18
mujunkumensis Obenberger, 1928b: 19

servistana Obenberger, 1929e: 60 A: IN IQ TM
dhiaahmedi Cobos, 1956: 29
kambyses Obenberger, 1930a: 104

tappesi Marseul, 1865: 387 E: AL GR MC A: IN IS JO LE SY TR
simulatrix Reitter, 1895a: 42

vanharteni Kalashian, 2011: 195 A: AE
genus Sphenoptera, nomen dubium,
parvula Gory & Laporte, 1839: 20 [HN] E: IT

subfamily Buprestinae Leach, 1815
tribe Actenodini Gistel, 1848

genus Belionota Eschscholtz, 1829: 9 type species Belionota sagittaria Eschscholtz, 1829
subgenus Belionota Eschscholtz, 1829: 9 type species Belionota sagittaria Eschscholtz, 1829
fallaciosa Deyrolle, 1864a: 81 A: FUJ TAI ORR
ceramensis Théry, 1935d: 254
intermedia van Lansberge, 1879: cxlviii
nigriventris Théry, 1929c: 144
purpurea Obenberger, 1924a: 82
purpurea Obenberger, 1932c: 370 [HN]
walkeri C.O. Waterhouse, 1894b: 67

prasina Thunberg, 1789: 90 (Buprestis) N: EGi A: FUJ SD TAI YUN AFRi AURi NTRi ORR
aenea Deyrolle, 1864a: 82
aruensis Obenberger, 1918c: 12
huebneri Fairmaire, 1883a: 16
obscuripennis Fisher, 1922: 7

495

BUPRESTIDAE
pyrotis Illiger, 1800b: 119 (Buprestis)
saundersii C.O. Waterhouse, 1894a: 285
scutellaris Weber, 1801: 72 (Buprestis) [HN]
strandi Obenberger, 1924a: 82
viridis Théry, 1929c: 141

tribe Anthaxiini Gory & Laporte, 1839

genus Anthaxia Eschscholtz, 1829: 9 type species Buprestis nitida P. Rossi, 1794 (=Buprestis fulgurans Schrank,
1789)
subgenus Anthaxia Eschscholtz, 1829: 9 type species Buprestis nitida P. Rossi, 1794 (=Buprestis fulgurans
Schrank, 1789)
Anthaxiola Richter, 1945b: 118, 128 type species Anthaxia anatolica Chevrolat, 1838
Callanthaxia Richter, 1945c: 157 type species Buprestis candens Panzer, 1792
Cyclanthaxia Richter, 1949: 58, 137 type species Buprestis croesus Villers, 1789 sensu Richter, 1949 (=Buprestis salicis
Fabricius, 1777)
Eoanthaxites Haupt, 1950: 68 type species Eoanthaxites unicolor Haupt, 1950
Euanthaxia Richter, 1945a: 67 type species Anthaxia discicollis Gory & Laporte, 1839
Homalanthaxia Richter, 1945c: 154 type species Buprestis funerula Illiger, 1803
Paranthaxia Gozis, 1886: 21 [RN] type species Buprestis nitida P. Rossi, 1794 (=Buprestis fulgurans Schrank, 1789)
Trichocratomerus Richter, 1945a: 62 type species Anthaxia intermedia Obenberger, 1913 (=Anthaxia mancatula Abeille
de Perrin, 1900)

alfierii Théry, 1929b: 139 N: EG
altaica Cobos, 1968: 359 A: MG
amasina amasina K. Daniel, 1903: 252 E: AR BU GG GR MC TR A: IN LE TR
amasina maceki Bílý, 1980c: 403 A: IQ IS JO
anadyomene Bílý & Kubáň, 2004: 3 A: CY
anatolica anatolica Chevrolat, 1838b: 75 E: AB AR BU GG GR MC RO TR A: CY IN IQ IS JO LE SY TR
hilaris Mulsant & Godard, 1855a: 7 [HN]
mulsanti Marseul, 1865: 243 [RN]

anatolica catei G. Novak, 1994b: 9 A: IN TM
anatolica ferulae Gené, 1839: 53 E: FR IT PT SP N: AG MO TU AFR
senegalensis Kerremans, 1898b: 302
vittaticollis Lucas, 1844c: 89

anatolica lucidiceps Gory, 1841: 289 A: AF IN KI KZ TD TM UZ
apollonii Ballion, 1871a: 349
auriceps Ménétriés, 1848: 29
gerneti Morawitz, 1867: 35

antoinei Baudon, 1955: 122 N: MO
berbera Bílý, 1982d: 224

ariadna Bílý, 1982a: 56 E: GR (Kríti)
aureoviridis Svoboda, 1994b: 213 A: CY

496

BUPRESTIDAE

auriventris Ballion, 1871a: 350 A: KI KZ UZ
alexandri Obenberger, 1938: 215
zarudniana Richter, 1945d: 146

baconis J. Thomson, 1879a: 27 A: AP BT HP KA NP PA SD UP ORR
afghanica Obenberger, 1913e: 126
birmanica Théry, 1910a: 33
bivulnerata Obenberger, 1914b: 12
nepalensis Cobos, 1966c: 248
radja Obenberger, 1922b: 96
splichali Obenberger, 1914c: 35

badghyzica Bílý, 1991b: 121 A: TM
bedeli bedeli Abeille de Perrin, 1893: 132 E: FR IT PT SP N: AG MO TU
bedeli peloponesiaca Obenberger, 1938: 209 E: GR
bercedoi Bílý, 2006b: 23 E: SP
bicolor bicolor Faldermann, 1835b: 149 E: AB AR BU CR GG GR MC RO ST TR UK (Krym) A: IN IQ IS JO LE SY
TM TR
taurica Mannerheim, 1837: 87

bicolor comptei Cobos, 1966b: 128 E: SP
bicolor uzbekistana Bílý, 2002: 195 A: UZ
blascoi F. Murria & Á. Murria, 2005: 75 E: SP
breviformis Kalashian, 1988: 137 E: AR A: IN
brevis brevis Gory & Laporte, 1839: 18 E: AB AL AR BU GR MC ST TR UK A: CY IN IS SY TR
achaia Obenberger, 1938: 196
cypriota Obenberger, 1938: 197
hellenica Obenberger, 1914b: 12, 13

brevis kovari Bílý, 1983a: 75 A: IN
brodskyi Bílý, 1982a: 58 E: GR (Kríti)
candens antoniocobosi Sautière, 2009: 271 E: SP
cobosi Sautière, 2009: 70

candens candens Panzer, 1792: 9 (Buprestis) E: AU BH BU CR CZ FR GE GG GR HU IT LU RO SK SL SZ UK
fulminans Schrank, 1789: 84 (Buprestis) [HN]
fulminatrix Herbst, 1801: 297 (Buprestis) [RN]
marchii Obenberger, 1925c: 55

candensiformis Novak, 2006: 83 E: GR
candiota Obenberger, 1938d: 199 E: GR (Kríti) A: CY TR
carmelita Abeille de Perrin, 1900a: 6 A: IS JO SY
carpetovetonica Sánchez, 2014: 49 E: SP
cashmirensis Obenberger, 1938: 174 A: KA PA
ceballosi Escalera, 1931a: 433 E: SP
chaerodrys Szallies, 2001: 125 A: TR
chevrieri Gory & Laporte, 1839: 33 E: AU BH BU BY CR CT CZ FR GG GR HU IT MC MD ME PL RO SB

BUPRESTIDAE

497

SK SL SP ST SZ TR UK A: KZ TR WS
aeneomicans Obenberger, 1930: 487 [RN]
aenescens Abeille de Perrin, 1909: 22 [HN]
alpina Obenberger, 1914b: 14
bannatica Gory, 1841: 297

chinensis Kerremans, 1898a: 132 A: FUJ HKG HUN JIA JIX SHG ZHE ORR
klebelsbergi Gebhardt, 1929a: 26
vagabunda Théry, 1910a: 35

cleopatra Obenberger, 1913c: 64 N: AG LB TU
freyi Théry, 1939c: 248

combusta Obenberger, 1922b: 96 A: HP NP SD UP
constricticollis Bílý, 2007a: 81 A: FE SHX
cupressi Bílý, 2005a: 5 A: SCH
cyanescens Gory, 1841: 284 E: FR SP N: AG MO TU
aeneiventris Küster, 1851: 31

cypraea Abeille de Perrin, 1900a: 6 A: CY
depressifrons Bílý, 2006b: 50 N: AG
dimidiata Thunberg, 1789: 91 (Buprestis) E: IT FR SP N: AG MO TU
saliceti Illiger, 1803: 254 (Buprestis)

discicollis discicollis Gory & Laporte, 1839: 25 E: AL BU CR GR MC ME TR A: TR
discicollis kanaanita Obenberger, 1912a: 9/17 E: AB AR GG A: CY IN IS LE SY TM TR
eggeri Brandl, 2003: 3 N: CI (Tenerife)
elberti Brandl, 1993b: 51 E: BH GR
ephippiata L. Redtenbacher, 1850: 47 E: AB AR GG A: AF IN IQ SY TM TR
edithae Reitter, 1890j: 194
kreuzbergi Richter, 1944a: 49
maschelii Kiesenwetter, 1880: 130

exigua Tolosa, Á. Murria & F. Murria, 2012: 29 E: SP
fageli Descarpentries, 1955: 30 N: AG TU
feloi Liberto, 2000: 5 N: CI
fernandezi Cobos, 1953b: 116 N: CI
fesana Bílý, 2006b: 54 N: MO
fritschi Heyden, 1887c: 446 N: AG EG LB MO A: IS
nigrofusca Obenberger, 1913c: 65

fuksai Obenberger, 1928c: 68 A: SY TR “Caucasus”
fulgurans Schrank, 1789: 85 (Buprestis) E: AL AR AU BE BH BU BY CR CZ FR GE GG GR HU IT MC MD ME PL
RO RU SB SK SL SP ST SZ UK
azurescens Gory & Laporte, 1839: 18
bifoveata Fisher, 1925b: 168
bipunctata A.G. Olivier, 1790a: no 32: 81 (Buprestis) [HN]
dorsalis Gory & Laporte, 1839: 17

498
formosa Townson, 1797: 466 (Buprestis)
foveolata Herbst, 1801: 243 (Buprestis) [RN]
fulgens Herbst, 1801: 298 (Buprestis) [HN] [RN]
lucidula Fabricius, 1794: 451 (Buprestis) [HN]
nitens Fabricius, 1801b: 215 (Buprestis) [RN]
nitida P. Rossi, 1792: 63 (Buprestis)

funerula Illiger, 1803: 251 (Buprestis) E: PT SP
gansuensis Bílý, 1991a: 85 A: GAN SCH XIN
gaditana F. Murria, Verdugo & Á. Murria, 2011: 17 E: SP
genistae Bílý, 2006b: 47 N: AG
ghazi Obenberger, 1938: 213 E: AB AR A: IS JO LE SY TR
khnzoriani Kalashian, 1988: 139
vseteckai Obenberger, 1940: 168

gianassoi Curletti & Magnani, 1992: 43 N: MO
glabrifrons Abeille de Perrin, 1900a: 8 N: AG MO TU
salambo Obenberger, 1913c: 65

gottwaldi Brandl & Mühle, 2008: 22 A: CY
granatensis Verdugo, 2013: 62 E: SP
guanche Liberto, 2000: 2 N: CI
hackeri J. Frivaldszky, 1884: 281 E: AU BU CR CZ HU IT RO SK TR A: TR
hepneri Bílý, 2006b: 46 N: MO
herbertschmidi G. Novak, 1992: 26 A: TR
holoptera Obenberger, 1914f: 115 E: AB AR A: IN TM
hozaki Bílý, 1973b: 427 A: CY
hypsigenia Obenberger, 1938: 217 A: HP NP SD UP
hyrcana Kirsch, 1880a: 132 E: AB A: IN
permira Reitter, 1898b: 120
seminigra Théry, 1925b: 176
semiusta Abeille de Perrin, 1895c: cxvii

idae Obenberger, 1938: 212 E: GR (Kríti)
impunctata Abeille de Perrin, 1909: 22 E: FR IT SP
nereis Schaefer, 1938a: 81

irregularis Abeille de Perrin, 1909: 24 A: SY TR
jenisi Bílý, 2006a: 32 A: IN
juliae Liberto, 1996: 369 N: CI (Gomera)
kervillei Théry, 1939a: 15 E: GR (Aegean Islands) A: TR
kmoniceki Bílý, 2015a: 482 A: SCH
kurdistana Obenberger, 1912b: 65 A: IQ TR
lauta Alexeev, 1964: 9 A: KI TD UZ
lecerfi Théry, 1930a: 322 N: MO
lucens Küster, 1852: 66 E: AL BH BU CR GR IT MA MC TR A: TR

BUPRESTIDAE

BUPRESTIDAE

499

lucia Bílý, 2006b: 35 E: SP
luctuosa Lucas, 1846e: pl. 16 N: AG LB MO TU
punica Obenberger, 1938: 211
sericea Abeille de Perrin, 1891: 286

lukjanovitshi Richter, 1949: 99 A: ES MG SHA SHX
rozhkovi Alexeev, 1964: 8

madridensis Bílý & Verdugo, 2014: 197 E: SP
magnanii Baiocci, 2011: 1 A: IN
magnifica Bílý, 1983a: 72 A: IN
majzlani Bílý, 1991b: 124 A: BEI SHA
malachitica Abeille de Perrin, 1893: 128 N: AG LB MO TU
carthaginensis Obenberger, 1912c: 160
cupreovirens Abeille de Perrin, 1893: 128
gastonis Desbrochers des Loges, 1901: 61

mamorensis Théry, 1930a: 340 N: AG MO TU
manca Linnaeus, 1767: 1067 (Buprestis) E: AB AL AR AU BE BH BU BY CR CT CZ DE FR GE GG GR HU IT MA
MC MD ME NL PL PT RO SB SK SL SP ST SZ TR UK N: AG A: TR
bistriata Fabricius, 1775: 222 (Buprestis)
elegantula Schrank, 1781: 195 (Buprestis)
rubina Geoffroy, 1785: 33 (Cucujus)

mancatula Abeille de Perrin, 1900a: 5 E: AB AR GG ST A: AF IN KA KI TD TM UZ XIN
intermedia Obenberger, 1913c: 64

margarita Bílý, 1995: 41 A: IS
marmottani hispanica Cobos, 1953c: 103 E: SP
marmottani marmottani Brisout de Barneville, 1883: 81 N: AG MO
medvedevi Alexeev, 1975: 133 A: MG
mendizabali Cobos, 1965b: 120 E: BE CR FR GE GR IT LU PT SL SP TR N: AG MO TU A: TR
meronica Bílý, 1995: 43 A: IS
midas midas Kiesenwetter, 1857: 90 E: CR FR IT
midas oberthuri Schaefer, 1938b: 208 E: FR IT SP N: AG MO
mogadorica Bílý, 2006b: 49 N: MO
morgani Théry, 1925b: 175 A: IN IQ
muliebris Obenberger, 1918b: 22 E: AB AR TR A: IN IS SY TM TR
myrmidon Abeille de Perrin, 1891: 264 E: BU GR MC A: IS LE SY TR
nanissima Alexeev, 1964: 5 A: KZ MG TD TM UZ
rubruki Cobos, 1972a: 21

nigricollis Abeille de Perrin, 1904a: 217 A: TR
nitidula Linnaeus, 1758: 140 (Buprestis) E: AL AU BE BH BU CR CZ DE EN FI FR GB GE GR HU IT KO LS LT LU
MC ME NL NT PL PT RO SB SK SL SP SV SZ N: AG
cyanipennis Gory, 1841: 286
grabowskii Obenberger, 1912a: 7

500

BUPRESTIDAE

laeta Fabricius, 1775: 223 (Buprestis)

nitiduliformis Bílý, 1995: 46 A: GAN HEB SCH SHX
oborili Bílý, 2006b: 53 N: MO
pardoi Cobos, 1966a: 73 N: MO
passerinii Pecchioli, 1837: 446 (Buprestis) E: AB AL AR BH BU CR GR IT MC ME TR A: IN TM TR
purpurea Abeille de Perrin, 1891: 262

patsyae Baiocchi, 2008: 323 A: IN
pengi Bílý, 1990a: 6 A: FUJ
permisa Abeille de Perrin, 1904a: 217 A: IN TR
perrini Obenberger, 1918b: 22 [RN] N: LB A: IS JO LE SY TR
hirticollis Abeille de Perrin, 1893: 131 [HN]

persuperba Obenberger, 1912c: 159 N: AG
ergensis Descarpentries, 1958b: 975

platysoma Abeille de Perrin, 1891: 261 A: SY TR
illustris K. Daniel, 1903: 252

plavilscikovi Obenberger, 1935g: 83 A: KI KZ TD UZ
plicata plicata Kiesenwetter, 1859b: 58 E: BU GR HU RO SB SK TR A: LE SY TR
biimpressa Marseul, 1865: 248
magdalenae Fairmaire, 1884h: cxlv

plicata vinogradovi Obenberger, 1938: 220 E: AB AR GG ST A: IN TR
podolica lucniki Richter, 1949: 160 E: AB AR GG ST
podolica podolica Mannerheim, 1837: 89 E: AL AU BH BU BY CR CT CZ FR GE GR HU IT KO MC MD ME PL
RO SB SK SL SZ TR UK A: TR
gramica Gory & Laporte, 1839: 19
grammica Kiesenwetter, 1857: 98 [unjustified emendation]

pontica Kwast, 2011: 1 A: TR
pseudofunerula Bílý, 2006b: 22 N: MO
pseudogenistae Bílý, 2006b: 48 N: AG
pseudonitidula Svoboda, 2007: 150 A: SY TR
pulex Abeille de Perrin, 1893: 129 N: AG EG LB MO A: IS
punjabensis Obenberger, 1928a: 236 A: PA
pinicola Obenberger, 1938: 183

pyrosoma Bílý, 2006a: 28 A: TR
reitteri Obenberger, 1913c: 65 A: TD UZ
retamae Bílý, 1995: 44 N: AG MO
richteri Stepanov, 1953: 46 A: KI TD UZ
rungsi Baudon, 1958b: 151 N: MO
rutilipennis Abeille de Perrin, 1891: 263 A: SY TR
fleischeri Obenberger, 1925c: 56

salicis discoidalis Obenberger, 1924b: 27 A: IS LE SY TR
salicis persica Théry, 1925b: 175 A: IN TR

BUPRESTIDAE

501

salicis salicis Fabricius, 1777: 237 (Buprestis) E: AB AL AU BE BH BU BY CR CZ FR GE GG GR HU IT LU MC MD
ME NL PL PT RO SB SK SL SP ST SZ TR UK N: AG MO TU A: IN TR
amplexa Rey, 1891a: 4
hirticollis Rey, 1891a: 5

schoenmanni G. Novak, 1984: 45 E: GR (Kríti)
mahri G. Novak, 1984: 46

sedilloti rabinovitchi Théry, 1938c: 267 N: EG A: IS
sedilloti sedilloti Abeille de Perrin, 1893: 129 N: AG LB MO TU
semicuprea Küster, 1851: 21 E: AL AU BH BU BY CR CT CZ FR GE GR HU IT MC MD ME PL RO SB SK SL SP ST
UK A: SY TR
longisquammis Obenberger, 1918b: 22

senicula cretica Brandl, 1993a: 17 E: GR (Kríti)
senicula senicula Schrank, 1789: 85 (Buprestis) E: AB AL AR AU BH BU BY CR CT CZ FR GE GG GR HU IT MC
MD ME PL RO SK SL SP ST TR UK N: AG MO A: TR
auricolor Herbst, 1801: 158 (Buprestis) [RN]
aurulenta Fabricius, 1787a: 182 (Buprestis) [HN]
deaurata Gmelin, 1790: 1934 (Buprestis) [RN]
hanakii Küster, 1851: 18

setipennis Obenberger, 1928a: 239 A: BT HP KA NP SD UP
babaulti Théry, 1930c: 151

shirasensis Obenberger, 1940k: 167 A: IN
signaticollis Krynicki, 1832: 92 E: AB AR BU CT GG GR HU MC MD RO SK ST TR UK A: TR
lichacevi Obenberger, 1934m: 139

simandli Baiocci, 2013: 462 A: IN
sobrinoi Tolosa, Á. Murria & F. Murria, 2012: 26 E: SP
spathuligera Obenberger, 1924b: 27 E: BH BU CR GR MC ME RO A: TR
muelleri Obenberger, 1925c: 56

spinolae helladis Bílý, 2006b: 31 E: CR GR MC SB TR A: TR
pygmaea Brullé, 1832: 134 [HN]

spinolae hesperica Obenberger, 1913e: 64 N: AG LB MO TU
spinolae rifensis Bílý, 2006b: 34 N: MO
spinolae spinolae Gory & Laporte, 1839: 36 E: FR IT PT SP
spinosa Abeille de Perrin, 1900a: 8 A: IN TM
anthocharea Obenberger, 1913g: 334

stateira Bílý, 1983a: 71 A: IN
stepanovi Richter, 1949: 50, 79 A: KZ
superba Abeille de Perrin, 1900a: 5 E: AR
suzannae Théry, 1942a: 92 E: AU BH BU CR CZ FR GE GR HU IT MC ME RO SB SK SL SP SZ UK A: TR
syriaca Bílý & Kubáň, 2004: 6 A: LE SY
tazaotana subatlassica Bílý, 2006b: 42 N: MO
tazaotana tazaotana Cobos, 1955b: 225 N: MO

502

BUPRESTIDAE

taddertensis Baudon, 1958c: 162

thalassophila iberica Cobos, 1986: 150 E: PT SP
thalassophila pseudokervillei Niehuis, 1990b: 57 E: GR
thalassophila thalassophila Abeille de Perrin, 1900a: 6 E: FR IT MA
togata Abeille de Perrin, 1882: cxlvii A: IS LE SY TR
togataeformis G. Novak, 2001a: 49 A: CY
tractata Abeille de Perrin, 1901a: 8 [RN] E: AR GG A: IN IQ TR
araxana Jakobson, 1913: 791
uniformis Abeille de Perrin, 1900a: 9 [HN]

tragacanthi Iablokoff-Khnzorian, 1957: 161 E: AR GG
truncata Abeille de Perrin, 1900a: 8 E: AB AR ST A: IN IQ IS SY TR
adiyamana Svoboda, 1994a: 66
lgockii Obenberger, 1917d: 38
talyshensis Bílý, 1991b: 123

tuerki Ganglbauer, 1886b: 87 E: AU BU CT CZ GR HU MC SK ST UK
turcica Svoboda, 1994a: 63 A: TR
ulmi Bílý, 2005a: 2 A: SHX
ungulata Bílý, 2015a: 487 A: SCH
urens Abeille de Perrin, 1882: cxlvi A: LE SY TR
phoenica Ganglbauer, 1882a: 68

verdugoi Bílý, 2006b: 51 N: MO
vittula Kiesenwetter, 1857: 91 E: GR
vladivostokana Obenberger, 1938: 201 A: NC SC FE
wittmeri Bílý, 1979b: 219 A: SA
subgenus Cratomerus Solier, 1833: 295 type species Buprestis cyanicornis Olivier, 1790 (=Mordella hungarica
Scopoli, 1772)
bonvouloiri Abeille de Perrin, 1869b: 401 N: AG MO TU
amabilis Obenberger, 1913c: 62
simplicipes Rey, 1890b: 172

brunneicolor Bílý, 1995: 47 A: KI UZ
diadema bazuftensis Obenberger, 1946: 31 A: IN
diadema diadema Fischer von Waldheim, 1824: 196 (Buprestis) E: BU GR MC ME RO ST TR UK A: CY IS JO LE
SY TR N: EG
adoxa Küster, 1852: 76 (Cratomerus)
divina Reiche & Saulcy, 1856: 415
juno Obenberger, 1913c: 62

diadema shelkovnikovi Obenberger, 1940h: 46 E: AB AR GG A: IN
shelkovnikovi Obenberger, 1940k: 166 [second description]

dives Obenberger, 1914f: 113 A: TD
unicolor Obenberger, 1914f: 113 [HN]

dundai Bílý, 1992: 357 A: SHA

BUPRESTIDAE

503

eugeniae eugeniae Ganglbauer, 1885: 317 E: BU GR TR A: LE SY TR
juvenilis Abeille de Perrin, 1895c: cxvi

eugeniae halperini Niehuis, 1993: 40 A: IS JO LE SY
farinigera Kraatz, 1882i: 314 (Cratomerus) A: AF IN KI KZ TD TM UZ
hungarica hungarica Scopoli, 1772: 104 (Mordella) E: AL AU BH BU CR CZ FR GR HU IT MC MD ME PT RO SB
SK SL SP ST SZ TR UK N: AG MO TU A: SY TR
cyanicornis Olivier, 1790a: no 32: 71 (Buprestis)
exaltata Fabricius, 1798: 136 (Buprestis)
femorata Villers, 1789: 338 (Buprestis)
stephanelli V. Petagna, 1787: 23 (Buprestis)
trochilus Fabricius, 1777: 235 (Buprestis)

hungarica sitta Küster, 1852: 75 E: AB AR GG ST UK A: IN TM TR
kondleri Svoboda, 1999: 23 A: TR
krueperi Ganglbauer, 1885: 319 E: AR GG A: IN TR
duo Semenov, 1898b: 599

kryzhanovskii Alexeev, 1978: 846 A: TD
margotana G. Novak, 1983a: 49 E: GR (Kríti)
meda Baiocci, 2011: 10 A: IN
mirabilis Zhicharev, 1918: 245 E: AB AR A: TR
nupta confinata Magnani, 1995a: 47 A: CY
nupta nupta Kiesenwetter, 1857: 82 A: SY TR
paphia G. Novak & Makris, 2002: 111 A: CY
pochoni Herman, 1969: 61 A: PA
scorzonerae I. Frivaldszky, 1837: 174 (Buprestis) E: BU GR MC TR A: TR
artemis K. Daniel, 1904: 344

sponsa Kiesenwetter, 1857: 82 E: AB AR BU GG GR MC RO TR A: IN IS JO LE SY TM TR
cyanescens Marseul, 1865: 213 [HN]
marseuli Obenberger, 1930c: 531 [RN]
violacea Bílý, 1977: 275

subgenus Haplanthaxia Reitter, 1911a: 187 type species Buprestris cichorii A.G. Olivier, 1790
Chrysanthaxia Richter, 1945b: 115 type species Anthaxia polychloros Abeille de Perrin, 1894 sensu Richter, 1945b
(=Buprestis millefolii Fabricius, 1801)
Cratomerella Richter, 1949: 37, 38 type species Anthaxia proteus E. Saunders, 1877
Cryptanthaxia Richter, 1945c: 153 type species Anthaxia rossica K. Daniel, 1903
Cryptocratomerus Richter, 1949: 180, 205 type species Anthaxia kiesenwetteri Marseul, 1865
Mesanthaxia Richter, 1945b: 124 type species Anthaxia olympica Kiesenwetter, 1880

abdita Bílý, 1982b: 113 A: AE IS JO OM SA SI
adenensis Bílý, 1973a: 17 A: YE
aeneocuprea Kerremans, 1914c: 103 A: TAI
agilis Obenberger, 1958a: 231 A: YUN ORR
akiyamai Bílý, 1989b: 383, 392 A: TAI

504

BUPRESTIDAE

aladin Obenberger, 1928c: 69 A: IN
alcmaeone Obenberger, 1938: 181 A: SCH YUN
angulaticollis Kurosawa, 1956: 37 A: TAI
angulinota Bílý, 1984b: 215 A: YE (Suqutra)
angustipennis Klug, 1829: [no 36] (Buprestis) N: AG EG LB MO TU A: IN IQ IS JO SI SY AFR
stupida Marseul, 1865: 224

antinoe Cobos, 1953a: 18 N: AG MO AFR
arakii Kurosawa, 1956: 35 A: JA (Ryukyus)
aspera Bílý, 1977: 278 A: IN IQ
astoreth Obenberger, 1934p: 175 A: IN IS LE SY TR
auricollis Kerremans, 1903: 174 A: HP NP SD UP ORR
achardi Obenberger, 1928a: 237
serenissima Obenberger, 1938: 207

beludjistana Bílý, 1983a: 59 A: IN
berytensis Abeille de Perrin, 1895c: cxvii A: IS LE SY TR
alauita Obenberger, 1938: 188

bettagi Niehuis, 1983: 86 E: GR A: TR
bicolorata Bílý & Svoboda, 2001: 38, 44 A: XIZ
boissyi Levey, 1985: 302 N: AG MO
bosdaghensis Obenberger, 1938: 178 A: TR
cavazzutii Bílý, 1980b: 107

brandli Niehuis & Baiocchi, 2014: 37 A: CY
buettikeri Bílý, 1979b: 220 A: SA
cadusiana Baiocchi 2015: 62, 71 A: IN
caudipennis Bílý, 1983a: 57 A: IN
cichorii A.G. Olivier, 1790a: no 32: 91 (Buprestis) E: AB AL AR AU BE BH BU CR CT CZ FR GE GG GR HU IT MC
MD ME PL RO SB SK SL SP ST SZ TR UK A: IN IQ IS JO LE SY TM TR
azurea Abeille de Perrin, 1909: 23
chamomillae Mannerheim, 1837: 90
gibbicollis Rey, 1891a: 4
parthica Obenberger, 1914a: 14
turcomanica Obenberger, 1938: 179 [HN]

clio Bílý, 1989b: 383, 390 A: TAI
collaris Kerremans, 1893b: 334 A: HP UP ORR
confusa baudii Obenberger, 1914g: 131 E: FR (Corse) IT (Sardegna)
confusa confusa Gory, 1841: 296 E: FR IT PT SP N: AG LB MO TU
congregata Klug, 1829: [no 35] (Buprestis) N: AG EG LB MO TU A: AE IN IQ IS JO SA SI AFR
arabs Marseul, 1865: 216
cairensis Obenberger, 1920a: 84
encaustiventris Abeille de Perrin, 1907a: xlix
obockiana Fairmaire, 1892a: 99

BUPRESTIDAE
cratomerella Bílý & Svoboda, 2001: 38, 45 A: SHA
croesus Villers, 1789: 339 (Buprestis) E: CR FR GR IT PT SP N: AG LB MO TU
atlasica Théry, 1930a: 325
cresus Villers, 1789: 339 (Buprestis) [unused original spelling]
ditescens Abeille de Perrin, 1872: xlii
scutellaris Gené, 1839: 52
semireducta Pic, 1911a: 97
viminalis Gory & Laporte, 1839: 14
viminalis Laporte, 1840: 218 [HN]

crotonivora Bílý, 2005b: 66 A: YE (Suqutra)
curlettii Magnani, 1993: 265 A: CY TR
cylindrica Abeille de Perrin, 1900a: 7 A: CY LE SY TR
dechambrei Bílý, 1991b: 121 A: PA
decima Bílý, 2015b: 50 A: GUA
denesi Svoboda, 2000: 137 A: TR
elaeagni Richter, 1945a: 64, 71 A: AF IN KZ TD TM UZ
elamita Baiocchi 2015: 63, 68 A: IN
erato Bílý, 1989b: 389 A: TAI
erichbettagi Svoboda & Niehuis, 2002: 75 N: MO
escalerina Obenberger, 1923c: 64 [RN] N: MO
superba Escalera, 1914a: 201 [HN]

escalerinella G. Novak, 1988: 41 N: MO
exsul Obenberger, 1914i: 256 A: CH
satanella Obenberger, 1914i: 257

farah Bílý, 1983a: 66 A: IN
fedtschenkoi Semenov, 1896d: 346 A: AF IN TD TM UZ
samarkandi Obenberger, 1921a: 95
scheibei Théry, 1936d: 188

flammifrons Semenov, 1891: 337 A: MG XIN
ignea Obenberger, 1913e: 127

flaviae Lo Cascio & Sparacio, 2011: 499 E: IT (Lipari, Panarea, Salina)
flavicomes Abeille de Perrin, 1900a: 7 E: AB AR A: IN IQ TR
erivana Obenberger, 1913c: 62

formosensis Bílý, 1980b: 110 A: TAI
francescoi Sparacio & Svoboda, 1999: 19 N: AG MO TU
fulgidipennis P.H. Lucas, 1846b: pl. 15 N: AG MO TU
fulgentipennis Abeille de Perrin, 1904a: 215

gedrosiana Bílý, 1983a: 61 A: IN
guizhouensis Bílý, 1996: 25 A: GUI
heyrovskyi Bílý, 1992: 359 A: FUJ JIA ORR
holynskii Bílý, 1990a: 10 A: UP

505

506

BUPRESTIDAE

houskai Obenberger, 1946: 34 A: IS JO LE SY
huashanica huashanica Bílý, 1993: 184 A: SHA
huashanica sculpticollis Bílý & Svoboda, 2001: 39, 42 A: HUN SCH
hypomelaena Illiger, 1803: 252 (Buprestis) E: AL BH BU CR FR GE GR IT MC MD ME PT RO SP ST TR UK N: LB
A: TM TR
nitidicollis Gory, 1841: 287

ignipennis Abeille de Perrin, 1882: cxlvi E: FR PT SP
flammeola Abeille de Perrin, 1907b: l

iliensis Obenberger, 1914f: 115 A: KI KZ NW UZ
iranica Richter, 1949: 212 (Cratomerus) A: IN
israelita Abeille de Perrin, 1882: cxlvi A: IS JO LE SY N: EG
bernardettae Herman, 1966b: 41

iveta Svoboda, 2003: 51 A: IN PA
jordanensis Bílý, 1984b: 213 A: IS JO SY
kabateki Bílý, 2005b: 68 A: YE (Suqutra)
kabyliana Obenberger, 1914b: 11 N: AG TU
kalalae Baiocci & Magnani, 2010: 465 A: IN IQ TR
kantneri Svoboda, 2000: 140 N: TU
karsanthiana Pic, 1917b: 5 E: GR (Ródos) A: CY SY TR
hamata Levey, 1985: 306

kaszabi Cobos, 1966d: 316 A: AF UP
kiesenwetteri Marseul, 1865: 228 [RN] E: BH BU CR GG GR MC ME RO A: IS LE SY TR
cupriventris Marseul, 1865: 215
melancholica Kraatz, 1862: 121 [HN]
rambouseki Obenberger, 1912a: 18

klessi Niehuis, 1991b: 79 A: IS
kneuckeri kneuckeri Obenberger, 1920a: 83 N: EG A: AE IS JO OM SA SI AFR
kneuckeri zabranskyi Bílý, 1995: 52 A: YE
kollari Marseul, 1865: 214 A: IN IQ
starkei Ganglbauer, 1886c: 87

krali Bílý & Kubáň, 2013: 288 A: SCH
kucerai Bílý & Svoboda, 2001: 38, 48 A: SHA
kuldjensis Obenberger, 1913e: 128 A: KI KZ UZ XIN
syrdarjensis Obenberger, 1934p: 176

kurosawai Bílý, 1989b: 383, 391 A: TAI
lameyi Théry, 1910a: 35 A: “China” ORR
lameyiformis Bílý, 1991b: 129 A: GUI SCH YUN
laticeps laticeps Abeille de Perrin, 1900a: 8 E: GR N: LB A: TR
minerva Obenberger, 1913c: 63

laticeps navratili Bílý, 1984b: 214 E: GR (Kríti) A: CY IS SY TR
laticollis Bílý & Kubáň, 2009: 92 A: GAN

BUPRESTIDAE

507

liuchangloi Obenberger, 1958a: 230 A: YUN ORR
ludovicae Abeille de Perrin, 1900a: 7 N: AG MO
lusitanica Obenberger, 1943a: 58 E: PT SP
lyciae Magnani, 1996: 87 A: TR
mannaea Baiocci, 2011: 6 A: IN
marani Obenberger, 1938: 184 A: CY
marginifera dhofarica Bílý, 2002: 200 A: OM YE
marginifera marginifera Abeille de Perrin, 1907a: xlix N: AG LB AFR
marginifera metallescens Bílý, 1999b: 231 [RN] A: IS SA SI
metallica Bílý, 1979b: 219 [HN]

marshalli Stebbing, 1914: 213 A: PA
martinhauseri Niehuis, 1996: 145 A: IS OM
maximiliani Brechtel, 2000: 227 A: YE AFR
medvedevorum Alexeev, 1978: 849 A: TM
millefolieta Svoboda, 2007: 152 E: GR (Kríti)
millefolii millefolii Herbst, 1801: 240 (Buprestis) E: AB AL AR AU BH BU BY CR CZ GE GG GR HU IT KO MC MD
ME PL RO SB SK SL ST TR UK A: CY IS JO LE SY TR
kusnezovi Richter, 1949: 65
millefolii Fabricius, 1801b: 215 (Buprestis) [HN]

millefolii polychloros Abeille de Perrin, 1894e: lxiv E: FR GE GR (Kríti) IT MA PT SP SZ
millefolii scutellata Obenberger, 1914b: 11 A: SY TR
millefolii smaragdifrons Marseul, 1865: 222 E: SP N: AG MO TU
moises Obenberger, 1921a: 96 A: IS JO LE SI SY
mokrzeckii Obenberger, 1927b: 83 A: ZHE
morosa Kerremans, 1892b: 173 A: NP “India” “Himalaya”
oresichaera Obenberger, 1940k: 167

moya ihanatumi Chûjô, 1970: 19 JA (Ryukyus)
moya moya Chûjô, 1970: 18 A: JA (Ryukyus)
mundula Kiesenwetter, 1857: 85 E: GR A: IS LE SY TR
nabataea Baiocci & Magnani, 2013: 375 A: JO
nasheri Bílý, 2008c: 50 A: YE
naviauxi Bílý, 1995: 50 A: IN
niehuisi Brandl, 1987: 14 A: TR
novickii Obenberger, 1938: 181 A: CY SY TR
octava Bílý, 2015b: 47 A: YUN
olivieri Gory & Laporte, 1839: 28 E: GR (Ródos) A: TR
olympica Kiesenwetter, 1880: 131 E: AB AL AR AU BH BU BY CR CZ GG GR HU MC MD ME RO SB SK SL ST TR
UK A: IN TR
ononidis Sánchez & Tolosa, 2006: 55 E: SP
oreas Peyerimhoff, 1919: 195 N: AG MO TU
osmastoni Stebbing, 1911: 80 A: HP UP

508

BUPRESTIDAE

cailloli Obenberger, 1926d: 102
semenovi Obenberger, 1926d: 101

ovaciki G. Novak, 1994a: 2 A: TR
palaestinensis Obenberger, 1946: 32 A: IS JO SI SY
parallela Gory & Laporte, 1839: 15 E: FR PT SP
basalis Küster, 1847b: 29
notaticollis Rey, 1891a: 4

parvula Baiocchi & Magnani, 2006: 219 A: IN
patrizii Théry, 1938c: 268 N: EG
pinda Bílý & Baiocchi, 2009: 63 A: AE OM
pleuralis pleuralis Fairmaire, 1883c: clvii N: AG
parapleuralis Obenberger, 1929e: 61

pleuralis aida Cobos, 1955b: 220 N: MO
praeclara creticola Brandl & Mühle, 1998: 39 E: GR (Kríti)
praeclara praeclara Mannerheim, 1837: 87 E: AL BH BU CR GR IT MC ME TR A: CY IS JO LE SY TR
bodoani Kerremans, 1911: 634
variipennis Küster, 1851: 23

prepsli Bílý, 1995: 51 A: PA
proteiformis Bílý, 1993: 180 A: SCH
proteus E. Saunders, 1873: 511 A: JA
protractipennis Obenberger, 1914i: 255 E: SP N: AG LB MO TU
chlorocephala P.H. Lucas, 1846e: pl. 16 [HN]

pseudocongregata Descarpentries & Bruneau de Miré, 1963: 184 N: AG LB MO AFR
psittacina nigrifrons Bílý & Svoboda, 2001: 36, 40 A: HUN
psittacina psittacina Heyden, 1887b: 303 A: FE NC NE NO SC
puella Bílý, 1980b: 108 A: IN TR
pumila Klug, 1829: [no 37] (Buprestis) N: AG EG LB MO TU A: IS AFR
nephthys Obenberger, 1916d: 260
nickerli Obenberger, 1923e: 83
strandi Obenberger, 1919b: 143

ringenbachi Baiocchi, 2004: 215 N: LB
robusticornis Bílý, 1990a: 12 A: UP
rossica K. Daniel, 1903: 252 E: BU CZ GR HU MC MD RO ST TR UK A: TR
horvathi Kaszab, 1940: 110

roxana Bílý, 1983a: 69 A: AE IN PA ORR
rubromarginata Miwa & Chûjô, 1935: 276 A: FE JA NC NE
egorovi Alexeev, 1979b: 123
primorjensis Obenberger, 1938: 206

rudis Kerremans, 1893b: 336 A: UP ORR
sagartiana Baiocci & Magnani, 2010: 475 A: IN
samai Curletti & Magnani, 1992: 40 N: MO TU

BUPRESTIDAE

509

schah Abeille de Perrin, 1904a: 215 A: IN IQ SY
lokayi Obenberger, 1925b: 32

scylla Levey, 1985: 301 E: IT MA
semiramis Obenberger, 1913e: 128 A: AE IQ IS SY
senilis palmensis Cobos, 1969: 48 N: CI (Palma)
senilis senilis Wollaston, 1864: 208 N: CI
septima Bílý, 2015b: 46 A: HUB
septimadecima Bílý, 2015b: 59 A: YUN ORR
serena K. Daniel, 1903: 252 A: TR
sichuanica Bílý, 1993: 183 A: SCH
sinica Bílý, 1994b: 42 A: SCH YUN
smaragdula Gebhardt, 1929a: 27 A: HEB HUB HUN JIA JIX SCH ZHE TAI YUN
monotona Gebhardt, 1929a: 28

socotrensis Bílý, 1984b: 217 A: YE (Suqutra)
sudana Obenberger, 1928a: 221 N: AG LB AFR
mirei Descarpentries & Mateu, 1965: 1039

taiwanensis Bílý, 1989b: 381, 394 A: TAI
tibetana Bílý, 2002: 198 A: XIZ
toyamai Bílý, 1989b: 383, 388 A: TAI
turana chorasanica Bílý, 1983a: 61 A: IN
turana turana Obenberger, 1914f: 115 A: TD TM
transcaspica Obenberger, 1917b: 56

turcomanica Obenberger, 1937a: 13 A: TR
turnai Bílý & Svoboda, 2001: 37, 47 A: YUN
umbellatarum domina Abeille de Perrin, 1900a: 22 N: AG
robustior Obenberger, 1913c: 63

umbellatarum umbellatarum Fabricius, 1787a: 183 (Buprestis) E: AL BH BU CR FR GE GG GR IT MA MC ME PT
RO SP TR UK N: AG LB MO TU A: TR
aerea Rey, 1891a: 4
inculta Germar, 1817: 217 (Buprestis)

ursulae Niehuis, 1989b: 98 A: IN IQ TR
vejdovskyi peyerimhoffi Obenberger, 1925c: 57 N: AG
vejdovskyi vejdovskyi Obenberger, 1914a: 19 N: AG TU
thais Normand, 1935: 291

vicesimatetria Bílý, 2015b: 69 A: SD
volkovitshi Bílý, 1979a: 203 A: TD
wethloi Obenberger, 1940l: 115 E: AR A: IN
winkleri farsica Bílý, 1983a: 53 A: IN
winkleri judinae Stepanov, 1954b: 1309 A: IN TD TM UZ
winkleri winkleri Obenberger, 1914i: 255 E: AB AR A: IN SY TR
armeniaca Obenberger, 1929c: 123

510

BUPRESTIDAE

yunnana Bílý, 1990a: 22 A: YUN
subgenus Melanthaxia Richter, 1949: 54, 102 type species Anthaxia godeti Gory & Laporte, 1839
acutangula Motschulsky, 1860e: 108 A: ES FE MG
acutiangula Motschulsky, 1860e: 108 [unused original spelling]
burjatica Obenberger, 1924b: 29
granulipennis Théry, 1928b: 77

aeneopicea Kerremans, 1900b: 71 A: HP KA PA
alziari Magnani, 1993: 267 A: CY
aresteni Baudon, 1959a: 106 N: MO
baicalensis Obenberger, 1938: 240 A: ES WS XIN
baiocchii Magnani & Izzillo, 1998: 335 A: LE SY
bilyi Curletti, 1984: 268 A: TR
carmen carmen Obenberger, 1912b: 67 E: PT SP
carmen maroccana Schaefer, 1938b: 254 N: MO
castiliana Obenberger, 1914f: 116 E: SP
caucasica Abeille de Perrin, 1900a: 9 E: AB AR GG A: TR
obenbergeri Roubal, 1913a: 35
parysatis Obenberger, 1913c: 67
vigens Hołyński, 1996: 142

cedri Obenberger, 1938: 216 A: LE
conradti Semenov, 1891: 335 A: KI KZ NMO TD UZ XIZ
bucharica Obenberger, 1913c: 67
canifrons Abeille de Perrin, 1894g: cccli [=1894a: 48]
ferghanensis Obenberger, 1938: 230
semirjetshica Obenberger, 1938: 230
strangulata Abeille de Perrin, 1900a: 9

corinthia Reiche & Saulcy, 1856: 414 E: GR TR A: CY IS JO LE SY TR
rudipennis Obenberger, 1938: 227

corsica corsica Reiche, 1861b: 208 E: FR (Corse)
binotata Rey, 1891a: 5 [HN]
binotatula Obenberger, 1930c: 478 [RN]

corsica maremmana Tassi, 1966: 23 E: IT
corsica tazzekensis Baudon, 1958c: 164 N: MO
cyaneonigra Svoboda, 1995: 245 A: YUN
escalerai Obenberger, 1913c: 68 [justified emendation] A: TR
espanoli Cobos, 1954a: 81 E: SP
frankenbergeri Obenberger, 1933b: 115 E: AL BH BU CR GR
muehlei Niehuis, 1983: 91

giorgioi Sparacio, 2002: 115 E: IT (Sicilia)
godeti Gory & Laporte, 1839: 31 E: AB AL AR AU BE BH BU BY CR CT CZ DE FR GE GG GR HU IT LA MC MD
NL NR NT PL PT RO SK SL SP ST SV SZ TR UK N: AG MO

BUPRESTIDAE

511

cupricolor Abeille de Perrin, 1909: 24
granulata Küster, 1851: 27
raclinae Baudon, 1959a: 109
submontana Obenberger, 1930c: 522

graeca Bílý, 1984c: 266 E: GR
griseocuprea Kiesenwetter, 1857: 99 A: CY LE
grafi Obenberger, 1925b: 31

helvetica appenina Obenberger, 1938: 231 E: IT
helvetica helvetica Stierlin, 1868: 345 E: AU BH BU CR CZ FR GE GR HU IT LS MC PL RO SB SK SL SP SZ UK
nigrocyanea Rey, 1891: 5
pecirkai Obenberger, 1913g: 335
teriolensis Obenberger, 1913c: 69

hemichrysis Abeille de Perrin, 1894g: cccliii [=1894a: 48] A: IN TM
demaisoni Abeille de Perrin, 1895e: ccxxx

heydeni Abeille de Perrin, 1894g: cccliii [=1894a: 48] A: KI KZ
zaitzevi Richter, 1945d: 145

hladili Bílý, 1984d: 436 E: GR
istriana Rosenhauer, 1847: 13 E: AL AU BH BU CR FR GR HU IT MC ME RO SK SL SP SZ TR A: CY SY TR
balatonica Bílý & Somorjai, 1989: 41
bulgarica Obenberger, 1913c: 68
mysteriosa Obenberger, 1917b: 71

kochi Obenberger, 1938: 230 E: IT
kubani Bílý, 1986: 223 E: CR IT RO SB
lencinai Arnáiz & Bercedo, 2003: 4 E: SP
liae Gobbi, 1983: 37 E: IT
malickyi Obenberger, 1925b: 30 A: PA “Kashmir”
mamaj Pliginskiy, 1924: 142 E: ST UK
masculina Bílý, 1984c: 265 E: GR (Ródos, Sporades) A: SY TR
meregallii Curletti & Magnani, 1985: 241 A: TR
montivaga Bílý, 1984d: 445 A: AF PA
morio Fabricius, 1792b: 210 (Buprestis) E: AL AU BE BH BU CT CZ FR GE HU IT LS MC ME NR PL PT RO SK SL
SP SV SZ UK
meridionalis Obenberger, 1913c: 67
similis E. Saunders, 1871: 54

negrei Cobos, 1953c: 99 A: TR
nigritula martini Brisout de Barneville, 1883: 82 N: AG MO TU
nigritula nigritula Ratzeburg, 1837: 52 (Buprestis) E: FR IT GE PT SP SZ
praticola La Ferté-Sénectère, 1841: 49
tatrensis Kaszab, 1940: 112

nigrojubata incognita Bílý, 1974b: 319 E: AL AU BH BU CR CZ FR GE GR HU IT MC ME PL RO SB SK SL SP SZ
UK A: TR UZi

512

BUPRESTIDAE

alexeevi Bílý, 1974b: 320

nigrojubata inexpectata G. Novak, 1986b: 111 E: GR (Kríti)
nigrojubata nigrojubata Roubal, 1913a: 33 E: AB AR GG ST A: TR
obesa Abeille de Perrin, 1900a: 9 E: GR A: SY TR
pinguis Kirsch, 1880a: 131 E: GR A: IS LE SY TR
astarte Obenberger, 1938: 224
mihoki Obenberger, 1938: 225
pohnerti Obenberger, 1925b: 31
schaeferi Cobos, 1953c: 101

potanini Ganglbauer, 1889: 23 A: GAN SCH
quadrifoveolata Solsky, 1871a: 359 A: ES FE MG NC NE
popovi Obenberger, 1938: 241

quadripunctata quadriimpressa Motschulsky, 1859a: 226 A: ES FE HEB HEI JIL KZ LIA MG NMO SHX WS XIN
attavistica Obenberger, 1918b: 22

quadripunctata quadripunctata Linnaeus, 1758: 410 (Buprestis) E: AB AL AN AR AU BE BH BU BY CR CT CZ
DE EN FI FR GB GE GG GR HU IT KO LA LS LT LU MC MD ME NL NR NT PL RO SB SK SL SP ST SV SZ TR
UK A: TR WS
angulata Küster, 1851: 30
angulicollis Küster, 1851: 28
punctata Ponza, 1805: 81 (Buprestis)

reticulata aino Kurosawa, 1963a: 94 A: JA (Hokkaido)
reticulata reticulata Motschulsky, 1860h: 490 A: ES FE MG NC NE NO
jureceki Obenberger, 1924b: 29
transbaicalica Obenberger, 1927b: 84
ussuriensis Obenberger, 1914f: 116

reticulata shinano Kurosawa, 1963a: 94 A: JA (Honshu)
reticulata yamato Kurosawa, 1963a: 94 A: JA (Honshu)
rugicollis atlantica Obenberger, 1938: 222 N: CI
rugicollis rugicollis P.H. Lucas, 1846b: pl. 15 E: FR IT (Sardegna) PT SP N: AG MO TU
rugicollis tassii Herman, 1966a: 241 E: GR A: TR
rugicollis zoufali Obenberger, 1925a: 101 E: SP (Baleares)
scurra Bílý, 1982c: 187 A: TR
segurensis Obenberger, 1924b: 28 E: SP
sepulchralis chobauti Abeille de Perrin, 1894e: lxiv N: AG MO
sepulchralis sepulchralis Fabricius, 1801b: 215 (Buprestis) E: AU CZ FR GE HU IT NL PL PT SP SZ UK
sepulchraloides Cobos, 1962c: 294 N: MO
sternalis Abeille de Perrin, 1895c: cxviii A: TR
sturanyii Obenberger, 1914g: 132 E: AL GR
tauricola Magnani, 2002: 99 A: TR
tenella tenella Kiesenwetter, 1858: 246 E: BH BU CR GR MC
insularis Obenberger, 1914b: 14

513

BUPRESTIDAE
tenella virgo Bílý, 1980c: 405 A: TR
thessalica Brandl, 1981: 1 E: AL BU GR ME
tianshanica Bílý, 1984d: 438 A: KI KZ XIN
tomyris Obenberger, 1913g: 333 A: KI
turkestanica Obenberger, 1912b: 68 A: KI KZ
venerabilis Marseul, 1869a: 122 [RN] A: “Orient”
verecunda Marseul, 1865: 256 [HN]

villiersi Baudon, 1959a: 111 N: MO
xiahensis Bílý, 1995: 53 A: GAN
zarazagai Arnáiz & Bercedo, 2003: 2 E: SP
subgenus Merocratus Bílý, 1989b: 384 type species Anthaxia insulaecola Obenberger, 1944
bellissima Bílý, 1990a: 4 A: SD ORR
insulaecola Obenberger, 1944f: 134 A: TAI
subgenus Thailandia Bílý, 1990a: 2 type species Thailandia paradoxa Bílý, 1990
capitata Kerremans, 1892b: 173 A: UP ORR
phyllanthi Obenberger, 1956b: 238 A: UP
genus Brachelytrium Obenberger, 1923a: 24 type species Brachelytrium transvalense Obenberger, 1923
subgenus Elytrobrachium Bílý & Bellamy, 2000: 104 type species Anthaxia blairi Obenberger, 1931
jemeni Brechtel, 2000: 229 A: YE
genus Chalcogenia E. Saunders, 1871: 51 type species Anthaxia contempta Mannerheim, 1837
Chalcogenia C.G. Thomson, 1879: 25 [HN] type species Anthaxia cupraea Laporte & Gory, 1839 (=Anthaxia contempta
Mannerheim, 1837)

halperini arabica Bílý, 2008a: 122, 137 A: OM SA YE
halperini halperini Volkovitsh & Bílý, 1997: 249 A: IS JO SI
nana Bílý, 2012: 210 A: YE (Suqutra)
theryi Abeille de Perrin, 1897b: 28 N: EG A: IS AFR

tribe Bubastini Obenberger, 1920

genus Bubastoides Kerremans, 1909b: 438 type species Bubastoides argodi Kerremans, 1909
kadleci Bílý, 2008b: 55 A: YE

tribe Buprestini Leach, 1815

genus Buprestis Linnaeus, 1758: 408 type species Buprestis octoguttata Linnaeus, 1758
subgenus Akiyamaia Kurosawa, 1988: 265 type species Buprestis (Cypriacis) mirabilis Kurosawa, 1969
Himalobuprestis Kurosawa, 1988: 266 type species Ancylochira costipennis Fairmaire, 1891

costipennis Fairmaire, 1891e: cxxvii (Ancylocheira) A: AF HP PA UP “India” “Kashmir”
impressicollis Kerremans, 1892b: 172 (Buprestis)
kashmirensis Stebbing, 1914: 212 (Buprestis)

intercostata Huang & Pan, 2015: 392 A: NP XIZ

514

BUPRESTIDAE

lebisi Descarpentries, 1956: 230 (Buprestis) A: YUN
mirabilis Kurosawa, 1969a: 191 (Buprestis) A: TAI
samanthae Hattori & Tanaka, 2007: 328 A: YUN ORR
subgenus Ancylocheira Eschscholtz, 1829: 9 type species Buprestis rustica Linnaeus, 1758
Ancylochira Eschscholtz, 1829: 9 [unused original spelling] type species Buprestis rustica Linnaeus, 1758
Anoplis Kirby, 1837: 151 type species Buprestis rustica Linnaeus, 1758
Gymnota Gistel, 1834: 10 type species Buprestis rustica Linnaeus, 1758

apicipennis Reitter, 1898c: 344 A: LE SY TR
severa Abeille de Perrin, 1904a: 213

atlas Szallies, 1994: 357 N: MO
bertheloti Laporte & Gory, 1837: 142 N: CI
bertholoti Laporte & Gory, 1837: 142 [incorrect original spelling]

cupressi Germar, 1836: 2 E: AL BH BU CR GR IT MC ME SB TR UK (Krym) N: EG A: IQ SY TR
cupressi Laporte & Gory, 1837: 126 [HN]
mutabilis O.G. Costa, 1839a: 9 (Buprestis)

dalmatina Mannerheim, 1837: 62 E: AL BH CR GG GR ME TR UK (Krym) N: EG A: CY SY TR
flavostrigata I. Frivaldszky, 1845: 179
ledereri Marseul, 1865: 179 (Ancylocheira)

decemspilota Hope, 1831: 25 A: HP KA NP PA SD UP
decemnotata Laporte & Gory, 1837: 133
geometrica Laporte & Gory, 1837: 141
sanguineosignata Théry, 1926c: 158

esakii Kurosawa, 1954a: 33 A: JA (Ryukyus) TAI
fairmairei Théry, 1910a: 16 [RN] A: SCH YUN ORR
marginicollis Fairmaire, 1902b: 268 (Ancylocheira) [HN]

flavoangulata baetica Verdugo, 2005: 12 E: SP
flavoangulata flavoangulata Fairmaire, 1856: 530 (Ancylocheira) N: AG MO
guttatipennis Abeille de Perrin, 1900a: 4 (Ancylocheira) A: LE TR
haardti Théry, 1934b: 317 A: GAN NIN QIN SCH XIZ YUN
haemorrhoidalis arakii Kurosawa, 1942: 421 A: JA (Ryukyus)
haemorrhoidalis araratica Marseul, 1865: 174 (Ancylocheira) E: AR GG ST UK (Krym) A: TR
haemorrhoidalis haemorrhoidalis Herbst, 1780: 97 E: AL AN AU BE BH BU BY CR CT CZ DE EN FI FR GE GR
HU IT KO LA LS LT MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: CI A: KZ WS AFRi
discoidea Laporte & Gory, 1837: 129
haemhorroidalis Herbst, 1780: 97 [incorrect original spelling]
marginicollis Laporte & Gory, 1837: 130
niphe Gistel, 1857a: 49 (Ancylocheira)
punctata Fabricius, 1787a: 176
quadristigma Herbst, 1801: 177

haemorrhoidalis japanensis E. Saunders, 1873: 511 A: FUI JA SC
coreana Obenberger, 1917e: 50

BUPRESTIDAE

515

haemorrhoidalis rifensis Kocher, 1964: 196 N: MO
haemorrhoidalis sachalinensis Miwa & Chûjô, 1935: 272 A: FE (Sakhalin)
haemorrhoidalis sibirica Fleischer, 1887: 236 [HN] A: ES FE GAN HEI HUN KI MG NC NMO SCH SHA WS
XIN YUN
brunneola Obenberger, 1919b: 143

humeralis Klug, 1829: [no 25] E: GR TR N: EG A: CY IN IQ IS LE SY TR
robertsi Cobos, 1972b: 30
tarsensis Marseul, 1865: 178 (Ancylocheira)

metallescens Abeille de Perrin, 1891: 261 A: TR
novemmaculata gravida Abeille de Perrin, 1904a: 214 (Ancylocheira) E: AL CR GG GR ME ST TR UK (Krym)
A: TR
novemmaculata novemmaculata Linnaeus, 1767: 662 E: AL AU BE BH BU BY CR CT CZ DE EN FI FR GE GR HU
IT LA LT MA MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: AG CI MO TU A: ES KI KZ UZ WS
AFRi NTRi
aurantiacoadspersa C.É. Blanchard, 1853: 95
flavopunctata De Geer, 1774: 129
maculata Fabricius, 1781: 275
maculosa Gmelin, 1790: 1929 [RN]
octomaculata Pallas, 1781: 72
rubromaculata Laporte & Gory, 1837: 132
tetrastichon Linnaeus, 1767: 656 (Elater)

rustica nickerli Obenberger, 1925c: 57 E: GG
rustica rustica Linnaeus, 1758: 409 E: AL AN AU BE BH BU BY CR CT CZ DE EN FI FR GE GR HU IT LA LS LT
LU MC MD ME NR NT PL PT RO SB SK SL SP ST SV SZ TR UK A: ES FE KZ TR WS
violacea De Geer, 1774: 130

strigosa Gebler, 1830: 78 A: ES FE KZ MG NC NMO WS
subgenus Buprestis Linnaeus, 1758: 408 type species Buprestis octoguttata Linnaeus, 1758
aetnensis Baviera & Sparacio, 2002: 93 E: IT (Sicilia)
magica doderoi Sparacio, 2015: 757 E: IT (Sardegna)
magica magica Laporte & Gory, 1837: 138 E: SP N: AG MO TU
corpulenta Fairmaire, 1884e: cxviii (Ancylocheira)
mauritanica Lucas, 1844a: 50

octoguttata corsica Schaefer, 1946a: 72 E: FR (Corse)
octoguttata octoguttata Linnaeus, 1758: 408 E: AL AU BH BU BY CR CT CZ EN FI FR GE GG GR HU IT LA LT
MC MD ME NR NT PL PT RO SB SK SL SP ST SV SZ TR UK A: KZ TR WS
albopunctata De Geer, 1774: 132

subgenus Cypriacis Casey, 1909: 116 type species Ancylocheira lauta LeConte, 1854 (=Buprestis aurulenta
Linnaeus, 1767)
splendens niponica Hoscheck, 1931: 727 A: JA
miwai Obenberger, 1943b: 110
niponica Miwa & Chûjô, 1935: 273

516

BUPRESTIDAE

splendens splendens Fabricius, 1775: 221 E: AL AU BH CR CZi DEi GBi GE GR HU IT NT PL RO SP SV UK A: CH
pretiosa Herbst, 1801: 127
splendida Paykull, 1799: 229

subgenus Orthocheira Richter, 1952: 33 type species Buprestis salomonii J. Thomson, 1878
salomonii J. Thomson, 1878a: 40 E: AB AR A: AF IN KZ PA TD TM TR UZ XIN
davidis Fairmaire, 1884g: clxviii (Ancylocheira)
denzleri Pochon, 1972: 103
nikolskii Semenov, 1889c: 682
vavrai Obenberger, 1930c: 388 [unjustified emendation]
wawrai Nonfried, 1894b: 45 (Ancylocheira)

subgenus Pseudyamina Richter, 1952: 31 type species Buprestis variegata Klug, 1829
bilyi G. Novak & Kubáň, 1993: 261 N: AG MO
douei P.H. Lucas, 1846b: pl. 15 E: IT (Sardegna) SP N: AG LB MO TU
amori Graells, 1858: 43
bellemarei P.H. Lucas, 1853b: lxviii
bellemaroei P.H. Lucas, 1853b: lxviii [incorrect original spelling]

hilaris Klug, 1829: [no 22] N: AG EG LB A: IS SI
variegata Klug, 1829: [no 21] [HN]

genus Eurythyrea Dejean, 1833: 78 type species Buprestis austriaca Linnaeus, 1767
aurata Pallas, 1776: 719 (Buprestis) E: AB AL AR BU GG GR HU KZ ME RO ST UK A: IN TM TR
inaurata Gmelin, 1790: 1939 (Buprestis) [RN]

austriaca Linnaeus, 1767: 661 (Buprestis) E: AL AU BH BU BY CR CZ FR GE GG GR HU IT MC ME PL RO SB SK
SL SP ST TR UK N: AG
eoa Semenov, 1895d: 256 A: FE JA (Hokkaido) NE
micans Fabricius, 1792b: 189 (Buprestis) E: FR IT PT SP N: AG MO TU
marginata A.G. Olivier, 1790a: no 32: 67 (Buprestis) [HN]
planicosta Théry, 1910a: 15

oxiana Semenov, 1895d: 254 A: TD TM UZ
quercus Herbst, 1780: 90 (Buprestis) E: AB AL AR AU BH BU BY CR CT CZ FR GE GG GR HU IT PL PT RO SK
SL SP ST SZ UK A: TR
carniolica Herbst, 1801: 122 (Buprestis)
oblita Faldermann, 1835a: 145
scutellaris A.G. Olivier, 1790a: no 32: 86 (Buprestis)
similis Schönherr, 1817: 221 (Buprestis) [RN]

tenuistriata Lewis, 1893: 330 A: JA
kawakamii Miwa & Chûjô, 1935: 271
obenbergeri Kurosawa, 1963a: 90

genus Eurythyrea, nomen dubium
fastidiosa P. Rossi, 1790a: 186 (Buprestis) E: IT

517

BUPRESTIDAE
genus Lamprocheila E. Saunders, 1871: 15 type species Chrysodema maillei Laporte & Gory, 1835
maillei Laporte & Gory, 1835: 12 (Chrysodema) A: SCH YUN ORR
splendida Akiyama, 1993b: 103 A: TAI
genus Sinokele Bílý, 1989a: 234 type species Sinokele mirabilis Bílý, 1989
mirabilis Bílý, 1989a: 234 A: SCH
genus Yamina Kerremans, 1903: 143 type species Buprestis sanguinea Fabricius, 1798
kruegeri Hoscheck, 1929: 390 (Yamina) N: LB
sanguinea calpetana Verdugo, 2006: 80 E: SP (Gibraltar)
sanguinea dumonti Théry, 1942b: 132 (Yamina) N: TU
sanguinea iberica Verdugo, 2006: 81 E: SP
sanguinea sanguinea Fabricius, 1798: 135 E: PT SP N: AG MO
levaillantii Lucas, 1844a: 50
margaripicta Marseul, 1865: 185 (Ancylocheira)

tribe Chrysobothrini Gory & Laporte, 1938

genus Chrysobothris Eschscholtz, 1829: 9 type species Buprestis chrysostigma Linnaeus, 1758
subgenus Abothris Semenov & Richter, 1934: 92 type species Chrysobothris jakovlevi Semenov, 1891
astartae Abeille de Perrin, 1895a: x N: AG MO TU
bicolorata Bílý, 2000: 20 A: IN
bumburetica Bílý, 2000: 22 A: PA
cuprina Klug, 1829: [no 39] (Buprestis) N: AG EG LB A: “Arabia” JO AFR
impressifrons Fairmaire, 1884g: cxlv

curlettii Magnani, 1995b: 102 N: AG MO
jakovlevi Semenov, 1891: 340 A: TM
jakowlewi Semenov, 1891: 340 [unused original spelling]

marina Abeille de Perrin, 1907a: xlviii N: AG EG A: OM YE AFR
nana Fairmaire, 1892d: clii A: AF IN KZ TD TM UZ
partha Obenberger, 1924b: 30
turkestanica Obenberger, 1924b: 29

samai Curletti & Magnani, 1988: 143 A: TR
yemenensis Bílý, 2000: 24 A: YE
subgenus Chrysobothris Eschscholtz, 1829: 9 type species Buprestis chrysostigma Linnaeus, 1758
Aglaura Gistel, 1848a: xi, 128 [SN]
Amblis Gistel, 1834: 10 type species Buprestis chrysostigma Linnaeus, 1758
Euocys Gistel, 1856: 336 [SN] type species Buprestis chrysostigma Linnaeus, 1758
Knowltonia Fisher, 1935: 117 type species Knowltonia biramosa Fisher, 1935
Odontomus Kirby, 1837: 156 type species Buprestis chrysostigma Linnaeus, 1758
Tamina Gistel, 1848b: [2] [SN] type species Buprestis chrysostigma Linnaeus, 1758

affinis adusta Abeille de Perrin, 1901: 8 [RN] E: GR

518

BUPRESTIDAE

carinithorax Obenberger, 1917e: 52
cavinithorax Obenberger, 1917e: 52 [unused original spelling]
tristis Abeille de Perrin, 1900a: 4 [HN]

affinis affinis Fabricius, 1794: 450 (Buprestis) E: AL AU BE BH BU BY CR CT CZ DE FI FR GE GG GR HU IT KO
KZ LA LS LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK A: CY IN IQ SY TR
congener Paykull, 1799: 222 (Buprestis)
materculae Hołyński, 1975: 357

affinis heliophila Abeille de Perrin, 1894g: ccclii [=1894a: 47] N: AG EG LB MO TU
affinis kostini Bellamy, 1998: 93 [RN] A: KZ
mali Kostin, 1973: 92 [HN]

affinis nevskyi Richter, 1944b: 72 A: IN KZ TD TM UZ
affinis tetragramma Ménétriés, 1832: 149 (Buprestis) E: AB AR GG A: IN
inaequalis Reitter, 1895c: 128 [HN]
reitteri Kerremans, 1900c: 304 [RN]

affinis tremulae Kostin, 1973: 92 A: KZ
amurensis amurensis Pic, 1904a: 25 A: FE HEI NC
amurensis ohmomoi Akiyama, 1993a: 35 A: JA
andamana Kerremans, 1891b: clxii A: UP ORR
boninensis boninensis Kurosawa, 1975a: 70 A: JA (Ogasawara)
boninensis suzukii Kurosawa, 1975a: 73 A: JA (Ogasawara)
bouddah Théry, 1940b: 144 A: YUN
brahma Obenberger, 1917h: 123 A: “India”
carinata Kerremans, 1892b: 197 A: “India”
cashmirensis Obenberger, 1934n: 149 A: NP PA SD “Kashmir” ORR
catei Barries, 2009: 55 A: IN
chalcophana Klug, 1829: [no 38] (Buprestis) N: AG AFR
cheni Théry, 1940b: 143 A: BEI JIA
chrysostigma chrysostigma Linnaeus, 1758: 409 (Buprestis) E: AU BH BU BY CR CT CZ DE EN FI FR GB GE GG
GR HU IT LA LS LT MD NR NT PL RO SK SL ST SV SZ UK A: ES FE KA KI KZ TR WS
jureceki Obenberger, 1913b: 29
stepaneki Obenberger, 1940k: 169

chrysostigma kerremansi Abeille de Perrin, 1894f: cxxxi [RN] A: ES FE GAN HEI JIL LIA MG NC NMO SC SCH
XIN YUN
quadraticollis Abeille de Perrin, 1894g: ccclii [=1894a: 47] [HN]

chrysostigma pekinensis Théry, 1940b: 146 A: BEI HEB HUB SHN
chrysostigma yezostigma Kurosawa, 1963a: 94 A: JA
costata Kerremans, 1895: 213 A: TAI ORR
cypria Magnani, 1993: 268 A: CY
delavayi Fairmaire, 1887b: 119 A: YUN
dorsata dorsata Fabricius, 1787a: 179 (Buprestis) E: PT SP N: AG MO AFR
dudichi Gebhardt, 1926: 50 A: AP ORR

BUPRESTIDAE

519

felixi Gebhardt, 1926: 51 A: AP ORR
gardneri Théry, 1930c: 152 A: PA ORR
gebhardti Théry, 1936c: 119 E: AR TR
grancanariae Niehuis & Gottwald, 1999: 111 N: CI
hexastigma Mannerheim, 1837: 78 A: KZ TD TM UZ
deserticola Semenov & Richter, 1934: 94

horvathi Gebhardt, 1926: 49 A: AP ORR
igai Kurosawa, 1948: 18 A: JA
karasawai Kurosawa, 1963a: 96
kumagaii Kurosawa, 1963a: 95
kurosawa Bellamy, 1996: 241 [RN]
monticola Kurosawa, 1963a: 96 [HN]
nikkoensis Kurosawa, 1963a: 95

igniventris Reitter, 1895c: 129 E: AU BU CT CZ GE HU IT PL RO SB SK ST UK
indica Laporte & Gory, 1837: 12 A: GUX HUN JIX NP UP SCH SD YUN ORR
delenifica Deyrolle, 1864b: 109
laosensis Obenberger, 1928a: 307

infranitens Kerremans, 1912c: 204 A: TAI
kabakovi Alexeev, 1991: 864 A: AF
kiangsuanus Théry, 1940b: 156 A: JIA
kotoensis Miwa & Chûjô, 1940: 68 A: TAI
kucerai Barries, 2008: 71 A: UP
leonhardi leonhardi Obenberger, 1916d: 261 E: BU CZ GR HU MC RO
leonhardi turcica Magnani, 1996: 92 A: TR
libanonica Obenberger, 1935p: 196 A: LE
manchurica Arakawa, 1932: 16 A: NC “Manchuria”
mandarina Théry, 1940b: 151 A: HUN JIX ZHE
ohbayashii Kurosawa, 1948: 15 A: JA
ohnoi Kurosawa, 1975a: 67 A: JA
parvipunctata Obenberger, 1914g: 133 A: AE AF IN IQ PA TM UP ORR
beesoni Obenberger, 1928a: 308
kheirii Cobos, 1970b: 191
zarudniana Semenov & Richter, 1934: 94

pulchripes Fairmaire, 1887d: 332 A: BEI FE GAN HEB HEI JA (Tsushima) JIA NC NIN NMO QIN SC SHA XIN
confucius Théry, 1940b: 158
sivah Théry, 1940b: 160
tientsinensis Théry, 1940b: 160
tsushimae Obenberger, 1936e: 71

quadraticollis Kerremans, 1892b: 198 A: SD ORR
rejzeki Niehuis, 2009a: 115 A: SA YE
saliaris saliaris Kurosawa, 1948: 26 A: JA

520

BUPRESTIDAE

saliaris yaeyamana Kurosawa, 1975a: 70 A: JA
samurai Obenberger, 1935p: 195 A: BEI FE HEB HEI HEN JA NC NMO SCH SHA SHX TIA
daisenensis Kurosawa, 1963a: 98
koreana Kurosawa, 1975a: 74
laevicollis Kurosawa, 1948: 20
licenti Théry, 1940b: 149
trassaerti Théry, 1939b: 153

sapphirina Swartz, 1817: 121 (Buprestis) A: “India”
sauteri Kerremans, 1912c: 204 A: GUA TAI
sexnotata Gory, 1841: 159 A: SD UP “China” ORR
shinanensis Kano, 1929: 94 A: FUI GAN HUN JIA JIX SHG
pieli Théry, 1940b: 155

shirakii Miwa & Chûjô, 1935: 286 A: TAI
sinensis Fairmaire, 1887b: 119 A: BEI FE JIX NMO SHX YUN
singalesa Obenberger, 1922b: 102 A: NP ORR
solieri Laporte & Gory, 1837: 10 E: AL BE BH BU CR FR GE GR IT MA MC ME NL PT RO SP SZ TR N: AG EG LB
MO TU A: CY IN IQ IS LE SY TR
bedeli Pic, 1930a: 10
multipunctata Companyo, 1863: 641
obenbergeriana Lotte, 1938b: 151 [RN]
pini Klingelhöffer, 1845: 349
quadridens Rey, 1890b: 172
sterbai Obenberger, 1935p: 196 [HN]

spuria Marseul, 1866: 411 E: GR IT (Sicilia) N: EG LB A: IQ IS JO SA YE
mediterranea G. Novak, 1986a: 37

succedanea E. Saunders, 1873: 512 A: FUI GAN GUX HEI HEN HKG HUB HUN JA JIA JIL JIX LIA NC NIN QIN
SC SCH SHN ZHE
tonkinensis Bourgoin, 1922a: 21 A: TAI ORR
tricolor Kerremans, 1892b: 196 A: “India”
violacea Kerremans, 1892b: 199 A: AP NP SCH XIZ YUN ORR
clermonti Obenberger, 1925c: 58
guerryi Théry, 1910a: 48

vitalisi Bourgoin, 1922a: 21 A: GUI SCH YUN ORR
yunnanensis Théry, 1940b: 153 A: NP YUN
znojkoi Semenov & Richter, 1934: 93 A: “Kashmir”
genus Sphaerobothris Semenov & Richter, 1934: 93 type species Chrysobothris globicollis Reitter, 1895
subgenus Hoggarobothris Théry, 1935c: 260 type species Hoggarobothris dinauxi Théry, 1935
dinauxi Théry, 1935c: 260 (Hoggarobothris) N: AG LB AFR
subgenus Sphaerobothris Semenov & Richter, 1934: 93 type species Chrysobothris globicollis Reitter, 1895

521

BUPRESTIDAE
aghababiani Volkovitsh & Kalashian, 1998: 307 E: AR
globicollis Reitter, 1895c: 130 (Chrysobothris) A: AF TD TM UZ
tschitscherini Semenov, 1899c: 652 (Chrysobothris)

tribe Coomaniellini Bílý, 1974

genus Coomaniella Bourgoin, 1924b: 178 type species Coomaniella modesta Bourgoin, 1924
subgenus Coomaniella Bourgoin, 1924b: 178 type species Coomaniella modesta Bourgoin, 1924
Coomaniellina Bílý, 1974a: 34 type species Coomaniella violaceipennis Bourgoin, 1924

chinensis Jendek & Kalashian, 1999: 21 A: SCH
copipes Jendek & Pham, 2013: 116 A: GUX ORR
janka Jendek, 2005: 1 A: HEN
marseuli Obenberger, 1940k: 165 A: ZHE
purpurascens Baudon, 1966a: 56 A: BT SD YUN ORR
kurosawai Bílý, 1974a: 32
michaeli Bílý, 1974a: 33

subgenus Tuberniella Jendek & Kalashian, 1999: 33 type species Coomaniella abeillei Obenberger, 1940
abeillei Obenberger, 1940k: 166 A: TAI
taiwanensis Baudon, 1966b: 10 A: TAI
formosensis Bílý, 1974a: 38

tribe Kisanthobiini Richter, 1949

genus Kisanthobia Marseul, 1865: 200 type species Anthaxia ariasi Robert, 1859
ariasi Robert, 1858: 220 (Anthaxia) E: BH BU CR FR GR HU IT SP ST N: AG TU A: CY LE SY TR
curta Pic, 1898a: 2

tribe Melanophilini Bedel, 1921

genus Melanophila Eschscholtz, 1829: 9 type species Buprestis acuminata De Geer, 1774
Apatura Laporte & Gory, 1837: 1 type species Buprestis appendiculata Fabricius, 1792 (=Buprestis acuminata De
Geer 1774)
Oxypteris Kirby, 1837: 160 type species Buprestis appendiculata Fabricius, 1792 (=Buprestis acuminata De
Geer 1774)

acuminata De Geer, 1774: 133 (Buprestis) E: AB AL AN AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE
GG GR HU IT KO LA LS LT LU MC MD NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG A: ES FE
GAN HUB IN KA KI KZ MG NE NMO NP SHA TR UZ WS XIN XIZ YUN NAR
acuminata Thunberg, 1787: 52 (Buprestis) [HN]
acuta Gmelin, 1790: 1939 (Buprestis) [RN]
amica Gistel, 1857a: 38 (Phaenops)
anthaxioides Marquet, 1870: 368
appendiculata Fabricius, 1792b: 210 (Buprestis)
assimilis LeConte, 1850: 227

522

BUPRESTIDAE

immaculata Mannerheim, 1837: 70
longipes Say, 1823: 164 (Buprestis)
opaca LeConte, 1860: 213
pecchiolii Gory & Laporte, 1939: 33 (Anthaxia)
rugata LeConte, 1857a: 7
summifex Gistel, 1857a: 6 (Phaenops)

cuspidata Klug, 1829: [no 34] (Buprestis) E: AB AL AR BH BU CR FR GG GR IT MA MC ME PT SL SP TR N: AG
EG LB MO TU A: AE AF CY IN IQ IS JO KI KZ LE SI SY TD TM TR UZ AFR
acutespina Reitter, 1893d: 175
aequalis Mannerheim, 1837: 71
aerata A. Costa, 1883: 42
jakovlevi Semenov, 1900d: 589
jakowlewi Semenov, 1900d: 589 [unused original spelling]
nigrita Fabricius, 1798: 134 (Buprestis) [HN]
oxyura Marquet, 1874: 32
regalis Fabricius, 1801b: 187 (Buprestis) [RN] [HN]

gestroi Obenberger, 1923e: 82 N: EG AFR
ignicola H.G. Champion, 1918: 199 A: AF NP UP ORR
obscurata Lewis, 1893: 331 A: FE JA NC NE SC
genus Phaenops Dejean, 1833: 79 [gender feminine, by Opinion 1826, 1996] type species Buprestis cyanea
Fabricius, 1775
aerea Ganglbauer, 1886e: 309 E: GR A: CY SY TR
chalcea Abeille de Perrin, 1895c: cxvi (Melanophila)

cyanea Fabricius, 1775: 223 (Buprestis) E: AB AL AN AR AU BE BH BU BY CR CT CZ DE EN FI FR GE GG GR
HU IT KO LA LS LT LU MC MD ME NL NR NT PL PT RO SK SL SP ST SV SZ TR UK N: AG MO A: ES FE MG
SY TR WS
azurea A.G. Olivier, 1790: Errata [3] (Buprestis)
calybea Villers, 1789: 339 (Buprestis) [unused original spelling]
chalybaea Villers, 1789: 339 (Buprestis)
clypeata Paykull, 1799: 223 (Buprestis)
tarda Fabricius, 1792b: 209 (Buprestis)
transbaicalica Obenberger, 1924b: 20 (Melanophila)

delagrangei cypria Magnani, 1993: 264 A: CY
delagrangei delagrangei Abeille de Perrin, 1891: 268 (Melanophila) A: LE SY TR
lycica Obenberger, 1944g: 147 (Melanophila)

formaneki bohemica Bílý, 1976: 33 E: AU CZ GE
formaneki formaneki Jakobson, 1913: 790 [RN] E: AU BH BU BY CR CT CZ FI GR HU IT MC PL RO SB SK ST SV
SZ TR UK A: ES FE MG TR WS
aerea Formánek, 1900: 168 [HN]

523

BUPRESTIDAE
laricicola Stepanov, 1949: 108
recticollis Pic, 1918c: 5
sibirica Pic, 1918c: 1
thessala Obenberger, 1924b: 20 (Melanophila)
viridescens Pic, 1909e: 169

formaneki lavagnei Théry, 1942a: 73 E: FR SP SZ
guttulata Gebler, 1830: 75 (Buprestis) E: NT A: ES FE MG WS
discopunctata Faldermann, 1833: 47 (Buprestis)

knoteki barbeyi Peyerimhoff, 1932: 125 (Melanophila) N: AG
knoteki knoteki Reitter, 1898c: 345 E: AU BH BU CR GR IT MC PL RO SK SL UK
knoteki ochsi Schaefer, 1947: 192 E: FR IT
knoteki turcica Niehuis, 1991a: 44 A: TR
marmottani Fairmaire, 1868a: 483 (Melanophila) N: AG MO
legrandi Mulsant & Pellet, 1872: 201 (Melanophila)

sumptuosa Abeille de Perrin, 1904a: 214 (Melanophila) E: FR SP
yang Kubáň & Bílý, 2009: 117 A: GUI HEB SCH XIZ YUN
yin Kubáň & Bílý, 2009: 115 A: GAN HEB SHA SHX
genus Trachypteris Kirby, 1837: 158 type species Buprestis decostigma Fabricius, 1787
picta decostigma Fabricius, 1787a: 180 (Buprestis) E: AB AL AR AU BH BU BY CR CZ FR GG GR HU IT MC MD
ME PT RO SB SK SL SP ST SZ TR UK N: AG MO TU A: CY IN IQ IS SY TR NTRi
consobrina Chevrolat, 1854a: 395 (Melanophila)
quatuordecimguttata A.G. Olivier, 1790a: no 32: 89 (Buprestis)
silphoides Schrank, 1789: 82 (Buprestis)

picta indica Théry, 1930c: 149 (Melanophila) A: UP ORR
picta picta Pallas, 1773: 719 (Buprestis) E: ST A: AF BEI GAN HEI HUN IN KA KI KZ MG NIN NMO PA SHA SHX
TD TM UZ WS XIN ORR
leonhardi Obenberger, 1924b: 25 (Melanophila)

tribe Thomassetiini Bellamy, 1987
subtribe Philanthaxiina Hołyński, 1988

genus Kurosawaia Tôyama & Ohmomo, 1985: 33 type species Philanthaxia yanoi Kurosawa, 1963
yanoi Kurosawa, 1963a: 92 (Philanthaxia) A: JA (Ogasawara)
genus Philanthaxia Deyrolle, 1864a: 72 type species Philanthaxia curta Deyrolle, 1864
Engycera E. Saunders, 1866: 308 type species Engycera rufimarginata E. Saunders, 1866

convexifrons Kurosawa, 1954a: 34 A: TAI
iriei Kurosawa, 1985b: 141 A: JA (Ryukyus)
sauteri Kerremans, 1913b: 110 A: TAI
simonae Bílý, 1983b: 215 A: NP

524

BUPRESTIDAE
subfamily Agrilinae Laporte, 1835
tribe Agrilini Laporte, 1835
subtribe Agrilina Laporte, 1835

genus Agrilus Curtis, 1825: 67 type species Buprestis viridis Linnaeus, 1758
Samboides Kerremans, 1900a: 16 type species Samboides viridana Kerremans, 1900
Sarawakita Obenberger, 1924c: 39 type species Sarawakita latifrons Obenberger, 1924
Teres Harris, 1829: 2 type species Buprestis granulata Say, 1823
Therysambus Descarpentries & Villiers, 1967b: 1007 type species Sambus apicalis Bourgoin, 1923
Wallaceilus Hołyński, 2003: 16 type species Agrilus scutellaris Deyrolle, 1864

subgenus Agrilomorpha Théry, 1909: 442 type species Agrilomorpha rothschildi Théry, 1909
Agrilophotus Bellamy & Westcott, 1992: 126 type species Amorphosoma acutipennis Fåhraeus, 1851

subgenus Agrilus Curtis, 1825: 67 type species Buprestis viridis Linnaeus, 1758
subgenus Anambus C.G. Thomson, 1864: 38 type species Buprestis biguttata Fabricius, 1777
subgenus Aridagrilus Alexeev, 1998: 382 type species Agrilus transversesulcatus Reitter, 1890
subgenus Austragrilus Alexeev, 1998: 374 type species Agrilus kurumi Kurosawa, 1957
subgenus Bubagrilus Curletti, 1993: 437 type species Agrilus embrikstrandellus Obenberger, 1935
subgenus Convexagrilus Alexeev, 1998: 377 type species Agrilus convexicollis Redtenbacher, 1847
subgenus Dentagrilus Alexeev, 1998: 377 type species Buprestis caerulea Rossi, 1792
subgenus Diplolophotus Abeille de Perrin, 1897a: 1 type species Buprestis deserta Klug, 1829
subgenus Duttus Curletti, 1993: 436, 438 type species Agrilus purpuratus Klug, 1829
subgenus Micragrilus Alexeev, 1998: 374 type species Agrilus ganglbaueri Semenov, 1891
subgenus Orientagrilus Alexeev, 1998: 379 type species Agrilus tempestivus Lewis, 1893
subgenus Paralophotus Curletti, 1998: 132 type species Agrilus baringanus Obenberger, 1931
subgenus Personatus Curletti, 1993: 436 type species Buprestis sexguttata Thunberg, 1789
subgenus Pseudoquercagrilus Alexeev, 1998: 378 type species Agrilus asiaticus Kerremans, 1898
subgenus Quercuagrilus Alexeev, 1998: 372 type species Buprestis angustula Illiger, 1803
subgenus Robertius Théry, 1947: 670 type species Agrilus robertii Chevrolat, 1838
Arquagrilus Alexeev, 1998: 369 type species Buprestis pratensis Ratzeburg, 1837

subgenus Rosagrilus Alexeev, 1998: 375 type species Agrilus roscidus Kiesenwetter, 1857
subgenus Sinagrilus Alexeev, 1998: 379 type species Agrilus sinensis J. Thomson, 1879
subgenus Sinuatiagrilus Alexeev, 1998: 372 type species Buprestis sinuata Olivier, 1790
subgenus Spiragrilus Alexeev, 1998: 381 type species Agrilus antiquus Mulsant & Rey, 1863
subgenus Uragrilus Semenov, 1935: 276 type species Agrilus guerini Lacordaire, 1835
subgenus Xeragrilus Alexeev, 1998: 375 type species Agrilus sericans Kiesenwetter, 1857
[species in alphabetical order, see Comments in New Acts]
abyssinicus Obenberger, 1917i: 209 N: EG
acastus Kerremans, 1913b: 114 A: TAI
horni Kerremans, 1914c: 104 (Sambus) [HN]
ohbayashii Tôyama, 1987: 308

achilleus Obenberger, 1935j: 121 [RN] A: TAI

BUPRESTIDAE

525

miwai Théry, 1936b: 61 [RN] [HN]
myrmido Kerremans, 1912c: 208 [HN]
myrmidonius Obenberger, 1936l: 1092 [RN]

acutus Thunberg, 1787: 52 (Buprestis) A: NP AUS ORR
armatus Weber, 1801: 74 (Buprestis)
cupreopunctatus Herbst, 1801: 248 (Buprestis)
frater Deyrolle, 1864a: 155
menadoensis Obenberger, 1924d: 26
pulchellus Kirby, 1819a: 380 (Buprestis)
spinosus Fabricius, 1801b: 214 (Buprestis)

adelphinus Kerremans, 1895: 222 A: ANH FE GUA HEB HUB JA NC SC SCH SHA SHN SHX XIZ YUN ORR
egorovi Alexeev, 1989: 480
nigrocoerulans Obenberger, 1924b: 39
nonfriedanus Obenberger, 1923c: 64 [RN]
nonfriedi Obenberger, 1914d: 44/49 [HN]
panhensis Baudon, 1968: 117

adlbaueri Niehuis, 1987: 58 A: SY TR
adspersus Kerremans, 1890c: cciv (Cisseis) A: NP ORR
aerosus Ganglbauer, 1889: 33 A: GAN SCH
agnatus Kerremans, 1892c: 820 A: GUI YUN ORR
alashanensis Obenberger, 1936h: 112 A: BEI ES MG NMO WS
rozhkovi Alexeev, 1964: 13
steppensis Cobos, 1968: 396

albogularis albogularis Gory, 1841: 235 E: AB AL AR AU BU BY CZ GG GR HU KZ MC MD RO SK SL ST TR UK
A: IN KZ NMO TM TR XIN
sieversi Abeille de Perrin, 1897a: 28

albogularis artemisiae Brisout de Barneville, 1863: 78 E: FR IT PT SP
albogularis dalmatinus Obenberger, 1937e: 87 E: BH CR IT ME SB
albogularis longepilosus Cobos, 1966d: 323 A: AF
albomarginatus A. Fiori, 1906: 260 E: IT
verres Théry, 1930d: 175

alesi Obenberger, 1935m: 164 A: JA SCH ORR
aritai Tôyama, 1985: 42
sacchari Obenberger, 1940k: 175

algiricus Obenberger, 1923e: 84 N: AG
alpenae Baudon, 1968: 83 A: AP HAI ORR
alutaceicollis Obenberger, 1930a: 110 A: BEI FE HEB SHA SHX
amamioshimanus Kurosawa, 1964a: 331 A: JA
amethystinus Solier, 1833: 304 E: FR
amulius Obenberger, 1936c: 35 A: TAI

526

BUPRESTIDAE

andresi Obenberger, 1920a: 86 N: EG A: IS JO SA YE
andrusi Baudon, 1968: 118 A: SHA ORR
pseudograminis Jendek, 2007: 34

angustulus angustulus Illiger, 1803: 240 [NP] (Buprestis) E: AB AL AR AU BE BH BU BY CR CT CZ DE FR GB
GE GG GR HU IT KZ LA LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: EG TU A:
FEi IN KZ SY TR
laetefrons Mannerheim, 1837: 114
laeticeps Semenov, 1891: 342
latisternum Théry, 1942a: 159
olivaceus Gyllenhal, 1808: 454 (Buprestis)
pavidus Fabricius, 1792: 214 [NO] (Buprestis)
rugicollis Ratzeburg, 1837: 55 (Buprestis)
semenovi Kerremans, 1892a: 271 [SN]

angustulus bicoloratus Abeille de Perrin, 1894g: ccclv E: SP N: AG MO TU
maroccanus Obenberger, 1923e: 86

antiquus antiquus Mulsant & Rey, 1863: 19 E: AU BU FR IT SP SZ N: AG TU
celticus Obenberger, 1916d: 278
linderi Marseul, 1866: 468
nemorivagus Obenberger, 1924d: 50
oxygonus Abeille de Perrin, 1907b: li
protractus Kerremans, 1903: 289 [RN]
proximus Bauduer, 1878: 80 [HN]
pruinosulus Abeille de Perrin, 1897a: 19

antiquus croaticus Abeille de Perrin, 1897a: 21 E: AL AU BH BU BY CR CT CZ GR HU IT ME PL RO SB SK ST UK
curtii Obenberger, 1913b: 9
hucul Obenberger, 1933c: 176
serbicus Obenberger, 1927f: 75
ukrainensis Obenberger, 1936h: 114

araxenus araxenus Iablokoff-Khnzorian, 1960: 251 E: AB AR A: IN TM TR
araxenus lopatini Alexeev, 1964: 11 A: KI KZ TD UZ
arisanus Miwa & Chûjô, 1940: 59 A: TAI
arsenevi Jendek, 2009b: 55 A: FE
asahinai Kurosawa, 1956: 38 A: FE HEB HEI JA NC SC
pseudorotundicollis Kurosawa, 1956: 40
stepanovi Alexeev, 1979b: 134

ashlocki Baudon, 1968: 67 A: YUN ORR
asiaticus Kerremans, 1898a: 178 A: BEI FE HEB HEI HEN HUB JA JIX NC NMO SC SCH SHA ORR
hauserellus Obenberger, 1935m: 171
igai Kurosawa, 1963a: 110
panphilovi Alexeev, 1959: 28
planefasciatus Obenberger, 1936h: 115

BUPRESTIDAE

527

ater Linnaeus, 1767: 663 (Buprestis) E: AL AU BE BH BU BY CR CT CZ DE FI FR GE GG GR HU IT KZ LA LT MC
MD ME NT PL RO SB SK SL SP ST SZ TR UK A: KZ TR
eutenes Obenberger, 1924b: 40
jureceki Obenberger, 1924: 39
sexguttatus Brahm, 1791: 141 (Buprestis) [HN]
sexguttatus Herbst, 1801: 265 (Buprestis)
subeuropeellus J. Thomson, 1878b: 88

atriplicis aegyptiacus Théry, 1930a: 424 N: EG
atriplicis atriplicis Théry, 1900a: 33 N: AG
auricollis Kiesenwetter, 1857: 149 [see New Acts] E: AL AU BH BU BY CR CT CZ FR GE GG GR HU IT LA LS MC
MD ME PL RO SB SK SL SP ST SZ UK A: TR
distinstipilis Obenberger, 1924b: 49
korbi Abeille de Perrin, 1897a: 7
madari Obenberger, 1924b: 49
tokatensis Obenberger, 1924b: 54

aurigaster Jendek, 2011: 44 A: YUN ORR
aurinotus Jendek, 2011: 44 A: GUI
auristernum Obenberger, 1924b: 34 A: YUN
aurithorax Jendek, 2011: 45 A: BEI FUJ GUX ORR
auriventris E. Saunders, 1873: 517 A: FUJ GUA GUX HKG HUB HUN JA JIX SCH TAI ZHE ORR
fleutiauxi Bourgoin, 1922a: 23
graptelytrus Obenberger, 1914d: 43
pidjinus Obenberger, 1924b: 53
samoensis Blair, 1928: 108

auroapicalis auroapicalis Kurosawa, 1957: 190 A: GUI TAI ORR
laurenconi Descarpentries & Villiers, 1963: 109

auroapicalis ishigakianus Tôyama, 1985: 33 A: JA
auropictus Kerremans, 1912c: 208 A: FUJ JA TAI ORR
chopardi Descarpentries & Villiers, 1963b: 52
kanohi Kurosawa, 1954b: 90
ryukyuensis Tôyama, 1985: 47

babai Tôyama, 1988: 758 A: JA
babaulti Théry, 1930c: 163 A: HP
bacchus Kerremans, 1913b: 114 A: TAI
bai Jendek, 2011: 48 A: YUN
baobdil Obenberger, 1913g: 338 N: MO
barrati Descarpentries & Villiers, 1963: 57 A: FUJ GUA HUB HUN YUN ORR
beauprei beauprei Théry, 1930a: 434 N: AG MO
adjunctus Théry, 1930a: errata behind title page [RN] [HN]
theryellus Obenberger, 1935c: 20 [RN]
vaulogeri Théry, 1930a: 473 [HN]

528

BUPRESTIDAE

beauprei mourguesi Schaefer, 1954: 148 E: FR PT SP
berberus Curletti, 2003: 289 N: TU
betuleti Ratzeburg, 1837: 57 (Buprestis) E: AU BE BH BY CR CT CZ DE FI FR GE HU IT LA LS LT ME NL NR NT
PL RO SB SK SL ST SV SZ UK A: ES FE KZ MG WS XIN
foveicollis Marseul, 1869a: 122 [RN]
impressicollis Marseul, 1866: 454 [HN]
saliceti Obenberger, 1924b: 42

bicuspidatus Jendek, 2011: 51 A: GUI
bidentulus Ganglbauer, 1889: 31 A: GAN SCH
bifoveolatus Kerremans, 1895: 219 A: AP NP SD ORR
biguttatus Fabricius, 1777: 237 (Buprestis) E: AB AL AR AU BE BH BU BY CR CT CZ DE FR GB GE GR HU IT KZ
LA LS LT LU MC MD ME NL NR PL PT RO SB SK SL SP ST SV SZ TR UK N: AG EG MO TU A: IN SY TR ORR
morosus Gory & Laporte, 1837: 35
pannonicus Piller & Mitterpacher, 1783: 37 (Buprestis)
subfasciatus Ménétriés, 1832: 153 (Buprestis)

bilineatus Weber, 1801: 74 (Buprestis) Ai: TR NAR
aurolineatus Gory, 1841: 248
bivittatus Kirby, 1837: 161
flavolineatus Mannerheim, 1837: 110

birmanicus Kerremans, 1892b: 211 A: PA ORR
giraudi Baudon, 1965: 198
legitimus Obenberger, 1936c: 42

biscrensis Pic, 1918c: 1 N: AG
bituberculatus Jendek, 2007: 39 A: BEI FUJ HEB HEN NMO SHA
blatteicollis Bourgoin, 1922b: 168 A: GUI HAI YUN ORR
bocaki Jendek, 2011: 53 A: GUI
bonadonai Descarpentries & Villiers, 1963a: 15 A: ANH ORR
boninensis Kurosawa, 1963a: 109 A: JA
bonnottei Bourgoin, 1925: 131 A: YUN ORR
brechteli Curletti & van Harten, 2004: 70 A: YE
brevinotus Jendek, 2011: 55 A: GAN SCH SHA SHX
bucephalus Daniel, 1903: 253 E: AR
buresi Obenberger, 1935d: 54 E: AB AR BU GG GR SB ST A: IN SY TR
businskorum Jendek, 2011: 55 A: SHA
caligans Bourgoin, 1925: 131 A: GUX HP NP SCH UP YUN ORR
assamensis Obenberger, 1936h: 117

carinihumeralis Kurosawa, 1964a: 327 A: JA
caucasicola Semenov, 1891: 341 E: GG
caudicula Jendek, 2011: 57 A: GUI SCH SHA
chaetocoxalis Jendek, 2011: 57 A: HUB SHA
chekiangensis Gebhardt, 1929a: 33 A: FUJ HUN JA JIX SCH SHA YUN ZHE

BUPRESTIDAE

529

semivittatus Tôyama, 1985: 45

chionochaetus Obenberger, 1913g: 337 N: MO A: IS
chlorophyllus Abeille de Perrin, 1904: 223 A: IN IS KI SY TD TM TR UZ
esfandiarinus Davatchi, 1958: 68
pistaciophagus Alexeev & Kulinitsh, 1962: 92

chobauti Abeille de Perrin, 1897b: 22 N: AG MO
chujoi Kurosawa, 1985b: 149 A: GUA JA LIA NC SC
cinctus A.G. Olivier, 1790a: no 32: 90 (Buprestis) E: AU BE BH FR GE LU PT SP SZ
cinctiformis Obenberger, 1924b: 51
escorialensis Obenberger, 1924b: 51
sinuatocollis Brisout de Barneville, 1883: 83

ciping Jendek, 2011: 61 A: JIX
compacticornis Jendek, 2011: 61 A: FE HEB HEN NIN SCH SHX YUN
conspersus Jendek, 2011: 62 A: YUN
constantini Obenberger, 1927: 17 E: CT KZ ST UK A: KZ
tamarinae Stepanov, 1952: 272

convexicollis L. Redtenbacher, 1847: 285 E: AB AL AR AU BE BH BU BY CR CT CZ FR GE GG GR HU IT LA LS
LU MC MD ME NL PL RO SB SK SL SP ST SV SZ TR UK A: TR
mancinii Obenberger, 1927b: 84
tesari Obenberger, 1935m: 167

convexifrons Kiesenwetter, 1857: 148 E: AU
convictor Descarpentries & Villiers, 1963: 112 A: YUN ORR
cordicollis Jendek, 2011: 64 A: SHA
coreanus Obenberger, 1935m: 162 A: HEB SC
croceivestis Marseul, 1866: 472 N: AG EG TU
sericellus Fairmaire, 1866a: 27

ctesias Kerremans, 1913b: 115 A: TAI
cupes Lewis, 1893: 336 A: JA NMO SHA SHX
cuprescens amethystopterus Semenov, 1891: 342 E: GG ST A: KZ TR
cuprescens chrysoderes Abeille de Perrin, 1891: 283 A: IS LE SY TR
cuprescens cuprescens Ménétriés, 1832: 154 (Buprestis) E: AB AL AR AU BE BH BU BY CR CT CZ DE FI FR GB
GE GG GR HU IT KZ LA LS LT LU MC MD ME NL NT PL PT RO SB SK SL SP ST SV SZ TR UK A: ES FE HEI
IN JA JIL KI KZ LIA MG NC SC TM TR WS NARi
altaicola Obenberger, 1935m: 168
aurichalceus Redtenbacher, 1847: 286
calcicola Obenberger, 1916d: 271
communis Obenberger, 1924b: 41
egenus Abeille de Perrin, 1895c: cxxii [HN]
epistomalis Abeille de Perrin, 1897a: 13
foveolatus Abeille de Perrin, 1897a: 15
krasai Obenberger, 1924b: 40

530

BUPRESTIDAE

kuznecovi Obenberger, 1935m: 169 [HN]
kuznecovinus Obenberger, 1936l: 1038 [RN]
lacrymans Abeille de Perrin, 1895f: cclxxiv [RN]
mokrzeckii Obenberger, 1927h: 204
obtusifera Schaefer, 1949: 402 [RN]
obtusus Abeille de Perrin, 1897a: 15 [HN]
proximus Rey, 1891b: 19 [HN]
rubicola Abeille de Perrin, 1897a: 15

curtulus Mulsant & Rey, 1863: 12 E: BU FR GE GR HU IT PT RO SP N: MO
panchlorus Abeille de Perrin, 1897: 24
planiceps Abeille de Perrin, 1897a: 18

cyaneoniger E. Saunders, 1873: 515 A: FE GUI HAI HEB HEN HUB JA JIL JIX KA NC NMO SC SCH SHA SHX
YUN ZHE ORR
ataman Obenberger, 1924b: 35
cyaneoniger J. Thomson, 1879a: 71 [HN]
impressifrons Kiesenwetter, 1879b: 254
jamesi Jakobson, 1913: 798 [RN]
marquardti Obenberger, 1914d: 41
melanopterus Solsky, 1876b: 277
mikado Obenberger, 1924b: 35

cyaneovirens Bourgoin, 1922a: 24 A: GUX YUN ORR
cyanescens cyanescens Ratzeburg, 1837: 54 (Buprestis) E: AL AR AU BE BH BU BY CR CT CZ DE FR GB GE GR
HU IT LA LS LT LU MC MD ME NL NR PL RO SB SK SL SP ST SV SZ UK A: FE HUN JIX KZ NC SC SHA SHX
TR WS NARi
acuticornis Abeille de Perrin, 1897a: 10
amabilis Gory & Laporte, 1837: 52
caeruleus Rossi, 1792: 62 (Buprestis) [HN]
cockerelli Fisher, 1925: 4
fissifrons Abeille de Perrin, 1897a: 3, 9 [HN]
italicus Obenberger, 1920: 212
kyselyi Obenberger, 1924b: 50 [RN]
pooli Théry, 1936c: 120
sulcaticeps Abeille de Perrin, 1869a: 79
teriolensis Obenberger, 1916d: 275

cyanescens johanidesi Niehuis, 1999a: 35 A: SY
cyanipennis Gory & Laporte, 1837: 18 A: NP PA YUN ORR
agilis Kerremans, 1894b: 471
amabilicolor Obenberger, 1936g: 91 [RN]
amabilis Kerremans, 1893b: 343 [HN]
dalbergiae Théry, 1930c: 158

BUPRESTIDAE

531

insignis E. Saunders, 1871: 122 [RN]
orientalis Harold, 1869d: 124 [RN]
ornativentris E. Saunders, 1866: 315

cygneus Abeille de Perrin, 1903: 334 N: AG TU
cytisi Baudi di Selve, 1870: 86 E: FR IT
baudii Bauduer, 1878: 81
pedemontanus Obenberger, 1917i: 210

daimio Obenberger, 1936i: 140 A: JA SC
dali Jendek, 2009b: 54 A: SCH YUN
darjiling Jendek, 2001: 9 A: NP SD ORR
daur Jendek, 2011: 82 A: NMO
decoloratus Kerremans, 1892b: 213 A: AF AP BT JA KA NC NP SC SCH TAI UP XIZ YUN ZHE ORR
afghanistanicus Alexeev, 1992: 378
ajax Kerremans, 1912c: 207
alazon Lewis, 1893: 333
lunlingensis Obenberger, 1958a: 237
octostictus Bourgoin, 1922a: 24
pyrrhus Kerremans, 1913b: 112

deguchii Tôyama, 1985: 36 A: JA
delphinensis Abeille de Perrin, 1897: 12 E: AU BE BH BY CT CZ DE FR GE HU IT LA LT ME NT PL RO SB SK ST
UK A: ES FE HEI JIL KZ LIA MG NC WS
eogenes Obenberger, 1930a: 112

densetomentosus Obenberger, 1940k: 184 A: FUJ HUN JIX
derasofasciatus Lacordaire, 1835: 613 E: AB AL AR AU AZ BH BU BY CR CZ FR GE GG GR HU IT MA MC MD
ME PL PT RO SB SK SL SP ST SZ TR UK N: AG EG MO MR TU A: CY IN IS JO LE SY TM TR NARi
hungerbuehleri Pochon, 1963: 69

desertus Klug, 1829: no. 14 (Buprestis) N: AG EG LB MO TU A: AE IN IS JO SA YE AFR
dianthus Kerremans, 1892c: 823 A: YUN ORR
coelicus Bourgoin, 1922a: 24
dorri Bourgoin, 1925: 131
nodosus Kerremans, 1894b: 470

diao Jendek, 2011: 88 A: SHA
diaolin Jendek, 2001: 5 A: YUN
dionides J. Thomson, 1879: 72 A: HP
kheilianus Obenberger, 1924b: 38

discalis E. Saunders, 1873: 514 A: FUJ GUI HAI HUB HUN JA JIA KA NC SC SCH TAI YUN
discicollis Deyrolle, 1864a: 189 A: TAI ORR
punctiventris Kerremans, 1900a: 31
simiolus Kerremans, 1896b: 143

discoloriformis Obenberger, 1920a: 86 N: AG AFR

532

BUPRESTIDAE

diversornatus Jendek, 2011: 89 A: FE
dureli Jendek, 2011: 90 A: SHX ORR
ecarinatus Marseul, 1866: 484 A: BEI ES FE GAN HEB HEI HEN JIL KI KZ LIA MG NC NMO QIN SC SCH SHA
SHX WS XIZ
gracilicornis Ganglbauer, 1889: 35
grassator Kerremans, 1903: 285 [SN]

elegans elegans Mulsant & Rey, 1863: 14 E: FR IT PT SP N: AG MO
cisti Brisout de Barneville, 1863: 77
elisicollis Abeille de Perrin, 1907c: lxx
roscidulus Abeille de Perrin, 1897a: 22

elegans teucrii Schaefer, 1949: 416 E: FR
emmerichi Obenberger, 1935m: 164 A: SCH
enriquei F. Murria Beltrán & Á. Murria Beltrán, 2007: 179 E: SP
erythrostictus Bourgoin, 1922b: 169 A: YUN ORR
barmensis Obenberger, 1936c: 44

esakii Kurosawa, 1964a: 323 A: JA
ethlius Gory, 1841: 238 A: HP UP
etruscus Curletti, 2013: 75 E: IT
eunuchus Jendek, 2011: 93 A: YUN
euonymi Tôyama, 1985: 28 A: FE HEN JA NC SC SHA SHG
yagii Tôyama, 1985: 30

fareastensis Jendek, 1995b: 177 A: FE HEB NMO
fasciatus Jendek, 2013b: 112 A: YUN ORR
fevreae Descarpentries & Villiers, 1963: 114 A: GUI ORR
filius Bílý, Curletti & van Harten, 2003: 82 A: YE
fissus Obenberger, 1917i: 214 A: BEI FE HEI NC SC SCH SHA SHN SHX
lubricus Théry, 1942c: 260

fleischeri Obenberger, 1925b: 33 A: BEI ES FE HEI JA KZ LIA MG NC SC SCH SHA WS
kochi Théry, 1942c: 253
kurosawai Obenberger, 1940c: 26
nipponicola Kurosawa, 1963a: 111
tscherepanovi Stepanov, 1954a: 115

foliatus Jendek, 2007: 40 A: GUI SCH SHA
foliicornis Abeille de Perrin, 1891: 284 A: SY TR
foramenifer Jendek, 2011: 98 A: SCH SHA TAI YUN
formosanus Kerremans, 1912c: 205 A: TAI
fornicatus Jendek, 2011: 98 A: FUJ JIX
fortunatus Lewis, 1893: 333 A: JA
friebi Obenberger, 1922a: 25 A: ANH BEI FE GAN HEI JA JIA JIL LIA MG NC SC SCH SHA SHX WS
ainu Miwa & Chûjô, 1940: 64
enisus Théry, 1942c: 264

BUPRESTIDAE

533

kiangsuanus Obenberger, 1935m: 171
nalajchanus Cobos, 1968: 391
vodaki Obenberger, 1936i: 141

fusciapex Jendek, 2011: 101 A: AP GUI ORR
fusiformis Jendek, 2011: 102 A: YUN
galiberti levantinus Curletti & van Harten, 2004: 70 A: YE
ganglbaueri Semenov, 1891: 344 A: AF KZ MG NMO TD TM UZ XIN
albifrons Reitter, 1897m: 249

gaoligong Jendek, 2000a: 152 A: YUN
gianassoi Magnani & Niehuis, 1994: 49 A: CY
globulifrons Obenberger, 1920c: 213 E: IT
graecus Obenberger, 1916d: 275 E: AU CZ FR GR HU IT ME SB SK SL SZ
viscivorus Bílý, 1991c: 371

graminis Kiesenwetter, 1857: 137 E: AB AL AR AU BH BU BY CR CT CZ FR GE GG GR HU IT MC MD ME PL PT
RO SB SK SL SP ST SZ TR UK N: AG EG MO TU A: IN SY TR
akbesinus Obenberger, 1924b: 50
armenus Obenberger, 1936h: 106
barbatus Abeille de Perrin, 1897c: 34
cervicornis Abeille de Perrin, 1897c: 35
disparicornis Bedel, 1921: 216 [RN]
mamorensis Théry, 1930: 446
reitteri Abeille de Perrin, 1897a: 16
talyschensis Obenberger, 1917i: 215
verticalis Abeille de Perrin, 1895c: cxxvi

grandiceps cyrenaicus Curletti, 2005: 233 N: LB
grandiceps grandiceps Kiesenwetter, 1857: 126 E: “Banat” A: CY
gregori Obenberger, 1936h: 109 N: MO
guerini Lacordaire, 1835: 608 E: AU BE CT CZ FR GE HU IT LT LU ME PL SB SK SL SP ST SV SZ UK
gussakovskiji Alexeev, 1981a: 3 A: FE SHA XIN
han Jendek, 2011:104 A: HEN
hani Jendek, 2011: 105 A: YUN
hasegawai Kurosawa, 1954b: 90 A: TAI
hastulifer Ratzeburg, 1837: 54 (Buprestis) E: AB AL AR AU BH BU BY CR CT CZ FR GE GR HU IT KZ MC ME PT
RO SB SK SL SP ST SZ TR UK N: AG MO TU A: IN IQ IS TR
aladaghensis Obenberger, 1918b: 24

hastuliformis Novak, 2003b: 70 A: CY
hattorii Nakane, 1983a: 111 A: JA
hemiphanes Marseul, 1866: 467 E: BH CR FR GR IT SP N: MO A: TR
mephistopheles Abeille de Perrin, 1897: 23
nigrivestis Abeille de Perrin, 1897a: 23

hirashimai hirashimai Kurosawa, 1964a: 329 A: JA

534

BUPRESTIDAE

hirashimai yaeyamaensis Tôyama, 1985: 21 A: JA
hittita Magnani, 1996: 93 A: TR
hmong Jendek, 2011: 106 A: YUN ORR
hokkaidoensis Tôyama, 1985: 45 A: JA
holzschuhi Jendek, 1994b: 31 A: YUN
hornianus Kerremans, 1912c: 206 A: TAI
huashanus Jendek, 2001: 3 A: SCH SHA
hui Jendek, 2011: 107 A: HEN SHA
hunanus Jendek, 2009a: 16 A: HUN
hyperici Creutzer, 1799b: 122 (Buprestis) E: AB AL AR AU BE BH BU BY CR CT CZ DE FR GE GG GR HU IT LA
LT LU MC MD ME NL PL PT RO SB SK SL SP ST SZ TR UK N: TU A: IN NMO TR NARi
albaniae Obenberger, 1922a: 66
hofferi Obenberger, 1936h: 111
impressus Rey, 1891b: 20 [HN]
modestulus Semenov, 1895c: 247
prasinus Mulsant & Rey, 1863: 17
reyellus Obenberger, 1936l: 1000 [RN]
trinacriae Obenberger, 1924b: 52

hypericicola Abeille de Perrin, 1893: 138 N: AG MO TU
ibericus Sánches Sobrino & Tolosa Sánchez, 2005: 99 E: SP
imasakai Tôyama, 1985: 23 A: JA
imitans Lewis, 1893: 332 A: JA
improcerus Jendek, 2011: 109 A: YUN
inadai Fukutomi, 2006: 158 A: JA
inamoenus Kerremans, 1892c: 824 A: FUJ GUA YUN ORR
ineptus Kerremans, 1892b: 218 A: YUN ORR
insulicola Kerremans, 1912c: 205 A: TAI ZHE
suensoni Obenberger, 1940k: 176

integerrimus Ratzeburg, 1837: 57 (Buprestis) E: AL AU BE BH BU BY CR CT CZ DE FI FR GE GG GR HU IT LA
LS LT LU MC ME NT PL RO SB SK SL SP ST SZ UK N: AG A: TR
bavaricus Obenberger, 1917i: 216
cupreus L. Redtenbacher, 1847: 286
nemeobius Obenberger, 1922a: 22

intrusus Kerremans, 1914c: 105 A: TAI
iriei Tôyama, 1985: 42 A: JA
jacetanus Sánches Sobrino & Tolosa Sánchez, 2004: 31 E: SP
jaminae Baudon, 1968: 80 A: AP BT SHA YUN ORR
japanocarinatus Ohmomo, 2002: 158 A: JA
jindrai Jendek, 2011: 111 A: SHX
jugurtha Abeille de Perrin, 1900a: 13 N: TU
kaluganus Obenberger, 1940i: 60 E: CT LT ST UK A: BEI ES FE HEB JA MG NMO WS

BUPRESTIDAE

535

pseudoussuricola Alexeev, 1979b: 130

kawarai Kurosawa, 1963a: 107 A: JA NMO
kettneri Curletti & van Harten, 2004: 71 A: YE
klapperichianus klapperichianus Cobos, 1966d: 321 A: AF
klapperichianus krupkai Alexeev & Bílý, 1980: 606 A: PA
kocheri Baudon, 1956b: 149 N: MO
komareki komareki Obenberger, 1926d: 102 A: ANH JA JIL NC SC
ronino Obenberger, 1935m: 167

komareki sapporoensis Obenberger, 1935m: 168 A: JA
emisi Ohmomo, 2005: 238

kostali Jendek, 2007: 42 A: SCH
kotoensis Miwa & Chûjô, 1940: 70 A: TAI
koyoi Ohmomo, 2002: 162 A: JA
kratochvili Obenberger, 1936h: 110 A: TAI
kuangcenensis Obenberger, 1940l: 114 A: JIL
kubani Bílý, 1991c: 374 E: AU CZ HU SK
kucerai Jendek, 2011: 115 A: HUB SHA
kurumi Kurosawa, 1957: 187 A: FE HUB JA SHA
kutahyanus Królik, 2002: 26 A: TR
lacrima Jendek, 2011: 116 A: YUN ORR
lacus Curletti & Ponel, 1994: 353 E: FR
laetecyanescens Obenberger, 1940k: 178 A: NP SCH YUN ORR
lafertei Kerremans, 1892b: 212 A: AP BT NP ORR
lahu Jendek, 2011: 117 A: YUN
lancrenoni Baudon, 1965: 225 A: NP YUN ORR
lanzarotensis Cobos, 1969: 49 N: CI
laticornis Illiger, 1803: 243 (Buprestis) E: AB AL AR AU BE BH BU BY CR CZ DE FI FR GB GE GG GR HU IT LA
LS LT LU MC MD ME NL NR PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO MR TU A: IS TR
asperrimus Marseul, 1866: 492
scaberrimus Ratzeburg, 1837: 55 (Buprestis)
sibiricola Obenberger, 1924b: 43

latipalpis Jendek, 2007: 39 A: YUN
lei Jendek, 2011: 120 A: SHA
leprieuri Pic, 1918c: 2 N: AG TU
limoniastri Bedel, 1886: cxxx N: AG TU
lineatomaculatus Jendek, 1994b: 34 A: AP SD ORR
lineola hermineus Abeille de Perrin, 1907b: li E: AB AR GG KZ ST TR UK A: IN IS KZ TM TR
shamyl Obenberger, 1922a: 68

lineola lineola Kiesenwetter, 1857: 147 E: AL AU BH BU BY CR CZ FR GR HU IT MC MD ME PL RO SB SK
SL SZ
rumanicus Obenberger, 1924b: 48

536

BUPRESTIDAE

lisu Jendek, 2011: 121 A: SCH
litura Kiesenwetter, 1857: 139 E: AL AU BH BU BY CR CZ GR HU IT MC MD ME RO SB SK SL ST UK A: TR
lituratus Klug, 1829: No. 13 (Buprestis) N: AG EG LB MO TU A: AE IN IS JO SA YE AFR
kiesenwetteri Pic, 1894a: 71

lubopetri Jendek, 2000b: 188 A: YUN ORR
lugu Jendek, 2011: 122 A: SCH YUN
lugubris Kerremans, 1914c: 107 A: TAI
lukesi Obenberger, 1936h: 117 A: NP SD ORR
modicus Kerremans, 1892c: 821 [HN]

macroderus Abeille de Perrin, 1897a: 15 E: AL AR AU BH BU CR CZ GG GR HU IT MC ME RO SB SK SL ST TR
UK A: IQ TR
angulifer Abeille de Perrin, 1897a: 15
fuscosericeus K. Daniel & J. Daniel, 1898: 66

macrotatopus Obenberger, 1930a: 110 A: KI
aridosus Obenberger, 1936h: 111

maculifer E. Saunders, 1873: 514 A: JA
madeci Baudon, 1968: 145 A: HAI HEN JA SC SHA TAI ORR
mushanus Kurosawa, 1976b: 136
rokuyai Kurosawa, 1976b: 134

majzlani Jendek, 2007: 42 A: YUN
makiharai Tôyama, 1987: 313 A: TAI
makiharaiellus Ohmomo, 2006: 248 [RN] A: JA
makiharai Ohmomo, 2005: 235 [HN]

mali Matsumura, 1924: 4 A: ES FE GAN GUX HEB HEI HEN HUB MG NC NMO QIN SC SCH SHN XIN XIZ
jenissejensis Obenberger, 1924b: 56

malicola Rungs & Schaefer, 1948: 121 N: MO
malloti Théry, 1930c: 168 A: SHA UP YUN ORR
diolaus Obenberger, 1958a: 238

mallotiellus Kurosawa, 1985b: 162 [RN] A: JA
malloti Kurosawa, 1957: 189 [HN]

manchu Jendek, 2011: 128 A: HEN SHA YUN ORR
mandala Jendek, 2011: 129 A: SCH
marginicollis E. Saunders, 1873: 516 A: FE HEN HUB JA NC SC SHA ZHE
beppuensis Obenberger, 1936i: 140
hastuliferoides Gebhardt, 1929b: 32

margotanae G. Novak, 2001b: 53 A: CY
marozzinii Gobbi, 1974: 255 E: AL BU CR FR GR IT ME SB N: AG A: TR
massanensis Schaefer, 1955: 34 E: FR SP
meridionalis Cobos, 1986: 241

masumotoi Ohmomo, 2011a: 211 A: JA
meloni Curletti, 1986: 129 E: IT

BUPRESTIDAE
mendax Mannerheim, 1837: 111 E: BE BY CT CZ DE FI GE LA LT NT PL RO SV SZ UK A: WS
faldermanni Gory & Laporte, 1837: 42

mercurini Théry, 1930a: 467 N: MO
miao Jendek, 2011: 130 A: SCH YUN
minor Deyrolle, 1864a: 171 A: TAI ORR
mio Jendek, 2011: 131 A: HUB SHA
miwai Obenberger, 1936c: 37 A: HEN SHA TAI ORR
lloydae Baudon, 1968: 162

moerens E. Saunders, 1873: 517 A: BEI FE GAN HEB HEI JA NC NMO SC SCH SHA SHX TAI
araxicola Abeille de Perrin, 1897a: 11
mandjuricus Obenberger, 1922a: 23
marinus Obenberger, 1922a: 24
paphius Théry, 1942c: 268
rotundicollis E. Saunders, 1873: 517

mogadoricus mogadoricus Escalera, 1914: 206 N: MO
mogadoricus talboti Théry, 1930a: 451 N: MO
molitor Abeille de Perrin, 1897a: 28 N: AG
mongolorum Obenberger, 1936h: 104 A: MG NMO
monogrammus J. Thomson, 1879: 72 A: AF HP KA NP PA UP ORR
andrewesi Obenberger, 1922a: 67
kabakovi Alexeev, 1992: 381
klapaleki Obenberger, 1924b: 55

morawitzi Obenberger, 1936h: 106 A: KI KZ MG TD WS
richteri Alexeev, 1975: 156

morio Kerremans, 1895: 220 A: BT SD ORR
moriscus Obenberger, 1913g: 339 E: SP N: MO
mucidus Jendek, 2013a: 49 A: HAI ZHE
munieri Brisout de Barneville, 1883: 82 N: AG TU
beduinus Obenberger, 1914d: 45/51

muongoides Jendek, 2011: 135 A: HUB HUN JIX YUN ORR
muscarius Kerremans, 1895: 224 A: NP SCH SHA YUN ORR
komiyai Tôyama, 1987: 305
seladon Obenberger, 1940k: 174

myopic Jendek, 2000a: 150 A: SCH
nagaoi Nakane, 1983a: 110 A: JA
naomii Tôyama, 1988: 756 A: JA
naxi Jendek, 2011: 138 A: YUN
nelsonodaimio Ohmomo, 2006: 245 A: JA NC
nemethi Théry, 1930a: 447 N: MO
nepalensis Tôyama, 1988: 759 A: AP BT NP SD ORR
nicolanus Obenberger, 1924b: 42 E: CT PL A: BEI FE HEB HUB JA NC NMO SC WS

537

538

BUPRESTIDAE

piscatorius Obenberger, 1935m: 169

niisatoi Tôyama, 1987: 318 A: HAI TAI
ningpoensis Obenberger, 1927b: 85 A: ZHE
nipponigena Obenberger, 1935m: 167 A: HEB HEN HUB JA LIA SHA SHN TAI ZHE
melanolucidus Ohmomo, 2005: 236

nishiyamai Tôyama, 1985: 30 A: TAI
niveoguttatus Kerremans, 1892b: 210 A: AP HUN JIX NP SD UP YUN ORR
nivosus Abeille de Perrin, 1900a: 13 E: KZ A: ES KI KZ MG NMO TD TM UZ WS XIN
candidiventris Obenberger, 1936h: 113
kuldjensis Obenberger, 1936h: 111

notoclavus Jendek, 2000a: 153 A: HEN SHA
nubeculosus deserticola Descarpentries & Bruneau de Miré, 1963: 196 N: AG AFR
nubeculosus nubeculosus Fairmaire, 1891c: 548 N: EG LB A: IS AFR
martini Abeille de Perrin, 1897a: 1 (Diplolophotus)

oasis Obenberger, 1922a: 28 N: AG
obnotatus Kerremans, 1912c: 208 A: TAI ORR
obscurecinctus Obenberger, 1935f: 59 [RN] A: JA TAI
motobuanus Fukutomi, 2006: 159
nigricinctus Kerremans, 1912c: 206 [HN]

obscuricollis Kiesenwetter, 1857: 144 E: AB AL AR AU BH BU BY CR CT CZ FR GE GR HU IT MA MC MD ME
NL PL RO SB SK SL SP ST SZ TR UK N: AG A: IN IS KZ TR WS
foveola Rey, 1891b: 19
reyi Bauduer, 1878: 83
savalae Stark, 1953: 300

obscurosericeus Obenberger, 1940k: 185 A: ANH FUJ
occipitalis Eschscholtz, 1822: 79 (Buprestis) A: TAI ORR
celebicola Obenberger, 1924a: 123
connexus Kerremans, 1900a: 22
cupricauda E. Saunders, 1867b: 520
evinadus Gory & Laporte, 1837: 30
korenskyi Obenberger, 1923d: 80
kurandae Obenberger, 1923d: 80
marmoreus Deyrolle, 1864a: 201
nirius Obenberger, 1924a: 123
nitidus Kerremans, 1898a: 179
oblatus Kerremans, 1900c: 340
occipitalis Gory, 1841: 222
tebinganus Obenberger, 1924: 120

odetteae Baudon, 1968: 96 A: AP FUJ HUN JIX TAI ORR
ogatai Ohmomo, 2002: 160 A: JA
oishii Tôyama, 1987: 320 A: TAI

BUPRESTIDAE

539

okinawensis okinawensis Miwa, 1933a: 7 A: JA
okinawensis shiozakii Tôyama, 1985: 33 A: JA TAI
olivaceidorsis Obenberger, 1917i: 211 A: FUJ GUI YUN ORR
duporti Bourgoin, 1922a: 23
ornamentifer Obenberger, 1940k: 181

oliveri Niehuis, 1989a: 33 A: TR
olivicolor Kiesenwetter, 1857: 135 E: AL AR AU BE BH BU BY CR CT CZ DE FR GE GG GR HU IT LA LS LU MC
ME NL NR NT PL RO SB SK SL SP ST SV SZ TR UK
sexualis Abeille de Perrin, 1897a: 17

orestes orestes Kerremans, 1913b: 113 A: TAI
orestes takaii Nakane, 1983a: 109 A: JA
ornatoides Jendek, 2011: 146 A: HEB SHA
ornatus Deyrolle, 1864a: 155 A: TAI ORR
paganettii Obenberger, 1913b: 7 E: IT
pisanus Curletti, 1980: 171

paludicola Krogerus, 1922: 111 E: CT DE FI LA NL NR NT SV
paralleloides Jendek, 2011: 147 A: YUN
pecirkai Obenberger, 1916d: 273 A: IN KI KZ TD TM TR UZ
pekinensis accolus Jendek, 1995a: 142 A: ANH SCH YUN
pekinensis pekinensis Obenberger, 1924b: 55 A: BEI ES FE FUJ HEB HEI JIA KZ MG NC NMO SC SHA SHN
XIN ZHE
boreooccidentalis Obenberger, 1936h: 107
cerskii Obenberger, 1936h: 104
charbinensis Théry, 1942c: 256
klapperichi Obenberger, 1940k: 183

peregrinus Kiesenwetter, 1879a: 145 A: FE HEB NC NMO SC
brodjaga Obenberger, 1914d: 46/52

perisi Cobos, 1986: 248 E: SP
perrini Pic, 1905g: 185 [RN] N: AG
divaricatus Abeille de Perrin, 1905: 207 [HN]
elzeari Obenberger, 1935j: 121 [RN]

perroti Descarpentries & Villiers, 1963c: 118 A: FUJ GUX YUN ORR
pharao Obenberger, 1923e: 85 N: EG
pilosovittatus E. Saunders, 1873: 515 A: GAN HUN JA JIA JIX NC NMO ZHE NARi
pusillipubis Gebhardt, 1929b: 94
spesivcevi Obenberger, 1927c: 16
zemani Obenberger, 1925b: 34

plagiatus Ganglbauer, 1889: 31 A: GAN SCH SHA TAI
mentor Kerremans, 1912c: 207

planipennis Fairmaire, 1888: 121 Ei: CT ST Ai: BEI FE HEB HEI JA JIL LIA MG NC NMO SC SCH SHN TAI TIA
NARi

540

BUPRESTIDAE

feretrius Obenberger, 1936c: 37
marcopoli Obenberger, 1930a: 108
ulmi Kurosawa, 1956: 40

plasoni dichrosomus Obenberger, 1924a: 122 A: TAI
plasoni plasoni Obenberger, 1917i: 212 A: FUJ GUI GUX HUB HUN JIX SC SHA YUN ZHE ORR
jeanvoinei Descarpentries & Villiers, 1963b: 54

plavilstshikovi Alexeev, 1989: 315 A: MG
pleuralis Jendek, 2011: 156 A: SCH
pleurostigmus Jendek, 2011: 157 A: SHA
pliculipennis Obenberger, 1940k: 182 A: FUJ GUI HEN HUN JIX SHA
pratensis Ratzeburg, 1837: 56 (Buprestis) E: AL AU BE BH BU BY CR CT CZ DE FI FR GB GE GG GR HU IT KZ
LA LS LT LU MC MD ME NL NR NT PL RO SB SK SP ST SV SZ TR UK A: ES FE HEB IN KZ MG NMO SHA
TR WS XIN
chankae Obenberger, 1935m: 169
djukini Obenberger, 1935m: 169
fennicus Obenberger, 1936l: 1029 [SN]
praeclarus Krogerus, 1925: 75
pseudocoeruleus Obenberger, 1930a: 112
robertii Chevrolat, 1838b: 89

presbys Jendek, 2011: 160 A: GUI
priamus Kerremans, 1912c: 207 A: TAI
proditor Obenberger, 1917e: 53 A: “China”
prolatus Jendek, 2011: 162 A: YUN
prosternalis Jendek, 2011: 162 A: YUN
proteus Abeille de Perrin, 1893: 137 E: SP N: AG MO TU
hassani Théry, 1930a: 471

przewalskii Obenberger, 1936h: 112 A: MG NMO
pseudobscuricollis Jendek, 2007: 38 A: SCH YUN
pseudoconstantini Alexeev, 1981a: 7 A: FE
pseudocyaneus Kiesenwetter, 1857: 150 E: AU BY CT FI PL ST SZ UK A: WS
bialowiezaensis Gutowski, 1993: 296
foveola Roubal, 1919: 37 [HN]
obrucevi Obenberger, 1936h: 106
roubalianus Obenberger, 1936l: 1032 [RN]
supremus Obenberger, 1917d: 40

pseudolimoniastri Cobos, 1968: 401 A: GAN KZ MG NMO WS
pseudolubopetri Jendek & Chamorro, 2012: 78 A: TAI ORR
pseudosericans Obenberger, 1936h: 107 A: KI KZ MG WS
angulicollis Cobos, 1968: 393

pterostigma Obenberger, 1927c: 17 A: ZHE
pubiventris Kiesenwetter, 1857: 126 N: AG

BUPRESTIDAE

541

mundus Abeille de Perrin, 1894c: 91

pulvereus Abeille de Perrin, 1895c: cxxiii E: PT SP
andalusiacus Obenberger, 1920c: 213
lusitanicus Obenberger, 1916d: 276
navasi Obenberger, 1925a: 103

pumi Jendek, 2011: 165 A: YUN
purpuratus Klug, 1829: no. 12 (Buprestis) N: AG EG LB MO TU A: IN IS JO SA SY YE AFR
pusillesculptus Obenberger, 1940k: 177 A: SHA ZHE
qiang Jendek, 2011: 166 A: SCH YUN
qinling Jendek, 2000b: 190 A: SHA
qinlingcola Jendek, 2011: 167 A: SHA
quadrisignatus Marseul, 1866: 444 A: ES FE GAN HEB JIA MG NC NMO SC SHA SHX
lama Obenberger, 1935m: 164
mongoliae Obenberger, 1922a: 26

quentini Descarpentries & Villiers, 1963a: 16 A: NP YUN ORR
wilsoni Baudon, 1968: 128

rabaticus Théry, 1930a: 463 N: MO
radiolus Kerremans, 1913b: 113 A: TAI ORR
recticollis Jendek, 2011: 169 A: YUN
reichardti Obenberger, 1935m: 165 A: SCH XIZ YUN ORR
relegatoides Novak, 2003b: 69 A: CY
relegatus alexeevi Bellamy, 1998: 94 [RN] E: AL BU GG GR IT ME SB ST A: IS JO LE SY TR
dualis Alexeev & Bílý, 1980: 600 [HN]

relegatus relegatus Curletti, 1990: 102 E: GR
rhoos Królik & Niehuis, 2003: 358 A: TR
ribbei Kiesenwetter, 1879b: 255 A: ES FE HEB HEI HEN HUB JA JIL LIA NC NMO SC SCH SHA SHX
bakinensis Obenberger, 1935m: 168
corax Obenberger, 1917d: 39
freyi Théry, 1939b: 153
gracilipes Lewis, 1893: 335 [HN]
ignoratus Obenberger, 1924b: 44
iturupicus Alexeev, 1979b: 133
japonicus Kerremans, 1898a: 178
lewisiellus Kerremans, 1903: 287 [RN]
prinadai Fisher, 1925a: 6
taigicola Obenberger, 1924b: 45
tibialis Lewis, 1893: 335

ribesi Schaefer, 1946b: 107 E: AU BU BY CT CZ FR GE GG IT MD PL RO SK ST SZ UK A: ES FE KZ NC SC WS
NARi
rivalieri Descarpentries & Villiers, 1963: 15 A: GUI HUN SHA ORR
goichetae Baudon, 1965: 229

542

BUPRESTIDAE

roscidus Kiesenwetter, 1857: 127 E: AB AL AR AU BH BU CR CT CZ FR GE GG GR HU IT KZ MA MC MD ME PT
RO SB SK SL SP ST SZ TR UK N: AG EG MO TU A: CY IN IS JO LE SY TR
affectans Obenberger, 1923e: 86
alticola Abeille de Perrin, 1897a: 8
ascendens Abeille de Perrin, 1895c: cxxv
perparvus Obenberger, 1918b: 23
roswitha Bellamy, 2003: 155 [RN]
rubi Kaszab, 1940: 113 [HN]
satunini Obenberger, 1936h: 110
schevyrevi Stark, 1955: 786
subroscidus Obenberger, 1924b: 54
willcocksi Théry, 1914: 129

rougeoti Descarpentries & Villiers, 1963: 115 A: NP UP YUN ORR
rudicollis Alexeev, 1979b: 136 A: FE HEI
sachalinicola Obenberger, 1940l: 115 A: ES FE HEI JA
tamanukii Stepanov, 1954a: 116

sagittifer Jendek, 2011: 177 A: HEN SHA
sahelicus Curletti, 1998a: 106 N: MO
salicis J. Frivaldszky, 1877: 319 E: AL AU BH BU BY CR CT CZ FR GE GR HU IT KZ MC MD ME PL RO SB SK SP
ST SZ UK A: KZ TR UZ WS
acutangulus Schaefer, 1949: 393
alutaceus Obenberger, 1916d: 273
filigranus Obenberger, 1916d: 274
lugovoi Stepanov, 1958b: 172

samai Magnani, 1995a: 49 A: CY
samuelsoni Tôyama, 1985: 41 A: JA
satoi Kurosawa, 1954b: 85 A: FUJ GUI JA JIX SHA TAI
saudita Curletti, 1998a: 110 A: SA
sauteri Kerremans, 1912c: 206 A: TAI
schillhammeri Jendek, 2011: 180 A: GUI
scythicus Królik & Janicki, 2005: 350 E: UK
sekii sekii Ohmomo, 2004: 138 A: JA
sekii yaeyamanus Ohmomo, 2004: 138 A: JA
semiaurovittatus Kurosawa, 1954b: 87 A: TAI
amphipyrus Pochon, 1965: 275

senilis Kerremans, 1914c: 107 A: TAI
senohi Tôyama, 1988: 761 A: TAI
sentis Jendek, 2011: 180 A: SHA
sericans Kiesenwetter, 1857: 127 E: AB BU CT GG GR HU KZ MD RO ST UK A: AF IN KI KZ MG NMO TD
TM TR
daghestanicus Obenberger, 1930a: 109

BUPRESTIDAE

543

sericarius Abeille de Perrin, 1897a: 25

shan Jendek, 2011: 181 A: SHA
shashamboe Kurosawa, 1963a: 104 A: JA
she Jendek, 2011: 182 A: FUJ JIX
shibatai Kurosawa, 1964a: 323 A: JA
shimabarensis Ohmomo, 2011a: 214 A: JA TAI
sibiricus fukushimenis Jendek, 1994a: 12 A: FE JA
sibiricus sibiricus Obenberger, 1912b: 70 A: FE HEI NC NMO SC
insuspectus Obenberger, 1924b: 43
turdus Théry, 1942c: 255

sichuanus Jendek, 2011: 185 A: SCH
sikkimensis Obenberger, 1928c: 69 A: GUX NP SD YUN ORR
damoiselli Baudon, 1968: 111

sinensis sinensis J. Thomson, 1879: 73 A: BEI FUJ GUI HAI HEI HUB HUN JA JIA JIX SCH SHG XIZ YUN ORR
fairmairei Kerremans, 1892a: 256 [RN]
obscuripennis Fairmaire, 1888b: 120 [HN]
pseudimitans Miwa & Chûjô, 1940: 54/60

sinensis splendidicollis Fairmaire, 1889b: 348 A: SCH XIZ YUN ORR
bayeri Obenberger, 1935m: 161
lameyi Obenberger, 1924b: 36

sinuatus sachalinensis Obenberger, 1935m: 165 A: ES FE GAN HEI JA MG
yokoyamai Iga, 1955: 79
zhelochovtsevi Alexeev, 1979b: 128

sinuatus sinuatus A.G. Olivier, 1790: 32: 74 (Buprestis) E: AB AL AU BE BH BU BY CR CT CZ DE FR GB GE GG
GR HU IT KZ LS LU MC ME NL PL PT RO SB SK SL SP ST SZ TR UK N: EG A: IN TR NARi
chryseis Curtis, 1825: 67
pyri Blanchard, 1845: 80

siskai Jendek, 2011: 189 A: YUN
smaragdifrons Ganglbauer, 1889: 29 A: BEI GAN SCH
kanssuanus Obenberger, 1930a: 109

smaragdinus habros Jendek, 2011: 190 A: SHA
smaragdinus smaragdinus Solsky, 1876b: 279 A: FE HEI JIL LIA NC SC
prasinulus Obenberger, 1924b: 48

smatanai Jendek, 2007: 41 A: TR
smetanai Jendek, 2011: 191 A: GAN
solieri confinatus Curletti, 1990: 102 N: AG MO TU
solieri solieri Gory & Laporte, 1837: 49 E: FR IT PT SP N: AG EG MO TU
catalonicus Pochon, 1963: 70

sospes Lewis, 1893: 334 A: HEN HUB JA LIA NC SC SCH SHA SHX
kinoshitae Obenberger, 1936i: 140
quadristictulus Obenberger, 1940: 179

544

BUPRESTIDAE

soudeki Obenberger, 1926d: 103 A: FE HEI NC SC SHA SHX
sericeus Théry, 1942c: 266

souliei Jendek, 1994b: 30 A: SCH XIZ YUN
spathulifer Jendek, 2011: 194 A: YUN
spectabilis Kerremans, 1895: 219 A: AP NP SD ORR
spiculum Jendek, 2011: 194 A: SCH SHA SHX
spinipennis Lewis, 1893: 332 A: GUA HUB JA SC ORR
dentiapex Obenberger, 1940k: 179

spiridion Kerremans, 1913b: 114 A: TAI
stigma Jendek, 2011: 198 A: YUN
strandi Obenberger, 1918b: 24 N: AG
subauratus amurensis Obenberger, 1922a: 21 A: ES FE HEB HEI JA JIL MG NC SC WS XIN
demuthi Théry, 1942c: 258

subauratus subauratus Gebler, 1833: 277 (Buprestis) E: AU BE BH BY CR CT CZ DE FI FR GE HU IT KZ LA LT
LU ME NL NT PL RO SB SK SL ST SV SZ UK A: ES KI KZ MG WS
auripennis Gory & Laporte, 1837: 46
coryli Ratzeburg, 1837: 55 (Buprestis)

subcuneiformis Kurosawa, 1954b: 84 A: TAI
subrobustus E. Saunders, 1873: 516 A: ANH FUJ GUI HUB HUN JA JIX MG NMO SC SCH SHA TAI YUN NARi
ORR
albizziae Kurosawa, 1963a: 101 [HN]
boisreymondi Alexeev & Bílý, 1980: 604
kumamotoensis Obenberger, 1935m: 170
kumamotoi Tôyama, 1987: 316
kurosawus Bellamy, 1998: 95 [RN]
malyi Obenberger, 1936h: 116
sorocinus Kerremans, 1914c: 106

sudai Kurosawa, 1985b: 160 A: JA NMO
suginoi Tôyama, 1985: 21 A: JA
sugiurai Jendek, 2007: 31 A: JA
sulcicollis Lacordaire, 1835: 614 E: AL AU BE BH BY CR CT CZ DE FI FR GB GE HU IT KZ LA LS LT LU MC ME
NL NR NT PL RO SB SK SL SP ST SV SZ UK N: AG A: WS NARi
sahlbergi Mannerheim, 1837: 113
tenuis Ratzeburg, 1837: 53 (Buprestis)

sulcifer Abeille de Perrin, 1895c: cxxii E: AB A: SY TR
hypericiformis Obenberger, 1936h: 115
lobiceps Abeille de Perrin, 1895c: cxxv

sulcinotus Jendek, 2007: 38 A: YUN ORR
suturicuspidatus Jendek, 2007: 40 A: ANH BEI GUI SHA
suvorovi Obenberger, 1935m: 165 E: AU BH BU BY CR CT CZ FI FR GE GG GR HU IT LA LS MC ME NR NT PL
RO SB SK SP ST SV SZ TR UK A: ES FE HEI IN JIL KZ MG NC NMO SC SHA SHX TR WS

BUPRESTIDAE
brussae Obenberger, 1956a: 45/47
populneus Schaefer, 1946a: 73

suzukii Kurosawa, 1985b: 158 A: JA
sylviae Niehuis, 1992: 209 A: IS TR
tai Jendek, 2011: 202 A: YUN
taiwanensis Kurosawa, 1954b: 88 A: TAI
takahashii Tôyama, 1988: 754 A: JA
tazoei Kurosawa, 1985b: 156 A: JA SHA
tempestivus Lewis, 1893: 334 A: FE HEB HEN HUN JA LIA NMO SC SHA
tenuissimus Abeille de Perrin, 1891: 284 A: IS JO LE SY
terracotta Jendek, 2011: 204 A: SHA ZHE ORR
thibetanus Obenberger, 1928c: 70 A: XIZ
tokyoensis Kurosawa, 1985b: 151 A: JA ORR
tomentipennis Jendek & Chamorro, 2012: 91 A: TAI ORR
tonkineus Kerremans, 1895: 222 A: FUJ HAI YUN ORR
blatteiceps Bourgoin, 1925b: 131

touareg Théry, 1930a: 458 N: AG
toyoshimai Tôyama, 1988: 752 A: JA
transversesulcatus transversesulcatus Reitter, 1890j: 195 E: AB AR GG KZ ST UK A: TR
grusinus Obenberger, 1917i: 209
juxtasuturalis Abeille de Perrin, 1897a: 21
suturisignatus Obenberger, 1924b: 52
tifliscus Obenberger, 1936h: 114

transversesulcatus validiusculus Semenov, 1891: 341 A: IN KZ TD TM UZ
ieiunulus Obenberger, 1936h: 113
niveosignatus Obenberger, 1914d: 44/50

trinotatus E. Saunders, 1873: 513 A: JA ZHE
truncatus Jendek, 2007: 39 A: BEI SHX
tschitscherini Semenov, 1895a: 135 A: ES KI KZ MG NMO TD UZ
attila Obenberger, 1930a: 111
gerschuni Stepanov, 1958b: 168
lubischevi Stepanov, 1958: 165
machnovskii Stepanov, 1958b: 170
uzbekistanus Stepanov, 1958: 166

tshapadarensis Alexeev, 1992: 379 A: AF
tsushimanus Kurosawa, 1963a: 106 A: GUI HEN JA SHA
tsutsumiuchii Ohmomo, 2006: 247 A: JA
tuber Jendek, 2011: 209 A: ZHE
turcicus Marseul, 1866: 467 A: TR
friwaldskyi Marseul, 1866: 438

tyndarus Kerremans, 1913b: 115 A: FUJ TAI ORR

545

546

BUPRESTIDAE

omissulus Obenberger, 1935j: 121 [RN]
omissus Kerremans, 1914c: 106 [HN]
therondi Descarpentries & Villiers, 1963a: 19

tyrtaeus Kerremans, 1913: 113 A: TAI
uenoi Kurosawa, 1963a: 102 A: JA TAI
uhagoni Abeille de Perrin, 1897a: 20 E: PT SP
bedeli Abeille de Perrin, 1905: 208
hispanicus Obenberger, 1913g: 336

ulaangomiensis Cobos, 1972a: 47 A: GAN MG NIN WS
emeljanovi Alexeev, 1975: 152
subelegans Cobos, 1972a: 4

ulmicola Jendek, 2011: 211 A: SHX
undulatipennis Kurosawa, 1976a: 56 A: JA TAI
unguiculosus Obenberger, 1936h: 105 A: KZ MG NMO WS
gobicola Cobos, 1968: 398

uromastyx Curletti & Magnani, 2006: 18 N: AG
ussuricola Obenberger, 1924b: 46 A: FE HEI JA JIL LIA NC SC SHA SHX XIZ YUN
anomalus Théry, 1942c: 262
lasiolus Obenberger, 1935m: 170

ustjurti Kostin, 1973: 110 A: KZ
vaginalis Abeille de Perrin, 1897a: 26 E: AB AR BU KZ MC ST A: IN KZ TM TR UZ
niveifrons Abeille de Perrin, 1897a: 27
philippovi Alexeev, 1965: 300
syrdarjensis Obenberger, 1928b: 20

varius Kerremans, 1895: 221 A: ANH FE HEN HUB SC SCH SHA SHN
vaucheri Abeille de Perrin, 1895c: cxxiv N: MO
venosus Gory & Laporte, 1837: 28 N: AG MO A: IN SA AFR
rothschildi Théry, 1909a: 442 (Agrilomorpha)
strandi Obenberger, 1931a: 6 (Agrilomorpha) [HN]
tassii Cobos, 1962a: 3 (Agrilomorpha) [HN]

ventricosus Fairmaire, 1888b: 120 A: BEI HEB JA NMO
kalganicus Obenberger, 1935m: 163
toyamai Kurosawa, 1985b: 153

viduus Kerremans, 1914c: 105 A: BEI ES FE GAN GUA HEB JA MG NC NMO SC SCH SHA SHN SHX TAI WS XIN
ZHE
cardaces Obenberger, 1936c: 38
chinganicus Obenberger, 1922a: 27
hoscheki Obenberger, 1916d: 273
subviduus Kurosawa, 1957: 192

villosostriatus J. Thomson, 1879a: 72 A: NP ORR
binominatus Kerremans, 1900c: 341 [RN]

BUPRESTIDAE

547

villosostriatus Kerremans, 1892b: 214 [HN]

viridicaerulans rubi Schaefer, 1937: 83 E: AL BH BU CR FR GE GR IT MC ME PT SB SK SP SZ N: AG A: TR
viridicaerulans viridicaerulans Marseul, 1868: 175 A: IS JO LE SY TR
viridiobscurus E. Saunders, 1873: 515 A: HEN JA
viridis Linnaeus, 1758: 410 (Buprestis) E: AB AL AR AU BE BH BU BY CR CT CZ DE FI FR GB GE GG GR HU IT
KZ LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG MO MR TU A: BEI ES
FE FUJ HEB HEI HEN HUB IN JA JIL KI KZ LIA MG NMO SC SHA SHN SHX TM TR UZ WS XIN XIZ YUN
aubei Gory & Laporte, 1837: 44
bicolor Schrank, 1798: 603 (Buprestis)
bicolor L. Redtenbacher, 1847: 287 [HN]
brevitarsis Lewis, 1893: 336
brydli Obenberger, 1924b: 41
caenus Obenberger, 1924b: 40
capreae Chevrolat, 1838a: 56 (Buprestis)
cernyi Obenberger, 1925c: 59
cojbalsanensis Cobos, 1968: 404
darwinii Wollaston, 1857: 82
distinguendus Gory & Laporte, 1837: 44
fagi Ratzeburg, 1837: 56 (Buprestis)
filiformis Herbst, 1801: 313 (Buprestis) [RN]
gebleri Obenberger, 1924b: 42
imbellis Kerremans, 1892b: 215 [HN]
importunus Kerremans, 1903: 286 [RN]
indulgens Kerremans, 1897: 76
kiricenkoi Obenberger, 1935m: 166
krogerusi Obenberger, 1924b: 41
lewisi Kerremans, 1903: 287 [RN]
linearis Fabricius, 1792b: 213 (Buprestis) [HN]
linearis Schrank, 1781: 197 (Buprestis) [HN]
lineatus Schrank, 1782: 168 (Buprestis) [RN] [HN]
littlei Curtis, 1840: 365
melantatus Obenberger, 1924b: 40
montanellus Obenberger, 1916d: 270, 272
nocivus Ratzeburg, 1837: 56 (Buprestis)
poppiusi Obenberger, 1924b: 40
pusztae Obenberger, 1936h: 108
quercinus L. Redtenbacher, 1847: 286
rosaceus Scopoli, 1763: 61 (Mordella)
rosei Niehuis & Bernhard, 2005: 1
rudis Abeille de Perrin, 1897a: 13
salicivola Kurosawa, 1963a: 106

548
serraticornis Scopoli, 1763: 61 (Mordella)
towadensis Miwa & Chûjô, 1940: 61
transbaicalensis Obenberger, 1935m: 166
vernadskii Obenberger, 1927c: 15
viridipennis Gory & Laporte, 1837: 45

volkovitshi Jendek, 2007: 33 A: YUN
voriseki Jendek, 1995b: 175 A: ES FE HEI KZ MG WS
watanabei Tôyama, 1985: 35 A: JA
watanukiorum Fukutomi, 2010: 59 A: JA
wittemani Jendek, 1994b: 33 A: NP ORR
xantippus Kerremans, 1913b: 115 A: TAI
yakushimensis Tôyama, 1985: 34 A: JA
yamabusi Miwa & Chûjô, 1940: 63 A: JA
nakanei Kurosawa, 1963a: 106
ohmomoi Tôyama, 1985: 31

yamajii Tôyama, 1985: 26 A: JA
yamawakii Kurosawa, 1957: 192 A: JA LIA NC NMO SC TAI
yami Kano, 1938: 114 A: TAI
yasumatsui Kurosawa, 1964a: 325 A: JA
yemenita Curletti & Harten, 2002: 54 A: AE YE AFR
yonahaensis Tôyama, 1985: 37 A: JA
yunnanicola Obenberger, 1936h: 116 A: YUN
yunnanus Obenberger, 1927f: 74 A: YUN
zanthoxylumi L.-M. Lou, 1989: 60 A: GAN HUB SHA SHN YUN ZHE
zhongdian Jendek, 2009b: 55 A: YUN
zhuang Jendek, 2011: 233 A: JIX
zhui Jendek, 2011: 234 A: SHA
ziegleri Niehuis, 2006: 23 E: BU
zigzag Marseul, 1866: 478 E: BU CT KZ ST UK A: KI KZ MG WS
clermontianus Roubal, 1925: 71
kusakievici Obenberger, 1936h: 107
rossicus Obenberger, 1916d: 276
scutellatus Abeille de Perrin, 1897a: 8
semirjeciae Obenberger, 1936h: 105
simbirensis Obenberger, 1925a: 102

genus Agrilus, nomina dubia
auricollis Betta, 1847: 20 E: “Ticino”
binotatus Gory, 1841: 236 E: IT
canaliculatus Bach, 1867: 493 E: AU GE
comolli Betta, 1847: 19 E: “Ticino”

BUPRESTIDAE

BUPRESTIDAE

549

confinis Faldermann, 1835a: 24 A: “Northern China”
sperkii Solsky, 1873: 159 E: “Kiew” “Volhynia”
elongatus Herbst, 1780: 99 (Buprestis) E: GE
wagleri Gistel, 1857: 3, 40 E: CR GE “Ticino”
aurichalceus Gistel, 1857: 3, 40 [HN]
exilistis Strand, 1917: 88 [RN]

genus Dorochoviella Jendek, 2006: 151 type species: Dorochoviella kunashirensis Jendek, 2006
kunashirensis Jendek, 2006: 152 A: FE (Kunashir)

tribe Agrilini, genera incertae sedis
genus Parasambus Descarpentries & Villiers, 1966b: 988 type species Parasambus aurosignatus
Descarpentries & Villiers, 1966
sauteri Kerremans, 1913b: 112 (Sambus) A: TAI
genus Pseudagrilus Laporte, 1835: 165 type species Pseudagrilus splendidus Laporte, 1835
arabicus arabicus Obenberger, 1925a: 102 A: “Arabia”
arabicus jemeni Brechtel, 1998: 91 A: YE
genus Sambus Deyrolle, 1864a: 110 type species Sambus lafertei Deyrolle, 1864
Arrowia Obenberger, 1923a: 28 type species Arrowia marshalli Obenberger, 1923
Brachycisseis Théry, 1931a: 26 type species Brachycisseis dohertyi Théry, 1931 (=Sambus barkeri Bellamy 2007)
Weyersiella Théry, 1930e: 202 type species Antaxomorphus viridiceps Kerremans, 1900

binhensis Descarpentries & Villiers, 1966b: 983 A: YUN ORR
caesar Obenberger, 1935b: 15 A: GUI SCH YUN
chalcosomus J. Thomson, 1879a: 55 A: SD “India” ORR
davidi Théry, 1926c: 185 A: SCH
deyrollei J. Thomson, 1878a: 82 A: GUI SD YUN XIZ “India” ORR
elegans Théry, 1926c: 182
fastidiosus Théry, 1926c: 183
tristis Théry, 1926c: 181
wendleri Obenberger, 1924a: 101
yunnanicola Obenberger, 1935b: 15

femoralis Kerremans, 1892c: 817 A: NP SD YUN ORR
beamishi Baudon, 1968: 53
coomani Descarpentries & Villiers, 1966b: 982
monticola Obenberger, 1924a: 101
oresitrophus Obenberger, 1924a: 102

formosanus Miwa & Chûjô, 1935: 280 A: TAI
fulgidicollis Kerremans, 1900b: 81 A: SD
strandiellus Obenberger, 1932b: 227

550

BUPRESTIDAE

gibbicollis Kerremans, 1892b: 206 A: NP SD ORR
gmelinae Théry, 1930c: 157 A: NP UP ORR
perroti Descarpentries & Villiers, 1966b: 981

kanssuensis Ganglbauer, 1889: 27 A: GAN SCH
melanoderus Kerremans, 1892b: 207 A: BT HP NP SD UP YUN
parryi Théry, 1926c: 180
simlae Obenberger, 1932b: 227

nickerli Obenberger, 1924a: 101 A: NP UP ORR
unicus Théry, 1926c: 184

nigritus Kerremans, 1892b: 207 A: YUN ORR
furvus Bourgoin, 1923b: 261

novus Théry, 1926c: 188 A: YUN ORR
optatus Théry, 1926c: 189 A: GUI GUX YUN ZHE ORR
bourgoini Descarpentries & Villiers, 1966b: 986
dessumi Descarpentries & Villiers, 1966b: 986
theryi Descarpentries & Villiers, 1966b: 986

prainae Hauser, 1900: 111 A: SD
quadricolor argenteus Kerremans, 1914c: 104 A: TAI
quadricolor quadricolor E. Saunders, 1873: 513 A: JA
quadricolor unifasciatus Kurosawa, 1985b: 148 A: JA (Ryukyus)
quinquefasciatus Miwa & Chûjô, 1935: 281 A: TAI
vitalisi Descarpentries & Villiers, 1966b: 983 A: YUN ORR
yaeyamanus Kurosawa, 1985b: 148 A: JA (Ryukyus)

tribe Aphanisticini Jacquelin du Val, 1859
subtribe Aphanisticina Jacquelin du Val, 1859

genus Aphanisticus Latreille, 1810: 169 type species Buprestis emarginata A.G. Olivier, 1790
Goniophthalma Chevrolat, 1838b: 106 type species Goniophthalma mitrata Chevrolat, 1838

aegyptiacus Théry, 1930a: 510 N: EG A: YE “Arabia” AFR
amblyderus Fairmaire, 1876b: 49 N: AG MO
angustatus P.H. Lucas, 1846e: pl. 16 E: GR FR IT PT SP TR N: AG MO TU A: SY TR
cupricolor Abeille de Perrin, 1893: 139

anniae Obenberger, 1932a: 220 A: TAI
antennatus E. Saunders, 1873: 518 A: JA NTRi ORRi
aureocupreus Kerremans, 1892b: 220 (Endelus) A: NP SD YUN ORR
aeneus Kerremans, 1893b: 349 (Endelus)
subviridis Kerremans, 1894b: 475 (Endelus)

bedeli Abeille de Perrin, 1893: 140 N: AG TU A: AE NP
binhensis Descarpentries & Villiers, 1963d: 158 A: YUN ORR
clavicornis Obenberger, 1918a: 4 A: “India or.”
congener E. Saunders, 1873: 518 A: JA SHN YUN
budhaicus Obenberger, 1944a: 8

BUPRESTIDAE

551

descarpentriesi Kalashian, 1993: 602 A: YUN ORR
dessumi Descarpentries & Villiers, 1963d: 157 A: GUX ORR
distinctus Perris, 1864: 283 E: FR IT PT SP N: AG MO TU
elongatus elongatus A. Villa & G.B. Villa, 1835b: 48 E: AB AL AU BE BH BU CR CZ FR GB GE GG GR HU IT LS
LU MC MD ME NL PL PT RO SB SK SL SP ST SZ TR UK N: AG EG MO TU A: IS SY TR
canaliculatus Schaufuss, 1882b: 554 [HN]
lamotei Guérin-Méneville, 1840b: 139

elongatus filum Schaefer, 1941: 76 E: FR SP
emarginatus A.G. Olivier, 1790a: no 32: 84 (Buprestis) E: AB AL AR AU BE BH BU CR CZ DE FR
GB GE GG GR HU IT LU ME NL PL PT RO SB SK SL SP ST SZ TR UK N: AG EG MO TU A: IN KZ TM
TR UZ
aetnensis Abeille de Perrin, 1897a: 16
coriaceus Rey, 1891c: 27

fossidiscus Obenberger, 1944a: 6 A: FUJ TAI
formosanus Obenberger, 1944a: 8

harlequin Obenberger, 1944a: 8 A: JIX
kanabei Kaszab, 1940: 113 E: HU RO
kaszabi Cobos, 1968c: 406 A: MG
mandarinus Gebhardt, 1929b: 96 A: ZHE
marseuli Tournier, 1868b: 144 N: EG AFR
metallescens Kerremans, 1892b: 219 A: SD
obsoletulus Obenberger, 1918a: 6 A: UP “India or.”
okinawanus Ohmomo, 2004: 136 A: JA (Ryukyus)
peninsulae Obenberger, 1924h: 625 A: FUJ GUX ORR
barbieri Descarpentries, 1958a: 64
klapperichi Obenberger, 1944a: 6

perpusillus Obenberger, 1918a: 5 A: “India or.”
perraudierei van de Pool, 1892: 19 A: IQ UP ORR
maerkeli Théry, 1934a: 84
georgi Obenberger, 1929f: 110

peyerimhoffi Descarpentries, 1953: 251 N: AG
punctulivertex Obenberger, 1944a: 7 A: JIX
pusillus A.G. Olivier, 1790a: no 32: 91 (Buprestis) E: AL AU BE BH BU CR CZ DE EN FR GB GE GR HU IT LA LS
LT NL PL PT RO SK SL SP ST SV SZ UK N: AG MO TU
lineola Germar, 1817b: 10 (Buprestis)

pygmaeus P.H. Lucas, 1846e: pl. 16 E: AB BU CR FR GR IT MA MD PT SP ST TR N: AG EG MO TU A: AF IS KZ
MG SA TD TM TR UZ
siculus Mulsant & Rey, 1863: 24

sinicola Obenberger, 1944a: 5 A: FUJ
sugonjaevi Alexeev, 1992: 382 A: AF PA
yasumatsui Kurosawa, 1954a: 37 A: JA

552

BUPRESTIDAE

genus Endelus Deyrolle, 1864a: 219 type species Endelus cupido Deyrolle, 1864
subgenus Endelus Deyrolle, 1864a: 219 type species Endelus cupido Deyrolle, 1864
acutiplicis Obenberger, 1944b: 44 A: FUJ
assamensis Obenberger, 1922b: 167 A: YUN “Assam” ORR
bicarinatus Théry, 1932a: 4 A: JA YUN ORR
eous Obenberger, 1944b: 42

collaris collaris E. Saunders, 1873: 518 (Aphanisticus) A: JA
collaris kerremansi Théry, 1927: 37 [RN] A: JA (Ryukyus) TAI
parallelus Kerremans, 1913b: 116 [HN]

continentalis Obenberger, 1944b: 43 A: FUJ JIX
formosae Obenberger, 1940f: 31 A: TAI
himalayanus Bílý, 1983b: 222 A: SD
japonicus Obenberger, 1944b: 42 A: TAI
klapperichi Obenberger, 1944b: 43 A: FUJ
lameyi Théry, 1932a: 13 A: YUN ORR
nepalensis Bílý, 1983b: 225 A: NP SD
pyrrosiae aokii Kurosawa, 1985b: 165 A: JA (Ryukyus)
pyrrosiae pyrrosiae Kurosawa, 1985b: 165 A: JA
sausai Kalashian, 2007: 660 A: GUI
sauteri Kerremans, 1912c: 209 A: TAI
sinensis Théry, 1932a: 13 A: TAI “China”
wittmeri Bílý, 1983b: 223 A: SD
subgenus Kubaniellus Kalashian, 1997: 73 type species Endelus kubani Kalashian, 1995
foveolatus Kalashian, 1995: 381 A: SCH
gyulnarae Kalashian, 1995: 380 A: SCH
jendeki Kalashian, 1997: 76 A: GUI
kareni Kalashian, 1997: 74 A: SCH SHA
opacipennis aeneopacus Kurosawa, 1985b: 163 A: TAI
opacipennis opacipennis Kurosawa, 1985b: 162 A: JA (Ryukyus)

subtribe Cylindromorphina Portevin, 1931

genus Cantonius Théry, 1929a: 57 type species Cantonius terryi Théry, 1929
subgenus Cantonius Théry, 1929a: 57 type species Cantonius terryi Théry, 1929
bolmi Kalashian, 2004: 128 A: GUI
fokienicus Kalashian, 2004: 126 A: FUJ
klapperichi Obenberger, 1940k: 188 A: FUJ
nitidifrons Kalashian, 2004: 127 A: SCH
obenbergeri Gebhardt, 1929b: 95 (Cylindromorphus) A: SHG
obesus Obenberger, 1940k: 188 A: FUJ
szechuanensis Obenberger, 1958a: 239 A: SCH

BUPRESTIDAE

553

terryi Théry, 1929a: 57 A: HKG
subgenus Procantonius Kalashian, 2004: 122 type species Cantonius khnzoriani Kalashian, 2004
khnzoriani Kalashian, 2004: 122 A: SCH
genus Cylindromorphus Kiesenwetter, 1857: 159 type species Buprestis filum Gyllenhal, 1817
araxidis Reitter, 1889a: 24 E: AB AR A: TR
bifrons Rey, 1889c: clxi E: AU BU CZ FR HU RO SK ST UK A: ES KZ MG TR UZ
acus Abeille de Perrin, 1897a: 31

bohemicus Obenberger, 1933c: 177 E: CZ
carpetanus Perris, 1864: 282 E: SP
cribratus Abeille de Perrin, 1897a: 30 N: AG MO
peyerimhoffi Abeille de Perrin, 1910: 222

dalmatinus Csiki, 1915: 103 E: BH CR IT
filum Gyllenhal, 1817: 124 (Buprestis) E: AL AR AU BE BH BU CR CZ FR GE GG GR HU IT KZ MC MD PL RO SB
SK SL ST TR UK A: KI KZ TR UZ
cylindrus A. Villa & G.B. Villa, 1835b: 48 (Aphanisticus)
opacus Abeille de Perrin, 1897a: 31
sulcatulus Pic, 1918c: 2

gallicus angelinii Curletti, 1990: 103 E: IT
gallicus gallicus Mulsant & Rey, 1863: 22 E: FR
parallelus Fairmaire, 1859: 49 E: FR GR (Kríti) PT SP
andalusiacus Cobos, 1950: 462
impressifrons Pic, 1918c: 2
strigatulus Rey, 1889c: clx

platiai Curletti, 1981: 93 E: IT (Sicilia)
popovii Mannerheim, 1853a: 103 (Aphanisticus) E: AU CT HU KZ SB ST UK A: ES KI KZ MG NMO WS
mongolicus Obenberger, 1928b: 21

pubescens Semenov, 1895d: 266 A: KZ TM UZ
pyrethri Stierlin, 1864b: 491 E: CT KZ MD ST UK A: ES KI KZ MG TD TM UZ WS
caspicus Obenberger, 1934a: 21
pyrethri Marseul, 1866: 501 [HN]

strictipennis Reitter, 1895d: 154 E: AB A: TR
talyshensis Kalashian, 1989: 39 E: AB
turkestanicus Abeille de Perrin, 1904a: 224 A: “Turkestan”
vedicus Kalashian, 2002: 81 E: AR
genus Paracylindromorphus Théry, 1930a: 493 type species Agrilus subuliformis Mannerhem, 1837
Agrilomorphus Portevin, 1931: 335 type species Agrilus subuliformis Mannerhem, 1837

assamensis Obenberger, 1947: 141 A: “Assam” “Himalaya”
birmanicus Obenberger, 1947: 140 A: YUN ORR

554

BUPRESTIDAE

bodongi Kerremans, 1914a: 88 (Aphanisticus) A: HP ORR
cairensis Théry, 1930a: 499 N: EG
cashmirensis Obenberger, 1935e: 30 A: “Kashmir”
chinensis Obenberger, 1927f: 77 (Cylindromorphus) A: YUN ZHE
fujianensis Kubáň, 2006: 407 A: FUJ
chinensis Obenberger, 1947: 142 [HN]

formosanus Miwa & Chûjô, 1935: 281 (Cylindromorphus) A: TAI
gebhardti Obenberger, 1927f: 76 (Cylindromorphus) A: ZHE
japanensis E. Saunders, 1873: 519 (Cylindromorphus) A: JA “China” “Korea”
klapperichi Obenberger, 1947: 142 A: FUJ
lebedevi Obenberger, 1928b: 21 (Cylindromorphus) A: KZ TD TM UZ
montanus Obenberger, 1924f: 110 (Cylindromorphus) A: “Assam”
pinguis pinguis Fairmaire, 1876b: 49 (Cylindromorphus) N: AG LB MO TU
pinguis rabinovitchi Théry, 1935b: 154 N: EG A: SI AFR
pinguis sefrensis Bonnaire, 1893: 317 (Cylindromorphus) N: AG MO
richteri Théry, 1937b: 249 A: FE HEI JA “Korea”
semenovi Théry, 1937a: 156 E: ST A: KZ TD TM
richteri Stark, 1952: 282 [HN]
starki Bellamy, 1998: 96 [RN]

simlaicus Obenberger, 1924f: 110 (Cylindromorphus) A: HP
simlaicus Obenberger, 1947: 141 [HN]

sinae Obenberger, 1947: 143 A: FUJ
sinuatus Abeille de Perrin, 1897c: 36 (Cylindromorphus) N: AG TU A: IS JO
spinipennis carmenae Coello & Verdugo, 2001: 17 E: SP
spinipennis spinipennis Bedel, 1890: cxxxvi (Cylindromorphus) N: AG MO
mariaeantoniae Abeille de Perrin, 1894b: 67 (Cylindromorphus)

subuliformis lethierryi Théry, 1930a: 496 N: AG MO
subuliformis subuliformis Mannerheim, 1837: 117 (Agrilus) E: AB AL AR AU BH BU CR CZ FR GG GR HU IT KZ
MC ME PL RO SK SL SP ST TR UK A: AF ES IN KI KZ LE MG NW SY TD TM TR UZ WS
tauricus Gory, 1841: 269 (Agrilus)

tokioensis Théry, 1937c: 322 A: JA (Honshu)
transversicollis kozlovi Alexeev, 1975: 160 A: MG NMO XIN
transversicollis transversicollis Reitter, 1913e: 657 (Cylindromorphus) E: AB AR GG KZ ST A: IN KZ TD
TM UZ
magnus Obenberger, 1929c: 123 (Cylindromorphus)

tribe Coraebini Bedel, 1921
subtribe Amorphosomatina Majer, 2000

genus Amorphosoma Laporte, 1835: 166 type species Buprestis penicillata Klug, 1825
aculeatum Ganglbauer, 1889: 24 (Coraebus) A: GAN SCH XIZ YUN
emmerichi Obenberger, 1934d: 43 (Coraebus)

afghanicum Alexeev & Volkovitsh, 1992: 372 (Vanroonia) A: AF PA SD

555

BUPRESTIDAE
bispinosum Wiedemann, 1823: 103 (Buprestis) A: NP SD ORR
cochinchinae Descarpentries & Villiers, 1967a: 489 A: YUN ORR
himalayanum Obenberger, 1958c: 510 (Vanroonia) A: NP UP
pectorale Kerremans, 1892b: 210 A: “Himalaya” ORR
perroti Descarpentries & Villiers, 1967a: 489 A: YUN ORR
rambouseki Obenberger, 1934n: 150 (Coraebus) A: SCH

genus Coroebina Obenberger, 1923a: 33 type species Coroebina maderi Obenberger, 1923 (=Coraebus gentilis
Kerremans, 1890)
beatricis Obenberger, 1944d: 70 A: “Himalaya” ORR
bilyi Akiyama & Ohmomo, 1993b: 137 A: YUN ORR
chloropicta Kerremans, 1892b: 202 (Coraebus) A: NP UP
balthasari Obenberger, 1934e: 73
srdinkoi Obenberger, 1924b: 32 (Coraebus)

gentilis Kerremans, 1890c: ccv (Coraebus) A: UP ORR
maderi Obenberger, 1923a: 34

gongis Gory, 1841: 278 (Coraebus) A: “Himalaya” ORR
kashmirensis Obenberger, 1934e: 71 A: “Kashmir”
specialis Obenberger, 1934e: 72 A: “Himalaya”
yunnanensis Obenberger, 1934e: 73 A: YUN ORR
genus Metasambus Kerremans, 1903: 237 type species Coraebus weyersi Kerremans, 1900
Pusilloderes Obenberger, 1916d: 262 type species Pusilloderes hoscheki Obenberger, 1916

hoscheki Obenberger, 1916d: 262 A: GUI SHN YUN
tonkinensis Descarpentries & Villiers, 1966d: 989 A: GUI HUN ORR
weyersi Kerremans, 1900a: 12 (Sambus) A: HAI ORR

subtribe Clematina Majer, 2000

genus Clema Semenov, 1900d: 590 type species Clema deserti Semenov, 1900
deserti deserti Semenov, 1900d: 592 A: AE IN KZ TM UZ XIN
candida Zubkov, 1833: 318 (Buprestis)

deserti rabinovitchi Théry, 1935b: 151 N: EG A: IS
freudei Cobos, 1963: 362 A: IQ IS
genus Niehuisia Curletti, 1995: 181 type species Niehuisia maghrebica Curletti, 1995
maghrebica Curletti, 1995: 182 N: TU (Djerba)

subtribe Cisseina Majer, 2000

genus Diphucrania Dejean, 1833: 81 type species Buprestis leucosticta Kirby, 1819
Cisseis Gory & Laporte, 1839: 1 type species Buprestis duodecimguttata Guérin-Méneville, 1830 (=Buprestis duodecimmaculata Fabricius, 1801)
Ethon Gory & Laporte, 1839: 1 type species Buprestis leucosticta Kirby, 1819

556

BUPRESTIDAE

viridipurpurea Carter, 1924: 27 (Cisseis) Ei: SP (Aranjuez Prov.) AUR
sapphira Carter, 1933: 163 (Cisseis)

subtribe Coraebina Bedel, 1921

genus Brachycoraebus Kerremans, 1903: 234 type species Coraebus viridis Kerremans, 1900
Neocoraebus Kerremans, 1903: 230 type species Amorphosoma abyssinicum Kerremans, 1899

arunensis Kubáň, 1996: 2 A: NP
klapperichi Obenberger, 1959a: 221 A: FUJ
punctatus Baudon, 1968: 36 (Cisseicoraebus) A: YUN ORR
yajimai Ohmomo, 2011b: 217 (Coraebus)

genus Coraebus Gory & Laporte, 1839: 1 type species Buprestis undata Fabricius, 1787
Coraegrilus Fairmaire, 1889a: 33 type species Coraegrilus amplithorax Fairmaire, 1889
Coroebus Agassiz, 1846: 100 [unjustified emendation] type species Buprestis undata Fabricius, 1787
Negreia Cobos, 1962b: 27 type species Negreia niveosparsa Cobos, 1962 (=Coraebus andrewesi Obenberger, 1922)

acutus J. Thomson, 1879a: 54 A: ANH FUJ GAN GUA GUI GUX HEN HUB HUN JIX NIN SCH SHA SHG YUN
ZHE ORR
bedeli Théry, 1895a: cxiii
chinensis Kerremans, 1895: 214
quadrispinosus Fairmaire, 1891f: ccvii

aeneopictus Kerremans, 1895: 217 (Meliboeus) A: SD YUN ORR
lienardae Baudon, 1968: 15 (Epimacha)

aequalipennis Fairmaire, 1888b: 121 A: BEI FE FUJ GAN HEB HEN JIA JIX NC SCH SHA SHG XIZ YUN ZHE
joannisi Théry, 1895a: cxi
nomurai Miwa & Chûjô, 1940: 53/58

aesopus Kerremans, 1913b: 111 A: FUJ TAI
amabilis Kerremans, 1895: 216 A: GUI GUX HAI HUB HUN JIX NP SCH SD SHA SHG XIZ YUN ZHE ORR
amamianus Kurosawa, 1985b: 143 A: JA (Ryukyus) ORR
amplithorax Fairmaire, 1889b: 33 (Coraegrilus) A: SCH XIZ YUN
vrazi Obenberger, 1934d: 41

andrewesi Obenberger, 1922b: 137 A: SD NTRi ORR
kanarae Obenberger, 1922b: 137
niveosparsus Cobos, 1962b: 30 (Negreia)

annamensis Descarpentries & Villiers, 1967a: 482 A: FUJ HUN ORR
blankae Kubáň, 1995a: 10

aurofasciatus Hope, 1831: 25 (Buprestis) A: NP SD UP “India” “Kashmir”
dorsalis Kerremans, 1892b: 200
wendleri Obenberger, 1924b: 31

baoshanensis Kubáň, 1996: 7 A: YUN
baylei Bourgoin, 1922a: 23 A: YUN ORR
becvari Kubáň, 1995a: 8 A: NIN SCH YUN

BUPRESTIDAE

557

bilyi Kubáň, 1996: 9 A: YUN ORR
bunun Miwa & Chûjô, 1940: 54/62 A: TAI
businskyorum Xu & Kubáň, 2013: 688 A: CHQ HUB SCH
cavifrons Descarpentries & Villiers, 1967a: 482 A: CHQ FUJ GUA GUX HAI SCH ZHE ORR
cervenkai Kubáň, 1995b: 93 A: FUJ
chrysogaster Kurosawa, 1953b: 96 A: FUJ TAI
cingulatus Hope, 1831: 25 (Buprestis) A: BT HP KA NP PA SCH SD UP XIZ “China” ORR
caschmirensis Kollar, 1844: 505 (Agrilus)
costatus J. Thomson, 1879a: 54
subtrifasciatus Obenberger, 1917f: 60
sulcicollis Kerremans, 1895: 215

cisseiformis Obenberger, 1913d: 90/99 A: CHQ “China”
clermonti Bourgoin, 1924b: 178 A: SCH YUN ORR
cloueti Théry, 1895a: cxii A: ANH FUJ GAN GUA GUI GUX HEN HUB HUN JIA JIX NIN SCH SHA SHG SHN SHX
TAI XIZ YUN ZHE ORR
aeneicollis Kerremans, 1895: 216
dollei Théry, 1895a: cxiv
heyrovskyi Obenberger, 1934i: 110
kerremansi Obenberger, 1916d: 265 [RN]
mushensis Miwa & Chûjô, 1935: 277
staudingeri Obenberger, 1914g: 135
szechuanensis Obenberger, 1934b: 25
tatsienluus Obenberger, 1934l: 129
tonkinensis Bourgoin, 1924b: 179

coelestiofulgens Akiyama & Ohmomo, 1993a: 64 A: HAI TAI
coeruleus Kerremans, 1892b: 200 A: AP PA SD ORR
monticola Obenberger, 1922b: 139

conspicuus J. Thomson, 1878a: 81 A: NP SD “India”
strandi Obenberger, 1922b: 136

cupreosplendens Akiyama, 1989: 207 A: GUX HUN TAI
hengshanensis Peng, 1992: 395/405

cyaneopictus Kerremans, 1892b: 201 A: HUN YUN ORR
daisenensis Miwa, 1940: 70 A: JA
daisensensis Miwa, 1940: 70 [unused original spelling]
montanus Miwa & Chûjô, 1940: 56/67 [HN]
susterai Obenberger, 1944e: 119

davidis Fairmaire, 1886: 332 A: FUJ GUI HUN JIX SCH YUN ORR
guerryi Théry, 1922b: 264

delepinei Baudon, 1960: 226 A: GUX YUN ORR
delicatus Kerremans, 1895: 214 A: SHG
denticollis E. Saunders, 1866: 313 A: FUJ GUA GUX HUB HUN JIX YUN ORR

558

BUPRESTIDAE

beauchenei Fairmaire, 1889b: 347
beauchenii Fairmaire, 1889b: 347 [incorrect original spelling]
fossulatus Kerremans, 1892c: 812
murinus Kerremans, 1888b: lxvii (Meliboeus)

diminutus Gebhardt, 1929a: 31 A: FUJ GUA GUI GUX HUB HUN JA JIA JIX SCH SHA SHG SHX TAI YUN ZHE ORR
grafi Obenberger, 1934d: 43

duodecimpunctatus Obenberger, 1940k: 173 A: FUJ
elatus Fabricius, 1787a: 184 (Buprestis) E: AB AL AR AU BE BH BU BY CR CT CZ FR GE GG GR HU IT KZ MC
MD ME NT PL PT RO SB SK SL SP ST SZ TR UK N: AG EG A: IN IQ IS KZ SY TR WS
aeruginosus Latreille, 1804a: 69 (Buprestis) [RN]
comari Marseul, 1866: 419
cylindraceus Abeille de Perrin, 1896b: 279 [HN]
lapsanae Bonelli, 1812: 176 (Buprestis)
metallicus Gory & Laporte, 1839: 14
protensulus Obenberger, 1935u: 826 [RN]
pruinosus Küster, 1846: 53
repletus Abeille de Perrin, 1893: 135
sinuatus Panzer, 1796b: 13 (Buprestis) [HN]
subfasciatus Küster, 1846: 54

elegantissimus Akiyama, 1989: 205 A: TAI
elongaticollis Akiyama & Ohmomo, 1993a: 63 A: TAI
elongatus Peng, 1998: 12 A: JIX
ephippiatus Théry, 1938b: 176 A: FUJ GUI GUX HEN HUB HUN JIX SHG TAI YUN ZHE
fallaciosus Bourgoin, 1925a: 112 A: GUX YUN ORR
fasciatus Villers, 1789: 339 (Buprestis) [on Off. List] E: AU BH BU CR FR GE GR HU IT MC ME PT RO SK SL SP
SZ TR UK N: AG TU A: IN LE SY TR
bifasciatus A.G. Olivier, 1790a: no 32: 46 (Buprestis)
florentinus Herbst, 1801: 178 (Buprestis)
trifasciatus Regimbeau, 1876: 65

femina Kubáň, 1997: 33 A: YUN ORR
fokienicus Obenberger, 1940k: 172 A: FUJ GUX YUN ORR
formosanus formosanus Miwa & Chûjô, 1935: 279 A: TAI
formosanus heikichii Akiyama, 1987: 80 A: JA (Ryukyus)
formosanus nishiyamai Akiyama, 1987: 79 A: JA (Ryukyus)
formosanus oshimensis Akiyama, 1987: 80 A: JA (Ryukyus)
frater Bourgoin, 1925a: 112 A: JIX ORR
fukamachii Akiyama, 1987: 77 A: JA (Ryukyus)
gagneuxi Baudon, 1963: 73 A: YUN ORR
gestroi Kerremans, 1892c: 813 A: YUN “Himalaya” ORR
gorkai Gebhardt, 1929a: 29 A: FUJ JIX ZHE
jiangxiensis Peng, 1990: 11/13

BUPRESTIDAE
gorkaianus Kubáň, 1997: 37 A: SCH YUN
hastanus Gory & Laporte, 1839: 10 A: BT FUI GUI JA (Ryukyus) NP SCH SD TAI YUN ORR
bajulus Deyrolle, 1864a: 122
oberthueri Lewis, 1896: 335

hauseri Obenberger, 1930a: 107 A: ZHE ORR
holzschuhi Kubáň, 1996: 11 A: NP
hoscheki Gebhardt, 1929a: 31 A: HUB HUN JIX ZHE ORR
houianus Kubáň, 1995a: 16 A: SCH YUN
ignifrons Fairmaire, 1895b: 174 A: YUN ORR
ignotus ignotus E. Saunders, 1873: 512 A: GUA JA TAI
dlabolai Obenberger, 1944e: 120

ignotus matsudai Kurosawa, 1953b: 102 A: JA (Tanega-shima, Yaku-shima)
ignotus okinawanus Kurosawa, 1985b: 142 A: JA (Ryukyus)
ignotus shibatai Kurosawa, 1963b: 151 A: JA (Ryukyus)
insulicola Kerremans, 1912c: 205 A: FUJ GUI GUX HUN JIX SCH TAI ORR
bechynei Obenberger, 1944e: 121
vimmeri Obenberger, 1934d: 39

intemeratus Obenberger, 1940k: 171 A: YUN ZHE ORR
bourgoini Descarpentries & Villiers, 1967a: 484

iriei iriei Kurosawa, 1985b: 145 A: JA (Ryukyus)
iriei yoshimotoi Kurosawa, 1985b: 147 A: JA (Ryukyus: Yaku-shima)
ishiharai Kurosawa, 1953b: 104 A: JA
jedlickai Obenberger, 1934d: 40 A: HUN SCH
jelineki Descarpentries & Villiers, 1967a: 483 A: YUN ORR
kiangsuanus Obenberger, 1934d: 41 A: FE HUB HUN JIA SCH SHA
kiyoshii Akiyama, 1989: 209 A: TAI
klapaleki Obenberger, 1924b: 32 A: GUA GUI GUX HAI HUN JIX SCH
klickai Obenberger, 1930a: 105 A: GAN HUB SHA
komiyai Akiyama, 1988: 59 A: TAI
kubani Peng, 1998: 13 A: FUJ GUA GUX JIX
kulti Obenberger, 1940k: 170 A: YUN
kurosawai Akiyama, 1988: 54 A: HAI HUN TAI ORR
larminati Kubáň, 1995a: 18 A: YUN ORR
lepidulus Obenberger, 1940k: 170 A: JIX ZHE
lesnei Bourgoin, 1922a: 22 A: YUN ORR
leucospilotus Bourgoin, 1922a: 22 A: FUJ GUI HUN JIX NP SCH YUN ORR
pruni Miwa & Chûjô, 1935: 278

levasseuri Bourgoin, 1925a: 111 A: GUX ORR
lienhwachiensis Akiyama, 1988: 58 A: TAI
linnei Obenberger, 1922b: 135 A: FUJ GUA HKG HUN JIX SCH TAI YUN ORR
loochooensis Kano, 1929: 96 A: JA (Ryukyus)

559

560

BUPRESTIDAE

lubopetri Kubáň, 1995a: 19 A: FUJ ORR
maculifer Abeille de Perrin, 1897c: 33 A: IN TR
meliboeoides Bílý, 1983b: 221 A: NP ORR
mianningensis Peng, 1991: 36 A: SCH YUN
mirus Kubáň, 1996: 13 A: SD
muehlei Kubáň, 1996: 13 A: FUJ
nelsoni Akiyama & Ohmomo, 1989: 197 A: NP YUN ORR
nigromaculatus Kurosawa, 1953b: 102 A: BEI JA
nikodymi Kubáň, 1995b: 97 A: FUJ
niponicus Lewis, 1894: 224 A: JA
oertzeni Ganglbauer, 1886e: 309 E: CR CZ GR RO SB SL TR A: TR
carinifer Abeille de Perrin, 1891: 280

olexai Bílý, 1983b: 218 A: NP SD
perroti Descarpentries, 1948b: 94 A: GUA ORR
pickai Kubáň, 1995a: 21 A: YUN ORR
probstorum Kubáň, 1995a: 22 A: NP ORR
pseudopurpura Kubáň, 1997: 45 A: SD YUN ORR
pulchellus Nonfried, 1895: 304 A: YUN ORR
purkynei Obenberger, 1934l: 129 A: TAI
purpura Kubáň, 1996: 16 [RN] A: GUX HUN JIX SCH YUN
purpuratus Obenberger, 1921a: 97 [HN]

purpuratiformis Kubáň, 1995a: 23 A: HAI YUN
quadriundulatus Motschulsky, 1866: 165 A: FE FUJ GAN GUI HUB HUN JA JIX SCH SHA XIZ YUN ZHE
“Himalaya” “Kashmir”
nipponicola Obenberger, 1914g: 135
obscurus Peng, 1991: 36

rubi Linnaeus, 1767: 661 (Buprestis) E: AB AL AR AU BH BU CR CZ FR GE GG GR HU IT LU MC MD ME PL PT
RO SB SK SL SP ST SZ TR UK N: AG MO A: CY IN IQ IS LE SY TR
nebulosus Scopoli, 1772: 104 (Mordella)

rusticanus fujiyamai Kurosawa, 1953b: 101 A: JA (Izu)
rusticanus rusticanus Lewis, 1893: 331 A: JA ZHE
kamikochianus Kano, 1929: 96

sakagutii imasakai Akiyama, 1988: 60 A: SHG TAI
sakagutii sakagutii Kurosawa, 1963a: 100 A: JA
salamandriformis Kubáň, 1995b: 98 A: YUN ORR
salvazai Bourgoin, 1922a: 22 A: FUJ GUA GUI JIX ORR
sauteri Kerremans, 1912c: 205 A: ANH CHQ FUJ GAN GUA GUI GUX HEN HUB HUN JIX NP SCH SD SHA SHX
TAI XIZ YUN ZHE ORR
auricomus Bourgoin, 1922a: 22
guangxiensis Peng, 1998: 14
komareki Obenberger, 1934d: 42

BUPRESTIDAE

561

semipurpureus Fairmaire, 1889b: 347 A: GUX JIX NP SD YUN ORR
feae Kerremans, 1892c: 811
montivagus Obenberger, 1922b: 139

sherpa Kubáň, 1996: 17 A: NP
sidae lucens Obenberger, 1958a: 235 A: GUA GUI SCH YUN ORR
sidae sidae Kerremans, 1888b: lxvii A: FUJ GUI GUX HAI HUN JIA JIX NP SCH SD YUN ORR
alternefasciatus Obenberger, 1914g: 134
intricatus Fairmaire, 1889b: 347

simplex Peng, 1991: 35 A: HUN YUN
sinomeridionalis Kubáň, 1995a: 23 A: FUJ GUA GUI GUX HAI JIX SCH YUN ORR
sonani Miwa & Chûjô, 1935: 278 A: GUX TAI
spathatus Kerremans, 1892c: 818 (Amorphosoma) A: XIZ YUN ORR
assamensis Obenberger, 1922b: 137
dentipennis Kerremans, 1900b: 78

spectabiliformis Kubáň, 1995a: 24 A: GUI SCH SHA
spectabilis Bílý, 1983b: 219 A: NP SD ORR
spevari Kubáň, 1995a: 25 A: SCH YUN ORR
splendens Peng, 1998: 12 A: GUX
stichai Obenberger, 1924a: 97 A: GUI GUX YUN ORR
signatulus Obenberger, 1934l: 129

taiwanus Akiyama, 1988: 60 A: TAI
tazoei Akiyama, 1988: 56 A: TAI
teres Kubáň, 1996: 20 A: FUJ GUA
thoracellus Kerremans, 1900c: 320 A: FUJ HUN
torigaii Akiyama & Ohmomo, 1993a: 60 A: TAI YUN
undatus Fabricius, 1787a: 182 (Buprestis) E: AU BE BH CR CZ FR GE GR HU IT MD PL PT RO SB SK SP ST SZ
UK N: MO A: TR
pruni Panzer, 1796a: 21 (Buprestis)

vagepictus Obenberger, 1934g: 90 [RN] A: SHN
nebulosus Obenberger, 1924b: 31 [HN]

vicarius Kubáň, 1995b: 100 A: YUN ZHE ORR
violaceipennis E. Saunders, 1866: 313 A: FUJ GAN GUI HUB HUN JIX NP SD YUN ZHE ORR
quadraticollis Fairmaire, 1895b: 174

viridimicans Akiyama & Ohmomo, 1993a: 57 A: TAI
volkovitshi Xu & Kubáň, 2013: 692 A: GUX YUN
vuilletae Bourgoin, 1925a: 112 A: GUA ORR
yanshanensis Peng, 1991: 37 A: HUB SCH SHX
yuanyunae Peng, 1998: 14 A: JIX ORR
zhangi Peng, 1990: 11, 13 A: HUN JIX
zhejiangensis Peng, 1998: 15 A: ZHE
zoufali Obenberger, 1930a: 106 A: GAN GUA HUN JIX SCH ZHE ORR

562

BUPRESTIDAE

genus Mundaria Kerremans, 1894b: 468 type species Mundaria typica Kerremans, 1894
typica Kerremans, 1894b: 469 A: NP SD ORR

subtribe Meliboeina Majer, 2000

genus Kamosia Kerremans, 1898b: 314 type species Kamosia duvivieri Kerremans, 1898
Gassneria Obenberger, 1919a: 17 type species Gassneria mirabilis Obenberger, 1919

duvivieri Kerremans, 1898b: 315 A: YE AFR
genus Meliboeus Deyrolle, 1864a: 132 type species Buprestis aeneicollis Villers, 1789 (=Coraebus fulgidicollis
P.H. Lucas, 1846)
subgenus Meliboeoides Théry, 1942a: 120 type species Buprestis amethystinus A.G. Olivier, 1790
adlbaueri Niehuis, 1989c: 121 A: IN IQ IS SY TR
alluaudi Théry, 1930a: 394 N: LB
amethystinus amethystinus A.G. Olivier, 1790a: no 32: 83 (Buprestis) E: FR IT PT SP SZ N: AG LB MO TU
amethystinus destefanii Sparacio, 1984: 14 E: IT (Sicilia)
cyaneus Ballion, 1871a: 350 (Coraebus) E: AR CT ST KZ A: IN KI KZ TD TM UZ
granulatus Gory & Laporte, 1839: 15 (Coraebus) E: SP N: AG LB MO TU
maceki Niehuis, 2014d:198 A: IQ
makrisi Mühle & Brandl, 2009: 54 A: CY
oliveri Niehuis, 2014c: 94 E: GR (Lesbos, Ródos) A: CY IN IQ IS JO LE SY TR
parvulus Küster, 1852: 63 (Coraebus) E: AB AL AR BH BU CR GG GR HU IT KZ MC ME RO SB ST TR UK A: CY
IN IQ IS KI KZ SY TD TM TR UZ
chalybaeus Küster, 1852: 64 (Coraebus)
ponticus Obenberger, 1924b: 33
violaceus Kiesenwetter, 1857: 115 (Coraebus)

robustus Küster, 1852: 62 (Coraebus) E: AB AR GG ST A: IN IQ TM TR
simandli Niehuis, 2014b: 85 A: IN
skalei Niehuis, 2011: 90 A: IN SY TM TR
ursulae Niehuis, 2014a: 76 A: IN
subgenus Meliboeus Deyrolle, 1864a: 132 type species Buprestis aeneicollis Villers, 1789 (=Coraebus fulgidicollis
P.H. Lucas, 1846)
Bourgoinia Obenberger, 1926c: 30 type species Bourgoinia achardi Obenberger, 1926
Chakriia Tongyai, 1935: 141 type species Chakriia ruficollis Tongyai, 1935 (=Coraebus aurofasciatus E. Saunders, 1866)
Lakhonia Descarpentries & Villiers, 1967b: 992 [HN] type species Meliboeus coomani Bourgoin, 1924
Meliacanthus Théry, 1942a: 120 type species Coraebus cupreomarginatus E. Saunders, 1866
Melicoraebus Théry, 1932b: 97 type species Meliboeus magnificus Kerremans, 1892
Melixes Schaefer, 1949: 341 type species Coraebus aeratus Mulsant & Rey, 1863
Nalanda Théry, 1904: 160 type species Nalanda horni Théry, 1904
Neosambus Fisher, 1921: 350 type species Neosambus cupricollis Fisher, 1921 (=Nalanda kurosawai Tôyama, 1986)
Tonkinula Obenberger, 1923a: 35 type species Coraebus aurofasciatus E. Saunders, 1866

BUPRESTIDAE
aeratus Mulsant & Rey, 1863: 10 (Coraebus) E: FR IT PT SP N: AG MO
alfierianus Théry, 1935b: 148 N: EG A: IN IS
hoberlandti Obenberger, 1944c: 58

algiricus Théry, 1930a: 408 N: AG
annon Gory, 1841: 279 (Coraebus) A: SD
anticerugosus Obenberger, 1944h: 305 A: JIX SHA
arrowi Bourgoin, 1924c: 214 A: YUN ORR
aureolus Abeille de Perrin, 1893: 136 (Coraebus) N: AG EG MO IS
latesculptus Obenberger, 1916d: 267

aurofasciatus E. Saunders, 1866: 312 (Coraebus) A: YUN ORR
mouhotii E. Saunders, 1871: 105 [RN]
ruficollis Tongyai, 1935: 142 (Chakriia)

babaulti Théry, 1930c: 156 A: HP
belzebuth Obenberger, 1944c: 58 A: CH
berbericus Théry, 1930a: 402 N: AG
bilyi Ohmomo & Akiyama, 1989: 6 (Nalanda) A: TAI YUN ORR
caucasicus Abeille de Perrin, 1896b: 282 (Coraebus) E: AB AR GG GR ST A: TM TR
angoranus Obenberger, 1935b: 18

chinensis Obenberger, 1927f: 19 A: FE HEI JA (Tsushima) JIA NC SC ZHE
shirozui Kurosawa, 1954a: 36
tscherskii Alexeev, 1979b: 125

chrysomelinus Kerremans, 1892b: 205 A: “Bengal” ORR
convexithorax Obenberger, 1944h: 304 A: FUJ
coraeboides Kerremans, 1892b: 220 (Endelus) A: SD
cryptocerus Kiesenwetter, 1858: 247 (Coraebus) E: GR IT
cupreomarginatus E. Saunders, 1866: 312 (Coraebus) A: YUN ORR
harmandi Baudon, 1962: 32
isanensis Ohmomo, 2011b: 223 (Meliacanthus)
prachadhipoka Tongyai, 1935: 142 (Chakriia)
transversus Kerremans, 1892c: 814
victor Obenberger, 1931e: 213

cupricollis E. Saunders, 1866: 314 A: TAI YUN ORR
balthasari Obenberger, 1935b: 17
saundersi Kerremans, 1892a: 233 [RN]

cyprius Zurcher, 1911: 254 (Coraebus) E: CY
episcopalis Mannerheim, 1837: 107 (Agrilus) E: AL BU CR GR IT MC ME TR N: EG A: SY TR
gemmeus Abeille de Perrin, 1896b: 280 (Coraebus)
purpureus Gory & Laporte, 1839: 13 (Coraebus)
saginatus Kiesenwetter, 1857: 115 (Coraebus)

fleutiauxi Bourgoin, 1924c: 215 A: YUN ORR

563

564

BUPRESTIDAE

formosanus Obenberger, 1944i: 436 A: TAI
fulgidicollis P.H. Lucas, 1846e: pl. 16 (Coraebus) E: AB AL AU BH BU CR CZ FR GR HU IT MC MD ME PT RO SK
SL SP ST SZ TR UK N: AG MO TU A: CY KZ SY TR
aeneicollis Villers, 1789: 340 (Buprestis) [HN]
bodemeyeri Kerremans, 1911: 634
villersii E. Saunders, 1870: 34 (Coraebus) [RN]

gibbicollis gibbicollis Illiger, 1803: 239 (Buprestis) E: FR IT PT SP N: AG MO
cupulariae Abeille de Perrin, 1891: 282 (Coraebus)
cylindraceus Gory & Laporte, 1839: 14 (Coraebus)

gibbicollis maddalenae Curletti & Magnani, 1987: 169 E: IT (Sardegna)
graminis graminis Panzer, 1799a: 8 (Buprestis) E: AB AL AR BH BU CR CT FR GR HU IT MD RO SL SP ST TR
UK N: TU A: SY TR
graminis Herbst, 1801: 262 (Buprestis) [HN]
helichrysi Abeille de Perrin, 1904b: 282 (Coraebus)

graminis mammutones Curletti, 2001: 132 E: IT (Sardegna)
graminis marrocanus Obenberger, 1916d: 269 N: AG MO
guyoti Obenberger, 1920a: 85 A: SI
halperini Niehuis, 1997a: 51 A: IS
helferi Obenberger, 1922b: 147 A: NP SD
heydeni Abeille de Perrin, 1897c: 34 (Coraebus) E: AR A: IN IS LE SY TM TR
bodenheimeri Théry, 1938d: 473
staneki Obenberger, 1935b: 16
xerophilus Obenberger, 1916d: 266

hideoi Ohmomo & Akiyama, 1989: 14 (Nalanda) A: TAI
hoscheki Obenberger, 1916d: 266 N: EG A: CY IS LE SY TR
alfierii Pic, 1924c: 54

impressithorax Pic, 1924c: 54 N: EG A: IQ IS
indicola Kerremans, 1892b: 204 A: “Bengal” “India” ORR
indignus Obenberger, 1931e: 214 A: SD
kabakovi Alexeev, 1992: 377 A: AF
kaszabi Cobos, 1966d: 319 A: AF PA
komiyai Ohmomo & Akiyama, 1989: 16 (Nalanda) A: TAI
kubani Niehuis, 1994: 353 N: CI
kurosawanus Ohmomo & Akiyama, 1989: 5 (Nalanda) A: TAI
lagerstraemiae Ohmomo & Akiyama, 1989: 16 (Nalanda) A: TAI
mandarinus Obenberger, 1927f: 19 A: ANH FUJ GUI GUX JIX SCH SHA SHN ZHE
margotanus Novak, 2010a: 111 (Kamosia) A: AE IN OM PA YE
minutus Kerremans, 1893b: 339 A: “Bengal” ORR
moghrebicus Théry, 1930a: 400 N: MO
morawitzi Semenov, 1905a: 132 (Coraebus) E: BU CT GR HU KZ SK ST UK “Caucasus” A: KI KZ TR WS

BUPRESTIDAE
proximus Obenberger, 1916d: 269
turkestanicus Obenberger, 1924b: 33

mukiensis Stepanov, 1958a: 118 A: TD
nickerli Obenberger, 1922b: 147 A: “Himalaya”
nigroscutellatus Obenberger, 1935b: 17 A: YUN ORR
niisatoi Ohmomo & Akiyama, 1989: 18 (Nalanda) A: TAI
ohbayashii Kurosawa, 1957: 185 (Nalanda) A: GAN JA JIX NC SCH SHA SHN
orientalis Abeille de Perrin, 1905: 207 (Coraebus) E: RO
pentacallosus Ohmomo & Akiyama, 1989: 7 (Nalanda) A: TAI
potanini Obenberger, 1929e: 61 A: SCH
primoriensis Alexeev, 1979b: 125 A: BEI FE NEI
princeps Obenberger, 1927f: 18 A: YUN
purpureicollis Théry, 1930f: 299 A: YUN
pygmaeolus Obenberger, 1917a: 34 A: CH
rajah Obenberger, 1944i: 436 A: “India ”
reitteri Semenov, 1890b: 211 (Coraebus) E: KZ ST A: AF IN KI KZ TD TM UZ
klapperichi Cobos, 1966d: 318
pamirensis Obenberger, 1935b: 15
zaisanensis Obenberger, 1935b: 16

romanovi Stepanov, 1958a: 119 A: TD
rutilicollis rutilicollis Obenberger, 1914g: 135 A: BEI JA TAI
rutilicollis ryukyuensis Kurosawa, 1985b: 147 (Nalanda) A: JA (Ryukyus)
santolinae Abeille de Perrin, 1894d: xxvi (Coraebus) E: FR PT SP N: MO
sculpticollis Abeille de Perrin, 1896b: 280 (Coraebus) E: BH BU CR GR TR A: TR
semenovi Obenberger, 1931e: 218 A: SD ORR
semenoviellus Obenberger, 1929e: 62 A: SCH
shimomurai Ohmomo & Akiyama, 1989: 18 (Nalanda) A: TAI
sikkimensis Obenberger, 1922b: 146 A: SD
sinae Obenberger, 1935b: 17 A: YUN
sinaiticus Théry, 1935b: 149 A: IS SI
solinghoanus Obenberger, 1935b: 16 A: YUN
subulatus Morawitz, 1861b: 284 (Coraebus) E: AR BU CT HU RO SK ST UK A: TR
eriwanensis Obenberger, 1935b: 18
graminioides Abeille de Perrin, 1896b: 281 (Coraebus)
hungaricus Kaszab, 1940: 112

suzukii Ohmomo & Akiyama, 1989: 10 (Nalanda) A: TAI
takashii Ohmomo & Akiyama, 1989: 9 (Nalanda) A: TAI
theryi Abeille de Perrin, 1893: 135 (Coraebus) N: AG TU
toyamai Ohmomo & Akiyama, 1989: 17 (Nalanda) A: TAI
transverserugatus Obenberger, 1935b: 16 A: SCH

565

566

BUPRESTIDAE

vitalisi Bourgoin, 1924c: 214 A: YUN ORR
hauserellus Obenberger, 1935b: 17

wenigi Obenberger, 1927f: 18 A: FUJ SC ZHE
fokienicus Obenberger, 1944c: 59

yunnanus Kerremans, 1895: 217 A: YUN
genus Meliboeus, nomen dubium
tribulis Faldermann, 1835a: 157 (Agrilus) E: “Transcaucasus”

subtribe Toxoscelina Majer, 2000

genus Cryptodactylus Deyrolle, 1864a: 130 type species Cryptodactylus lugubris Deyrolle, 1864
cyaneoniger Kerremans, 1892a: 207 A: NP SD ORR
laosensis Baudon, 1961: 81 A: YUN ORR
scutellaris Kerremans, 1892a: 209 A: NP ORR
vavrai Obenberger, 1917g: 88

genus Dessumia Descarpentries & Villiers, 1966a: 88 type species Discoderes vitalisi Bourgoin,
1922
vitalisi Bourgoin, 1922a: 22 (Discoderes) A: YUN ORR
genus Discoderoides Théry, 1936a: 12 type species Discoderoides priesneri Théry, 1936
Acanthocephalon Théry, 1955: 397 type species Acanthocephalon vansoni Théry, 1955 (=Amorphosoma immunitum
Fåhraeus, 1851)

priesneri Théry, 1936a: 13 N: EG
genus Indiadactylus Bellamy, 1992: 5 type species Indiadactylus fasciatus Bellamy, 1992 (=Evagora pulchella
Gory & Laporte, 1839)
chinensis Obenberger, 1924a: 105 (Cryptodactylus) A: HKG ORR
theryi Descarpentries & Villiers, 1966b: 992 (Cryptodactylus)

indicus Obenberger, 1924a: 104 (Cryptodactylus) A: UP ORR
nigricans Kerremans, 1890c: ccvi (Cryptodactylus) A: SD ORR
genus Neotoxoscelus Fisher, 1921: 418 type species Neotoxoscelus bakeri Fisher, 1921
Metatoxoscelus Hattori, 1990: 221 type species Metatoxoscelus kurosawai Hattori, 1990
Paracryptodactylus Alexeev, 1975: 148 type species Cryptodactylus kerzhneri Alexeev, 1975

kerzhneri Alexeev, 1975: 149 (Cryptodactylus) A: LIA MG SHG SHX
manchuricus Kurosawa, 1977: 172 (Cryptodactylus)
shirahatai Kurosawa, 1977: 171 (Cryptodactylus)

kurosawai Hattori, 1990: 223 (Metatoxoscelus) A: TAI
genus Polyonychus Chevrolat, 1838b: 102 type species Polyonychus mucidus Chevrolat, 1838

567

BUPRESTIDAE
mucidus Chevrolat, 1838b: 102 A: KA ORR
smei Gory & Laporte, 1839: 3 (Coraebus)

tricolor E. Saunders, 1866: 310 A: YUN ORR
genus Toxoscelus Deyrolle, 1864a: 127 type species Toxoscelus undatus Deyrolle, 1864
akiyamai Kurosawa, 1977: 178 A: JA (Ryukyus)
amamiensis Kurosawa, 1963b: 151 A: JA (Ryukyus: Yaku-shima)
angustimaculatus Hattori, 2014: 16 A: TAI
auriceps auriceps E. Saunders, 1873: 513 (Cryptodactylus) A: JA “Korea”
gracilis Schönfeldt, 1888b: 209 (Cryptodactylus)
japonicus Obenberger, 1934n: 151

auriceps tokarensis Kurosawa, 1977: 182 A: JA (Ryukyus)
carbonarius Obenberger, 1958a: 232 A: SCH YUN
gracilitibialis Hattori, 2014: 18 A: TAI
inadai Hattori, 2014: 2, 4 A: JA (Ryukyus)
mandarinus Obenberger, 1917e: 52 (Cryptodactylus) A: SHN ZHE
matobai Tôyama, 1985: 19 A: JA SC
miwai miwai Kurosawa, 1977: 174 A: JA (Ryukyus)
miwai makiharai Kurosawa, 1977: 176 A: JA (Ryukyus)
nakajimai Ohmomo, 2002: 156 A: JA (Ryukyus)
nigrogracilis Hattori, 2014: 11 A: TAI
purpureihumeralis Hattori, 2014: 9 A: TAI
reticulimaculatus Hattori, 2014: 14 A: TAI
sasakii Kurosawa, 1957: 183 A: JA
similis Gebhardt, 1929a: 28 A: JIA ZHE
sterbai Obenberger, 1934n: 151 A: YUN
sugiurai Hattori, 2014: 8 A: TAI
yakushimensis gressitti Kurosawa, 1977: 178 A: JA (Ryukyus)
yakushimensis hatayamai Kurosawa, 1977: 179 A: JA
yakushimensis yakushimensis Kurosawa, 1957: 184 A: JA (Ryukyus: Yaku-shima)
yokoyamai Kurosawa, 1977: 184 A: TAI

tribe Tracheini Laporte, 1835
subtribe Pachyschelina Böving & Craighead, 1931

genus Pachyschelus Solier, 1833: 313 type species Pachyschelus scutellatus Solier, 1833 (=Trachys terminans
Fabricius, 1801)
Metonius Say, 1833: 68 type species Metonius purpureus Say, 1833

bedeli Obenberger, 1921b: 237 A: GUI GUX TAI ORR
brancuccii Bílý, 1983b: 227 A: NP SD
mandarinus Théry, 1936f: 346 A: NP YUN ORR
similis Bílý, 1983b: 229

568

BUPRESTIDAE

sinicus Obenberger, 1940k: 188 A: FUJ ZHE
tonkinensis Théry, 1936f: 346 A: YUN ORR

subtribe Tracheina Laporte, 1835

genus Habroloma C.G. Thomson, 1864: 42 type species Buprestis nana Paykull, 1799
subgenus Habroloma C.G. Thomson, 1864: 42 type species Buprestis nana Paykull, 1799
aureum Semenov, 1890b: 212 (Trachys) E: AB AR A: IN KI KZ TD TM TR
reticulatum Abeille de Perrin, 1900a: 15 (Trachys)

bifrons Kiesenwetter, 1879b: 256 (Trachys) A: FE JA SC
amurense Obenberger, 1922a: 69 (Trachys)

breiti Obenberger, 1918a: 29 (Trachys) E: ST UK A: ES KI KZ WS
benedictum Roubal, 1920: 215 (Trachys)
lukianovici Obenberger, 1927f: 20 (Trachys)

mongolicum Cobos, 1968c: 408 A: ES MG
nanum Paykull, 1799: 233 [ICZN 1985: 180] (Buprestis) E: AU BE BH BU CR CT CZ DE EN FI FR GE GR HU IT
LA LT NL NR NT PL PT RO SB SK SL SP SV SZ UK
geranii Silfverberg, 1977: 93 (Trachys) [RN]

rungsi Baudon, 1959b: 213 N: MO
subalutaceum Pic, 1921: 3 (Trachys) N: AG EG
theryi Rungs, 1936: 297 N: MO
triangulare Lacordaire, 1835: 616 (Trachys) E: FR IT PT SP N: AG LB MO TU
pandellei Fairmaire, 1852: 79 (Trachys)
tournieri Pic, 1894a: 71 (Trachys)

tyli Obenberger, 1930a: 113 (Trachys) A: KI
viridipenne Théry, 1940a: 61 A: KZ UZ
subgenus Parahabroloma Kurosawa, 1959: 248 type species Trachys eximia Lewis, 1893
aeneocupreum Kerremans, 1890c: ccviii (Trachys) A: “Bengal” ORR
alphaxia Obenberger, 1929a: 64 (Trachys) A: TAI
anchiale Obenberger, 1929a: 57 (Trachys) A: FUJ JIX SD “Assam”
antonia Obenberger, 1929a: 58 (Trachys) A: “India or.”
asahinai Kurosawa, 1959: 257 A: JA (Ryukyus) TAI
astraea Obenberger, 1929a: 19 [1929c: 116] (Trachys) A: FUJ JIX
atronitidum Gebhardt, 1929a: 98 (Trachys) A: JA YUN ZHE
borchsenii Alexeev, 1989: 477 A: NC SHN
brachycephalum Gebhardt, 1929b: 103 (Trachys) A: ZHE
canace Obenberger, 1929a: 68 (Trachys) A: “Assam”
cheni Théry, 1935a: 133 A: JIX
clymene Obenberger, 1929a: 53 (Trachys) A: “Assam”
elissa Obenberger, 1929a: 62 (Trachys) A: YUN ORR
eximium eupoeta Obenberger, 1929a: 61 (Trachys) A: FUJ HUN JA (Ryukyus) TAI

BUPRESTIDAE
eximium eximium Lewis, 1893: 337 (Trachys) A: JA
euchariessum Gebhardt, 1929b: 100 (Trachys) A: ZHE
griseonigrum E. Saunders, 1873: 520 (Trachys) A: JA TAI “China”
chionochaetum Obenberger, 1929a: 54 (Trachys)

hikosanense Kurosawa, 1959: 255 A: JA (Kyushu)
hoscheki Obenberger, 1917n: 217 A: SHN
integrum Kerremans, 1892b: 221 (Trachys) A: “Himalaya”
konbirense Kerremans, 1900b: 92 (Trachys) A: “Bengal” ORR
lewisii E. Saunders, 1873: 519 (Trachys) A: FUJ HAI HUN JA SCH YUN “Korea”
liukiuense Obenberger, 1940g: 39 (Trachys) A: JA (Ryukyus) TAI
marginicolle Fairmaire, 1888a: 24 (Trachys) A: JA “Central China” “Korea”
kagosimanum Obenberger, 1940g: 39 (Trachys)
shirozui Kurosawa, 1959: 256

minotaurum Hołyński, 2003: 25 (Trachys) A: YUN
miwai Obenberger, 1929a: 65 (Trachys) A: TAI
nixilla insulicola Kurosawa, 1959: 260 A: JA (Ryukyus)
nixilla nixilla Obenberger, 1929a: 67 (Trachys) A: TAI
sinna Obenberger, 1937b: 35 (Trachys)

orestea Obenberger, 1929a: 65 (Trachys) A: “Assam”
praxilla Obenberger, 1929a: 63 (Trachys) A: TAI
protracticolle Obenberger, 1918a: 32 (Trachys) A: “Bengal” ORR
septimia Obenberger, 1929a: 57 (Trachys) A: YUN ORR
sinnicum Obenberger, 1929c: 124 (Trachys) A: HKG
subbicorne Motschulsky, 1860a: 8 (Brachys) A: HUB JA (Ryukyus) JIX SCH YUN “Korea”
elegantulum E. Saunders, 1873: 520 (Trachys)
formaneki Obenberger, 1930a: 112 (Trachys)
ronino Obenberger, 1918a: 63 (Trachys)
suensoni Gebhardt, 1929b: 102 (Trachys)

tecmessa Obenberger, 1929a: 54 (Trachys) A: SD ORR
wagneri Gebhardt, 1929a: 97 (Trachys) A: JIA YUN
yuasai Kurosawa, 1976b: 130 A: JA
genus Trachys Fabricius, 1801b: 218 type species Buprestis minuta Linnaeus, 1758
Phytotera Gistel, 1856: 366 [SN] type species Buprestis minuta Linnaeus, 1758

abeillei Obenberger, 1940g: 43 A: FUJ
aeneiceps Obenberger, 1929d: 12 A: YUN
ahenatus Mulsant & Rey, 1863: 26 E: BU RO UK (Krym)
alpinus Obenberger, 1918a: 60 E: AU (Kärnten)
althaea Obenberger, 1929a: 84 A: SD
alticola Cobos, 1966c: 249 A: NP

569

570

BUPRESTIDAE

amplus Roubal, 1920: 214 E: ST
andrewesi Kerremans, 1893b: 353 A: “Bengal” ORR
anthrenoides J. Thomson, 1879a: 77 A: “India”
antistes Obenberger, 1937b: 37 A: YE AFR
araxicola Obenberger, 1918a: 58 E: “Araxestal” [in AB or AR]
arhema Obenberger, 1929a: 77 A: YUN ORR
asiaticus Kerremans, 1900b: 87 A: “Kashmir”
astarte Obenberger, 1929a: 91 A: “Bengal”
atrus Kerremans, 1893b: 354 A: “Bengal” ORR
auricollis E. Saunders, 1873: 519 A: FUJ JA TAI ORR
sauteri Kerremans, 1912c: 209

aurifluus Solsky, 1876b: 280 A: FE HEI JA NC
freyi Théry, 1942c: 272
orichalceus Kiesenwetter, 1879b: 255 (Brachys)
tiliae Kurosawa, 1959: 245

barnevillei Tournier, 1868b: 145 N: AG
bedfordi Théry, 1929b: 143 N: EG A: YE AFR
bodenheimeri Théry, 1934a: 85 A: IS
broussonetiae Kurosawa, 1985b: 167 A: JA “China”
chinensis Kerremans, 1898a: 191 A: HKG
commixtus J. Thomson, 1879a: 76 A: UP
compressus Abeille de Perrin, 1891: 286 E: CZ FR GB GE PT SK SP SZ
cuneiferus Kurosawa, 1959: 215 A: JA
cupricolor E. Saunders, 1873: 521 A: JA TAI
curia Obenberger, 1929a: 81 A: “Bengal”
danae Obenberger, 1929a: 79 A: “Bengal”
davidis Fairmaire, 1888a: 23 A: JIX
dichrous Obenberger, 1918a: 51 E: IT (Sicilia) N: AG MO TU
tricuspidatus Théry, 1919: 235

dilaticeps Gebhardt, 1929a: 103 A: FUJ JA TAI ZHE ORR
mixtipilis Obenberger, 1929a: 94
robustissimus Kurosawa, 1959: 241

duplofasciatus Gebhardt, 1929a: 101 A: ZHE
elvira Obenberger, 1929a: 96 A: TAI
erythreae Obenberger, 1937d: 93 A: AE
eurynome Obenberger, 1929a: 99 A: “Assam”
excavatus Kerremans, 1893b: 355 A: “Bengal” ORR
fallax Kerremans, 1893b: 350 A: “Bengal” ORR
flaviceps Kerremans, 1892a: 223 A: TAI “India”
formosanus Kerremans, 1912c: 209 A: TAI ORR
fragariae Brisout de Barneville, 1874: lxx E: AU BH BU CT CZ FR GE GR HU IT PL PT RO SK SP ST SZ UK

BUPRESTIDAE
laevipennis Obenberger, 1918a: 62

furia Obenberger, 1929a: 86 A: “India or.”
goberti Gozis, 1889: 204 E: FR SP
clermonti Roger, 1900: 32
coloreus Abeille de Perrin, 1900a: 14

griseofasciatus E. Saunders, 1873: 521 A: JA “China” “Korea”
helena Obenberger, 1929a: 97 A: “Assam”
hera Obenberger, 1929a: 76 A: “Assam”
hermione Obenberger, 1929a: 90 A: “India or.”
hipponensis Marseul, 1866: 513 N: AG TU
hornianus Obenberger, 1918b: 25 [RN] A: TAI
horni Kerremans, 1913b: 116 [HN]

houskai Obenberger, 1946: 37 A: IS
ida Obenberger, 1929a: 91 A: PA “India”
impressus Boheman, 1858: 65 A: HKG
inconspicuus E. Saunders, 1873: 522 A: FUJ HUN JA SC TAI
galloisi Obenberger, 1940g: 41
ogumensis Obenberger, 1924b: 57

ine Obenberger, 1918a: 42 A: “Bengal” ORR
ineditus E. Saunders, 1873: 522 A: HUN JA
oviformis Kurosawa, 1959: 229

isis Obenberger, 1937c: 45 N: EG
jakobsoni Obenberger, 1929c: 125 A: HKG
janthe Obenberger, 1929a: 75 A: “India or.”
jo Obenberger, 1918a: 41 A: “Bengal” ORR
juno Obenberger, 1918a: 42 A: “Bengal” ORR
klapaleki Obenberger, 1930a: 114 A: “China or.”
kocheri Baudon, 1958a: 88 N: MO
koenigi Reitter, 1890e: 146 E: ST
utschderensis Obenberger, 1918a: 56

koshunensis Obenberger, 1940g: 43 A: SCH TAI
kurosawai Bellamy, 2003: 157 [RN] A: TAI
mirabilis Kurosawa, 1954b: 82 [HN]

latifrons Kerremans, 1907a: 104 A: AE SA YE AFR
libanonicus Obenberger, 1937c: 44 A: LE
lichtensteini Buysson, 1918: 30 E: BU CR FR GR IT SP
maculatus Kerremans, 1894b: 478 A: “Bengal” ORR
maia Obenberger, 1929a: 100 A: “Assam”
mandarinus Obenberger, 1917n: 217 A: SCH SHN
mansuetus Kerremans, 1894b: 479 A: “Bengal” ORR
menthae Bedel, 1921: 225 E: BE CR FR IT NL PT SL SP SZ

571

572

BUPRESTIDAE

subacuminatus Pic, 1922c: 29

minimus Wiedemann, 1823: 104 (Buprestis) A: “Bengal”
minutus minutus Linnaeus, 1758: 410 (Buprestis) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE
GG GR HU IR IT LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK A: ES FE IN
MG NE NO SY TR WS NARi
mandjuricus Obenberger, 1917n: 218
reflexiformis Obenberger, 1918a: 46

minutus salicis Lewis, 1893: 337 (Brachys) A: JA TAI
nemethi Théry, 1930a: 536 N: MO
nitidus Kerremans, 1893b: 351 A: “Bengal” ORR
nodulipennis Obenberger, 1940g: 40 A: YUN
nudus Abeille de Perrin, 1894g: ccclv [=1894a: 48] A: TR
obesulus Obenberger, 1918a: 44 A: “Bengal” ORR
obliquus Kerremans, 1892b: 223 A: “Bengal” ORR
ohbayashii Kurosawa, 1954b: 83 A: TAI
olmyra Obenberger, 1929a: 86 A: “Assam”
opulentus Abeille de Perrin, 1894g: ccclv [=1894a: 48] A: KZ UZ
orientalis J. Thomson, 1879a: 77 A: “India”
pacificus Kerremans, 1894b: 478 A: “Bengal” ORR
parvulus Kerremans, 1893b: 352 A: “Bengal” ORR
pecirkai Obenberger, 1925a: 100 A: ES FE JA ZHE “Korea”
amuricola Obenberger, 1929d: 11
semenovi Obenberger, 1929c: 126

perparvus Obenberger, 1918a: 47 N: MO
phlyctaenoides Kolenati, 1846: 35 E: AB AR ST UK A: IN TM TR
iranicus Obenberger, 1937c: 43
phlyctaenodes Kolenati, 1846: 35 [unused original spelling]
transcaucasicus Obenberger, 1924b: 57

pilosulus Kerremans, 1893b: 354 A: “Bengal” ORR
plebejus Kerremans, 1894b: 476 A: “Bengal” ORR
polyhymnius Obenberger, 1929a: 85 A: PA “India or.”
problematicus Obenberger, 1918a: 60 E: AB AL AR AU BH BU CR CT CZ DE GE GR HU IT MC MD PL RO SB SK
SL ST TR UK
proserpina Obenberger, 1929a: 85 A: “India or.”
pseudoscrobiculatus pseudoscrobiculatus Obenberger, 1940g: 41 A: JA
pseudoscrobiculatus shirozui Kurosawa, 1959: 209 A: FUJ HUN JA
pumilus Illiger, 1803: 257 E: GR PT SP A: IS LE SY TR
creticus Obenberger, 1918a: 60
intermedius Gory & Laporte, 1840: 7
major Perris, 1864: 284
tristis Abeille de Perrin, 1900a: 14 [HN]

BUPRESTIDAE

573

puncticollis puncticollis Abeille de Perrin, 1900a: 14 E: AB AR ST A: IN TR
puncticollis rectilineatus Abeille de Perrin, 1900a: 14 E: AU BH BU CR CZ GE GR HU IT MC RO SK SL ST UK
obscurellus Obenberger, 1918a: 49

quercicola Marseul, 1871: 80 [=1872: 414] E: FR IT PT SP SZ
marseulii Brisout de Barneville, 1874a: lxx
subglaber Rey, 1891c: 27

reflexus Gené, 1839: 54 E: IT (Sardegna) PT SP N: AG MO TU
reitteri Obenberger, 1930a: 114 A: FE JA SC SHT TAI
falcatae Kurosawa, 1959: 223

robustus E. Saunders, 1873: 521 A: JA
rotundatus Kerremans, 1893b: 352 A: “Bengal” ORR
rufopubens Fairmaire, 1888a: 24 A: JIX
saundersi Lewis, 1893: 337 A: FUJ HUN JA SCH TAI YUN
jakovlevi Obenberger, 1929c: 126
mariola Obenberger, 1929a: 74
obscuripennis Obenberger, 1923c: 65
opsigonus Obenberger, 1937c: 43
vimmeri Obenberger, 1930a: 113
yunnanus Obenberger, 1929d: 11

scrobiculatus Kiesenwetter, 1857: 171 E: AU CT CZ DE FR GB GE GR HU NL PL SK SL ST SZ UK
convergens Abeille de Perrin, 1900a: 14

splendidulus Reitter, 1890e: 146 E: AB AR ST A: IN TR
cavatus Abeille de Perrin, 1895b: xxiii
ebenipterus Obenberger, 1918a: 52
morgani Théry, 1925b: 176

serapis Obenberger, 1929a: 86 A: “Assam”
simulans Kerremans, 1893b: 349 A: “Bengal” ORR
sordidulus Obenberger, 1918a: 39 A: “Bengal” ORR
sororculus Obenberger, 1929d: 10 A: HKG
sussamyrensis Obenberger, 1937c: 44 A: KI
suspectatrix Obenberger, 1918a: 43 A: “Bengal” ORR
taiwanensis Obenberger, 1929d: 9 A: TAI
tokyoensis Obenberger, 1940g: 42 A: JA
toringoi Kurosawa, 1951: 73 A: HUN JA
transversus Kerremans, 1892b: 224 A: “Bengal” ORR
tristiculus Obenberger, 1918a: 39 A: “Bengal” ORR
tristis Fairmaire, 1888a: 23 A: “China bor.”
troglodytes Gyllenhal, 1817: 125 E: AL AU BE BH BU CR CT CZ DE EN FR GB GE GR HU IT LA LT LU MD NL NT
PL RO SB SK SL ST SV SZ UK
aeneus Mannerheim, 1837: 122
arbustorus Gistel, 1857a: 25

574

BYRRHIDAE

puncticeps Abeille de Perrin, 1900a: 13
vehemens Gistel, 1857a: 43

troglodytiformis Obenberger, 1918a: 54 E: AL AU BE BH BU CR CZ FR GE GR HU IT MA NL PT RO SB SK SL SP
ST TR UK N: AG LB MO TU A: TR NARi
coruscus Ponza, 1805: 81 (Buprestis) [HN]
fabricii Schaefer, 1949b: 472 [RN]
indigopterus Obenberger, 1918a: 54
pygmaeus Fabricius, 1787a: 183 (Buprestis) [HN]
theresae Pic, 1922c: 29

tsushimae Obenberger, 1922a: 54 A: HUN JA SHN TAI
japonicus Obenberger, 1923c: 65
ornatus Théry, 1934a: 86

turanicus Semenov, 1893c: 497 A: AF KZ TD UZ
bactrianus Semenov, 1896d: 351
lopatini Stepanov, 1958a: 116

variolaris E. Saunders, 1873: 521 A: FUJ HEN HUN JA SHN TAI YUN “Korea”
clavicornis Obenberger, 1919b: 144

vaulogeri Abeille de Perrin, 1900a: 15 N: AG
vavrai Obenberger, 1918a: 37 A: YUN ORR
hauseri Obenberger, 1937c: 42

vicarians Obenberger, 1918a: 38 A: “Bengal” ORR
convolvuli Descarpentries, 1958a: 69
laodamea Obenberger, 1929a: 102

violae Kurosawa, 1959: 209 A: JA
yanoi Kurosawa, 1959: 233 A: JA “Northern China” “Korea”
yoshidai Kurosawa, 1959: 222 A: JA
genus Trachys, nomen dubium
viridis Samouelle, 1819b: 42 E: GB
family Buprestidae, nomina dubia
Buprestis dubia Piller & Mitterpacher, 1783: 38, 113 E: SL
Buprestis duodevigintipunctata Piller & Mitterpacher, 1783: 38 E: SL
Cucujus dentatus Geoffroy, 1785: 34

superfamily BYRRHOIDEA Latreille, 1804
family BYRRHIDAE Latreille, 1804
subfamily Byrrhinae Latreille, 1804
tribe Simplocariini Mulsant & Rey, 1869

genus Chrysosimplocaria Paulus, 1982: 319 type species Chrysosimplocaria nepalensis Paulus, 1982
ahrensi Jäger, 1998: 258 A: NP

BYRRHIDAE
alticola Fabbri, 2005: 14 A: NP
arborea Jäger, 1998: 260 A: NP
bowaki Jäger, 1998: 262 A: NP
dobbertini Jäger, 1998: 263 A: NP
fiorii Pütz & Fabbri, 1997: 300 A: NP
ganeshi Jäger, 1998: 259 A: NP
jaegeri Pütz, 2000b: 181 A: NP
lamjungana Jäger, 1997: 62 A: NP
nepalensis Paulus, 1982: 320 A: NP
nivalis Jäger, 1997: 62 A: NP
philippi Jäger, 2000: 373 A: NP
pisangana Jäger, 1998: 261 A: NP
schawalleri Pütz & Fabbri, 1997: 304 A: NP
schmidti Jäger, 1997: 59 A: NP
silvestris Jäger, 2000: 374 A: NP
varia Jäger, 1997: 64 A: NP
genus Himalayoligus Fabbri, 2002a: 50 type species Himalayoligus daccordii Fabbri, 2002
ahrensi Jäger & Pütz, 2003: 224 A: NP
daccordii Fabbri, 2002a: 52 A: NP
franzi Jäger & Pütz, 2003: 223 A: NP
gunnari Jäger & Pütz, 2003: 227 A: NP
hartmanni Jäger & Pütz, 2003: 224 A: NP
letelaensis Jäger & Pütz, 2003: 228 A: NP
nepalensis Jäger & Pütz, 2003: 226 A: NP
notabilis Jäger & Pütz, 2003: 226 A: NP
similis Jäger & Pütz, 2003: 228 A: NP
telburgensis Jäger & Pütz, 2003: 225 A: NP
genus Horiella Takizawa, 1983: 7 type species Simplocaria shikokensis Takizawa, 1983
aikoae Kitano & Matsumoto, 2008: 49 A: JA
bicornis Kitano & Sakai, 2007: 253 A: JA
franzi Pütz, 2003a: 93 A: JA
hisamatsui Kitano & Sakai, 2009: 238 A: JA
japonica Pütz, 2002c: 186 A: JA
kannoi Kitano & Sakai, 2009: 242 A: JA
loebli Pütz, 2002c: 187 A: JA
maruyamai Pütz, 2003a: 95 A: JA
matsunoi Kitano & Sakai, 2009: 239 A: JA
shikokensis Takizawa, 1983: 8 A: JA
shodoshimaensis Pütz, 2002c: 188 A: JA

575

576
yoshidai Kitano & Sakai, 2009: 240 A: JA
genus Shanna Pütz, 2007a: 21 type species Shanna wuyishanensis Pütz, 2007
wuyishanensis Pütz, 2007a: 23 A: JIX
genus Lasiomorychus Ganglbauer, 1902b: 93 type species Pedilophorus apfelbecki Reitter, 1889
apfelbecki Reitter, 1889f: 279 (Pedilophorus) E: BH BU GR
genus Simplocaria Stephens, 1829b: 9 type species Byrrhus semistriatus Fabricius, 1794
subgenus Simplocaria Stephens, 1829b: 9 type species Byrrhus semistriatus Fabricius, 1794
acuminata Erichson, 1847a: 496 E: AU CR IT PL RO SK SL SZ UK
arctica Poppius, 1904a: 7 E: NT
atayal Pütz, 2003b: 238 A: TAI
basalis J.R. Sahlberg, 1903b: 22 A: ES
basistriata Nakane, 1964a: 29 A: JA
brevistriata Reitter, 1900c: 83 E: SP N: AG MO
battandieri Peyerimhoff, 1918: 371

bunun Pütz, 2003b: 239 A: TAI
deubeli Ganglbauer, 1899: 534 E: RO UK
elongata J.R. Sahlberg, 1903b: 23 A: ES NAR
erlangshanensis Pütz, 2007a: 14 A: SCH
frigida Krogerus, 1921: 12 E: FI
glacialica Jäger & Pütz, 2003: 220 A: NP
hakoensis Takizawa, 1983: 5 A: JA
hispidula Fairmaire, 1886: 319 A: FUJ GUX HUN JA JIX SCH SHG ZHE
apicalis Pic, 1935: 3
bicolor Pic, 1935: 3

indica Pütz, 2002e: 325 A: UP
ivani Pütz, 2003b: 241 A: TAI
ivanloebli Pütz, 2002d: 252 A: JA
jaechi Jäger & Pütz, 2003: 221 A: NP
jaisulica Jäger & Pütz, 2003: 219 A: NP
jugicola Baudi di Selve, 1889: 151 E: IT
kamtshatensis Lafer, 1989: 456 A: FE
kurilensis Lafer, 1989: 456 A: FE
limbuorum Jäger & Pütz, 2003: 217 A: NP
longistriata G.C. Champion, 1923b: 277 A: UP
macularis Reitter, 1896h: 285 A: ES
maculosa Erichson, 1847a: 495 E: AU CZ GB GE PL RO ST SZ UK A: ES
bernhaueri Ganglbauer, 1899: 533

maritima Lafer, 1989: 456 A: FE

BYRRHIDAE

BYRRHIDAE

577

metallica Sturm, 1807: 111 (Byrrhus) E: AU BU CZ FI GE IT NR NT PL RO SV SZ NAR
picipes Gyllenhal, 1808: 200 [HN]

meybohmi Pütz, 2009d: 165 A: TR
munetoshii Pütz, 2002d: 254 A: JA
naganoensis Pütz, 2004a: 39 A: JA
nebulosa Poppius, 1904a: 9 A: ES
nenkaoshan Pütz, 2003b: 242 A: TAI
nivalis Ganglbauer, 1904b: 198 E: IT
obscuripes Poppius, 1904a: 5 A: ES
oharai Pütz, 2002d: 256 A: JA
paiwan Pütz, 2003b: 243 A: TAI
palmeni Poppius, 1904a: 6 E: NT
qinlingensis Pütz, 2007a: 16 A: SHA
raiorum Jäger & Pütz, 2003: 218 A: NP
rukai Pütz, 2003b: 244 A: TAI
saysiat Pütz, 2003b: 245 A: TAI
schillhammeri Pütz, 2007a: 13 A: HUN
schmidti Jäger & Pütz, 2003: 221 A: NP
schuelkei Pütz, 2007a: 17 A: SHA
semistriata Fabricius, 1794: 437 (Byrrhus) E: AB AR AU BE BH BU BY CZ DE EN FI FR GB GE GG
GR HU IR IT LA LS LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: AG A:
FE NAR
minuta Thunberg, 1794: 68 (Byrrhus)
picea Marsham, 1802: 106 (Cistela)
picipes A.G. Olivier, 1790a: no 13: 9 (Byrrhus)
rufipes Kugelann, 1792b: 485 (Byrrhus)

sherpaorum Jäger & Pütz, 2003: 218 A: NP
sichuanensis Pütz, 2007a: 18 A: SCH
smetanai Pütz, 2003b: 246 A: TAI
striata Brisout de Barneville, 1866: 372 E: FR PT SP
taiwanica Pütz, 2003b: 247 A: TAI
taroko Pütz, 2003b: 248 A: TAI
tsou Pütz, 2003b: 249 A: TAI
wolongensis Pütz, 2007a: 15 A: SCH
xueshanensis Pütz, 2007a: 20 A: YUN
yulongshanensis Pütz, 2007a: 19 A: YUN
subgenus Trirania Mroczkowski, 1983: 178 [RN] type species Simplocaria carpathica Hampe, 1853
Trinaria Mulsant & Rey, 1869a: 158 [HN] type species Simplocaria carpathica Hampe, 1852

carpathica Hampe, 1853: 223 E: AU BH BU CR IT SL RO
montenegrina Obenberger, 1917a: 30 E: YU

578

BYRRHIDAE

genus Trichobyrrhulus Ganglbauer, 1902b: 94 type species Pedilophorus variolosus Perris, 1864
algiricus Chobaut, 1898: 75 (Pedilophorus) N: AG
arragonicus Reitter, 1911a: 159 E: SP
assingi Pütz, 2004b: 386 E: SP
fabbrii Allemand, 1998: 187 E: SP
franzi Pütz, 2004b: 390 E: SP
meybohmi Pütz, 2009c: 103 E: SP
nevadensis Pütz, 2004b: 391 E: SP
piochardi Heyden, 1870a: 109 (Morychus) E: FR PT SP
puetzi Allemand, 1998: 190 E: SP
rondaensis Pütz, 2004b: 392 E: SP
stierlini Gozis, 1882: 193 (Pedilophorus) E: FR SP
variolosus championi Reitter, 1905a: 90 E: SP
variolosus variolosus Perris, 1864: 281 (Morychus) E: SP
championi Reitter, 1905a: 90
rufipes Mulsant & Rey, 1869a: 150 (Pedilophorus)

wunderlei Pütz, 2004b: 394 E: SP
zercheorum Pütz, 2002a: 48 E: SP

tribe Exomellini Casey, 1914

genus Exomella Casey, 1914: 378 [RN] type species Exoma pleuralis Casey, 1908
Exoma Casey, 1908: 282 [HN] type species Exoma pleuralis Casey, 1908

sikhotealinensis Pütz & Lafer, 1995: 341 A: FE

tribe Morychini El Moursy, 1961

genus Byrrhochomus Fabbri, 2003b: 54 type species Byrrhochomus ameliae Fabbri, 2003
ameliae Fabbri, 2003b: 55 A: SCH
crassicornis Fabbri, 2003b: 59 A: SCH
rugifrons Fabbri, 2003b: 60

longiusculus Fabbri, 2003b: 58 A: SCH
pinguiculus Fabbri, 2003b: 57 A: SCH
genus Morphobyrrhulus Pütz, 2007a: 26 type species Byrrhochomus shaanxianus Fabbri, 2003
shaanxianus Fabbri, 2003b: 73 (Byrrhochomus) A: SHA
genus Morychus Erichson, 1847a: 491 type species Byrrhus aeneus Fabricius, 1775
Byrrhobolus G. Fiori, 1965: 215 type species Pedilophorus rutilans Motschulsky, 1845
Chlorobyrrhulus Fabbri, 2002b: 108 type species Chlorobyrrhulus sinicus Fabbri, 2002
Hypolorus Mulsant & Rey, 1869a: 143 type species Pedilophorus modestus Kiesenwetter, 1850
Steffahnyellus Portevin, 1931: 286 type species Pedilophorus modestus Kiesenwetter, 1850

579

BYRRHIDAE

aeneus Fabricius, 1775: 60 (Byrrhus) E: AU BE BH BU BY CZ DE EN FI FR GB GE HU IR IT LA LT NL NR NT PL
RO RU SK SL SV SZ UK YU A: ES FE KZ WS
modestus Kiesenwetter, 1850: 223 (Morychus)

biacuminatus Jäger & Pütz, 2009: 287 A: NP
constanzeae Jäger & Pütz, 2009: 286 A: NP
dudkorum Tshernyshev, 1997: 123 A: WS
nepalensis Paulus, 1982: 323 (Byrrhobolus) A: NP SD
ostasiaticus Tshernyshev, 1997: 118 A: WS
rolwalingensis Jäger & Pütz, 2009: 286 A: NP
rutilans Motschulsky, 1845a: 51 (Pedilophorus) A: ES FE
selaensis Pütz, 2009a: 289 A: AP
sinicus Fabbri, 2002b: 109 (Chlorobyrrhulus) A: SCH
subparallelus Motschulsky, 1859a: 231 A: ES
tibetanicus Tshernyshev, 2000: 134 A: QIN SCH
seiunctus Fabbri, 2003b: 53

viridis Kuzmina & Korotyaev, 1987: 344 A: FE
yamalus Tshernyshev, 1999: 18 A: ES
genus Sinorychomus Fabbri, 2003b: 74 type species Sinorychomus albovillosus Fabbri, 2003
albovillosus Fabbri, 2003b: 76 A: SCH
alesi Pütz, 2007a: 36 A: YUN
allemandi Fabbri, 2003b: 61 (Byrrhochomus) A: SCH
angelinii Fabbri, 2003b: 73 (Byrrhochomus) A: SCH
campadelli Fabbri, 2003b: 69 (Byrrhochomus) A: YUN
chromatus Fabbri, 2003b: 63 (Byrrhochomus) A: SCH
fulgidus Fabbri, 2003b: 61 (Byrrhochomus) A: SCH
hemisphaericus Fabbri, 2003b: 67 (Byrrhochomus) A: YUN
inauratus Fabbri, 2003b: 70 (Byrrhochomus) A: GAN
jindingshanus Fabbri, 2003b: 65 (Byrrhochomus) A: SCH
kalabi Pütz, 2007a: 35 A: SCH
mandibulatus Fabbri, 2003b: 71 (Byrrhochomus) A: GAN
panda Fabbri, 2003b: 66 (Byrrhochomus) A: YUN
venustus Fabbri, 2003b: 64 (Byrrhochomus) A: SCH
yunnanus Fabbri, 2003b: 69 (Byrrhochomus) A: YUN

tribe Pedilophorini Casey, 1912

genus Carpathobyrrhulus Ganglbauer, 1902b: 92 type species Pedilophorus transsylvanicus Suffrian, 1848
tatricus Mroczkowski, 1957: 216 E: PL SK
transsylvanicus Suffrian, 1848: 100 (Byrrhus) E: RO UK

580

BYRRHIDAE

genus Chrysobyrrhulus Reitter, 1911a: 159 type species Morychus metallicus Chevrolat, 1865
asturiensis Allemand, 1997a: 155 E: SP
metallicus Chevrolat, 1865: 350 (Morychus)

metallicus Chevrolat, 1865: 350 (Morychus) E: SP
moltonii moltonii G. Fiori, 1964: 199 E: SP
moltonii montanensis Allemand, 1997a: 154 E: SP
triangularis Allemand, 1997a: 156 E: SP
genus Lamprobyrrhulus Ganglbauer, 1902b: 94 type species Byrrhus nitidus Schaller, 1783
hayashii G. Fiori, 1967: 49 A: JA
aino Nakane, 1983b: 142

nitidus Schaller, 1783: 252 (Byrrhus) E: AR AU BE BH BU CR CZ DE EN FI FR GE HU IT LT MD NL NT PL RU
RO SK ST SZ UK YU A: ES FE NC WS
niger Kugelann, 1792b: 485 (Byrrhus)
nitens Panzer, 1795a: 4 (Byrrhus)
punctatus Germar, 1817: 187 (Byrrhus)

genus Pedilophorus Steffahny, 1843: 35 type species Byrrhus auratus Duftschmid, 1825
Hypolorus Mulsant & Rey, 1869a: 143 type species Pedilophorus modestus Kiesenwetter, 1850

auratus Duftschmid, 1825: 20 (Byrrhus) E: AU BH CR CZ GE HU IT MC ME PL SK SL
obscurus Fleischer, 1908: 212

coerulescens Fabbri & Allemand, 2003: 11 E: AL BH CR ME
colonnellii colonnellii Fabbri, 1999: 10 E: GR
pindicus Pütz, 2000a: 124

colonnellii schneideri Pütz, 2000a: 126 E: GR
inexspectatus Fabbri & Allemand, 2003: 20 E: BH
macedonicus Schubert, 1969: 74 (Carpathobyrrhulus) E: AL BH MC
zelenosivus Fabbri & Allemand, 2002: 62

monicae amicorum Pütz, 2000a: 124 E: GR
monicae monicae Fabbri, 1999: 13 E: GR
graecus Pütz, 2000a: 122

moraveci Fabbri & Allemand, 2002: 64 E: AL MC ME
obscurus inflexus Fabbri & Allemand, 2003: 16 E: AL ME
obscurus obscurus Fabbri & Allemand, 2003: 14 E: AL BH CR HU ME RO SB
permixtus Fabbri & Allemand, 2003: 19 E: AL KO MC
rhodopensis Paulus, 1972b: 341 E: BH BU CR IT ME RO SB SL
sennii Fabbri & Allemand, 2003: 13 E: BH CR HU KO ME SL
shqiperianus Fabbri & Allemand, 2002: 68 E: AL
szeli Fabbri & Allemand, 2002: 65 E: AL MC
zierisi Fabbri & Allemand, 2002: 66 E: GR MC

581

BYRRHIDAE
tribe Byrrhini Latreille, 1804

genus Arctobyrrhus Munster, 1902: 90 type species Arctobyrrhus dovrensis Munster, 1902
dovrensis Munster, 1902: 91 E: FI NR NT A: ES WS
speciosus J.R. Sahlberg, 1903b: 20 (Pedilophorus)

genus Byrrhus Linnaeus, 1767: 568 type species Dermestes pilula Linnaeus, 1758
subgenus Aeneobyrrhus Pütz, 1998a: 31 type species Byrrhus bajankolensis Pütz, 1998
bajankolensis Pütz, 1998a: 32 A: KZ
bermani Korotyaev, 1990: 129 A: WS
mordkovitshi Tshernyshev & Dudko, 1997: 134 A: WS
aenescens Korotyaev, 1997: 170

subaeneus sayanicus Tshernyshev, 1999: 16 A: ES
subaeneus subaeneus Reitter, 1896d: 172 (Seminolus) A: ES
talgarensis Pütz, 1998a: 33 A: KZ
subgenus Asiatobyrrhus Paulus, 1971: 167 type species Byrrhus tibetanus Paulus, 1971
asiaticus Solsky, 1876a: 283 (Curimus) A: KI TD UZ
seravshanensis Pütz, 1999a: 226

breviuncinatus Fabbri & Zhou, 2003a: 60 A: GAN
busii Fabbri, 2000: 48 A: PA
dacatrai Fabbri, 2000: 45 A: PA
degiovannii Fabbri, 2000: 49 A: KA
desioi G. Fiori, 1957a: 28 (Seminolus) A: PA
fortii Fabbri, 2000: 48 A: PA
fulvosetosus Reitter, 1899c: 199 (Seminolus) A: TM
kaszabi G. Fiori & A. Fiori, 1986: 5 A: KA
kazakhstanus Fabbri, 2000: 45 A: KZ
linxiaensis Pütz, 2007a: 83 A: GAN
lajishanensis Pütz, 2007a: 80 A: QIN
macrosetosus Paulus, 1971: 168 A: PA
nodulosus G.C. Champion, 1923b: 220 A: HP
pakistanus Fabbri, 2000: 47 A: PA
punjabanus Fabbri, 2000: 51 A: PA
rugipennis Fabbri, 2005: 17 A: PA
singularis Pic, 1923c: 4 A: HP
smetanai Pütz, 2007a: 82 A: GAN
staveni Fabbri, 2000: 50 A: KA
subverrucosus G.C. Champion, 1923b: 219 A: HP
tagliaferrii Fabbri, 2000: 53 A: HP
tibetanus Paulus, 1971: 167 A: QIN
wittmeri Pütz, 1999a: 227 A: NP

582

BYRRHIDAE

zoiai Fabbri, 2000: 51 A: PA
subgenus Byrrhocaulus Fairmaire, 1901b: 266 type species Byrrhus allemandi Fabbri & Pütz, 1997
allemandi Fabbri & Pütz, 1997: 22 A: SD
inaequalis Fairmaire, 1901b: 266 [HN]

lisae Jäger, 2002: 322 A: XIZ
subgenus Byrrhus Linnaeus, 1767: type species Dermestes pilula Linnaeus, 1758
altaesayanus Tshernyshev & Dudko, 2000: 250 A: WS
arietinus arietinus Steffahny, 1842: 17 E: AB AL AR AU BE BH BU BY CR CZ DE EN FI FR GB GE GG GR HU IT
LA LS LT LU MC MD NL NR PL RO RU SK SL SP SV SZ UK YU A: CH ES FE IN KZ WS
cinctus Sturm, 1807: 98 [HN]
imafukui Nakane, 1963b: 44
montanus Czwalina, 1887: 495

arietinus jaegeri Pütz, 2007a: 45 A: XIZ
arietinus konoi Takizawa & Nakane, 1977: 59 A: FE
arietinus obscurus G. Fiori, 1948: 16 E: IT
arietinus safraneki Pütz, 2007a: 44 A: XIZ
arietinus zadoi Pütz, 2007a: 45 A: XIZ
bamda bamda Pütz, 2007a: 47 A: XIZ
bamda tryznai Pütz, 2007a: 48 A: XIZ
belloi Fabbri, 2001: 115 A: SCH
cavazzutii Fabbri, 2000: 35 A: SCH
cholashanensis Fabbri, 2000: 32 A: SCH
daxueshanensis Fabbri, 2000: 41 A: SCH
danieli Paulus, 1974: 217 E: AR
eniseyensis Tshernyshev & Dudko, 2000: 252 A: ES
ekashi Kôno, 1934b: 130 A: JA
fasciatus Forster, 1771: 12 (Cistela) E: AB AL AR AU BE BH BU BY CR CZ DE EN FA FI FR GB GE GG GR HU IC
IR IT LA LS LT LU MC ME NL NR PL PT RO RU SB SK SL SP SV SZ TR UK A: ES FE JA JIL KI KZ MG TR WS
XIN NAR
arcuatus Rey, 1889b: 66
auratofasciatus Duftschmid, 1825: 11
bellus Reitter, 1881a: 77
cinctus Kugelann, 1794: 520
cinctus Thunberg, 1794: 68
complicans Reitter, 1881a: 78
daisetsuzanus Kôno, 1934b: 129
decipiens Fairmaire, 1863: 74
dianae Kugelann, 1794: 520
dorsalis Kugelann, 1792a: 354 [HN]
fabricii Reitter, 1881a: 78

BYRRHIDAE
fusculus Reitter, 1881a: 78
inornatus Reitter, 1881a: 77
montivagus Grimmer, 1841: 39
nebulosus Motschulsky, 1859a: 230
nigrolineatus Nakane, 1963b: 44
nigrosignum Reitter, 1916: 221
niveus Reitter, 1881a: 78
reyi Plavilshchikov, 1924: 233
ruficornis J.R. Sahlberg, 1889: 122
shinanensis Nakane, 1963b: 44
sibiricus Motschulsky, 1859a: 230
stoicus O. Fabricius, 1780: 184
subornatus Reitter, 1881a: 77

franzi Pütz, 1998b: 208 A: ES
geminatus LeConte, 1854c: 114 E: IT NT SZ A: ES FE JA JIL KZ MG NC WS NAR
sachalinensis Matsumura, 1911: 117

hongzhangi Pütz, 2007a: 67 A: SCH
indicus Fairmaire, 1901b: 266 A: SD
insperatus Pütz, 2007a: 72 A: SCH
imstriatus Reitter, 1896d: 172 (Seminolus) A: KI
janschneideri janschneideri Fabbri, 2000: 37 A: SCH
janschneideri nangquensis Pütz, 2007a: 77 A: QIN
janschneideri sertarensis Pütz, 2007a: 77 A: SCH
katrinae Pütz, 2007a: 74 A: SCH
kozlovi kozlovi Tshernyshev, 2000: 134 A: QIN
kozlovi manzhangensis Pütz, 2007a: 59 A: QIN
kozlovi maquensis Pütz, 2007a: 60 A: GAN
kozlovi zekogensis Pütz, 2007a: 57 A: QIN
kusnetzovi Pütz, 1998b: 206 A: FE
letiziae Fabbri, 2000: 35 A: QIN
litangensis Fabbri, 2000: 36 A: SCH
luhuoensis Fabbri, 2000: 38 A: SCH
maltonii Fabbri, 2001: 114 A: SCH
maomaoshanensis Pütz, 2007a: 49 A: GAN
markamensis markamensis Fabbri, 2000: 41 A: XIZ
markamensis oriunduss Pütz, 2007a: 70 A: XIZ
mellonii Fabbri, 2000: 30 A: SCH
numidicus Normand, 1935: 301 N: TU
grandii G. Fiori, 1948: 20

pieraccinii Fabbri, 2000: 31 A: SCH

583

584

BYRRHIDAE

pilula depilis Graëlls, 1858: 59 E: FR SP
pubipennis Mulsant & Rey, 1869a: 96

pilula herculeanus Ganglbauer, 1902a: 42 E: RO
pilula kartschchalensis Ganglbauer, 1902a: 42 E: GG A: TR
pilula pilula Linnaeus, 1758: 356 (Dermestes) E: AB AR AU BE BH BU BY CR CZ DE FI FR GB GE GG GR HU IT
LA LS LT LU MC MD ME NL NR PL RO RU SB SK SL SP SV SZ UK A: ES FE IN KZ TR WS
albopunctatus Fabricius, 1792a: 84
alpinus Newman, 1833a: 53
argentofasciatus Duftschmid, 1825: 14
auratopunctatus Reitter, 1881a: 76
aurofuscus Reitter, 1881a: 77
chinensis Paulus, 1971: 167
dennii Curtis, 1826: 135
ferrugineus Marsham, 1802: 104 (Cistela)
fuscolineatus Pic, 1948: 12 (Seminolus)
kamtschaticus Motschulsky, 1860e: 123
notatus Rey, 1889b: 66
oblongus Sturm, 1807: 97
quadrifasciatus Mulsant & Rey, 1869a: 107
rufiventer Newman, 1833b: 508
striatus Forster, 1771: 15 (Cistela)
sulcatus Zetterstedt, 1828: 130
tuscanus Dohrn, 1872: 482
viridescens Geoffroy, 1785: 28 (Cistela)

pilula regalis Steffahny, 1842: 13 E: RO
laevigatus Ganglbauer, 1902a: 42

ponticus piluloformis Paulus, 1974: 215 A: TR
ponticus ponticus Paulus, 1974: 214 E: GG A: TR
pustulatus minor G. Fiori, 1948: 18 E: IT
pustulatus pustulatus Forster, 1771: 13 (Cistela) E: AB AL AN AR AU BE BH BU BY CR CZ DE EN FI FR GB GE
GR HU IT LA LT MC MD NL NR PL RO RU SK SL SP SV SZ UK YU A: ES FE IN JIL KZ MG NC TR WS
albopunctatus Thunberg, 1794: 69 [HN]
ater Fabricius, 1781: 69 [HN]
dorsalis Fabricius, 1787a: 38
dubius Czwalina, 1887: 496
fasciatus Herbst, 1783: 26 [HN]
miyakei Nakane, 1963b: 45
morio Illiger, 1798: 93
niger Forster, 1771: 14 (Cistela)
ornatus Sulzer, 1776: 27 (Cistela)
osanaii Nakane, 1963b: 44

BYRRHIDAE
rufipennis Illiger, 1802b: 44
starcki Reitter, 1889c: 98 (Seminolus)

qinghaicus heka Pütz, 2007a: 53 A: QIN
qinghaicus ngola Pütz, 2007a: 56 A: QIN
qinghaicus qinghaicus Fabbri, 2000: 29 A: QIN
qinghaicus qingshuihe Pütz, 2007a: 57 A: QIN
reitteri G. Fiori, 1957b: 94 A: GAN
shagula Pütz, 2007a: 50 A: XIZ
shannae Pütz, 2007a: 73 A: SCH
sichuanus Fabbri, 2000: 39 A: SCH
sochondensis Tshernyshev, 1999: 18 A: ES
tongshanensis Pütz, 2007b: 205 A: HUB
turnai Fabbri, 2000: 33 A: SCH
vallei pseudoimstriatus Pütz, 2007a: 61 A: GAN
vallei vallei Fabbri, 2000: 28 A: SCH
waqensis Pütz, 2007a: 65 A: SCH
xiangchengensis Pütz, 2007a: 71 A: SCH
subgenus Ornatobyrrhus Fabbri, 2000: 53 type species Byrrhus luiginegrettoi Fabbri, 2000
Rotundobyrrhus Fabbri, 2000: 42 type species Byrrhus gansuensis Fabbri, 2000

artoprocerus Fabbri & Zhou, 2003a: 56 A: HUB
brunoi Fabbri, 2001: 118 A: SHX
dabashanensis Pütz, 2007a: 94 A: SHX
gansuensis gansuensis Fabbri, 2000: 43 A: GAN
gansuensis minshanensis Pütz, 2007a: 92 A: GAN
houzhenziensis Pütz, 2007a: 89 A: SHX
hubeianus Fabbri, 2000: 44 A: HUB
luiginegrettoi Fabbri, 2000: 54 A: SHX
plutenkoi Pütz, 2007a: 88 A: SHX
qin Pütz, 2007a: 90 A: SHX
qinlingicus crispisulcans Fabbri & Zhou, 2003a: 58 A: SCH SHX
qinlingicus jaroslavturnai Pütz, 2007b: 206 A: HUB
qinlingicus qinlingicus Fabbri, 2001: 116 A: SHX
shang Pütz, 2007a: 86 A: SHX
taibaishanensis Pütz, 2007a: 85 A: SHX
zhou Pütz, 2007a: 90 A: SHX
subgenus Pseudobyrrhus Jäger & Pütz, 2006 type species Byrrhus pilosellus A. Villa & G.B. Villa, 1833
derrei Allemand, 1987: 204 E: FR (Vercors)
focarilei Fabbri & Pütz, 1997: 23 E: IT
glabratus Heer, 1841: 447 E: AU BU CR CZ FR GE IT PL RO SK SL SP SZ
ater Dalla Torre, 1879: 101
austriacus Ganglbauer, 1902a: 40

585

586
fuscus Dalla Torre, 1879: 101
hormuzachii Marcu, 1939: 38
ornatus Panzer, 1794b: 1 [HN]
seminator Dalla Torre, 1879: 101
similaris Mulsant & Rey, 1869a: 82
striatus Steffahny, 1842: 11 [HN]
thuringiacus Giebel, 1855: 129

luniger Germar, 1817: 186 E: AU CR CZ GE IT NL PL RO SK UK
lineatus Sturm, 1823: 10
luniger Duftschmid, 1825: 10 [HN]

nigrosparsus Chevrolat, 1866: 101 E: FR SP
kiesenwetteri Mulsant & Rey, 1869a: 78

picipes amphibolus Ganglbauer, 1902a: 41 E: IT
picipes judicarius Ganglbauer, 1904b: 200 E: IT
picipes orobianus G. Fiori, 1952: 380 E: IT
picipes picipes Duftschmid, 1825: 9 E: AU
pilosellus doderoi G. Fiori, 1952: 386 (Seminolus) E: IT
pilosellus pilosellus A. Villa & G.B. Villa, 1833: 33 E: FR IT
pilosellus similis G. Fiori, 1952: 386 (Seminolus) E: IT
subgenus Seminolus Mulsant & Rey, 1869a: 50 type species Byrrhus pyrenaeus Dufour, 1834
Sinobyrrhus Fabbri, 2000: 56 type species Byrrhus pederzanii Fabbri, 2000

auromicans Kiesenwetter, 1851b: 582 E: FR SP
melanostictus Fairmaire, 1861: 581

businskyorum Fabbri, 2001: 119 A: SCH
espanoli G. Fiori, 1960: 105 E: FR SP
gigas Fabricius, 1787a: 38 E: AU BH CR CZ GE HU IT SK SL
gonggashanensis Pütz, 2007a: 95 A: SCH
inopinatus Pütz, 2007a: 96 A: SCH
lisellae G. Fiori, 1953: 99 E: FR (Pyrénées)
muliensis Fabbri, 2000: 57 A: SCH
nicolasi G. Fiori, 1966: 213 E: AN FR
occidentalis G. Fiori, 1953: 97 E: FR (Pyrénées)
pederzanii Fabbri, 2000: 56 A: SCH
pyrenaeus Dufour, 1834: 60 E: AN FR SP
bigorrensis Kiesenwetter, 1851b: 581
lobatus Kiesenwetter, 1851b: 580
soreziacus Fairmaire, 1860b: 338
suffriani Kiesenwetter, 1851b: 580

reuteri Pütz, 2009a: 290 A: AP
scabripennis Steffahny, 1842: 8 E: AU IT

BYRRHIDAE

BYRRHIDAE

587

alpinus Gory, 1844: 69 [HN]

signatus Sturm, 1823: 9 E: AU GE IT SL
inaequalis Erichson, 1846: 477
pennicoronatus Grimmer, 1841: 39

genus Curimus Erichson, 1846: 472 type species Byrrhus erinaceus Duftschmid, 1825
Norosus Mulsant & Rey, 1869a: 44 type species Byrrhus insignis Steffahny, 1843

abbreviatus J.R. Sahlberg, 1903b: 19 E: GR
anatolicus Ganglbauer, 1905: 275 A: TR
brenskei Reitter, 1884b: 71 E: GR
caucasicus Reitter, 1881a: 72 E: ST
circassicus Reitter, 1890g: 168 E: ST
decorus Steffahny, 1842: 26 (Byrrhus) E: RO
erichsoni Reitter, 1881a: 72 E: AU CZ GE HU PL RO SK
erinaceus Duftschmid, 1825: 22 (Byrrhus) E: AU BH CR GE HU IT SL
hispidus Erichson, 1846: 474
lariensis Steffahny, 1842: 27 (Byrrhus) [HN]
ruficornis Obenberger, 1917a: 31

insignis Steffahny, 1842: 26 (Byrrhus) E: BU A: TR
submaculosus Fairmaire, 1875b: cxciii
tauricus Reitter, 1884b: 72

interstitialis Reitter, 1881a: 71 E: “Eur. m.”
lariensis A. Villa & G.B. Villa, 1833: 34 (Byrrhus) E: FR IT SZ
montenegrinus Reitter, 1881b: 218 E: ME
maglicensis Obenberger, 1917a: 31

petraeus Gredler, 1863: 173 E: AU IT
alpinus Halbherr, 1890: 7 (Syncalypta)

rudis Fairmaire, 1876a: 25 E: GR
parnassius Reitter, 1884b: 71
taygetanus Reitter, 1884b: 70
terrifer Reitter, 1884b: 71

terminatus Reitter, 1884b: 72 E: GR
genus Cytilus Erichson, 1847a: 489 type species Byrrhus varius Fabricius, 1775
avunculus Fairmaire, 1887b: 98 A: SCH SHA YUN
kanoi Takizawa & Nakane, 1977: 64 A: TAI
sericeus Forster, 1771: 16 (Cistela) E: AB AL AN AR AU AZ BE BH BU BY CR CZ DE EN FI FR GB GE GG GR HU
IC IT LA LS LT LU MC MD ME NL NR PL RO RU SB SK SL SP SV SZ UK N: AG A: ES FE GAN IN JA KI KZ
MG NC QIN SC SCH WS XIN NAR
albotessellatus Letzner, 1884: 303

588

BYRRHIDAE

auricomus Duftschmid, 1825: 16 (Byrrhus)
bicolor Marsham, 1802: 105 (Cistela)
detritus Letzner, 1884: 303
fuscotesselatus Letzner, 1884: 303
maculatus Herbst, 1783: 25 (Byrrhus)
nigroviridis Letzner, 1884: 303
pilula De Geer, 1774: 213 (Dermestes)
pulchellus Heer, 1841: 448 (Byrrhus)
rufescens Letzner, 1884: 303
scutellaris Steffahny, 1842: 30 (Byrrhus)
stoicus Kugelann, 1792b: 484 (Byrrhus)
stoica O.F. Müller, 1776: 58
tessellatus Dalla Torre, 1879: 101
tessellatus Reitter, 1881a: 80 [HN]
varius Fabricius, 1775: 60
viridis Letzner, 1884: 303

genus Porcinolus Mulsant & Rey, 1869a: 94 type species Byrrhus murinus Fabricius, 1794
murinus mongoliensis Paulus, 1970b: 250 A: ES FE MG
murinus murinus Fabricius, 1794: 437 (Byrrhus) E: AR AU BE BY CZ DE FI FR GB GE HU IT LA LT NL NR PL RO
RU SV SZ UK
alternans Mulsant & Rey, 1869a: 133 (Byrrhus)
cinereoalbus Fleischer, 1900: 179
complicans Reitter, 1881a: 78
fuscus Marsham, 1802: 105 (Cistela)
niger Fleischer, 1900: 179
pulverulentus Thunberg, 1794: 68 (Byrrhus)
rubidus Kugelann, 1792b: 484 (Byrrhus)
undulatus Kugelann, 1792b: 484 (Byrrhus)
transversalis Fleischer, 1900: 179

subfamily Syncalyptinae Mulsant & Rey, 1869
tribe Syncalyptini Mulsant & Rey, 1869

genus Chaetophora Kirby, 1823: 258 type species Georyssus cretifer Spence, 1823 (=Byrrhus spinosus P. Rossi,
1794)
Syncalypta Dillwyn, 1829: 25 type species Byrrhus arenarius Sturm, 1807

godavariensis Pütz, 1999b: 258 A: NP
minuta Reitter, 1884c: 119 (Syncalypta) E: AB GR
pilosella Motschulsky, 1858d: 49 (Syncalypta) A: NP
spinosa P. Rossi, 1794: 81 (Byrrhus) E: AU BE BU CZ DE FI FR GB GE HU IT NL NR PL RO RU SK SL SP SV SZ
UK A: ES FE WS NAR

BYRRHIDAE

589

arenarius Sturm, 1807: 117 (Byrrhus)
cretifera Stephens, 1830: 133 (Syncalypta)
japonica Nakane, 1963b: 45 (Syncalypta)
oblonga Motschulsky, 1858d: 50 (Syncalypta)
pusillus Sturm, 1807: 110 (Byrrhus)

genus Curimopsis Ganglbauer, 1902a: 50 type species Syncalypta striatopunctata Steffahny, 1842 (=Cistela
maritima Marsham, 1802)
subgenus Atlantopsis Franz, 1967: 140 type species Syncalypta canariensis Franz, 1967
brancomontis Pütz, 2002b: 109 N: MR
canariensis Franz, 1967: 142 (Syncalypta) N: CI
capitata Wollaston, 1854: 207 (Syncalypta) N: MR
granulosa Wollaston, 1865: 28 (Syncalypta) N: CI
horrida Wollaston, 1854: 208 (Syncalypta) N: MR
integra Wollaston, 1864: 162 (Syncalypta) N: CI
madeirensis Pütz, 2002b: 111 N: MR
ovuliformis Wollaston, 1854: 207 (Syncalypta) N: MR
senicis Pütz, 2002b: 112 N: MR
tenerifensis Palm, 1976: 97 (Syncalypta) N: CI
wollastoni Pütz, 2002b: 113 N: MR
subgenus Curimopsis Ganglbauer, 1902a: 50 type species Syncalypta striatopunctata Steffahny, 1842 (=Cistela
maritima Marsham, 1802)
Humerimopsis Fabbri, 2003a: 33 type species Humerimopsis planicorpis Fabbri, 2003

abdita Jäger & Pütz, 2003: 233 A: NP
afghanica afghanica Pütz, 1990: 85 A: AF
afghanica tadjikistanica Pütz, 1999b: 253 A: TD
ainuorum Pütz, 2003a: 95 A: BEI JA
orientalis Fabbri & Zhou, 2003d: 82

assingi Pütz, 2006: 134 E: SP
austriaca Franz, 1967: 143 (Syncalypta) E: AU BH CR CZ FR GE HU IT RU SK SL SZ YU
cabrerensis Pütz, 2006: 136 E: SP
cantabrica Franz, 1967: 154 (Syncalypta) E: SP
carnicolica Ganglbauer, 1902c: 105 (Syncalypta) E: AU CR IT SL
curimoides G.C. Champion, 1923b: 221 (Syncalypta) A: AP HP NP UP
cyclolepidia Munster, 1902: 87 (Syncalypta) E: AU FI NR NT SV SZ A: ES FE
densa Jäger & Pütz, 2003: 234 A: NP
dhaulagirica Jäger & Pütz, 2003: 229 A: NP
ehimensis Kitano & Sakai, 2006: 267 A: JA
eos Fabbri, 2003b: 83 A: YUN
saeticirrata Fabbri, 2003b: 86

ezoensis Pütz, 2003a: 96 A: JA

590

BYRRHIDAE

franzi Paulus, 1973: 358 E: IT SZ
ganglbaueri Plavilshchikov, 1924: 233 (Syncalypta) E: PT
oblonga Ganglbauer, 1902a: 52 (Syncalypta) [HN]

ghorepanica Jäger & Pütz, 2003: 230 A: NP
giorgiofiorii Fabbri, 2003c: 87 (Humerimopsis) A: YUN
hokkaidoensis Pütz, 2003a: 97 A: JA
horii Pütz, 2003a: 97 A: JA
incisa Obenberger, 1917a: 30 (Syncalypta) E: GR IT
paganettii Franz, 1967: 145 (Syncalypta)

incognita Normand, 1935: 303 (Syncalypta) N: AG
erratrix Normand, 1935: 303 (Syncalypta)

indica Pütz, 2000c: 198 A: NP PA
italica Franz, 1967: 144 (Syncalypta) E: FR IT SZ
savoiensis Paulus, 1973: 355
vicentina Paulus, 1973: 358

japonica Nakane, 1964a: 30 A: JA
jordai Reitter, 1910b: 181 (Syncalypta) E: SP
karachina G.C. Champion, 1923b: 222 (Syncalypta) A: PA
klapperichi Pütz, 2000c: 199 A: AF HP
kolovi Tshernyshev, 2009: 637 A: KZ
lewisi Pütz, 2000c: 199 A: JA
longisetis G.C. Champion, 1923b: 221 (Syncalypta) A: AP HP NP UP
magna Paulus, 1971: 163 (Syncalypta) A: AP HP NP UP
maritima andalusiaca Franz, 1967: 151 (Syncalypta) E: PT SP
maritima maritima Marsham, 1802: 105 (Cistela) E: FR GB GR IT NL PT SP N: AG MO TU A: CY IN LE TR
hirsuta Sharp, 1871b: 151 (Syncalypta)
striatopunctata Steffahny, 1843: 34 (Syncalypta)
syriaca Baudi di Selve, 1870: 63 (Syncalypta)

maritima phoenicia Allemand, 1997b: 442 A: LE
medvedevi Tshernyshev, 2002: 80 A: TD
moosilauke Johnson, 1986: 38 A: ES FE WS NAR
uralensis Pütz, 1992: 240

mongoliensis Paulus, 1970b: 251 (Syncalypta) A: MG
monticola Franz, 1967: 152 (Syncalypta) E: AU GE IT PL SK UK A: ES KI KZ WS
motoyamaensis Pütz, 2003a: 99 A: JA
nepalica Jäger & Pütz, 2003: 230 A: NP
nigrita Palm, 1934: 314 (Syncalypta) E: DE FR GB GE IT NL PL SV
nordensis Tshernyshev, 2013: 4 A: ES
obenbergeri Paulus, 1970b: 252 (Syncalypta) A: ES
olgae Tshernyshev, 2013: 6 A: ES

591

ELMIDAE

paleata Erichson, 1846: 470 (Syncalypta) E: AU BY CZ DE FI FR GE GG HU IT NR NT PL SK SV SZ A: ES MG WS
palmi palmi Franz, 1967: 146 (Syncalypta) E: FR IT
palmi vicina Franz, 1967: 147 (Syncalypta) E: FR IT
planicorpis Fabbri, 2003a: 35 (Humerimopsis) A: NP
provencalis Franz, 1967: 147 (Syncalypta) E: FR IT
pseudocurimoides Pütz, 1999b: 255 A: NP
pustulifer Pütz, 2007a: 101 A: SCH
setica Jäger & Pütz, 2003: 232 A: NP
setigera Illiger, 1798: 95 (Byrrhus) E: AU BY CZ DE FI GB GE IT NL NT PL SV SZ
histrix Baudi di Selve, 1889: 150 (Syncalypta)

setosa Waltl, 1838: 273 (Byrrhus) E: FR GE IT SZ
reichei Mulsant & Rey, 1868: 282 (Syncalypta)

sibirica Paulus, 1970b: 253 (Syncalypta) A: ES
stefanozoiai Fabbri, 2003a: 31 A: PA
taibaishanensis Pütz, 2007a: 99 A: SHA
takizawai Pütz, 2002c: 189 A: JA
taurica Paulus, 1973: 360 A: TR
teuriensis Pütz, 2005: 173 A: JA
tibetana Franz, 1967: 149 (Syncalypta) A: QIN
tsurugisana Kitano & Pütz & Sakai, 2008: 316 A: JA
ussuriensis Lafer, 1989: 463 A: FE
weiperti Jäger & Pütz, 2003: 234 A: NP
yoshitomii Pütz, 2004a: 40 A: JA
yulongshanensis Pütz, 2007a: 102 A: YUN

family ELMIDAE Curtis, 1830
subfamily Larainae LeConte, 1861
tribe Potamophilini Mulsant & Rey, 1872

genus Dryopomorphus Hinton, 1936: 165 type species Dryopomorphus extraneus Hinton, 1936
amami Yoshitomi & Satô, 2005b: 466 A: JA (Ryukyus)
extraneus Hinton, 1936: 166 A: JA
nakanei Nomura, 1958b: 45 A: JA
yaku Yoshitomi & Satô, 2005b: 470 A: JA
genus Laorina Jäch, 1997: 393 type species Laorina schillhammeri Jäch, 1997
schillhammeri Jäch, 1997: 396 A: UP YUN ORR
genus Potamodytes Grouvelle, 1896c: 78 type species Potamophilus abdominalis C.O. Waterhouse, 1880
Gridelliana Bollow, 1939c: 149 type species Gridelliana zavattarii Bollow, 1939

subrotundatus Pic, 1939b: 143 N: EG A: AE IS SA YE AFR

592

ELMIDAE

bispinosus Pic, 1950: 23 [HN]
ochus Hinton, 1948: 137
pici Delève, 1967: 431 [RN]

genus Potamophilinus Grouvelle, 1896c: 78 type species Potamophilus longipes Grouvelle, 1892
Freyiella Bollow, 1938: 168 type species Freyiella foveicollis Bollow, 1938

bispinosus Bollow, 1938: 172 (Freyiella) A: YUN
foveicollis Bollow, 1938: 169 (Freyiella) A: GUA ORR
genus Potamophilus Germar, 1811: 41 type species Parnus acuminatus Fabricius, 1792
Hydera Latreille, 1816: 268 type species Parnus acuminatus Fabricius, 1792

acuminatus Fabricius, 1792a: 246 (Parnus) E: AB AL AU BU BY CR CZ FR GE GR HU IT NL PL RO RU SB SK SL
SP TR UK N: TU A: AF IN IS LE SY TM TR WS

subfamily Elminae Curtis, 1830
tribe Ancyronychini Ganglbauer, 1904

genus Ancyronyx Erichson, 1847: 522 type species Macronychus variegatus Germar, 1824
yunju Bian, Guo & Ji, 2012: 58 A: JIX ORR

tribe Elmini Curtis, 1830
subtribe Elmina Curtis, 1830

genus Dupophilus Mulsant & Rey, 1872a: 41 type species Dupophilus brevis Mulsant & Rey, 1872
brevis Mulsant & Rey, 1872a: 42 E: AR FR GG IT PT SP A: TR
gigas Sharp, 1872b: 263 (Limnius)
insignis Reitter, 1885b: 364 (Latelmis)

genus Elmis Latreille, 1802b: 398 type species Elmis maugetii Latreille, 1802
Elminus Rafinesque, 1815: 112 type species Elmis maugetii Latreille, 1802
Helmis Bedel, 1878: lxxv type species Elmis maugetii Latreille, 1802
Lareynia Jacquelin du Val, 1859: 276 type species Elmis maugetii Latreille, 1802
Philydrus Duftschmid, 1805: 304 type species Philydrus megerlei Duftschmid, 1805 (=Elmis maugetii Latreille, 1802)

aenea P.W.J. Müller, 1806a: 202 (Limnius) E: AL AN AU BE BH BU BY CR CZ DE DK EN FI FR GB GE HU IR IT
LA LS LT LU ME NL NR PL PT RO RU SK SL SP SV SZ UK
affinis Rey, 1889b: 67
alpina Kuwert, 1890a: 30 (Lareynia)

atlantis Alluaud, 1922: 34 (Helmis) N: MO
bosnica assyriae Olmi, 1981: 338 A: TR
bosnica bosnica Zaitzev, 1908: 303 (Helmis) [RN] E: AL AR BH BU CR GR MC ME SB ST A: IN TR
longicollis Kuwert, 1890a: 48 (Lareynia) [HN]
lousisi Mařan, 1939: 41 (Helmis)

bosnica tenuis Berthélemy, 1979: 25 E: GR (Lésvos) A: TR

ELMIDAE

593

latreillei Bedel, 1878: lxxv (Helmis) E: AU BH BU CR CZ FR GE HU IT LU PL PT RO SK SL SP SZ UK
interrupta Kuwert, 1890a: 48 (Lareynia)

maugetii fossulata Kuwert, 1890a: 30 (Lareynia) E: FR (Corse) IT (Sardegna)
maugetii maugetii Latreille, 1802b: 400 E: AR AU BE BH BU BY CR CZ EN FR GE GR HU IT LA LT LU MC NL
PL PT RO RU SB SK SL SP SZ UK A: TR
confusa Laporte, 1840: 43
kirschii Gerhardt, 1869: 261
maugei Bedel, 1878: lxxv (Helmis)
media Rey, 1889b: 67
megerlei Duftschmid, 1805: 305 (Philydrus)
similis Flach, 1882: 253 (Lareynia)

maugetii velutina Reiche, 1879: 238 N: AG
obscura P.W.J. Müller, 1806a: 204 (Limnius) E: AU BH BU CR CZ FR GE GR HU IT LA MC NL PL RU
SK SL SZ
caliginosa Laporte, 1840: 43
croatica Kuwert, 1890a: 49 (Lareynia)

perezi Heyden, 1870a: 110 E: FR PT SP
quadricollis Reitter, 1887c: 258 A: AF IN KI KZ TM UZ
lindbergi E. Janssens, 1959b: 7

rietscheli Steffan, 1958: 130 E: AU FR GE HU IT SL SZ
rioloides Kuwert, 1890a: 49 (Lareynia) E: AL AU BE BH BU CR CZ FR GE GR HU IT LU NL PT RO SK SP ST SZ
A: IS LE TR
coiffaiti Berthélemy & Clavel, 1961: 243
minuta Knie, 1975: 141

robusta Jäch, 1984c: 136 A: IN TR
syriaca syriaca Kuwert, 1890a: 49 (Lareynia) A: IS LE SY
syriaca zoufali Reitter, 1910a: 36 (Helmis) E: AL BH BU GR SB A: TR
genus Esolus Mulsant & Rey, 1872a: 36 type species Limnius parallelepipedus P.W.J. Müller, 1806
angustatus P.W.J. Müller, 1821: 187 (Limnius) E: AN AR AU BE BH BU CR CZ FR GE GR HU IT LA LU MC NL PL
PT RO RU SB SK SL SP SV SZ UK A: TR
carpetanus Kuwert, 1889b: 33
galloprovincialis Abeille de Perrin, 1900b: 137
solarii Ganglbauer, 1904a: 116

berthelemyi Olmi, 1975b: 236 E: FR IT
bicuspidatus Alluaud, 1922: 36 N: MO
brevis Kuwert, 1890a: 46 E: FR (Corse) IT (Sardegna)
czwalinae Kuwert, 1889b: 33 [=1890a: 46] E: BH CR FR IT SL
doderoi Olmi, 1975b: 232

filum Fairmaire, 1871a: 373 (Elmis) N: AG MO TU
nepalensis Jäch, 1982: 91 A: NP

594

ELMIDAE

parallelepipedus P.W.J. Müller, 1806a: 200 (Limnius) E: AU BE BH BU CR CZ FR GB GE GR HU IR IT LU MC NL
PL PT RO SB SK SL SP SZ UK N: MO A: IN LE TR
dossowi Kuwert, 1889b: 33 [=1890a: 47]
kuenowi Kuwert, 1889b: 33 [=1890a: 47]
nicariae Kuwert, 1890a: 46
perparvulus Kuwert, 1890a: 48
politus Kuwert, 1889b: 33
rugosus Babington, 1832: 329

pygmaeus P.W.J. Müller, 1806a: 201 (Limnius) E: AU BE BU CR CZ FR GE GR LU NL PL PT RO SK SP SZ N: AG
MO A: IS LE TR
maroccanus Alluaud, 1922: 35
subparallelus Kuwert, 1889b: 33

taurus Jäch, 1982: 90 A: TR
theryi Alluaud, 1922: 36 N: MO
genus Graphelmis Delève, 1968: 169 type species Graphelmis scapularis Delève, 1968
clermonti Pic, 1923b: 4 (Stenelmis) A: YUN ORR
delevei Čiampor, 2001: 27 A: SCH
jaechi Čiampor, 2001: 20 A: ANH FUJ GUA GUX HKG HUN JIX YUN ORR
schillhammeri Čiampor, 2004: 12 A: YUN ORR
shirahatai Nomura, 1958a: 42 (Stenelmis) A: JA
genus Grouvellinus G.C. Champion, 1923a: 168 [RN] type species Macronychus caucasicus Motschulsky, 1839
Grouvelleus Zaitzev, 1908: 311 [HN RN] type species Macronychus caucasicus Motschulsky, 1839
Microdes Motschulsky, 1860b: 48 [HN] type species Macronychus caucasicus Motschulsky, 1839

aerosus Jung, Jäch & Bae, 2015: 94 A: SC
babai babai Nomura, 1963b: 54 A: TAI
babai satoi Jeng & P.-S. Yang, 1998: 538 A: JA (Ryukyus)
brevior Jäch, 1984b: 110 A: NP XIZ
carinatus Jäch, 1984b: 111 A: NP
carus Hinton, 1941: 69 A: FUJ
caucasicus Motschulsky, 1839: 70 (Macronychus) E: AR GG GR (Sámos) ST A: IN IQ IS LE SY TR
coyei Allard, 1869: 466 (Elmis)

chinensis Mařan, 1939: 42 A: SCH
duplaris G.C. Champion, 1923a: 168 A: HP UP
hadroscelis Jäch, 1984b: 115 A: NP
hercules Jäch, 1984b: 113 A: NP XIZ
hygropetricus Jeng & P.-S. Yang, 1998: 533 A: TAI
marginatus Kôno, 1934a: 127(Grouvelleus) A: JA
montanus Jeng & P.-S. Yang, 1998: 530 A: TAI
nepalensis Delève, 1970a: 319 A: NP XIZ

ELMIDAE

595

nitidus Nomura, 1963b: 52 A: JA
pilosus Jeng & P.-S. Yang, 1998: 527 A: TAI
rioloides Reitter, 1887c: 259 (Macronychus) A: AF KI KZ TD TM UZ XIN
arius E. Janssens, 1959b: 3 (Macronychus)
flavibasis Reitter, 1887c: 260 (Macronychus)

sinensis Grouvelle, 1906: 125 (Microdes) A: YUN
subopacus Nomura, 1962: 48 A: JA (Ryukyus)
tibetanus Jäch, 1984b: 114 A: NP XIZ
unicostatus G.C. Champion, 1923a: 169 A: HP UP
genus Heterlimnius Hinton, 1935: 178 type species Helmis koebelei Martin, 1927
Cyclolimnius Nomura, 1958b: 54 type species Optioservus kubotai Nomura, 1958

amabilis Kamite, 2011: 409 A: GUI SCH
ater Nomura, 1958b: 56 (Optioservus) A: JA
hayashii Nomura, 1960a: 34 (Optioservus)

ennearthrus Kamite, 2009: 216 A: ES KZ
hasegawai Nomura, 1958b: 52 (Optioservus) A: FE (Kuril’skie ostrova Sakhalin) JA JIL LIA NC SC
kubotai Nomura, 1958b: 54 (Optioservus)
saghaliensis Nomura, 1958b: 55 (Optioservus)

hisamatsui Kamite, 2009: 219 A: SCH YUN
horii Kamite, 2012: 291 A: SCH
ikedai Kamite, 2011: 411 A: GUI
jaechi Kamite, 2009: 218 A: BT UP
quadrigibbus Kamite, 2012: 292 A: XIZ
shepardi Kamite, 2009: 221 A: GUI SCH SHA
trachys E. Janssens, 1959b: 5 (Stenelmis) A: AF
yokoii Kamite, 2012: 294 A: YUN
genus Limnius Illiger, 1802c: 297 type species Dytiscus volckmari Panzer, 1793
Latelmis Reitter, 1883c: 75 type species Dytiscus volckmari Panzer, 1793
Lathelmis Zaitzev, 1908: 307 type species Dytiscus volckmari Panzer, 1793

colchicus Delève, 1963: 7 E: AR BU GG GR A: TR
gibbosus Jäch, 1984a: 35 A: TR
intermedius Fairmaire, 1881b: xi E: AL AU BH BU BY CR CZ DE FR GR HU IT MC PL PT RO RU SB SK SL SP TR
UK N: AG MO TU A: IS LE TR
horioni Illies, 1953: 179 (Lathelmis)
longior Pic, 1905d: 145 (Elmis)
palaestinus Pic, 1901: 3 (Elmis)
rufiventris Kuwert, 1890a: 45 (Elmis)
subcrenatus Rey, 1889b: 67 (Latelmis)

letourneuxi Pic, 1894b: 195 (Elmis) A: IS SY

596

ELMIDAE

muelleri Erichson, 1847a: 529 (Elmis) E: AU CZ FR GE HU SZ
oblongus Rey, 1889b: 67 (Latelmis)

opacus jahandiezi Alluaud, 1922: 42 (Latelmis) N: MO
opacus opacus P.W.J. Müller, 1806a: 197 E: AR AU BE BH BU CR CZ FR GE GR HU IT LU NL PL PT RO SB SK SL
SP ST SZ UK N: AG MO A: IS LE TR
carreti Pic, 1895c: cxxviii (Elmis)
dissidens Alluaud, 1922: 41 (Latelmis)
latiusculus Zaitzev, 1947: 90 (Lathelmis)
lepidopterus Kuwert, 1890a: 45 (Elmis)
liouvillei Alluaud, 1922: 42 (Latelmis)
subopacus Ganglbauer, 1904a: 118(Latelmis)

perrisi carinatus Pérez Arcas, 1865b: 413 (Elmis) E: PT SP
mediocarinatus Berthélemy, 1964: 261
subcarinatus Sharp, 1872b: 263 (Elmis)

perrisi perrisi Dufour, 1843: 56 (Elmis) E: AL AN AU BE BH BU CR CZ FR GE GR HU IT LU MC NL PL RO SB SK
SL SP SZ UK A: TR
germari Erichson, 1847a: 528 (Elmis)
rambouseki Mařan, 1939: 41 (Latelmis)

satanus Jäch, 1982: 93 A: TR
stygius Hernando, Aguilera & Ribera, 2001: 69 N: MO
sulcipennis damryi Fairmaire, 1881b: xii E: FR (Corse)
sulcipennis sulcipennis Fairmaire, 1881b: xii E: IT (Sardegna)
surcoufi Pic, 1905d: 145 (Elmis) N: AG
swayambhu Jäch, 1982: 92 A: NP
volckmari Panzer, 1793b: 4 (Dytiscus) E: AR AU BE BH BU BY CR CZ DE EN FI FR GB GE GR HU IR IT LA LT LU
MC ME NL NR PL PT RO RU SB SK SL SP SV SZ UK A: IN TR
cecconii Reitter, 1906c: 239 (Latelmis)

wewalkai Jäch, 1984a: 36 A: TR
genus Macrelmis Motschulsky, 1860b: 52 type species Macrelmis dentata Motschulsky, 1859
Elsianus Sharp, 1882: 131 type species Elsianus striatus Sharp, 1882
Uralohelmis Roubal, 1940: 152 type species Uralohelmis jureceki Roubal, 1940

jureceki Roubal, 1940: 154 (Uralohelmis) E: CT
genus Neoriohelmis Nomura, 1958b: 45 type species Neoriohelmis kurosawai Nomura, 1958
kurosawai Nomura, 1958b: 46 A: JA
kuwatai Satô, 1963b: 132 A: JA
genus Nomuraelmis Satô, 1964a: 11 type species Nomuraelmis amamiensis Satô, 1964
amamiensis Satô, 1964a: 12 A: JA (Ryukyus)
genus Optioservus Sanderson, 1953b: 155 type species Limnius trivittatus W.J. Brown, 1930

ELMIDAE

597

gapyeongensis Jung, Kamite & Bae, 2011: 179 A: FE LIA NC SC
hagai Nomura, 1958b: 56 A: JA
inahatai Kamite 2015: 203 A: JA
maculatus Nomura, 1958b: 50 A: JA
masakazui Kamite, 2015: 206 A: JA
nitidus Nomura, 1958b: 53 A: JA
occidens Kamite, 2015: 207 A: JA
ogatai Kamite, 2015: 209 A: JA
rugulosus Nomura, 1958b: 53 A: JA
sakaii Kamite, 2015: 212 A: JA
variabilis Nomura, 1958b: 51 A: JA
yoshitomii Kamite, 2015: 216 A: JA
genus Orientelmis Shepard, 1998: 289 type species Orientelmis sinensis Shepard, 1998
sinensis Shepard, 1998: 291 A: GUX HUN JIX
parvula Nomura & Baba, 1961: 4 (Cleptelmis) A: JA
genus Oulimnius Gozis, 1886: 9 type species Limnius tuberculatus P.W.J. Müller, 1806
aegyptiacus Kuwert, 1890a: 44 (Limnius) N: EG A: SI
lineatus Kuwert, 1890a: 20 (Limnius)

bertrandi Berthélemy, 1964: 256 E: PT SP
cyneticus Berthélemy, 1980: 421 E: PT SP
echinatus Berthélemy, 1979: 77 E: SP (Baleares)
fuscipes Reiche, 1879: 238 (Limnius) E: SP N: AG MO TU
interruptus Fairmaire, 1884c: lxi (Limnius)
kebir Alluaud, 1926: 21 (Limnius)

hipponensis Berthélemy, 1979: 82 N: AG TU
jaechi Hernando, Ribera & Aguilera, 1998: 254 N: MO
major Rey, 1889b: 67 (Limnius) E: FR GB NL PT SP
falcifer Berthélemy, 1962: 218

maurus Berthélemy, 1979: 85 N: AG
perezi Sharp, 1872b: 264 (Limnius) E: PT SP
reygassei Peyerimhoff, 1929b: 171 (Limnius) N: AG
rivularis Rosenhauer, 1856: 113 (Limnius) E: FR GB GE IT NL PT SP N: AG MO TU
neuter Kuwert, 1890a: 20 (Limnius)
subparallelus Fairmaire, 1863: 74 (Elmis)

troglodytes Gyllenhal, 1827: 395 (Limnius) E: BE DE FR GB GE IT LT NL PL PT SP SV N: MO TU
brevis Sharp, 1872b: 264 (Limnius)
fluviatilis Stephens, 1828: 107 (Elmis)
thermarius Sainte-Claire Deville, 1919: 263 (Limnius)

tuberculatus P.W.J. Müller, 1806a: 199 (Limnius) E: AL AR AU BE BU BY CZ DE EN FI FR GB GE GG GR HU IR
IT LA LT LU MC NL NR PL RU SB SK SL SP SV SZ UK A: TR

598

ELMIDAE

dargelasi Latreille, 1807: 51 (Elmis)
formosus Kuwert, 1890a: 44 (Limnius)
lacustris Stephens, 1828: 107 (Elmis)
variabilis Stephens, 1828: 107 (Elmis)

villosus Berthélemy, 1979: 88 N: AG MO
genus Pseudamophilus Bollow, 1940a: 9 type species Pseudamophilus malaisei Bollow, 1940
japonicus Nomura, 1957a: 3 A: JA
tibiaferus J. Yang & Z. Zhang, 2002: 812 A: FUJ
genus Riolus Mulsant & Rey, 1872a: 29 type species Limnius cupreus P.W.J. Müller, 1806
Aptyktophallus Steffan, 1958: 165 type species Limnius nitens P.W.J. Müller, 1817
Normandia Pic, 1900b: 267 type species Limnius villosocostatus Reiche, 1879

cupreus P.W.J. Müller, 1806a: 205 (Limnius) E: AU BE BH BU CR CZ DE EN FR GB GE GR HU IT LA LT LU NL
NT PL RO SB SK SL SP SV SZ UK A: TR
argolicus E. Janssens, 1959a: 16
erichsoni Kuwert, 1889a: 25
lentzi Kuwert, 1889a: 25 [=1890: 52]
mulsanti Kuwert, 1889a: 25
senex Kuwert, 1889a: 24 [=1890: 51]
steineri Kuwert, 1889a: 24 [=1890: 51]

fontinalis Jäch, 1984c: 140 A: TR
illiesi Steffan, 1958: 159 E: BE FR GE SP SZ
nitens P.W.J. Müller, 1817: 273 (Limnius) E: AU BE BH BY CR CT EN FI FR GB GE GR IT LA LT MC NL NT PT SL
SP SV SZ N: AG MO A: IS LE TR
sauteri Kuwert, 1889a: 24 [=1890a: 50]
seidlitzi Kuwert, 1889a: 24 [=1890a: 50]

robustior Pic, 1900b: 266 N: AG TU
sodalis Erichson, 1847a: 532 (Elmis) E: FR GE IT SP SZ
meridionalis Grouvelle, 1900: 137

somcheticus Kolenati, 1846: 64 (Elmis) E: GG GR (Sámos) ST UK A: IN LE SY TR
anatolicus Delève, 1963: 9
syriacus Allard, 1869: 466 (Elmis)

substriatus Grouvelle, 1889: lxxx N: AG MO TU
punctatostriatus Alluaud, 1922: 38

subviolaceus P.W.J. Müller, 1817: 273 (Limnius) E: AU BE BH BU CR CZ FR GB GE GR HU IT LU NL PL RO SB SK
SL SP SZ
apfelbecki Ganglbauer, 1904c: 354
auronitens Kuwert, 1889a: 26
australis Rey, 1889b: 67
bosnicus Kuwert, 1889a: 26

599

ELMIDAE
nigricans Rey, 1889b: 67
wichmanni Zimmermann, 1908: 341

villosocostatus Reiche, 1879: 238 (Limnius) N: AG MO TU

subtribe Stenelmina Mulsant & Rey, 1872

genus Leptelmis Sharp, 1888: 243 type species Leptelmis gracilis Sharp, 1888
Lephthelmis Zaitzev, 1910: 21 type species Leptelmis gracilis Sharp, 1888

brunnelineata Z. Zhang, Su & C. Yang, 2003b: 189 A: GUX
flavicollis Bollow, 1941: 86 A: “South China”
formosana Nomura, 1962: 48 A: TAI ORR
fracticollis G.C. Champion, 1923a: 167 A: UP
fujiana J. Yang & Z. Zhang, 2002: 812 A: FUJ
gracilis gracilis Sharp, 1888: 244 A: JA SC
coreana Jung & Bae, 2012: 255
ochra Jung & Bae, 2012: 256
parallela Nomura, 1962: 46

gracilis impubis Z. Zhang & Ding, 1995: 17 A: BEI
guangxiana Z. Zhang, Su & C. Yang, 2003b: 191 A: GUX
obscura Delève, 1968: 153 A: GUX ORR
vittata Z. Zhang, Su & C. Yang, 2003b: 190 A: GUX
genus Ordobrevia Sanderson, 1953a: 159 type species Stenelmis nubifer Fall, 1901
amamiensis amamiensis Nomura, 1957a: 4 (Stenelmis) A: JA (Ryukyus)
amamiensis okinawana Nomura, 1959a: 34 A: JA (Ryukyus)
foveicollis Schönfeldt, 1888a: 193 (Stenelmis) A: JA
flavovittata Kôno, 1934a: 126 (Stenelmis)
freyi Bollow, 1941: 21 (Stenelmis)
japonica E. Janssens, 1956: 2 (Stenelmis)

gotoi Nomura, 1959a: 33 A: JA
maculata Nomura, 1957b: 43 (Stenelmis) A: JA
genus Sinelmis Satô & Kishimoto, 2001: 76 type species Sinelmis uenoi Satô & Kishimoto, 2001
uenoi Satô & Kishimoto, 2001: 77 A: GUI
genus Stenelmis Dufour, 1835: 158 type species Limnius canaliculatus Gyllenhal, 1808
accessoria J. Yang & Z. Zhang, 2002: 818 A: ANH FUJ JIA
angustisulcata Z. Zhang & J. Yang, 1995: 103 A: ZHE
anytus Hinton, 1941: 82 A: FUJ
aria E. Janssens, 1961: 5 A: AF
aritai Satô, 1964b: 34 A: JA (Ryukyus)

600

ELMIDAE

auriculata J. Yang & Z. Zhang, 2002: 820 A: FUJ GUI
beijingana Z. Zhang & Ding, 1995: 15 A: BEI
canaliculata Gyllenhal, 1808: 552 (Limnius) E: AU BE CR FI FR GB GE IT LU NL NR PL PT SK SP SV SZ UK
bollovotiva Steffan, 1961: 307
elongata Motschulsky, 1860b: 51
stoeckleini Bollow, 1941: 12

capys Hinton, 1941: 78 A: FUJ
casius Hinton, 1941: 80 A: FUJ
centridivisa Z. Zhang, C. Yang & L. Zhang, 2003: 121 A: GUX
consobrina consobrina Dufour, 1835: 161 E: AR BH CR CZ FR GE GG GR IT SP ST SZ N: AG MO TU A: IN IS SY
TM TR
consobrina peropaca Reitter, 1907e: 483 A: AF KZ UZ XIN
convexa Z. Zhang, C. Yang & L. Zhang, 2003: 120 A: GUX
debilis G.C. Champion, 1927: 51 A: HP
decipiens Bollow, 1941: 39 A: FUJ
elfriedeae Bollow, 1941: 64 A: GUA
euronotana J. Yang & Z. Zhang, 1995: 102 A: FUJ GUX ZHE
exarata Motschulsky, 1860b: 49 A: “continent Indien”
fissicollis G.C. Champion, 1923a: 166 A: UP
formosana Jeng & P.-S. Yang, 1991: 242 A: TAI
fukiensis Bollow, 1941: 75 A: FUJ
fursovi Zaitzev, 1951: 73 A: TD
gauglerae Bollow, 1941: 28 A: “South China”
granulosa Z. Zhang, C. Yang & L. Zhang, 2003: 118 A: GUX
grossepunctata Bollow, 1941: 56 A: FUJ
grossimarginata J. Yang & Z. Zhang, 1995: 102 A: FUJ ZHE
guangxinensis Z. Zhang, C. Yang & L. Zhang, 2003: 122 A: GUX
gutianshana J. Yang & Z. Zhang, 1995: 104 A: ZHE
hayashii Satô, 1999: 121 A: JA (Ryukyus)
heteromorpha Z. Zhang & J. Yang, 1995: 106 A: GUI
hisamatsui Satô, 1960: 253 A: JA (Ryukyus)
huangkengana J. Yang & Z. Zhang, 1995: 105 A: FUJ
inaccessoria J. Yang & Z. Zhang, 2002: 819 A: FUJ
indepressa J. Yang & Z. Zhang, 1995: 103 A: FUJ ZHE
insufficiens Jäch & Kodada, 2006: 60 [RN] A: ZHE
sinuata Z. Zhang & J. Yang, 1995: 107 [HN]

iranensis Delève, 1970b: 701 A: IN
ishiharai Satô, 1964b: 33 A: JA
kaihuana Z. Zhang & J. Yang, 1995: 105 A: CHQ ZHE
klapperichi Bollow, 1941: 42 A: FUJ
kochi Bollow, 1941: 26 A: “South China”

601

ELMIDAE
koreana Satô, 1978: 147 A: FE NC SC
kuntzeni Bollow, 1941: 71 A: GUA
leblanci Peyerimhoff, 1929b: 171 N: AG
lutea Z. Zhang, Su & C. Yang, 2003a: 107 A: FUJ GUX ZHE
magnofoveola Bollow, 1941: 48 A: FUJ
montana J. Yang & Z. Zhang, 1995: 102 A: FUJ GUX ZHE
nematodes E. Janssens, 1959b: 6 A: AF
nipponica Nomura, 1958a: 41 A: JA NC SC
obscurifusca Z. Zhang, C. Yang & L. Zhang, 2003: 119 A: GUI GUX
orthotibiata J. Yang & Z. Zhang, 2002: 814 A: FUJ GUX
parvifoveola Z. Zhang, Su & C. Yang, 2003a: 107 A: GUX
peyerimhoffi Bollow, 1941: 15 N: MO
priapus Hinton, 1941: 76 A: FUJ
puberula Reitter, 1887c: 259 E: AR BH GG SK ST UK A: IN IS LE SY TM TR
apfelbeki Kuwert, 1890a: 52
damascena Reitter, 1907e: 484

punctulata Bollow, 1940a: 24 A: FUJ ORR
roiae Bollow, 1941: 60 A: “South China”
sauteri Kôno, 1936: 122 A: TAI
scutellicarinata Z. Zhang & J. Yang, 1995: 106 A: ZHE
sinica J. Yang & Z. Zhang, 1995: 102 A: ANH FUJ GAN GUI HUB HUN SCH ZHE
subtilis Zaitzev, 1951: 72 A: KZ
sulcaticarinata Z. Zhang & J. Yang, 1995: 102 A: FUJ ZHE
sulmo Hinton, 1941: 73 A: FUJ
szechuanensis Mařan, 1939: 40 A: SCH
testacea Grouvelle, 1892: 867 A: “South China” ORR
trisulcata Fairmaire, 1889b: 337 A: BEI “South China” ORR
troilus Hinton, 1941: 75 A: FUJ
tros Hinton, 1941: 77 A: FUJ
venticarinata Z. Zhang & J. Yang, 1995: 102 A: FUJ GUI GUX ZHE
ventiplana Z. Zhang & C. Yang, 2003: 276 A: GUX
vulgaris Nomura, 1958a: 44 A: JA LIA NC SC
miyamotoi Nomura & Nakane, 1958: 81

wongi Jeng & P.-S. Yang, 1991: 240 A: TAI
yangi Z. Zhang & Ding, 1995: 16 A: BEI
zhangi Jäch & Kodada, 2006: 60 [RN] A: GUX
subsulcata Z. Zhang, Su & C. Yang, 2003a: 107 [HN]

tribe Macronychini Gistel, 1848

genus Cuspidevia Jäch & Boukal, 1995: 301 type species Cuspidevia velaris Jäch & Boukal, 1995
brevis Bian & Ji, 2010: 54 A: JIX

602

ELMIDAE

jaechi Bian & Ji, 2010: 56 A: JIX
velaris Jäch & Boukal, 1995: 302 A: HKG
genus Eonychius Jäch & Boukal, 1996: 179 [RN] type species Eonychus dudgeoni Jäch & Boukal, 1995
Eonychus Jäch & Boukal, 1995: 303 [HN] type species Eonychus dudgeoni Jäch & Boukal, 1995

dudgeoni Jäch & Boukal, 1995: 304 (Eonychus) A: FUJ GUA HKG
genus Indosolus Bollow, 1940a: 28 type species Esolus nitidus Bollow, 1940
acutangulus G.C. Champion, 1923a: 171 (Zaitzevia) A: UP
nitidus Bollow, 1940a: 28 (Esolus) A: YUN ORR
genus Jilanzhunychus Jäch & Boukal, 1995: 304 type species Jilanzhunychus costatus Jäch & Boukal, 1995
costatus Jäch & Boukal, 1995: 306 A: GUX
genus Macronychus P.W.J. Müller, 1806b: 207 type species Macronychus quadrituberculatus P.W.J. Müller,
1806
indicus Hinton, 1940: 118 A: NP UP ORR
jaechi Čiampor & Kodada, 1998: 240 A: HAI
jendeki Čiampor & Kodada, 1998: 245 A: ANH GUA HUN JIX
kubani Čiampor & Kodada, 1998: 243 A: SCH YUN
levanidovae Lafer, 1980: 50 A: FE SC
quadrituberculatus P.W.J. Müller, 1806b: 215 E: AU BY CR CZ EN FI FR GB GE GR HU IT LA LT PL PT RO RU SB
SK SP SZ UK N: MO
reticulatus Čiampor & Kodada, 1998: 237 A: GUA HUN YUN ORR
genus Paramacronychus Nomura, 1958b: 47 type species Paramacronychus granulatus Nomura, 1958
granulatus Nomura, 1958b: 48 A: JA
genus Sinonychus Jäch & Boukal, 1995: 306 type species Sinonychus lantau Jäch & Boukal, 1995
lantau Jäch & Boukal, 1995: 308 A: HKG
satoi Yoshitomi & Nakajima, 2007: 97 A: JA (Ryukyus)
tsujunensis Yoshitomi & Nakajima, 2012: 55 A: JA
genus Urumaelmis Satô, 1963a: 39 type species Zaitzevia uenoi Nomura, 1961
uenoi tokarana Satô, 1963a: 40 (Zaitzevia) A: JA (Ryukyus)
uenoi uenoi Nomura, 1961: 2 (Zaitzevia) A: JA (Ryukyus)
amamiana Satô, 1963a: 41 (Zaitzevia)

genus Vietelmis Delève, 1968: 174 type species Vietelmis brevicornis Delève, 1968
sinensis Kodada & Čiampor, 2000: 73 A: GUX HAI HUN

603

DRYOPIDAE
genus Zaitzevia G.C. Champion, 1923a: 170 type species Zaitzevia solidicornis G.C. Champion, 1923
Awadoronus Kôno, 1934a: 127 type species Awadoronus awanus Konô, 1934

aritai Satô, 1963a: 40 A: JA (Ryukyus)
awana Kôno, 1934a: 128 (Awadoronus) A: JA SC
babai Nomura, 1963b: 43 A: TAI
bhutanica Satô, 1977: 202 A: BT
crassipes G.C. Champion, 1927: 50 A: HP
elongata Nomura, 1962: 49 A: JA (Ryukyus)
formosana Nomura, 1963b: 50 A: TAI
nitida Nomura, 1963b: 46 A: JA
parallela Nomura, 1963b: 45 A: TAI
rivalis Nomura, 1963b: 48 A: JA
rufa Nomura & Baba, 1961: 5 A: JA
solidicornis G.C. Champion, 1923a: 170 A: UP
tsushimana Nomura, 1963b: 47 A: FE JA (Tsushima) JIL SC
yaeyamana Satô, 1963a: 40 A: JA (Ryukyus)
genus Zaitzeviaria Nomura, 1959a: 35 type species Zaitzevia brevis Nomura, 1958
atratula Grouvelle, 1911b: 252 (Microdes) A: YUN
brevis Nomura, 1958b: 49 (Zaitzevia) A: JA
gotoi Nomura, 1959a: 36 (Zaitzevia) A: FE (Kunashir) JA
kuriharai Kamite, Ogata & Satô, 2006: 149 A: JA (Tsushima)
kyungseoki Jung, Jäch & Bae, 2015: 116 A: SC
obesa Jung, Jäch & Bae, 2015: 117 A: SC
ovata Nomura, 1959a: 35 (Zaitzevia) A: JA
sotai Hayashi & Yoshitomi, 2015: 71 A: JA

family DRYOPIDAE Billberg, 1820 (1817)

genus Ahaggaria Bollow, 1938: 176 type species Dryops foleyi Peyerimhoff, 1929
acutangula Bollow, 1938: 181 A: SA YE AFR
foleyi Peyerimhoff, 1929b: 170 (Dryops) N: AG
genus Ceradryops Hinton, 1937: 79 type species Ceradryops punctatus Hinton, 1937
Uenodryops Satô, 1981: 54 type species Uenodryops himalayanus Satô, 1981

himalayanus Satô, 1981: 55 (Uenodryops) A: NP
matei Kodada & Boukal, 2003: 475 A: HKG
genus Dryops A.G. Olivier, 1791: 297 type species Dermestes auriculatus Geoffroy, 1785
Parnus Fabricius, 1792a: 245 type species Parnus prolifericornis Fabricius, 1792 (=Dermestes auriculatus Geoffroy, 1785)
Yrdops Steffan, 1961: 286 type species Parnus striatopunctatus Heer, 1841

604

DRYOPIDAE

algiricus Lucas, 1846c: pl. 23 (Parnus) E: AZ BU CR FR GR IT MA PT SP N: AG MO TU A: IN IQ IS LE SY TR
antonii Dodero, 1918: 109
discretus Bollow, 1936: 156
freyi Bollow, 1939a: 356
hispanus Olmi, 1972: 99
hydrobates Kiesenwetter, 1850: 223 (Parnus)

anglicanus Edwards, 1909: 218 E: AU CT CZ DE EN FI FR GB GE GR HU IT LA LT NL NR NT PL SK SV SZ
auriculatus Geoffroy, 1785: 20 (Dermestes) E: AU BU BY CT CZ DE EN FI FR GB GE GR HU IT LA LT LU ME NL
NR NT PL SK SL ST SV SZ “Caucasus” A: ES FE KI “Northwest China”
argentipilosus Poppius, 1907: 12 (Parnus)
bicolor Curtis, 1825: 80 (Parnus)
dermestoides Fabricius, 1787a: 175 (Elater)
hirsutus Seidlitz, 1888a: 126 (Parnus)
impressus Curtis, 1825: 80 (Parnus)
niveus Heer, 1841: 467 (Parnus)
prolifericornis Fabricius, 1792a: 245 (Parnus)
rudolfi Rusek, 1973: 86
sericeus Samouelle, 1819: 185 (Parnus)

caspius Ménétriés, 1832: 172 (Parnus) E: AB GR (Kríti) “Caucasus” A: TM TR
caucasicus Kuwert, 1890a: 42 (Parnus)

cassius Hinton, 1948: 135 A: SA YE
championi Dodero, 1918: 109 E: PT SP
costae Heyden, 1891: 76 (Parnus) [RN] E: FR (Corse) IT (Sardegna)
bicolor A. Costa, 1883: 49 (Parnus) [HN]

doderoi Bollow, 1936: 156 E: FR IT SP N: AG MO TU
ernesti Gozis, 1886: 9 E: AU BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR IT LA LT LU NL NR NT PL RO
SK SL SP SV SZ
schneideri Reitter, 1897a: 46 (Parnus)

fiorii Olmi, 1986: 589 A: XIN
friebi Bollow, 1939a: 353 A: WS
fuscipennis Solsky, 1876a: 284 (Parnus) A: AF KI UZ
goidanichi Olmi, 1975a: 216 A: AF IN IQ NP PA TM
gracilis Karsch, 1881: 45 (Parnus) E: FR GR IT PT SP N: AG CI MO A: SI YE AFR
distinctus Bollow, 1939b: 124
ignotus Bollow, 1939b: 121
subspoliatus Rey, 1889b: 67 (Parnus)

griseus Erichson, 1847a: 513 (Parnus) E: AU BY CR CT CZ DE EN FI FR GB GE GR HU LA LU ME NL NR NT PL
SK SV
kaszabi Olmi, 1976: 95

iranensis Olmi, 1986: 592 A: IN
jaechi Olmi, 1986: 590 A: NP

DRYOPIDAE

605

jeanneli Bollow, 1939a: 337 A: TR UZ
luridus Erichson, 1847a: 513 (Parnus) E: AU AZ BE BU BY CR CZ DE EN FR GB GE GR HU IR IT LA LT LU NL
NR PL PT RO SK SP SV SZ N: AG CI EG MR TU A: SI TR AFR
fulvago Rey, 1889b: 67 (Parnus)
fuscipes Rey, 1889b: 67 (Parnus)
intermedius Kuwert, 1890a: 53 (Parnus)
pubescens Walker, 1871: 11 (Parnus)
subseriatus Rey, 1889b: 67 (Parnus)

lutulentus Erichson, 1847a: 514 (Parnus) E: AL BU CR CZ EN FR GE GR HU IT LT ME PL SK SL SP ST SZ
“Caucasus” N: MO A: AE AF IN SA TM TR XIN YE “Turkestan”
complanatus J. Müller, 1908: 316

mesatlanticus Peyerimhoff, 1930: 256 N: MO
pauliani Delève, 1940: 43

monticola Grouvelle, 1913: 115 A: AP
nepalensis Olmi, 1986: 592 A: NP
nitidulus Heer, 1841: 467 (Parnus) E: AU BE BU BY CT CZ DE EN FR GB GE GR HU IT LA LT LU NL NR NT PL
RO SK SL SP SV SZ UK A: TR
osellai Olmi, 1978: 381 A: PA
peyerimhoffi Bollow, 1939b: 126 N: AG MO TU
villiersi Delève, 1940: 42

rufipes Krynicki, 1832: 115 (Parnus) E: AB AU BU CR CT CZ FR (Corse) GE GR HU IT RO SK SL ST SZ UK N: EG
A: AF IN IS LE SY TD TM TR UZ
corsicus Kuwert, 1890a: 41 (Parnus)
pilosellus Erichson, 1847a: 515 (Parnus)
puberulus Reiche & Saulcy, 1856: 368 (Parnus)
punctatellus Bollow, 1936: 56

seurati Bollow, 1939b: 128 N: AG
similaris Bollow, 1936: 191 E: AU CR CT CZ DE EN FI FR GB GE GR HU IR IT LU NL NR NT PL RO SK SP ST SV
“Caucasus” A: FE TR
omissus Bollow, 1939a: 354

striatellus Fairmaire & Brisout de Barneville, 1859: 46 (Parnus) E: AU FR GB GE GR IT NL PL PT SK SP N: AG
MO TU
seriatus Rey, 1889b: 67 (Parnus)

striatopunctatus Heer, 1841: 466 (Parnus) E: AU BH BU CZ FR GE GR IT PL RO SK SL SZ UK
impressus Mulsant & Rey, 1872: 22 (Parnus)
kourilii Roubal, 1934: 43

subincanus Kuwert, 1890a: 38 (Parnus) E: AL BH BU CR FR GE GR IT ST N: “North Africa”
sulcipennis A. Costa, 1883: 49 (Parnus) E: BU FR GR IT PT SP N: AG MO TU A: IN IS SA YE AFR
brittoni Hinton, 1948: 135

viennensis Laporte, 1840: 44 (Parnus) E: AL AU BH BU CR CT CZ FR GE GR HU IT LT NL PL RO SK SL SP ST SZ
UK “Caucasus” A: TR NAR

606

DRYOPIDAE

italicus Dodero, 1918: 105
obscurus Duftschmid, 1805: 308 (Parnus)

genus Elmomorphus Sharp, 1888: 242 type species Elmomorphus brevicornis Sharp, 1888
Dryopidius Grouvelle, 1896b: 33 type species Dryopidius castaneus Grouvelle, 1896
Elmomorphellus Chûjô & Satô, 1964: 193 type species Elmomorphus sarawakensis Chûjô & Satô, 1964

amamiensis Nomura, 1959a: 33 A: JA (Ryukyus)
brevicornis Sharp, 1888: 243 A: JA SC
montanus Grouvelle, 1913: 116 (Dryopidius) A: AP
nepalensis Satô, 1981: 52 A: NP
genus Helichus Erichson, 1847a: 510 type species Elmis lithophila Germar, 1824
crenulatus Kodada & Jäch, 1995: 333 A: YUN
haraldi Kodada & Jäch, 1995: 332 A: YUN
lareynioides G.C. Champion, 1924b: 235 A: UP YUN
tenuis G.C. Champion, 1924b: 235 A: UP
ussuriensis Lafer, 1980: 47 A: FE JA LIA
hasegawai Satô, 1985: 53

genus Pachyparnus Fairmaire, 1889b: 338 type species Pachyparnus tonkineus Fairmaire, 1888
dicksoni C.O. Waterhouse, 1878: 491 (Dryops) A: TAI
formosanus Bollow, 1940b: 55 (Helichus) A: TAI
gressitti Hinton, 1936: 164 (Helichus) A: TAI
indicus C.O. Waterhouse,1876: 19 (Parygrus) A: TAI ORR
konoi Bollow, 1940b: 52 (Helichus) A: TAI
whiteheadi C.O. Waterhouse, 1900b: 310 (Parygrus) A: HAI
genus Parahelichus Löbl & Smetana, 2006: 61 type species Pomatinus angulicollis Reitter, 1887
angulicollis Reitter, 1887c: 257 (Pomatinus) E: AB GG ST A: AF IN IQ TM TR
fenyesi Reitter, 1894h: 313 (Dryops) N: EG AFR
hintoni Bollow, 1940b: 32 (Helichus) E: AB
koltzei Bollow, 1940b: 34 (Helichus) A: XIN
genus Pomatinus Sturm, 1853: 62 type species Parnus substriatus P.W.J. Müller, 1806
substriatus P.W.J. Müller, 1806b: 219 (Parnus) E: AL AU BE BH BU CR CZ FR GB GE GR HU IT LT LU NL PL RO
SB SK SL SP ST SV SZ UK “Caucasus” N: AG MO TU A: IS LE SY TR “Turkestan”
dumerilii Latreille, 1807: 56 (Dryops)
longipes G. Redtenbacher, 1842: 14 (Parnus)

genus Praehelichus Löbl & Smetana, 2006: 61 type species Dryops sericatus C.O. Waterhouse, 1881
asiaticus Motschulsky, 1845a: 33 (Parnus) E: GG ST A: IN TM UZ XIN

607

LIMNICHIDAE
longus Solsky, 1876a: 285 (Dryops)

sericatus C.O. Waterhouse, 1881a: 410 (Dryops) A: BEI JIX LIA NMO SCH TIA XIN
sinensis Fairmaire, 1889b: 115 (Helichus)

solskyi Zaitzev, 1908: 417 (Dryops) [RN] E: ST “Caucasus” A: AF TM UZ
asiaticus Solsky, 1876a: 286 (Helichus) [HN]

genus Sostea Pascoe, 1860: 40 type species Sostea westwoodii Pascoe, 1860
Malaiseianus Bollow, 1940a: 6 type species Malaiseianus tuberculatus Bollow, 1940

diversa Pic, 1924a: 1 A: “Asia Minor”
genus Stenomystax Kodada, Jäch & Čiampor, 2003: 434 type species Stenomystax depressus Kodada, Jäch &
Čiampor, 2003
jengi Kodada, Jäch & Čiampor, 2003: 450 A: TAI
saetosus Kodada, Jäch & Čiampor, 2003: 445 A: HAI

family LIMNICHIDAE Erichson, 1846
subfamily Limnichinae Erichson, 1846

genus Bothriophorus Mulsant & Rey, 1852: 19 type species Bothriophorus atomus Mulsant & Rey, 1852
atomus Mulsant & Rey, 1852: 20 E: FR IT NL SP N: EG A: TR
venetus Ferrari, 1864: 108

genus Byrrhinus Motschulsky, 1858d: 50 type species Byrrhinus latus Motschulsky, 1858
Byrrhininus Pic, 1922a: 6 type species Byrrhinus gracilicornis Pic, 1922
Cyphonichus Sharp, 1902b: 679 type species Cyphonichus panamensis Sharp, 1902
Notiocyphon Blackburn, 1896: 272 type species Notiocyphon convexus Blackburm, 1896
Pelocherops Pic, 1923b: 3 type species Pelocherops atrolucidus Pic, 1923

flavicornis G.C. Champion, 1923b: 251 A: SD
formosanus Pic, 1935: 2 A: TAI
helicophallus Hernando & Ribera, 2014a: 180 A: YE
hirsutulus G.C. Champion, 1923b: 250 A: SD UP ORR
inlineatus Pic, 1923e: 15 A: “India”
marginatus G.C. Champion, 1923b: 249 A: UP
monilicornis G.C. Champion, 1923b: 251 A: UP
peguensis Pic, 1923e: 15 A: “India”
sikkimensis G.C. Champion, 1923b: 249 A: SD
socotrensis Hernando & Ribera, 2014a: 182 A: YE (Suqutra)
genus Caccothryptus Sharp, 1902a: 63 type species Caccothryptus compactus Sharp, 1902
Macrobyrrhinus Pic, 1922a: 4 type species Macrobyrrhinus rouyeri Pic, 1922

chayuensis Yoshitomi, 2015a: 10 A: XIZ
fujianensis Hernando & Ribera, 2014b: 291 A: FUJ

608

LIMNICHIDAE

maculosus Pic, 1923b: 3 (Macrobyrrhinus) A: HAI ORR
nepalensis Hernando & Ribera, 2014b: 292 A: NP
orion Yoshitomi, 2015a: 6 A: JA (Okinawa)
ripicola G.C. Champion, 1923b: 223 A: NP UP
sinensis Hernando & Ribera, 2014b: 294 A: FUJ
taiwanus Yoshitomi, 2015a: 3 A: TAI
testudo G.C. Champion, 1923b: 222 A: NP UP ORR
tibetanus Yoshitomi, 2015a: 7 A: XIZ
genus Chibidoronus Satô, 1966b: 60 type species Chibidoronus aureus Satô, 1966
aureus Satô, 1966b: 60 A: JA (Ryukyus)
genus Limnichomorphus Pic, 1922a: 4 type species Limnichomorphus curtulus Pic, 1922
ohbayashii Satô, 1966b: 58 A: JA (Ryukyus)
puetzi Hernando & Ribera, 2004: 45 A: NP
genus Limnichus Dejean, 1821: 49 type species Byrrhus pygmaeus Sturm, 1807
Limnichus Latreille, 1829: 510 [HN] type species Byrrhus pygmaeus Sturm, 1807

angustulus J. Weise, 1877: 300 E: FR (Corse) IT (Sardegna) N: TU
arabicus Hernando & Ribera, 2014a: 185 A: YE
astruci Pic, 1922a: 3 A: “India”
aurosericeus Jacquelin du Val, 1857: 268 E: FR IT PT SP N: AG MO TU
leprieuri Perris, 1864: 282

cuneatus G.C. Champion, 1923b: 275 A: UP
fulvopubens Pic, 1922a: 3 A: CH
globosus G.C. Champion, 1923b: 276 A: UP
incanus Kiesenwetter, 1851b: 584 E: AL AU BU BY CZ FR GR HU IT PT SK SP SZ “Caucasus” N: MO A: “Turkestan”
inornatus J. Weise, 1877: 301 N: MO A: CY TR
latiusculus Reitter, 1901a: 72 A: KZ
lederi J. Weise, 1877: 302 E: ST “Caucasus”
lewisi Nakane, 1963b: 43 A: FE JA JIL
paululus Lewis, 1879a: 12

mendax J.R. Sahlberg, 1913: 31 A: TR
micraspis G.C. Champion, 1923b: 276 A: PA ORR
obliteratus G.C. Champion, 1923b: 275 A: UP
pruinosus Fleischer, 1909: 244 A: UZ
pumilio Obenberger, 1917a: 30 E: GR (Kérkira)
punctipennis Kraatz, 1858: 148 E: GR (Peloponnísos) IT (Sicilia) N: MO A: IS LE TR
pygmaeus Sturm, 1807: 114 (Byrrhus) E: AU BE BY CZ DE FI FR GB GE GG HU IT LA LT NL NT PL RO SK SP ST
SV SZ UK A: “Turkestan”
auricomus Reitter, 1878: 180

609

LIMNICHIDAE
minutus Rey, 1889b: 66
variegatus Guillebeau, 1886: 3

sericeus Duftschmid, 1825: 24 (Byrrhus) E: AU BE BY CT CZ EN FI FR GE HU IT LA LT NL NT PL PT RO SK SL
SP SV SZ N: AG MO TU A: TR WS
subchalybeus Reitter, 1881a: 85 E: AB
subnotatus G.C. Champion, 1923b: 273 A: UP
trachyformis Pic, 1922a: 3 A: SD UP ORR
turkestanicus Pic, 1905a: 97 A: UZ
unistriatus G.C. Champion, 1923b: 274 A: UP
genus Pelochares Mulsant & Rey, 1869b: 361 type species Pelochares emarginatus Mulsant & Rey, 1869
(=Limnichus versicolor Waltl, 1838)
acuminatus G.C. Champion, 1923b: 271 A: SD UP
murinus Baudi di Selve, 1870: 64 (Limnichus) E: GR “Caucasus” A: CY IS LE SY TR
oblongulus G.C. Champion, 1925: 263 A: UP
rugiventris G.C. Champion, 1923b: 270 A: UP ORR
ceylanicus Delève, 1971: 370

ryukyuensis Satô, 1966b: 57 A: JA (Ryukyus) TAI
sabaeanus Hernando & Ribera, 2014a: 178 A: JO YE
sinbad Hernando & Ribera, 2014a: 174 A: OM QA
versicolor Waltl, 1838: 274 (Limnichus) E: AB AU BE BU BY CZ FR GE HU IT LT NL PL PT RO SK SP ST SZ UK
N: EG MO A: CY TR
emarginatus Mulsant & Rey, 1869a: 161
opacus J. Weise, 1877: 299

genus Platypelochares G.C. Champion, 1923b: 271 type species Platypelochares trifidus G.C. Champion,
1923
Wooldridgeus Spangler, 1999: 181 type species Wooldridgeus perforatus Spangler, 1999

chinensis Yoshitomi, 2011: 386 A: FUJ
latimargo G.C. Champion, 1923: 272 A: UP ORR
genus Pseudothryptus Hernando & Ribera, 2005: 132 type species Caccothryptus multiseriatus G.C.
Champion, 1923
multiseriatus G.C. Champion, 1923b: 224 (Caccothryptus) A: SD ORR

subfamily Hyphalinae Britton, 1971

genus Hyphalus Britton, 1971: 88 type species Hyphalus insularis Britton, 1971
taekoae Satô, 1997: 110 A: JA (Ryukyus) TAI

subfamily Cephalobyrrhinae G.C. Champion, 1925

genus Cephalobyrrhus Pic, 1923c: 4 type species Cephalobyrrhus latus Pic, 1923

610

HETEROCERIDAE

bertiae Pütz, 1998c: 344 A: YUN
brevipalpis Pütz, 1998c: 346 A: YUN
emeishanensis Pütz, 1998c: 347 A: SCH
gibbicollis G.C. Champion, 1925: 175 A: UP
jaechi Pütz, 1998c: 351 A: SCH
japonicus G.C. Champion, 1925: 176 A: JA
jiangxiensis Pütz, 1998c: 354 A: JIX
jinggangshanensis Pütz, 1998c: 355 A: JIX
latus Pic, 1923c: 4 A: TAI
longipalpis Pütz, 1998c: 359 A: JIX
nepalensis Pütz, 1998c: 360 A: NP
schillhammeri Pütz, 1998c: 362 A: SCH
schuelkei Pütz, 1998c: 363 A: SCH
sichuanensis Pütz, 1998c: 365 A: SCH
subelongatus Pic, 1923c: 4 A: NP SD
genus Erichia Reitter, 1895b: 79 type species Erichia longicornis Reitter, 1895
Jaechobyrrhinus Pütz, 1991: 132 type species Jaechobyrrhinus amanosius Pütz, 1991

longicornis Reitter, 1895b: 80 A: TR
amanosius Pütz, 1991: 132 (Jaechobyrrhinus)

subfamily Thaumastodinae G.C. Champion, 1924

genus Acontosceles G.C. Champion, 1924a: 27 type species Acontosceles hydroporoides G.C. Champion, 1924
hydroporoides G.C. Champion, 1924a: 29 A: UP
siwalikensis Pütz, 2008: 314 A: NP
yorioi Satô, 1966b: 60 A: JA (Ryukyus) TAI
yunnanensis Pütz, 2008: 315 A: YUN
genus Babalimnichus Satô, 1994: 174 type species Babalimnichus taiwanus Satô, 1994
masamii Satô, 1994: 175 A: JA (Honshu, Ryukyus)
taiwanus Satô, 1994: 174 A: TAI

family HETEROCERIDAE W.S. Macleay, 1825
subfamily Heterocerinae W.S. Macleay, 1825
tribe Augylini Pacheco, 1964

genus Augyles Schiödte, 1866: 141 type species Heterocerus hispidulus Kiesenwetter, 1843
subgenus Augyles Schiödte, 1866: 141 type species Heterocerus hispidulus Kiesenwetter, 1843
Littorimus Gozis, 1885: 120 type species Scarabaeus flavidus P. Rossi, 1794
Taenheterocerus Kuwert, 1890a: 14 type species Heterocerus crinitus Kiesenwetter, 1850

albineus Reitter, 1901a: 66 (Heterocerus) A: UZ
andalusiacus Breit, 1916: 58 E: SP

HETEROCERIDAE

611

atratus Grouvelle, 1896a: 53 A: PA ORR
barborae Skalický, 2012: 311 A: HKG
bellus Grouvelle, 1911: 317 (Heterocerus) A: PA ORR
cantus W.V. Miller, 1995: 40 A: NP PA UP
crinitus Kiesenwetter, 1850: 224 (Heterocerus) E: AU CR GE HU IT NL RO SK SL SZ
dilutissimus Reitter, 1887b: 209 (Heterocerus) E: AB A: CH IN IS TD TM
euphraticus Kiesenwetter, 1843: 210 (Heterocerus) A: IN IQ TM
feae Grouvelle, 1896b: 51 (Heterocerus) A: NP PA UP ORR
variabilis Mamitza, 1933: 85 (Heterocerus)

flavidus P. Rossi, 1794: 79 (Scarabaeus) E: AB AL AR AU BH BU CR CT CZ FR GE GG GR HU IT KZ MD ME SL
SP ST SZ TR UK N: AG LB MO TU A: AE ES FE IN IQ KI LE PA SA SY TD TM UZ WS AFR
albipennis Kuwert, 1890b: 540 (Heterocerus)
campestris Motschulsky, 1854a: 17 (Heterocerus)
flavescens Schaufuss, 1861: 48 (Heterocerus)
minimus Kiesenwetter, 1843: 214 (Heterocerus)
minutus Kiesenwetter, 1835: 213 (Heterocerus)
pallescens Schaufuss, 1861: 48 (Heterocerus)

fornicatus Mamitza, 1933: 88 (Heterocerus) A: NP UP
gabriellae Mascagni, 1993: 3 (Littorimus) A: NP ORR
gracilis Motschulsky, 1858d: 25 (Heterocerus) A: UP
castaneus Mamitza, 1933: 91 (Heterocerus)
exiguus Mamitza, 1933: 90 (Heterocerus)

gravidus Kiesenwetter, 1850: 224 (Heterocerus) E: FR IT PT SP N: MO TU
curtinigripes Kuwert, 1890b: 533 (Heterocerus)
curtulus Fairmaire, 1868a: 477 (Heterocerus)
curtus Rosenhauer, 1856: 116 (Heterocerus)

grohmanni Mascagni, 1987: 51 (Heterocerus) A: NP UP ORR
haldwaniensis Skalický, 2003b: 298 A: UP
hartmanni Mascagni, 2003: 291 A: NP
hispidulus Kiesenwetter, 1843: 211 (Heterocerus) E: AU BE BH CR CZ DE FI FR GB GE HU IT KZ LA LT NL PL
RO SB SK SL SP ST SV SZ TR A: IQ IS TM TR UZ
holdhausi Mamitza, 1907: 28 (Heterocerus) A: FE NE
indicus Motschulsky, 1858d: 26 (Heterocerus) A: PA ORR
infuscatus Grouvelle, 1911a: 107 (Heterocerus) A: YUN
intermedius Kiesenwetter, 1843: 209 (Heterocerus) E: BU CT DE FI FR GE KZ LA LT NL PL SV A: ES ES FE
KZ MG
maritimus Motschulsky, 1845b: 353 (Heterocerus)
motschulskyi Reiche, 1879: 239 (Heterocerus)

interspidulus R. Charpentier, 1979: 233 (Heterocerus) A: MG SHA
ivojenisi Mascagni, 1995a: 8 (Littorimus) A: NP UP ORR
japonicus Kôno, 1931b: 3 (Heterocerus) A: JA NE

612

HETEROCERIDAE

kaalo Mascagni, 1995a: 4 (Littorimus) A: NP
kolibaci Skalický, 1998b: 91 A: NP
koreanus Skalický, 1994: 2 (Littorimus) A: SC
kulabensis Reitter, 1900e: 227 (Heterocerus) A: UZ
laticollis Mamitza, 1933: 87 (Heterocerus) A: NP UP ORR
martensi Mascagni, 1990: 107 (Heterocerus)

luciae Mascagni, 1993: 5 (Littorimus) A: XIZ ORR
manfredjaechi Mascagni, 1995a: 2 (Littorimus) A: CE HKG NP PA UP ORR
maritimus Guérin-Méneville, 1844b: 69 (Heterocerus) E: FR GB GE GR IR IT MA NL NT SP N: AG MO TU A:
SY TR
basalis Mamitza, 1929: 203 (Heterocerus)
britannicus Kuwert, 1890b: 546 (Heterocerus)
burchanensis Schneider, 1896: 177 (Heterocerus)
nanus Gené, 1836: 183 (Heterocerus)

marmota Kiesenwetter, 1850: 224 (Heterocerus) E: AR FR GR IT RO SP N: AG TU A: MG TR
funebris Schaufuss, 1863b: 117 (Heterocerus)
maritimus Kuwert, 1890b: 523 (Heterocerus)
unicolor Dufour, 1851: 333 (Heterocerus)

marshalli Mamitza, 1928: 107 (Heterocerus) A: NP PA “India” ORR
arrowi Mamitza, 1928: 108 (Heterocerus)
repexus W. V. Miller, 1995: 38
schawalleri Mascagni, 1990: 107 (Heterocerus)

mascagnii Skalický, 1994: 3 (Littorimus) A: SC
modicus W.V. Miller, 1995: 38 A: NP
muelleri Breit, 1916: 54 (Heterocerus) E: PT
nebulosus Kuwert, 1890b: 544 (Heterocerus) A: TM UZ
niloticus Grouvelle, 1896b: 54 (Heterocerus) N: AG EG MO A: SA YE AFR
obliteratus Kiesenwetter, 1843: 219 (Heterocerus) E: BU CT GG ST A: AF IN IQ KZ SY TM TR UZ YE
pruinosus Kiesenwetter, 1851a: 286 (Heterocerus) E: AU BH BU BY CR CZ ES FR GE GG GR HU IT KZ PL RO SK
SL SP SZ A: AF IN TM TR UZ
pucholti Skalický, 2001a: 53 A: NP PA UP ORR
robustus Mascagni, 2003: 287 A: HAI
royi Skalický, 2005: 134 A: PA
saano Mascagni, 1995a: 10 (Littorimus) A: NP UP ORR
scharlottae Skalický, 2005: 136 A: PA
schmidtjaegeri Mascagni, 1998: 4 A: NP
scutellatus Motschulsky, 1854a: 17 (Heterocerus) E: AB GG
vitticollis Reitter, 1884a: 3 (Heterocerus)

senescens Kiesenwetter, 1865: 368 (Heterocerus) E: FR NR PT SP SV N: AG MO A: JO
punctatus Brisout de Barneville, 1866: 373 (Heterocerus)

613

HETEROCERIDAE

sericans Kiesenwetter, 1843: 212 (Heterocerus) E: AL AU BU CR CT CZ DE FR GB GE HU IT NL PL RO SK SL SP
SV SZ UK A: IS SY
pusillus Stephens, 1835: 395 (Heterocerus)

sinensis Grouvelle, 1896b: 55 (Heterocerus) A: NO
siyo Mascagni, 1995a: 7 (Littorimus) A: NP PA
skalei Skalický, 2001c: 211 A: NP PA
skalickyi Mascagni, 1993: 2 (Littorimus) A: IS JO TM YE
stipebozici Mascagni & Rada, 2012: 307 A: PA
taiwanensis Mascagni, 1995b: 343 (Littorimus) A: TAI
terzanii Mascagni, 1995a: 5 (Littorimus) A: NP
tokejii Nomura, 1958: 45 (Heterocerus) A: JA
torretassoi Mamitza, 1937: 193 (Heterocerus) A: JO
turanicus Reitter, 1887d: 502 (Heterocerus) E: GG KZ N: AG A: AE IN IQ IS KZ SA SY TM TR UZ
ahngeri J.R. Sahlberg, 1900: 207 (Heterocerus)

uncis W.V. Miller, 1995: 40 A: NP PA
viklundi Mascagni, 2010: 11 A: SCH
weigeli Skalický, 2003b: 241 A: NP
subgenus Phyrites Schiödte, 1866: 159 type species Phyrites aureolus Schiödte, 1866
aureolus Schiödte, 1866: 159 (Phyrites) E: BE DE FI FR NL

tribe Heterocerini W.S. Macleay, 1825

genus Heterocerus Fabricius, 1792a: 262 type species Apate marginatus Fabricius, 1787
aragonicus Kiesenwetter, 1850: 223 E: FR GE IT PT SP N: AG MO TU
amoenus Reiche, 1879: 239
arragonicus Kiesenwetter, 1850: 223 [unused original spelling]
biskrensis Fairmaire, 1879c: 169
coxaepilus Kuwert, 1890b: 545
mendax Kuwert, 1890b: 545
pictus Mulsant & Rey, 1873: 433

astoni Skalický, 2010: 155 A: HKG
dayremi Peyerimhoff, 1921: 231 N: AG TU
dubius Fabricius, 1801: 356 A: PA ORR
fenestratus Thunberg, 1784: 3 (Dermestes) E: AL AR AU BE BH BU BY CR CT CZ FI FR GB GE GG GR HU IR IT
KZ LA LT MD ME NL NT PL PT RO SK SL SP ST SV SZ TR UK N: AG MO TU A: AF CE CY ES FE HKG IN IQ
IS JA JO KZ LE MG NC NE NO SA SC SE SY TAI TD TM TR UZ WS XIZ YE NAR ORR
asiaticus Nomura, 1958: 58
beckeri Kuwert, 1892: 311
completus Hatch, 1965: 13
confinis Rey, 1890a: 163
corsicus Kuwert, 1890b: 538

614

HETEROCERIDAE

croaticus Kuwert, 1890b: 538
laevigatus Panzer, 1794a: 12
lanatus Rey, 1890a: 163
latus Grouvelle, 1915: 134
marshami Stephens, 1829a: 101
molleri Varenius, 1891: 22
obscurus Rey, 1890a: 163
pusillus Waltl, 1839: 221
multimaculatus Motschulsky, 1854a: 18
siculus Kuwert, 1890b: 547
tonkinensis Grouvelle, 1896a: 5
tristis Mannerheim, 1853b: 218

flexuosus Stephens, 1828: 101 E: AB AL AU BU CR CT CZ DE FI FR GB GE GG GR HU IR IT KZ LT ME NL NR
RO SK SL SP ST SV SZ TR UK N: AG EG MO TU A: CY ES IN IQ IS JO KZ MG PA SA SY TD TM TR UZ
WS AFR
apfelbecki Kuwert, 1890b: 543
arenarius Kiesenwetter, 1851a: 284
corsicus Rey, 1890a: 163 [HN]
damryi Kuwert, 1890b: 542
dentifasciatus Kuwert, 1890b: 542
fausti Reitter, 1879: 545
femoralis Krynicki, 1832: 115
hamifer Gené, 1836: 182
heydeni Kuwert, 1890b: 543
maculosus Fairmaire, 1868a: 476
marginatus Gyllenhal, 1808: 137
reyi Heyden, 1906: 376 [RN]
senegalensis Fairmaire, 1894d: 652
syrticus Peyerimhoff, 1924: 159

fossor Kiesenwetter, 1843: 204 E: AU BE BU CT CZ FI FR GB GE GR HU IR IT KZ NL PT RO SK SP ST SV SZ A:
ES IN KZ TD TR WS
bifasciatus Küster, 1849a: 37
cincticollis Rey, 1890a: 163
parallelus Dufour, 1852: 456
rectus G.W. Waterhouse, 1859: 168

fusculus etruscus Mascagni, 1986: 41 E: IT (Toscana, Lazio)
fusculus fusculus Kiesenwetter, 1843: 220 E: AU BE BH BY CR CT CZ FI FR GB GE HU IT KZ LA LT MD ME NL
PL PT RO SK SL ST SV SZ UK A: ES IN KZ TR WS
oblongulus Kuwert, 1890b: 548
opacus Rey, 1890a: 163

HETEROCERIDAE

615

pulchellus Kiesenwetter, 1843: 220
pulchellus Kuwert, 1890b: 537
similis Kuwert, 1890b: 547

hankae Skalický, 1998a: 4 (Littorimus) A: IN
harteni Mascagni, 2009: 154 A: AE
holosericeus Rosenhauer, 1856: 114 E: CR FR IT PT SP N: AG MO TU A: IN PA TR UP
fluviatilis A. Fiori, 1906: 251
lineatus Kuwert, 1890b: 546
panormitanus Kuwert, 1890b: 545
pustulatus Schilsky, 1890: 177
pustulatus Schilsky, 1892: 254
ragusae Kuwert, 1890b: 546
subfossor A. Fiori, 1906: 259
villiger A. Fiori, 1906: 260
villosus A. Fiori, 1908: 239

humilis Guillebeau, 1896: 243 N: AG
guillebeaui Zaitzev, 1908: 319

infuscatus Grouvelle, 1911a: 107 A: CH
kaszabi R. Charpentier, 1979: 230 A: MG
kamtschaticus Egorov, 1989: 453 A: FE
lorenzevae Mascagni, 1993: 7 A: NP PA ORR
magnus Mamitza, 1933: 83 A: AE IN NP OM UP ORR
jelineki Skalický, 1998b: 7 (Heterocerus)

marginatus Fabricius, 1787a: 33 (Apate) E: AR AU BE BH BU CR CT CZ DE FI FR GB GE GG GR HU IR IT LA LT
NL NR PL RO SK SL SP ST SV SZ UK N: AG A: ES NC SC WS
scutellaris Guillebeau, 1897: 163
sulcatus Kuwert, 1890b: 544

medius R. Charpentier, 1965: 238 N: EG AFR
morgani Grouvelle, 1907: 506 A: IN
mus R. Charpentier, 1965: 257 A: AE AF IN KZ PA TD TM UZ AFR ORR
nepalensis Mascagni, 1993: 2 A: CH HKG NP PA TM ORR
obsoletus Curtis, 1828: 224 E: AB AU BE CR CT CZ DE FI FR GB GE GG GR HU IT KZ LA LT NL PL SK SP ST SV
SZ TR A: ES IN MG TD TM TR WS
marginatus Marsham, 1802: 400
quadrimaculatus Hochhuth, 1873a: 319

ottomerkli Skalický, 2001b: 143 N: TU
pallidivestis Guillebeau, 1893: 326 N: AG
major Pic, 1893b: 122

parallelus Gebler, 1830: 102 E: AU BU CT CZ FR GE GG GR HU IT KZ PL SK SP ST A: ES LE MG TM
UZ WS

616

PSEPHENIDAE

cornutus Motschulsky, 1854a: 15
hauseri Kuwert, 1893: 17
maxillosus Motschulsky, 1854a: 15
salinus Kiesenwetter, 1843: 204

persicus Mascagni, 1989: 2 N: TU A: IN IQ PA SA
philippensis javanicus Grouvelle, 1896: 3 A: NP PA UP ORR
parvulus Grouvelle, 1896: 4
fryi Grouvelle, 1896: 4
thieleni Mascagni, 1993: 9

philippensis philippensis Grouvelle, 1896: 6 A: SA ORR
sauteri Grouvelle, 1911a: 109 A: TAI
schatzmayri Mamitza, 1936: 233 N: TU
sichuanensis Mascagni, 2003: 287 A: SCH
stultus Grouvelle, 1915: 105 A: CH
thebaicus thebaicus Grouvelle, 1896b: 55 N: EG AFR
soudanicus Grouvelle, 1909: 134

virgatus Mamitza, 1933: 81 A: CH HKG NP PA ORR
fulvipes Miller, 1995: 41

wolfgangi Skalický, 2014: 306 A: SA

tribe Micilini Pacheco, 1964

genus Micilus Mulsant & Rey, 1872a: 15 type species Heterocerus murinus Kiesenwetter, 1843
Micromicilus J.R. Sahlberg, 1900: 205 type species Heterocerus minutissimus J.R. Sahlberg, 1900

minutissimus J.R. Sahlberg, 1900: 205 (Heterocerus) A: TD TM UP UZ
murinus Kiesenwetter, 1843: 22 (Heterocerus) E: AL AU BH CR FR GE HU IT SL SZ A: NP

family PSEPHENIDAE Lacordaire, 1854
subfamily Eubrianacinae Jakobson, 1913

genus Eubrianax Kiesenwetter, 1874: 246 type species Eubrianax ramicornis Kiesenwetter, 1874
Heibrianax Lee, Satô & P.-S. Yang, 1999a: 20 type species Eubrianax granicollis Lewis, 1895
Placonycha Horn, 1880: 111 type species Dicranopselaphus edwardsii LeConte, 1874

albomaculatus Pic, 1926a: 30 A: CH
amamiensis amamiensis Satô, 1965: 92 A: JA (Ryukyus)
amamiensis kimurai Lee, P.-S. Yang & Satô, 2001: 358 A: JA (Ryukyus)
brunneicornis Nakane, 1952b: 40 A: JA
granicollis Lewis, 1895b: 104 A: JA SC
ihai Chûjô & Satô, 1970: 14 A: JA (Ryukyus)
insularis Nakane, 1952b: 41 A: JA (Ryukyus: Yaku-shima)
loochooensis Nakane, 1952b: 40 A: JA (Ryukyus)
maai Lee, Satô & P.-S. Yang, 1999a: 21 (Heibrianax) A: FUJ
manakikikuse Satô, 1964b: 35 A: JA (Ryukyus) TAI

617

PSEPHENIDAE
wulaiensis Lee & P.-S. Yang, 1990: 80

meridianus Lee, Satô & P.-S. Yang, 1999a: 17 A: TAI
niger Lee & P.-S. Yang, 1990: 81 A: TAI
alishanensis Lee & P.-S. Yang, 1990: 83

nobuoi Satô, 1965: 92 A: JA (Ryukyus)
pellucidus Lewis, 1895b: 104 A: JA SCH
ramicornis Kiesenwetter, 1874: 247 A: FE JA NC SC
secretus Lee, Satô & P.-S. Yang, 1999a: 23 (Heibrianax) A: FUJ TAI
serratus Lee, P.-S. Yang & Satô, 2001: 358 A: TAI
sichuanus Lee, Satô & P.-S. Yang, 1999a: 21 (Heibrianax) A: SCH SHA
tarokoensis Lee & P.-S. Yang, 1990: 81 A: TAI
flavus Lee & P.-S. Yang, 1990: 82

genus Jaechanax Lee, Satô & P.-S. Yang, 2000b: 120 type species Eubrianax insignis Fairmaire, 1904
major Pic, 1913d: 172 (Eubrianax) A: NP ORR
limbatithorax Pic, 1923c: 9

genus Jinbrianax Lee, Satô &P.-S.Yang, 1999b: 170 type species Eubrianax javanus Pic, 1913
incompositus Lee, Satô & P.-S. Yang, 1999b: 181 A: NP
rotundatus Lee, Satô & P.-S. Yang, 1999b: 182 A: NP
tenuis Lee, Satô & P.-S. Yang, 1999b: 185 A: JIX
genus Odontanax Lee, Satô & P.-S.Yang, 2000a: 151 type species Eubrianax maculicollis Fairmaire, 1888
chinensis Lee, Satô & P.-S. Yang, 2000a: 156 A: HKG HUB JIX
dudgeoni Lee, Satô & P.-S. Yang, 2000a: 158 A: HKG
laosensis Pic, 1923c: 9 (Eubrianax) A: NP ORR

subfamily Eubriinae Lacordaire, 1857

genus Dicranopselaphus Guérin-Méneville, 1861: 531 type species Dicranopselaphus lesueurii GuérinMéneville, 1861
Alabameubria H. P. Brown, 1980: 188 type species Alabameubria starki H. P. Brown, 1980
Furcipalpus Guérin-Méneville, 1861: 533 type species Dicranopselaphus lesueurii Guérin-Méneville, 1861
Spineubria Nakane, 1952b: 39 type species Spineubria reticulata Nakane, 1952

bicolor Lee & P.-S. Yang, 1996: 190 A: YUN
gressitti Lee, P.-S. Yang & Satô, 2000: 564 A: HUB
jiangxiensis Lee & P.-S. Yang, 1996: 183 A: JIX
nantai Lee & P.-S. Yang, 1996: 183 A: TAI
taiwanus Lee & P.-S. Yang, 1996: 186

nepalensis Lee & P.-S. Yang, 1996: 194 A: NP SD
raii Lee, P.-S. Yang & Satô, 2000: 566 A: NP
reticulatus Nakane, 1952b: 39 (Spineubria) A: JA (Amami-Ôshima)

618

PSEPHENIDAE

rufus Pic, 1916e: 3 (Grammeubria) A: YUN ORR
yamasakii Lee & P.-S. Yang, 1996: 189
yasumatsui Chûjô & Satô, 1975: 319 (Spineubria)

sichuanensis Lee & P.-S. Yang, 1996: 194 A: SCH
similis Lee & P.-S. Yang, 1996: 185 A: FUJ HKG
spadix Lee, P.-S. Yang & Satô, 2000: 560 A: NP
testaceicornis Pic, 1923c: 10 (Grammeubria) A: SHN
genus Ectopria LeConte, 1854b: 351 type species Eubria thoracica Ziegler, 1845
Eurea LeConte, 1854b: 352 type species Eubria nervosa Melsheimer, 1845
Grammeubria Kiesenwetter, 1874: 248 type species Grammeubria opaca Kiesenwetter, 1874
Urumaeubria Satô, 1968: 140 type species Urumaeubria tachikawai Satô, 1968

chikuni Lee, P.-S. Yang & Satô, 1998: 297 A: ANH SCH
hsui Lee & P.-S. Yang, 1994b: 387 A: FUJ TAI
iranensis Lee & Jäch, 2007: 231 A: IN
lobata Lee, P.-S. Yang & Satô, 1998: 298 A: SCH
opaca opaca Kiesenwetter, 1874: 249 (Grammeubria) A: JA
nitidula Kiesenwetter, 1874: 249
testaceipes Pic, 1944b: 1 (Drupeus)

opaca tomokunii Lee, P.-S. Yang & Satô, 1998: 299 A: JA (Ryukyus)
tachikawai Satô, 1968: 142 (Urumaeubria) A: JA (Ryukyus)
genus Eubria Latreille, 1829: 462 type species Cyphon palustris Germar, 1818
palustris Germar, 1818: 238 (Cyphon) E: AU BE CR CZ DE FI FR GB GE GR HU IT LA LT NL PL RO RU SK SL SP
SZ UK YU A: WS
marchantiae Jacquelin du Val, 1854: xxxvii

genus Granuleubria Jäch & Lee, 1994: 225 type species Drupeus atriceps Pic, 1916
kurdistanica Jäch & Lee, 1994: 228 A: TR
marginata Pic, 1944b: 1 (Drupeus) A: UP
pakistanica Jäch & Lee, 1994: 228 A: IN NP PA
genus Homoeogenus C.O. Waterhouse, 1880: 565 type species Homoeogenus punctatum C.O. Waterhouse, 1880
chinense Lee & P.-S. Yang, 1993: 352 A: SCH
elongatum Lee & P.-S. Yang, 1995: 355 A: YUN
laurae Lee & P.-S. Yang, 1993: 353 A: TAI
maai Lee & P.-S. Yang, 1999: 8 A: FUJ
obscurum Pic, 1923c: 12 (Drupeus) A: SHN
punctatum C.O. Waterhouse, 1880: 565 A: CH
genus Macroeubria Pic, 1916e: 4 type species Macroeubria striatipennis Pic, 1916
bicolor Lee, P.-S. Yang & Satô, 1997: 139 A: FUJ

PSEPHENIDAE

619

diffusa Lee, P.-S. Yang & Satô, 1999: 183 A: BT NP SD YUN ORR
lewisi Nakane, 1952b: 37 A: JA
amamiana Satô, 1964b: 34

luei Lee, P.-S. Yang & Satô, 1997: 148 A: TAI
okinawana Lee, P.-S. Yang & Satô, 1997: 145 A: JA (Ryukyus)
quadrimaculata Lee, P.-S. Yang & Satô, 1997: 140 A: GUA HKG
similis Lee, P.-S. Yang & Satô, 1997: 141 A: JA
taiwana Lee, P.-S. Yang & Satô, 1997: 152 A: TAI
testacea Pic, 1954: 64 A: FUJ
genus Microeubria Lee & P.-S. Yang, 1994a: 325 type species Microeubria jaechi Lee & Yang, 1994
minima G.C. Champion, 1924c: 250 (Eubria) A: SD UP
reticulata Lee & Jäch, 1996: 49 A: NP SD
semistriata G.C. Champion, 1921: 204 (Eubria) A: SD UP
wittmeri Lee & P.-S. Yang, 1998: 371 A: SD
genus Nipponeubria Lee & Satô, 1996: 123 type species Nipponeubria yoshitomii Lee & Satô, 1996
yoshitomii Lee & Satô, 1996: 126 A: JA
genus Schinostethus C.O. Waterhouse, 1880: 563 type species Schinostethus nigricornis C.O. Waterhouse,
1880
subgenus Schinostethus C.O. Waterhouse, 1880: 563 type species Schinostethus nigricornis C.O. Waterhouse,
1880
Cophaesthetus C.O. Waterhouse, 1880: 566 type species Cophaesthetus opacus C.O. Waterhouse, 1880
Drupeubria Nakane, 1952b: 37 type species Drupeus brevis Lewis, 1895

brevis Lewis, 1895b: 103 (Drupeus) A: FUJ JA
takeuchii Nakane, 1952b: 37 (Drupeubria)

jii Lee, Jäch & P.-S. Yang, 1998: 307 A: GUA GUX SCH
medius Lee, Jäch & P.-S. Yang, 1998: 308 A: NP
niger Lee, P.-S. Yang & H.P. Brown, 1993: 687 A: NP SD ORR
nigricornis C.O. Waterhouse, 1880: 564 A: BT FUJ GUA HKG KA NP SD UP YUN ORR
diversipes Pic, 1923c: 10 (Grammeubria)
indicus Pic, 1916e: 3 (Drupeus)

satoi Lee, P.-S. Yang & H.P. Brown, 1993: 688 A: TAI
junghuaensis Lee, P.-S.Yang & H.P. Brown, 1993: 692

sichuanensis Lee, Jäch & P.-S. Yang, 1998: 312 A: SCH
subgenus Sundodrupeus Pic, 1916e: 4 type species Drupeus pruinosus Pic, 1916
nepalensis Lee, Jäch & P.-S. Yang, 1998: 315 A: NP
notatithorax Pic, 1923c: 12 (Drupeus) A: YUN ORR
theresae Pic, 1944b: 1 (Drupeus)

transversus Lee & Jäch, 2007: 239 A: UP

620

PSEPHENIDAE
subfamily Psepheninae Lacordaire, 1854

genus Mataeophephus C.O. Waterhouse, 1876: 16 type species Mataeopsephus nitidipennis C.O. Waterhouse,
1876
Betelmis Matsumura, 1915: 214 type species Betelmis japonicus Matsumura, 1915
Sinopsephenus Nakane, 1964c: 50 type species Psephenus chinensis Nakane, 1964

chinensis Nakane, 1964c: 50 (Psephenus) A: FUJ GUA HKG
dentatus Lee, Jäch & Satô, 2003: 486 A: GUI JIX
esakii Nakane, 1964c: 49 A: TAI
japonicus Matsumura, 1915: 214 (Betelmis) A: FE HUB JA LIA NC SC
coreanicus Delève, 1967: 414
granulicollis Hinton, 1935: 169
sasajii Satô, 1970: 28

maculatus Nomura, 1957a: 2 A: JA
minimus Lee, Jäch & Satô, 2003: 489 A: YUN
nitidipennis C.O. Waterhouse, 1876: 17 A: GUI HKG
quadribranchiae Lee, Jäch & Satô, 2003: 490 A: TAI
sichuanensis Lee, Jäch & Satô, 2003: 491 A: HUB SCH
taiwanicus Lee, P.-S. Yang & H. P. Brown, 1990: 74 A: JA (Ryukyus) TAI
tenuipes G.C. Champion, 1921: 202 (Psephenus) A: UP
vietnamensis Lee, Jäch & Satô, 2003: 494 A: GUI ORR

subfamily Psephenoidinae Bollow, 1938

genus Afropsephenoides Basilewsky, 1959: 30 type species Afropsephenoides marlieri Basilewsky, 1959
volatilis G.C. Champion, 1924c: 251 (Psephenoides) A: NP PA UP
genus Malacopsephenoides Jeng & Satô, 2006b: 112 type species Psephenoides japonicus Masuda, 1935
eureka Jeng & Jäch, 2006b: 113 A: SCH
japonicus Masuda, 1935: 9 (Psephenoides) A: JA SC SHA
genus Micreubrianax Pic, 1922d: 5 type species Eubrianax (Micreubrianax) flabellicornis Pic, 1922d
nepalensis Jeng & P.-S. Yang, 2006a: 71 A: NP
genus Psephenoides Gahan, 1914: 189 type species Psephenoides immsi Gahan, 1914
fluviatilis J. Yang, 1995: 111 A: ZHE
gahani G.C. Champion, 1920: 194 A: NP UP ORR
immsi Gahan, 1914: 189 A: BT NP UP
magnioculus J. Yang, 1993: 374 [= J. Yang, 2002: 824] A: FUJ HKG
rajouri Chowdhari, 1977: 50 A: KA
subopacus Pic, 1954a: 64 (Micreubrianax) A: FUJ JA (Ryukyus) NP TAI UP ZHE
wuhongi J. Yang, 1997: 231

621

PTILODACTYLIDAE

genus Sinopsephenoides J. Yang, 1993: 374 [=J. Yang, 2002: 825] type species Sinopsephenoides filitarsus J.
Yang, 1993
Nematopsephus Jäch & Jeng, 1995: 159 type species Nematopsephus malickyi Jäch & Jeng, 1995

filitarsus J. Yang, 1993: 375 [=J. Yang, 2002: 825] A: FUJ ORR

family PTILODACTYLIDAE Laporte, 1838
subfamily Anchytarsinae G.C. Champion, 1897

genus Anchycteis Horn, 1880: 87 type species Anchycteis velutina Horn, 1880
brunneicornis Lewis, 1895b: 100 (Epilichas) A: JA
usori Nakane, 1958: 91 (Epilichas)

miyatakei Nakane, 1952b: 36 (Epilichas) A: JA
monticola Nakane, 1952b: 36 (Epilichas) A: JA
brunneipennis Nakane, 1963b: 42

genus Epilichas White, 1859a: 290 type species Paralichas candezii White, 1859
atricolor Lewis, 1895b: 101 A: JA
candezii White, 1859a: 289 (Paralichas) A: CH
testaceipes Pic, 1927d: 132

flabellata amamiana Nakane, 1963b: 42 A: JA (Ryukyus)
flabellata flabellata Kiesenwetter, 1874: 242 (Octoglossa) A: JA
flabellata muneaka Nakane, 1963b: 42 A: JA (Ryukyus)
flabellata mutsuensis Nakane, 1958: 90 A: JA
flabellata okinawana Satô, 1968: 135 A: JA (Ryukyus)
flabellata rubricollis Nakane, 1952b: 35 A: JA
flabellata tamaii Satô, 1964b: 33 A: JA (Ryukyus)
flabellata taniuchiensis Nakane, 1985b: 153 A: JA
otsukai Nakane, 1985b: 153 A: JA
taiwanensis Nakane, 1996: 129 A: TAI
yakushimensis iriomotensis Satô, 1968: 132 A: JA (Ryukyus)
yakushimensis oshimana Nakane, 1952b: 35 A: JA (Ryukyus)
yakushimensis yakushimensis Nakane, 1952b: 35 A: JA (Yaku-shima)
genus Pseudoepilichas Armstrong & Nakane, 1956: 212 type species Epilichas niponica Lewis, 1895
niponica Lewis, 1895b: 101 (Epilichas) A: FE (Kunashir) JA
robustior Nakane, 1963b: 42 A: JA

subfamily Cladotominae Pic, 1914

genus Drupeus Lewis, 1895b: 102 type species Drupeus laetabilis Lewis, 1895
hygropetricus Lee, Lawrence & Satô, 2005: 381 A: TAI
laetabilis Lewis, 1895b: 102 A: JA
ogatai Yoshitomi & Hayashi, 2013: 10 A: JA (Ryukyus)
vittipennis Lewis, 1895b: 103 A: FE JA

622

PTILODACTYLIDAE

genus Paralichas White, 1859a: 287 type species Paralichas guerinii White, 1859
Eucteis Guérin-Méneville, 1861: 539 type species Eucteis bimaculata Guérin-Méneville, 1861
Odontonyx Guérin-Méneville, 1843c: 194 type species Dasytes trivittis Germar, 1824

bicoloripes Pic, 1911b: 143 (Eucteis) A: TAI
guerinii White, 1859a: 287 A: SHG
bimaculata Guérin-Méneville, 1861: 539 (Eucteis)
disconiger Pic, 1939a: 3
diverselineata Pic, 1939a: 3
juncta Pic, 1939a: 3
submaculata Pic, 1939a: 3

griseolineata Pic, 1939a: 3 A: [distribution unknown, probably CH]
higoniae gyotokui Nakane, 1963b: 42 A: JA
higoniae higoniae Lewis, 1895b: 99 A: JA
pectinata Kiesenwetter, 1874: 242 (Odontonyx) A: CH JA
rufolimbata Pic, 1886: 336 (Eucteis) A: JIX
apicicornis Pic, 1939a: 3

striolata Fairmaire, 1897f: 249 A: SCH
subnitida Pic, 1911b: 143 (Eucteis) A: CH
genus Pseudocladotoma Pic, 1918a: 3 type species Pseudocladotoma maculata Pic, 1918
piceiceps Pic, 1930b: 271 (Paralichas) A: GUI
vitaticollis Pic, 1930b: 271 (Epilichas) A: GUI

subfamily Ptilodactylinae Laporte, 1838

genus Microdrupeus Nakane, 1993b: 4 type species Microdrupeus insularis Nakane, 1993
insularis Nakane, 1993b: 5 A: JA (Yaku-shima)
genus Ptilodactyla Illiger, 1807b: 342 type species Ptilodactyla elaterina Illiger, 1807 (=Pyrochroa nitida De
Geer, 1775)
Falsodaemon Pic, 1913a: 4 type species Falsodaemon madecassus Pic, 1913
Hypselothorax Kirsch, 1866: 188 type species Ptilodactyla cruciata Kirsch, 1866
Stenactyla Fairmaire, 1896c: 346 type species Stenactyla pallipes Fairmaire, 1896
Theriomorphus Pic, 1913a: 1 type species Theriomorphus niasensis Pic, 1913

amamioshimana Nakane, 1963b: 43 A: JA (Ryukyus)
angustatipennis Pic, 1916f: 4 A: HP ORR
ardua Satô, 1979: 61 A: HP
baroniurbanii Satô, 1979: 65 A: NP
bhutanica Satô, 1979: 66 A: BT
brancuccii brancuccii Satô, 1979: 57 A: NP
brancuccii darjeelingiana Satô, 1979: 58 A: HP
diversepunctata Pic, 1924b: 9 A: CH

623

CHELONARIIDAE
exotica Chapin, 1927: 246 Ei: CZ FR GB GE IT SK SZ AFR NAR ORR
formosana Nakane, 1996: 130 A: TAI
furcata Pic, 1954b: 65 A: FUJ
godavarensis Satô, 1979: 63 A: NP
ishigakiana Satô, 1968: 138 A: JA (Ryukyus)
japonensis Nakane, 1977a: 87 [RN] A: JA
macrophtalma Nakane, 1963b: 43 [HN]

jiriensis Satô, 1979: 59 A: NP
klapperichi Pic, 1954b: 65 A: CH
miscella Satô, 1979: 60 A: BT
multifaria Satô, 1979: 67 A: NP
nitidissima Pic, 1927c: 249 A: TAI
ramea Lewis, 1895b: 107 A: JA
satoi Stribling, 2006: 61 [RN] A: BT
bicolor Satô, 1979: 61 [HN]

sinensis sauteri Pic, 1927c: 249 A: TAI
sinensis sinensis Pic, 1923c: 11 A: CH
takahashii Satô, 1968: 136 A: JA (Ryukyus)
wittmeri Satô, 1979: 64 A: BT

family CHELONARIIDAE C.É. Blanchard, 1845

genus Chelonarium Fabricius, 1801a: 101 type species Chelonarium atrum Fabricius, 1801
indicum Grouvelle, 1894: 464 A: TAI ORR
obscuripenne Pic, 1917a: 3 A: TAI
ohbayashii Satô, 1964b: 36 A: JA (Ryukyus)
vartianae Mandl, 1967a: 443 A: AF
genus Pseudochelonarium Pic, 1916d: 1 type species Pseudochelonarium hirsutum Pic, 1916
subgenus Pseudochelonarium Pic, 1916d: 1 type species Pseudochelonarium hirsutum Pic, 1916
assamensis Méquignon, 1935: 68 A: HP
aureovestitum Méquignon, 1935: 68 A: HP
dohertyi Méquignon, 1935: 68 A: BT ORR
japonicum Nakane, 1963b: 43 (Chelonarium) A: JA
maculosum Pic, 1927a: 10 A: HAI ORR
subhirtum Méquignon, 1935: 68 A: HP
yakushimanum Nakane, 1963b: 43 (Chelonarium) A: JA

family EULICHADIDAE Crowson, 1973

genus Eulichas Jakobson, 1913: 727 [RN] type species Lichas funebris Westwood, 1853
subgenus Eulichas Jakobson, 1913: 727 [RN] type species Lichas funebris Westwood, 1853
Lichas Westwood, 1853: 237 [HN] type species Lichas funebris Westwood, 1853

624

CALLIRHIPIDAE

alesbezdeki Hájek, 2009: 6 A: YUN ORR
bertiae Jäch, 1995: 374 A: FUJ GUI HUB HUN
dudgeoni Jäch, 1995: 372 A: FUJ GUA GUX HKG HUB JIX SCH SHA ZHE “North China”
funebris Westwood, 1853: 238 (Lichas) A: FUJ GUA GUX HKG JIA ZHE
davidis Deyrolle & Fairmaire, 1878: 111 (Lichas)
impressicollis Pic, 1939a: 2

gigantea Fairmaire, 1891e: cxxviii (Lichas) A: HP
jendeki Jäch, 1995: 373 A: YUN ORR
siamensis Hájek, 2009: 20 A: YUN ORR
sikkimensis Pic, 1913b: 108 (Lichas) A: NP SD UP
seillierei Pic, 1915b: 7 (Lycas)

similis Hájek, 2009: 21 A: YUN ORR
tenuicornis Jäch, 1995: 369 A: HAI
tonkinensis Jäch, 1995: 366 A: GUX ORR
trapezicollis Fairmaire, 1891e: cxxix (Lichas) A: SD
undulata Pic, 1911b: 142 (Lichas) A: YUN ORR
uniformis Pic, 1911b: 142 (Lichas) A: NP SD UP ORR
wewalkai Hájek, 2009: 36 A: NP

family CALLIRHIPIDAE Emden, 1924

genus Callirhipis Latreille, 1829: 459 type species Callirhipis dejeanii Latreille, 1829
subgenus Callirhipis Latreille, 1829: 459 type species Callirhipis dejeanii Latreille, 1829
dissimilis C.O. Waterhouse, 1877d: 380 A: FUJ ORR
maculosa Fairmaire, 1887c: 146
pinguis Fairmaire, 1887c: 151

formosana Pic, 1912: 5 A: TAI
kojimai Nakane, 1996: 132 A: TAI
kurosawai Satô, 1995: 17 A: JA (Ryukyus)
miwai Nakane, 1985: 36 A: JA (Ryukyus)
robusta indica Emden, 1936: 154 A: NP UP ORR
subgenus Ennometidium Emden, 1929: 118 type species Ennometes horni Emden, 1924
horni Emden, 1924a: 30 (Ennometes) A: TAI
genus Ennometes Pascoe, 1866: 445 type species Ennometes lacordairei Pascoe, 1866 (=Callirhipis ruficornis
Gray, 1832)
incertus Emden, 1936: 151 (Parennometes) A: UP
genus Simianus C.É. Blanchard, 1853: 52 type species Callirhipis bicolor C.É. Blanchard, 1845 (=Callirhipis
separata Gemminger, 1869)
Homoeorhipis Fairmaire, 1887c: 152 type species Homoeorhipis mesomelaena Fairmaire, 1887
Horatocera Lewis, 1895a: 35 type species Horatocera niponica Lewis, 1895

CALLIRHIPIDAE
Simianellus Emden, 1924a: 33 type species Simianellus melanocephalus Emden, 1924

cyaneicollis C.O. Waterhouse, 1877d: 391 (Callirhipis) A: NP SD ORR
globicollis Emden, 1924b: 94 (Simianellus) A: NP SD
inapicalis Pic, 1925: 4 A: XIZ
melanocephalus Emden, 1924a: 33 (Simianellus) A: TAI
similis Miwa, 1928: 373 (Horatocera)

niponicus Lewis, 1895a: 36 (Horatocera) A: JA
galloisi Pic, 1932: 1 (Horatocera) A: JA

obscurus sikkimensis Emden, 1932: 200 (Simianellus) A: SD
oshimanus Nakane, 1973: 177 (Horatocera) A: JA (Ryukyus)
rubricollis Pic, 1916c: 8 (Horatocera) A: XIZ

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Balthasar V. 1936a: Neue Aphodius-Arten aus China und Syrien. Entomologisches Nachrichtenblatt 10: 1–6.
Balthasar V. 1936b: Einige neue palaearktische Scarabaeiden. Entomologische Blätter 32: 94–98.
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Balthasar V. 1936d: Monographie der Subfam. Troginae der palaearktischen Region. Festschrift für Prof.
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Balthasar V. 1937: Neue coprophage Scarabaeiden von der Ukerewe-Inseln in Ost-Afrika. Arbeiten über
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Balthasar V. 1938a: Neue palaearktische Aphodius-Arten. Časopis Československé Společnosti Entomologické
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Keith D. 2006f: Sur le genre Tocama Reitter, 1902 (Col. Scarabaeoidea Melolonthidae Melolonthinae).
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Keith D. 2007b: Description de nouvelles espèces et d’un nouveau genre de Melolonthidae asiatiques
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Keith D. & Li J. 2009: Lamellicornes Pleurosticti du mont Wushizan (île d’Hainan, République populaire de
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Keith D. & Li J. 2013: Description d’une nouvelle espèce du genre Bunbunius Nomura, 1970 du Xizang,
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Keith D. & Miessen G. 2009: Une nouvelle espèce de Maladera de Chypre (Coleoptera, Scarabaeoidea,
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Keith D. & Montreuil O. 2014: Révision des genres Otoclinius Brenske 1896, Falsotoclinius n. gen. et
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Keith D. & Moretto P. 2008: Un nouvelle espèce du genre Onitis d’Arabie Saoudite (Coleoptera, Scarabaeoidea,
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Obenberger J. 1929e: Buprestidarum supplementa palaearctica V. Časopis Československé Společnosti
Entomologické 26: 59–63.
Obenberger J. 1929f: Insecta Baumiana I. 1. Buprestidae. Acta Entomologica Musaei Nationalis Pragae 7:
107–119.
Obenberger J. 1929g: Revision der Sphenopteren-Untergattung Deudora B. Jakovlev. Coleopterologisches
Centralblatt 4: 111–130.
Obenberger J. 1930a: Buprestidarum supplementa palaearctica VI. Časopis Československé Společnosti
Entomologické 27: 102–115.
Obenberger J. 1930b: Insecta Davidiana I. 1. Buprestidae. Časopis Československé Společnosti Entomologické 27:
116–118.
Obenberger J. 1930c: Buprestidae II. Pp. 215–568. In: Schenkling S. (ed.): Coleopterorum Catalogus. Pars 111.
Berlin: W. Junk, pp. 213–568. [Volumen XII.].
Obenberger J. 1930d: Revision der Sphenopteren-Untergattung Deudora B. Jakovlev. Coleopterologisches
Centralblatt 4: 233–255.
Obenberger J. 1931a: Révision des espèces du genre Agrilomorpha Théry (Col. Bupr.). Přehled druhů rodu
Agrilomorpha Théry (Col. Bupr.). Acta Entomologica Musaei Nationalis Pragae 9: 5–8.
Obenberger J. 1931b: Příspěvek k poznání javanských krasců. De nonnullis insulae Javae Buprestidarum
speciebus. Časopis Československé Společnosti Entomologické 28: 35–39.
Obenberger J. 1931c: O krascích z rodu Agrilus z příbuzenstva afrického druhu A. variegatus Klug. Sur les
Agrilus de l’Afrique de voisinage de l’A. variegatus Klug (Col. Bupr.). Acta Entomologica Musaei Nationalis
Pragae 9: 84–98.
Obenberger J. 1931d: Studien über die aethiopischen Buprestiden I. Folia Zoologica et Hydrobiologica 2:
175–201.
Obenberger J. 1931e: Specierum generis Meliboeus H. Deyrolle diagnoses praeliminares (Coleoptera,
Buprestidae). Folia Zoologica et Hydrobiologica 2 (2): 202–219.

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Obenberger J. 1932a: Neue Beiträge zur Kenntnis der orientalischen Prachtkäfer (Col., Bupr.). Folia Zoologica
et Hydrobiologica 4 (2): 205–221.
Obenberger J. 1932b: Elf neue Sambus-Arten (Col., Bupr.) aus der orientalischen Region. Folia Zoologica et
Hydrobiologica 4: 221–228.
Obenberger J. 1932c: Fauna Buruana. Coleoptera, Fam. Buprestidae. Treubia Supplement 7: 361–375.
Obenberger J. 1933a: Několik poznámek o krascích. Quelques notes et rectifications sur les buprestides (Col.).
Časopis Československé Společnosti Entomologické 30: 13–17.
Obenberger J. 1933b: Catalogue raisonné des buprestides de Bulgarie II. Partie. Bulletin des Institutions Royales
d’Histoire Naturelle (Sofia) 5 [1932]: 49–115.
Obenberger J. 1933c: O některých zajímavých krascích republiky československé (Col.). Sur quelques espèces
intéressantes de la famille des buprestides de la république tchécoslovaque. Časopis Československé
Společnosti Entomologické 30: 173–180.
Obenberger J. 1934a: O některých zajímavých krascích republiky československé (Col.). Sur quelques espèces
intéressantes de la famille des buprestides de la république tchécoslovaque (Dokončení). Časopis
Československé Společnosti Entomologické 21: 20–23.
Obenberger J. 1934b: Nový Coroebus z Číny (Col. Bupr.). Časopis Československé Společnosti Entomologické
31: 25.
Obenberger J. 1934c: Fragmenta buprestologica I. Časopis Československé Společnosti Entomologické 31: 38.
Obenberger J. 1934d: Nové druhy rodu Coroebus Cast. et Gory (Col. Bupr.). De generis Coroebus Cast. et Gory
speciebus novis. Časopis Československé Společnosti Entomologické 31: 39–44.
Obenberger J. 1934e: O rodu Coroebina Obenb. – De genere Coroebina Obenb. (Col. Bupr.). Časopis
Československé Společnosti Entomologické 31: 71–74.
Obenberger J. 1934f: Nový krasec z ČSR. Časopis Československé Společnosti Entomologické 31: 81.
Obenberger J. 1934g: Fragmenta buprestologica II. Časopis Československé Společnosti Entomologické 31: 90.
Obenberger J. 1934h: Studie o palaearktických druzích rodu Lampra (Col. Bupr.). Sur quelques rares espèces
paléarctiques du genre Lampra (Col. Bupr.). Acta Entomologica Musaei Nationalis Pragae 12: 105–115.
Obenberger J. 1934i: Nový čínský Coroebus (Col. Bupr.). Časopis Československé Společnosti Entomologické
31: 110.
Obenberger J. 1934j: Noví krasci z palaearktické oblasti (Col. Bupr.). Časopis Československé Společnosti
Entomologické 31: 112.
Obenberger J. 1934k: Noví palaearktičtí krasci (Col. Bupr.). Časopis Československé Společnosti Entomologické
31: 123.
Obenberger J. 1934l: Noví palaearktičtí Coroebové (Col. Bupr.). Časopis Československé Společnosti
Entomologické 31: 129.
Obenberger J. 1934m: Poznámky k rodu Anthaxia (Col. Bupr.). Časopis Československé Společnosti
Entomologické 31: 139.
Obenberger J. 1934n: Buprestidarum supplementa palaearctica VII. Časopis Československé Společnosti
Entomologické 31: 148–152.
Obenberger J. 1934o: Studien über die palaearktischen Buprestiden I. Folia Zoologica et Hydrobiologica 5:
158–290.

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Obenberger J. 1935a: Revise palaearktických druhů rodu Chalcophora Sol. Révision des espèces paléarctiques
du genre Chalcophora Solier (Col. Bupr.). Acta Entomologica Musaei Nationalis Pragae 13: 5–12.
Obenberger J. 1935b: Buprestidarum supplementa palaearctica VIII. Časopis Československé Společnosti
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Obenberger J. 1935c: Systematické a synonymické poznámky. I. Notes systématiques et synonymiques sur les
buprestides (Col. Bupr.). Časopis Československé Společnosti Entomologické 32: 20.
Obenberger J. 1935d: Catalogue raisonné des buprestides de Bulgarie. III Partie. Bulletin des Institutions
Royales d’Histoire Naturelle (Sofia) 8: 23–96.
Obenberger J. 1935e: Fragmenta buprestologica VI. Časopis Československé Společnosti Entomologické 32: 30.
Obenberger J. 1935f: Synonymia Agrilorum. (Col. Bupr.) I. Časopis Československé Společnosti Entomologické
32: 59.
Obenberger J. 1935g: Anthaxia plavilščikovi n. sp. (Col. Bupr.). Časopis Československé Společnosti
Entomologické 32: 83.
Obenberger J. 1935h: Nové druhy rodu Acmaeodera (Col. Bupr.). De generis Acmaeodera speciebus novis
(Col. Bupr.). Časopis Československé Společnosti Entomologické 32: 105–108.
Obenberger J. 1935i: Anthaxia aurulenta Fabr. (Col. Bupr.). Časopis Československé Společnosti Entomologické
32: 111.
Obenberger J. 1935j: Synonymia Agrilorum (Col. Bupr.) II. Časopis Československé Společnosti Entomologické
32: 121.
Obenberger J. 1935k: Fragmenta buprestologica X. Časopis Československé Společnosti Entomologické 32: 125.
Obenberger J. 1935l: Perotis orientalis var. lábleri n. Časopis Československé Společnosti Entomologické 32: 140.
Obenberger J. 1935m: De regionis palaearcticae generis Agrili speciebus novis (Col. Bupr.). O nových palaearktických druzích krasců z rodu Agrilus. Časopis Československé Společnosti Entomologické 32: 161–171.
Obenberger J. 1935n: Acmaeodera brancsiki n. sp. Časopis Československé Společnosti Entomologické 32: 178.
Obenberger J. 1935o: De generis Agrilis Africae speciebus novis (Col. Bupr.). Folia Zoologica et Hydrobiologica
8: 191–226.
Obenberger J. 1935p: Nové druhy palaearktických krasců z rodu Chrysobothris. De generis Chrysobothris
regionis palaearcticae speciebus novis (Col. Bupr.). Časopis Československé Společnosti Entomologické 32:
195–197.
Obenberger J. 1935q: Dvě nové formy rodu Acmaeodera (Col. Bupr.). Časopis Československé Společnosti
Entomologické 32: 202.
Obenberger J. 1935r: Nový krasec z Buchary (Col. Bupr.). Časopis Československé Společnosti Entomologické
32: 207.
Obenberger J. 1935s: Nový zajímavý druh rodu Acmaeodera z Číny (Col. Bupr.). Časopis Československé
Společnosti Entomologické 32: 211.

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Obenberger J. 1935t: Julodis aeneipes var. Štěrbai n. var. Časopis Československé Společnosti Entomologické 32:
215.
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ʼs-Gravenhage: W. Junk, pp. 783–934. [Volumen XII.].
Obenberger J. 1936a: Nové formy krasce Steraspis speciosa Klug (Col. Bupr.). Časopis Československé
Společnosti Entomologické 33: 5.
Obenberger J. 1936b: Oprava. Correction. Časopis Československé Společnosti Entomologické 33: 30.
Obenberger J. 1936c: De novis regionis orientalis Agrili generis speciebus (Col. Bupr.) II. Nové druhy orientálních krasců rodu Agrilus. Časopis Československé Společnosti Entomologické 33: 33–44.
Obenberger J. 1936d: Acmaeodera plavilščikovi sp. n. Časopis Československé Společnosti Entomologické
33: 46.
Obenberger J. 1936e: Sur les especes japonaises du genre Chrysobothris (Col. Bupr.). O japonských druzích
rodu Chrysobothris (Col. Bupr.). Časopis Československé Společnosti Entomologické 33: 70–71.
Obenberger J. 1936f: Acmaeodera aleši n. sp. Časopis Československé Společnosti Entomologické 33: 74.
Obenberger J. 1936g: Synonymia Agrilorum (Col. Bupr.) II. Časopis Československé Společnosti Entomologické
33: 91–92.
Obenberger J. 1936h: De speciebus novis palaearcticis generis Agrilus (Col. Bupr.). Nové palaearktické druhy
rodu Agrilus (Col. Bupr.). Časopis Československé Společnosti Entomologické 33: 104–118.
Obenberger J. 1936i: Nové druhy krasců z rodu Agrilus z Japonska (Col. Bupr.). De novis Japoniae generis
Agrili speciebus (Col. Bupr.). Časopis Československé Společnosti Entomologické 33: 140–141.
Obenberger J. 1936j: Tři nové zajímavé formy palaearktických krasců (Col. Bupr.). De novis generis
Chalcophorella formis (Col. Bupr.). Časopis Československé Společnosti Entomologické 33: 175.
Obenberger J. 1936k: Nová Acmaeodera z Nubie (Col. Bupr.). Časopis Československé Společnosti Entomologické
33: 185.
Obenberger J. 1936l: Buprestidae V. Pars 152. Pp. 935–1246. In: Junk W. & Schenkling S. (eds) 1936–1937:
Coleopterorum Catalogus. Volumen XIII. Gravenhage: W. Junk, Verlag für Naturwissenschaften,
pp. 935–1714.
Obenberger J. 1937a: Nový druh Anthaxie ze střední Asie (Col. Bupr.). Časopis Československé Společnosti
Entomologické 34: 13.
Obenberger J. 1937b: De novis generis Trachys speciebus orientalibus (Col. Bupr.). Nové orientální druhy rodu
Trachys (Col. Bupr.). Acta Entomologica Musaei Nationalis Pragae 15: 35–39.
Obenberger J. 1937c: Nové druhy rodu Trachys z palaearktické oblasti (Col. Bupr.). De novis generis Trachys
regionis palaearcticae speciebus (Col. Bupr.). Acta Entomologica Musaei Nationalis Pragae 15: 42–45.
Obenberger J. 1937d: Révision des espèces exotiques du genre Trachys Fabr. du continent africain. Přehled
exotických druhů rodu Trachys africké pevniny. Acta Entomologica Musaei Nationalis Pragae 15: 46–101.
Obenberger J. 1937e: Dvě nové palaearktické formy rodu Agrilus (Col. Bupr.). Časopis Československé
Společnosti Entomologické 34: 87.
Obenberger J. 1937f: Nová palaearktická Sphenoptera (Bupr.). Časopis Československé Společnosti
Entomologické 34: 91.
Obenberger J. 1937g: Nová varieta Sphenoptera sancta Rtt. (Bupr.). Časopis Československé Společnosti
Entomologické 34: 92.

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Obenberger J. 1937h: Nové variety palaearktických Sphenopter (Col. Bupr.). Časopis Československé Společnosti
Entomologické 34: 110.
Obenberger J. 1937i: Buprestidae VI. Pp. 1249–1714. In: Schenkling S. (ed.): Coleopterorum Catalogus. Pars 157.
Verlag für Naturwissenschaften, ʼs-Gravenhage: W. Junk, pp. 1247–1714. [Volumen XIII.].
Obenberger J. 1938: Studie o druzích rodu Anthaxia Eschsch. (Col. Bupr.). Etudes sur les espèces du genre
Anthaxia Eschsch. (Col. Bupr.). Sborník Národního Musea v Praze, Zoologia No. 4 (1B): 171–249.
Obenberger J. 1939: Dva noví indičtí krasci. Časopis České Společnosti Entomologické 36: 35.
Obenberger J. 1940a: Studie o rodu Sternocera Eschsch. (Col. Bupr.) Ad nonnularum generis Sternocera
Eschsch. specierum cognitionem (Col. Bupr.). Acta Entomologica Musaei Nationalis Pragae 18: 5–55.
Obenberger J. 1940b: Agrilus olivicolor var. Nitrae n. var., Agrilus olivicolor ab. Richteri n. Časopis České
Společnosti Entomologické 37: 17.
Obenberger J. 1940c: Nová Acmaeodera ze Syrie (Col. Bupr.). Časopis České Společnosti Entomologické 37: 20.
Obenberger J. 1940d: Nové formy středoevropských Agrilů (Col. Bupr.). Časopis České Společnosti Entomologické
37: 23.
Obenberger J. 1940e: Agrilus fleischeri ssp. kurosawai n. Časopis České Společnosti Entomologické 37: 26.
Obenberger J. 1940f: Nové zajímavé druhy rodu Endelus H. Deyr. (Col. Bupr.). De novis generis Endelus H.
Deyr. speciebus (Col. Bupr.). Časopis České Společnosti Entomologické 37: 29–32.
Obenberger J. 1940g: Nové formy palaearktických krasců z rodu Trachys. De novis regionis palaearcticae
generis Trachys formis (Col. Bupr.). Časopis České Společnosti Entomologické 37: 38–44.
Obenberger J. 1940h: Anthaxia diadema var. shelkovnikovi n. var. Časopis České Společnosti Entomologické 37:
46.
Obenberger J. 1940i: Nový krasec z Ruska. Časopis České Společnosti Entomologické 37: 60.
Obenberger J. 1940j: Nové formy rodu Chrysobothris Eschsch. (Col. Bupr.). De generis Chrysobothris Eschsch.
formis novis (Col. Bupr.). Acta Entomologica Musaei Nationalis Pragae 18: 79–106.
Obenberger J. 1940k: Ad regionis palaearcticae Buprestidarum cognitionem additamenta. Studie o palaearktických krascích (Col. Bupr.). Sborník Národního Musea v Praze, Zoologia No. 3 (2B): 111–189.
Obenberger J. 1940l: Nové druhy palaearktických krasců (Col. Bupr.). De novis regionis palaearcticae
Buprestidis (Col. Bupr.). Časopis České Společnosti Entomologické 37: 113–116.
Obenberger J. 1940m: Note su alcune Buprestidi dell’ A.O.I. conservate al Museo di Trieste. Atti del Museo
Civico di Storia Naturale di Trieste 14: 251–273.
Obenberger J. 1941: Revision der palaearktischen Buprestis-Arten (Col. Buprest.). Mitteilungen der Münchener
Entomologischen Gesellschaft 31: 460–554.
Obenberger J. 1942: De genere Chalcophorella Kerr. et generibus vicinis (Col. Bupr.). O rodu Chalcophorella
Kerr. a rodech příbuzných (Col. Bupr.). Časopis České Společnosti Entomologické 39: 2–12.
Obenberger J. 1943a: Nová evropská anthaxie (Col. Bupr.). De nova Europae specie generis Anthaxia Eschsch.
(Col. Bupr.). Časopis České Společnosti Entomologické 40: 58.
Obenberger J. 1943b: Palaearktické druhy podrodu Cypriacis Casey. De subgeneris Cypriacis Casey speciebus
palaearcticis. (Genus Buprestis L., Col. Bupr.). Časopis České Společnosti Entomologické 40: 106–110.
Obenberger J. 1944a: Nové palaearktické druhy rodu Aphanisticus Latr. (Col. Bupr.). De novis regionis
palaearcticae generis Aphanisticus Latr. speciebus (Col. Bupr.). Acta Entomologica Musaei Nationalis
Pragae 21–22 [1943–1944]: 5–9.

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Obenberger J. 1944b: Nové palaearktické druhy rodu Endelus H. Deyrolle (Col. Bupr.). De generis Endelus H.
Deyr. speciebus palaearcticis novis (Col. Bupr.). Časopis České Společnosti Entomologické 41: 42–44.
Obenberger J. 1944c: Nové palaearktické druhy rodu Meliboeus H. Deyr. De novis regionis palaearcticae
generis Meliboeus H. Deyr. speciebus (Coleoptera, Buprestidae). Časopis České Společnosti Entomologické
41: 57–59.
Obenberger J. 1944d: Coraebina beatricis n. sp. Časopis České Společnosti Entomologické 41: 70.
Obenberger J. 1944e: Tři nové druhy rodu Coroebus Lap. (Col. Bupr.). De tribus generis Coroebus Lap. speciebus novis (Col. Bupr.). Časopis České Společnosti Entomologické 41: 119–122.
Obenberger J. 1944f: Anthaxia insulaecola n. sp. Časopis České Společnosti Entomologické 41: 134.
Obenberger J. 1944g: O některých palaearktických druzích rodu Melanophila Eschsch. (Col. Bupr.). De
­nonnullis generis Melanophila Eschsch. speciebus palaearcticis (Col. Bupr.). Acta Entomologica Musaei
Nationalis Pragae 21–22 [1943–1944]: 147–150.
Obenberger J. 1944h: De duabus Chinae generis Meliboeus speciebus novis (Col. Bupr.). Dva nové čínske
druhy rodu Meliboeus H. Deyr. (Col. Bupr.). Acta Entomologica Musaei Nationalis Pragae 21–22 [1943–
1944]: 304–305.
Obenberger J. 1944i: Nové orientální druhy rodu Meliboeus H. Deyr. (Col. Bupr.). De generis Meliboeus H.
Deyr. regionis orientalis speciebus novis (Col. Bupr.). Acta Entomologica Musaei Nationalis Pragae 21–22
[1943–1944]: 435–437.
Obenberger J. 1945: Studie o krascích ze skupiny Capnodis tenebricosa A. Oliv. Une étude sur les espèces du
groupe de Capnodis tenebricosa A. Oliv. (Col. Buprestidae). Acta Entomologica Musaei Nationalis Pragae
23: 5–22.
Obenberger J. 1946: Insecta Houškeana: Buprestidae (Col.). Acta Entomologica Musaei Nationalis Pragae 24:
5–38.
Obenberger J. 1947: De novis regionis orientalis et palaearcticae generis Paracylindromorphus Théry speciebus (Col. Bupr.). Nové palaearktické a orientální druhy rodu Paracylindromorphus Théry. Acta
Entomologica Musaei Nationalis Pragae 25: 139–143.
Obenberger J. 1950: Contribution à l’étude de l’Aïr. (Mission L. Chopard et A. Villiers.) Coléoptères Buprestidae.
Mémoires de l’Institut Français d’Afrique Noire, Zoologie 10: 191–198.
Obenberger J. 1952a: Druhy Sphenopter podrodu Paradeudora Obenb. (Col. Bupr.). Sur les sphénoptères du
sous-genre Paradeudora Obenb. (Col. Bupr.). Acta Entomologica Musaei Nationalis Pragae 26 (No. 348)
[1948–1950]: 1–5.
Obenberger J. 1952b: De subgeneris Chrysoblemma B. Jak. generis Sphenopterae Sol. speciebus novis
(Col. Buprestidae). Nové druhy podrodu Chrysoblemma B. Jak. rodu Sphenoptera Sol. (Col. Buprestidae).
Acta Entomologica Musaei Nationalis Pragae 26 (No. 350) [1948–1950]: 1–5.
Obenberger J. 1952c: De subgeneris Deudora B. Jak. Sphenopterarum speciebus novis (Col. Buprestidae).
O nových druzích Sphenopter podrodu Deudora B. Jak. (Col. Buprestidae). Acta Entomologica Musaei
Nationalis Pragae 26 (No. 357) [1948–1950]: 1–23.
Obenberger J. 1952d: Monografie Sphenopter podrodu Chilostetha B. Jak. (Col. Buprestidae). Monographie
des Sphénoptères du sous-genre Chilostetha B. Jak. (Col Buprestidae). Acta Entomologica Musaei
Nationalis Pragae 26 No. 359 [1948–1950]: 1–115.
Obenberger J. 1952e: Monographie du genre Lampra Sol. (Col. Buprestidae) de la région paléarctique. Acta
Entomologica Musaei Nationalis Pragae 27 [1951]: 279–374.

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l’expédition scientifique zoologique du Muséum National de Praha en Turquie 15. Coleoptera IV.
Buprestidae. Acta Entomologica Musaei Nationalis Pragae 29 [1953–1954]: 5–23.
Obenberger J. 1955b: Nový druh krasce z rodu Sphenoptera, škůdce broskví v Pakistanu. A new species of
Sphenoptera, damaging the peach-trees in Pakistan. Acta Entomologica Musaei Nationalis Pragae 29
[1953–1954]: 89–91.
Obenberger J. 1956a: 1. Příspěvek k poznání palaearktických krasců (Col. Buprestidae). Matériaux pour servir
à la connaissance des buprestides paléarctiques – I. Acta Entomologica Musaei Nationalis Pragae 30
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Obenberger J. 1956b: On some buprestid beetles from the coll. of Forest Research Institute in Dehra Dun in
India, I. Acta Entomologica Musaei Nationalis Pragae 30 [1955]: 243–252.
Obenberger J. 1958a: Buprestides, trouvés par l’expédition chino-soviétique 1955 en Chine méridionale (Col.
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Petrovitz R. 1959d: Phyllopertha (Blitopertha) lineata Fbr. ist eine Mischart. (Col. Scarab.). Fragmenta
Balcanica Musei Macedonici Scientiarum Naturalium 2: 185–190.
Petrovitz R. 1961a: Scarabaeidae. (Col.) II. Contribution à l’étude de la faune d’Afghanistan. Kungliga
Fysiografiska Sällskapets i Lund. Förhandlingar 31: 31–45.
Petrovitz R. 1961b: Neue und verkannte Aphodiinae aus allen Erdteilen (Col. Scarab.). Entomologische
Arbeiten aus dem Museum Georg Frey 12: 99–135.
Petrovitz R. 1961c: Zoologische Ergebnisse der Österreichischen Karakorum-Expedition. 1958. II. Teil.
Coleoptera: Scarabaeidae. Sitzungsberichte der Österreichischen Akademie der Wissenschaften.
Mathematisch-Naturwissenschaftliche Klasse, Abteilung 1 170: 101–118.
Petrovitz R. 1961d: Eine neue Aegialiinae-Art aus Japan (Coleoptera, Scarabaeidae). Entomologische Arbeiten
aus dem Museum Georg Frey 12: 136–137.
Petrovitz R. 1961e: Neue und bemerkenswerte Aphodiinae aus dem Museo entomologico del Pontificio
Istituto Missioni estere, Monza. Bollettino della Società Entomologica Italiana 91: 144–149.
Petrovitz R. 1961f: Neue und verkannte Aphodiinae aus allen Erdteilen. II Teil. Entomologische Arbeiten aus
dem Museum Georg Frey 12: 344–356.
Petrovitz R. 1961g: Pleurophorus caesus Creutz. und seine nächsten Verwandten. Atti della Società Italiana di
Scienze Naturali e del Museo Civico di Storia Naturale in Milano 100: 389–396.
Petrovitz R. 1961h: Drei neue Aphodius-Arten aus dem Senckenberg-Museum. Senckenbergiana Biologica 42:
443–446.

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Petrovitz R. 1962a: Neue und verkannte Aphodiinae aus allen Erdteilen. 3. Entomologische Arbeiten aus dem
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Petrovitz R. 1962b: Neue und interessante Scarabaeidae aus dem vorderen Orient. I. Teil. Reichenbachia 1:
107–124.
Petrovitz R. 1962c: Neue interessante Scarabaeidae aus dem vorderen Orient II. Teil. Reichenbachia 1:
235–267.
Petrovitz R. 1963a: Neue Arten der Gattung Rhyssemus Mulsant (Coleoptera, Scarabaeidae, Aphodiinae).
Reichenbachia 2: 33–45.
Petrovitz R. 1963b: Neue Aegialiinae, Orphninae und Hybosorinae. Entomologische Arbeiten aus dem Museum
Georg Frey 14: 118–125.
Petrovitz R. 1963c: Neue und interessante Scarabaeidae aus dem vorderen Orient. II. Teil. Reichenbachia 2:
235–267.
Petrovitz R. 1963d: Neue und verkannte Aphodiidae aus allen Erdteilen. 4. Entomologische Arbeiten aus dem
Museum Georg Frey 14: 630–647.
Petrovitz R. 1964a: Eine neue Onthophagus-Art aus Jugoslavien (Scarabaeidae, Coleoptera). Fragmenta
Balcanica Musei Macedonici Scientiarum Naturalium 5: 25–26.
Petrovitz R. 1964b: Neue Arten der Gattung Hybalus Brulle (Col,. Scarab., Orphnidae) (mit 20 Abbildungen
und einer Bestimmungstabelle). Bulletin de la Société des Sciences Naturelles et Physiques du Maroc 44:
49–63.
Petrovitz R. 1964c: Amphicoma-Studien. Zur Kenntnis einiger Amphicoma-Arten. Reichenbachia 3 Nr. 5:
99–105.
Petrovitz R. 1964d: Neue Melolonthidae und Dynastidae aus Evropa und Asien. Reichenbachia 3: 127–131.
Petrovitz R. 1964e: Neue afrikanische und australische Aphodiidae. Reichenbachia 4: 171–207.
Petrovitz R. 1965a: Neue Aphodiinae (Coleopt., Scarab.). Mitteilungen aus dem Zoologischen Museum in Berlin
41: 161–176.
Petrovitz R. 1965b: Drei neue palaearktische Melolonthinae. Reichenbachia 5: 191–195.
Petrovitz R. 1965c: Österreichische entomologische Expeditionen nach Persien und Afghanistan. Beiträge
zur Coleopterologie. Teil II. Lamellicornia. Annalen des Naturhistorischen Museum in Wien 68: 671–694.
Petrovitz R. 1967a: Scarabaeidae aus Marokko. Bulletin de la Société des Sciences Naturalles et Physiques du
Maroc 47: 227–235.
Petrovitz R. 1967b: Ergebnisse zoologischer Sammelreisen in der Türkei, Lamellicornia, Coleoptera. Annalen
des Naturhistorischen Museum in Wien 70: 325–343.
Petrovitz R. 1967c: Neue und verkannte Aphodiinae aus allen Erdteilen (Col., Scarab.). Entomologische
Arbeiten aus Museum G. Frey 18: 388–403.
Petrovitz R. 1967d: Neue Acanthocerinae, Hyposorinae und Ochodaeinae (Scarab., Col.). Entomologische
Arbeiten aus dem Museum G. Frey 18: 425–431.
Petrovitz R. 1967e: Österreichische entomologische Expeditionen nach Persien und Afghanistan.
Beiträge zur Coleopterologie Teil XI. Lamellicornia. Annalen des Naturhistorischen Museums in Wien 70:
479–490.
Petrovitz R. 1968a: Fünf neue Trichiorhyssemus-Arten aus der Orientalischen Region. Entomologisches
Nachrichtenblatt 15: 6–12.

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Petrovitz R. 1968f: Neue Tanyproctus- und Atanyproctus-Arten (Pachydemini, Melolonthinae, Scarabaeidae).
Entomologische Arbeiten aus dem Museum G. Frey 19: 237–247.
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Folge. Annalen des Naturhistorischen Museum in Wien 72: 465–491.
Petrovitz R. 1968h: Die afrikanischen Arten der Gattung Glaresis Erichson nebst einer mit dieser nahe
­verwandten neuen Gattung (Troginae, Scarabaeidae, Coleoptera). Entomologische Arbeiten aus dem
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Petrovitz R. 1969a: Zwei neue Calamosternus-Arten. Koleopterologische Rundschau 46: 81–84.
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Mitteilungen aus dem Zoologischen Museum Berlin 45: 215–219.
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Folge. Die mediterranen Arten der Gattung Maladera Mulsant. Lamellicornia, Coleoptera. Annalen des
Naturhistorischen Museums Wien 73: 383–400.
Petrovitz R. 1970a: Eine neue Paracoptochirus-Art aus Libien. Fragmenta Entomologica 7: 1–4.
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Scienze Naturali e del Museo Civico di Storia Naturale di Milano 110: 387–390.
Petrovitz R. 1971a: Neue Aphodiidae der äthiopischen und australischen Region (Col.). Entomologische
Arbeiten aus dem Museum G. Frey 22: 99–104.
Petrovitz R. 1971b: Die nordafrikanischen Aphodius-Arten der Artengruppe Anomius Mulsant. Bollettino
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Petrovitz R. 1971c: Drei neue Anoxia-Arten (Scarabaeidae, Melolonthini). Koleopterologische Rundschau 49:
183–187.
Petrovitz R. 1971d: Scarabaeidae from the Near East. Israel Journal of Entomology 6: 215–237.
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des Naturhistorischen Museum in Wien 75: 565–589.
Petrovitz R. 1972a: Paläarktische und orientalische Glaphyrinae-Formen (Coleoptera, Scarabaeidae).
Mitteilungen aus dem Zoologischen Museum in Berlin 48: 21–30.
Petrovitz R. 1972b: Colobopterus indagator Mann. und apicalis Harold. Koleopterologische Rundschau 50:
31–33.

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Entomologica 16: 141–202.
Petrovitz R. 1973d: Neue Arten aus der Unterfamilie Geotrupinae (Coleoptera, Scarabaeidae). Folia
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in Wien 83: 597–638.
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Reiche L.J. 1850: Entomologie. Ordre de coléoptères. Pp. 259–532, 18 pls. In: Ferret A. & Galinier: Voyage en
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Reiche L.J. 1853a: Notes synonymiques sur les espèces de la famille des pectinicornes décrites dans le 5e vol.
de l’Handbuch der Entomologie par M.H. Burmeister. Annales de la Société Entomologique de France (3) 1:
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Reiche L.J. 1853b: Note sur l’Hybosorus arator Auctorum, etc. Annales de la Société de Entomologique France
(3) 1: 87–90.
Reiche L.J. 1856a: Description d’une espèce nouvelle du genre Onitis, Fabricius, suivie d’un catalogue synoptique des espèces de ce genre qui se trouvent en Europe et sur les bords des bassins méditerranées, de la
Méditerranée, de l’Euxin et de la mer Caspienne. Revue et Magasin de Zoologie Pure et Appliquée (2) 8:
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Reiche L.J. 1859a: Notes synonymiques sur le cinquième volume de l’Handbuch der Entomologie, Par
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Reiche L.J. 1861b: Coléoptères nouveaux reccueillis en Corse par M.E. Bellier de la Chavignerie. Annales de la
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4: 233–246.
Reiche L.J. 1871: Examen de quelques espèces de cétonides d’Europe et pays limitrophes et description de
quatre espèces nouvelles Annales de la Société Entomologique de France (5) 1: 83–87.
Reiche L.J. 1877: Les descriptions de quatre nouvelles espèces de buprestides. Bulletin de la Société
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Reiche L.J. 1879: Descriptions de quelques nouvelles espèces de géorissides, parnides et hétérocérides propres
à la faune européenne. Annales de la Société Entomologique de France (5) 9: 237–239.
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membre de l’Institut, dans son voyage en Orient (Suite). Annales de la Société Entomologique de France (3)
4: 353–422.
Reitter E. 1876: Neue transcaucasische Coleopteren, gesammelt von Hans Leder. Deutsche Entomologische
Zeitschrift 20: 289–294.
Reitter E. 1878: [new taxa]. In: Schneider O. & Leder H.: Beiträge zur Kenntniss der kaukasischen Käferfauna.
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Reitter E. 1880a: [new taxa]. In: Leder H.: Beitrag zur kaukasischen Käferfauna. Unter Mitwirkung hervorragender Fachgenossen. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in
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Reitter E. 1880b: Neue Coleopteren aus dem südöstlichen Russland, aufgefunden von Herrn J. Faust aus
Helsingfors. Verhandlungen der Kaiserlich-Königlichen Zoologische-Botanischen Gesellschaft in Wien 29
[1879]: 543–546.
Reitter E. 1881a: Bestimmungs-Tabellen der europäischen Coleopteren. IV. Enthaltend die Familien:
Cistelidae, Georyssidae un Thorictidae. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien
31: 67–96, pl. II.
Reitter E. 1881b: Neue und seltene Coleopteren im Jahre 1880 in Süddalmatien und Montenegro gesammelt.
Deutsche Entomologische Zeitschrift 25: 177–230.
Reitter E. 1883a: [new names]. In: Heyden L.F.J.D. von, Reitter E. & WeiseJ.: Catalogus Coleopterorum Europae
et Caucasi. London, Berlin, Paris: Edw. Janson, Libraria Nicolai, Luc. Buquet, 228 pp.
Reitter E. 1883b: Neue Coleopteren aus Russland und Bemerkungen über bekannte Arten. Revue Mensuelle
d’Entomologie 1: 40–44, 70–74, 111–114.
Reitter E. 1883c: Coleopterologische Notizen. Deutsche Entomologische Zeitschrift 27: 74–75.
Reitter E. 1884a: Diagnosen neuer Coleopteren aus Lenkoran. Verhandlungen des Naturforschenden Vereins in
Brünn 22 [1883]: 3–10.
Reitter E. 1884b: [new taxa]. In: Brenske E. & Reitter E.: Neuer Beitrag zur Käferfauna Griechenlands. Deutsche
Entomologische Zeitschrift 28: 17–100, pls I–II.
Reitter E. 1884c: Resultate einer coleopterologischen Sammel-Campagne während den Monaten Februar bis
April 1883 auf den jonischen Inseln. Deutsche Entomologische Zeitschrift 28: 101–122.
Reitter E. 1885a: Bestimmungs-Tabellen des europäischen Coleopteren 12. Necrophaga. (Platypsyllidae,
Leptinidae, Silphidae, Anisotomidae und Clambidae). Verhandlungen des Naturforschenden Vereins in
Brünn 23 [1884]: 3–122.
Reitter E. 1885b: Neue Coleopteren aus Europa und den angrenzenden Ländern, mit Bemerkungen über
bekannte Arten. Deutsche Entomologische Zeitschrift 29: 353–392.
Reitter E. 1885c: Uebersicht der Phyllopertha-Arten aus Europa und den angrenzenden Ländern. Deutsche
Entomologische Zeitschrift 29: 393–397.
Reitter E. 1887a: Tabelle zur Bestimmung der europäischen Arten der Coleopteren-Gattung Homaloplia
Steph. Wiener Entomologische Zeitung 6: 135–139.
Reitter E. 1887b: Insecta in itinere Cl.N. Przewalskii in Asia centrali novissime lecta. VI. Clavicornia,
Lamellicornia et Serricornia. Horae Societatis Entomologicae Rossicae 21: 201–234.
Reitter E. 1887c: Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen
über bekannte Arten. 3. Teil. Deutsche Entomologische Zeitschrift 31: 241–288.
Reitter E. 1887d: Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen
über bekannte Arten. Vierter Theil. Deutsche Entomologische Zeitschrift 31: 497–528.
Reitter E. 1887e: Zur Species-Kenntniss der Maikäfer aus Europa und den angrenzenden Ländern. Deutsche
Entomologische Zeitschrift 1887: 529–542.
Reitter E. 1888a: Vier neue Coleopteren. Wiener Entomologische Zeitung 7: 68–71.

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Reitter E. 1888b: Coleoptera aus Circassien, gesammelt von Hans Leder im Jahre 1887. V. Theil. Wiener
Entomologische Zeitung 7: 207–214.
Reitter E. 1888c: Uebersicht der Arten der Coleopteren-Gattung Phyllopertha Kirby, aus Europa, den angrenzenden Ländern, Sibirien mit Central-Asien. Entomologische Nachrichten 14: 289–294.
Reitter E. 1888d: Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen
über bekannte Arten. Fünfter Theil. Deutsche Entomologische Zeitschrift 1888: 417–432.
Reitter E. 1889a: Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen
über bekannte Arten. Sechster Theil. Deutsche Entomologische Zeitschrift 1889: 17–44.
Reitter E. 1889b: Coleopterologische Ergebnisse der im Jahre 1886 und 1887 in Transcaspien von Dr. G. Radde,
Dr. A. Walter und A. Konchin ausgeführten Expedition. Verhandlungen des Naturforschenden Vereins in
Brünn 27 [1888]: 95–133.
Reitter E. 1889c: Neue Coleopteren aus Circassien, gesammelt von A. Starck. Wiener Entomologische Zeitung
8: 97–104.
Reitter E. 1889d: Uebersicht der Arten der Coleopteren-Gattung Anisoplia aus Europa und den angrenzenden
Ländern. Deutsche Entomologische Zeitschrift 1889: 99–111.
Reitter E. 1889e: Uebersicht der mir bekannten Adoretus-Arten der palaearktischen Fauna. Entomologische
Nachrichten 15: 267–270.
Reitter E. 1889f: Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen
über bekannte Arten. Siebenter Theil. Deutsche Entomologische Zeitschrift 1889: 273–288.
Reitter E. 1889g:Uebersicht der mir bekannten Coleopteren-Gattungen der Melolonthini im Erichson’schen
Sinne, aus der paläarktischen Fauna. Wiener Entomologische Zeitung 8: 275–279.
Reitter E. 1889h:Uebersicht der mir bekannten Arten der Coleopteren-Gattung Triodonta Muls. Wiener
Entomologische Zeitung 8: 283–285.
Reitter E. 1889i: Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen
über bekannte Arten. Deutsche Entomologische Zeitschrift 1889: 369–376.
Reitter E. 1890a: Uebersicht der Arten der Coleopterengattung Polyphylla Harr. aus der paläarctischen
Region. Wiener Entomologische Zeitung 9: 21–22.
Reitter E. 1890b: Analytische Revision der Coleopteren-Gattung Amphicoma. Deutsche Entomologische
Zeitschrift 1890: 53–64.
Reitter E. 1890c: Revision der Arten der Coleopteren-Gattung Triodonta Muls. aus der palaearctischen Fauna.
Entomologische Nachrichten 16: 65–69.
Reitter E. 1890d: Uebersicht der Arten der Coleopteren-Gattung Anoxia Lap. aus Europa und der angrenzenden Ländern. Wiener Entomologische Zeitung 9: 105–107.
Reitter E. 1890e: Coleopterologische Notizen XXXVII. Wiener Entomologische Zeitung 9: 142–146.
Reitter E. 1890f: Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen
über bekannte Arten. Neunter Theil. Deutsche Entomologische Zeitschrift 1890: 145–164.
Reitter E. 1890g: Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen
über bekannte Arten. Zehnter Theil. Deutsche Entomologische Zeitschrift 1890: 165–176.
Reitter E. 1890h: Zusätze und Ergänzungen zu mainer “Uebersicht der Arten der Coleopteren-Gattung Anoxia
Lap. aus Europa und der angrenzenden Ländern”. Wiener Entomologische Zeitung 9: 173–176.
Reitter E. 1890i: Zur Synonymie einiger Polyphylla-Arten. Entomologische Nachrichten 16: 185–190.

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Turkestan. Wienter Entomologische Zeitung 9: 189–198.
Reitter E. 1890k: Uebersicht der bekannten Hymenoplia-Arten. Wiener Entomologische Zeitung 9: 259–263.
Reitter E. 1890l: Coleopterologische Notizen XXXIX. Wiener Entomologische Zeitung 9: 264–267.
Reitter E. 1890m: Uebersicht der trispinosen Sphenoptera-Arten aus Europa und dem Kaukasus.
Entomologisches Nachrichtenblatt 16: 276–283.
Reitter E. 1890n: Analytische Uebersicht der bekannten Lethrus-Arten. Deutsche Entomologische Zeitschrift
1890: 289–295.
Reitter E. 1890o: Uebersicht der mir bekannten Arten der Coleopteren-Gattung Acmaeodera Eschsch. aus
Europa und den angrenzenden Ländern. Entomologische Nachrichten 16: 337–347.
Reitter E. 1890p: [new taxa]. In: Heyden L.F.J.D. von.: XII. Beitrag zur Coleopteren-Fauna von Turkestan. Unter
Mitwirkung der Herren Reitter und Weise zusammengestellt von Major z. D. Dr. L. von Heyden. Deutsche
Entomologische Zeitschrift 1890: 353–367.
Reitter E. 1890q: Revision der Arten der Coleopteren-Gattung Hoplia Ill. aus der palaearctischen Fauna.
Deutsche Entomologische Zeitschrift 1890: 375–383.
Reitter E. 1890r: Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen
über bekannte Arten. Elfter Theil. Deutsche Entomologische Zeitschrift 1890: 385–396.
Reitter E. 1891a: [new names]. In: Heyden L.F.J.D. von, Reitter E. & Weise J.: Catalogus Coleopterorum Europae,
Caucasi et Armeniae rossicae. Berlin, Modling, Caen, 420 pp.
Reitter E. 1891b: Neue Coleopteren aus Europa, den angrenzenden Ländern und Sibirien, mit Bemerkungen
über bekannte Arten. Zwölfter Theil. Deutsche Entomologische Zeitschrift 1891: 17–36.
Reitter E. 1891c: Drei neue Coleopterern. Wiener Entomologische Zeitung 10: 33–34.
Reitter E. 1891d: Darstellung der echten Cetoniden-Gattungen und deren mir bekannten Arten aus Europa
und den angrenzenden Ländern. Deutsche Entomologische Zeitschrift 35: 49–74.
Reitter E. 1891e: Erster Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische Zeitung
10: 138–142.
Reitter E. 1891f: Uebersicht der Onthophagus-Arten aus dem nächsten Verwandtschaftskreise des O. Amyntas
Ol. Wiener Entomologische Zeitung 10: 241–245.
Reitter E. 1891g: Ueber die mit Mendidius Er. verwandten Gattungen. Wiener Entomologische Zeitung 10:
253–255.
Reitter E. 1892a: Bestimmungs-Tabellen der Lucaniden und coprophagen Lamellicornen des palaearctischen
Faunengebietes. Brünn: Edmund Reitter, 230 pp. [Separate publication with both parts of the
Bestimmungstabellen, issued before the volume 31 of the Verhandlungen des Naturforschenden Vereins in
Brünn].
Reitter E. 1892b: Fünfter Beitrag zur Coleopterenfauna des russischen Reiches. Wiener Entomologische
Zeitung 11: 59–68.
Reitter E. 1892c: Ueber eine Lasiopsis-Art aus dem Kaukasus. Wiener Entomologische Zeitung 11: 101–102.
Reitter E. 1892d: Sechster Beitrag zur Coleopteren-Fauna des Russischen Reiches. Wiener Entomologische
Zeitung 11: 133–136.
Reitter E. 1892e: Bestimmungs-Tabellen der Lucaniden und coprophagen Lamellicornen des palaearctischen
Faunengebietes. Verhandlungen des Naturforschenden Vereins in Brünn 30 [1891]: 141–262.

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Reitter E. 1892g: Siebenter Beitrag zur Coleopterenfauna des russischen Reiches. Wiener Entomologische
Zeitung 11: 151–154.
Reitter E. 1893a: Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen
Faunengebietes. Verhandlungen des Naturforschenden Vereins in Brünn 31 [1892]: 1–109.
Reitter E. 1893b: Coleopterologische Notizen. Wiener Entomologische Zeitung 12: 73.
Reitter E. 1893c: Achter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische
Zeitung 12: 109–114.
Reitter E. 1893d: Dritter Beitrag zur Coleopteren-Fauna von Europa und den angrenzenden Ländern. Wiener
Entomologische Zeitung 12: 172–176.
Reitter E. 1893e: Neunter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische
Zeitung 12: 219–224.
Reitter E. 1894a: Übersicht der mir bekannten palaearctischen Arten der Coleopteren-Gattung Trichius F.
Wiener Entomologische Zeitung 13: 5–6.
Reitter E. 1894b: [new taxa]. In: Hauser F.: Beitrag zur Coleopteren-Fauna von Transcaspien und Turkestan.
Deutsche Entomologische Zeitschrift 1894: 17–74.
Reitter E. 1894c: Vierter Beitrag zur Coleopteren-Fauna von Europa und den angrenzenden Ländern. Wiener
Entomologische Zeitung 13: 101–107.
Reitter E. 1894d: Zehnter Beitrag zur Coleopteren-Fauna des Russischen Reiches. Wiener Entomologische
Zeitung 13: 122–128.
Reitter E. 1894e: Nachträge und Berichtungen zu meiner Bestimmungs-Tabelle der coprophagen
Lamellicornen. Entomologische Nachrichten 20: 183–190.
Reitter E. 1894f: Elfter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische Zeitung
13: 238–248.
Reitter E. 1894g: Fünfter Beitrag zur Coleopteren-Fauna von Europa und den angrenzenden Ländern. Wiener
Entomologische Zeitung 13: 299–306.
Reitter E. 1894h: Uebersicht der Arten der Gattung Dryops Oliv. Wiener Entomologische Zeitung 13: 313.
Reitter E. 1895a: Uebersicht der trispinosen Sphenoptera-Arten (Oplistura und Chrysoblemma) aus der
palaearctischen Fauna. Wiener Entomologische Zeitung 14: 32–42.
Reitter E. 1895b: Beschreibung neuer oder wenig gekannter Coleoptera aus der Umgebung von Akbes in
Syrien. Wiener Entomologische Zeitung 14: 79–88.
Reitter E. 1895c: Synoptische Uebersicht der Chrysobothris-Arten mit Gruben auf den Flügeldecken, aus der
palaearctischen Fauna. Wiener Entomologische Zeitung 14: 127–130.
Reitter E. 1895d: Zwölfter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische
Zeitung 14: 149–162.
Reitter E. 1895e: Nachträgliches über Cyphonotus Fisch. und Beschreibung einer neuer Art aus Transkaspien.
Wiener Entomologische Zeitung 14: 205–207.
Reitter E. 1895f: Einige neue Coleopteren aus Korea und China. Wiener Entomologische Zeitung 14: 208–210.
Reitter E. 1895g: Anisoplia Koenigi n. sp. Deutsche Entomologische Zeitschrift 1895: 309–310.

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Reitter E. 1896a: Tribax certus und Cetonia chrysosoma, n. sp. Entomologische Nachrichten 22: 4–5.
Reitter E. 1896b: Beschreibungen neuer Coleopteren aus dem russischen Reiche. Deutsche Entomologische
Zeitschrift 1896: 33–48.
Reitter E. 1896c: Dreizehnter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische
Zeitung 15: 64–77.
Reitter E. 1896d: Zwei neue Seminolus-Arten aus Ostsibirien. Deutsche Entomologische Zeitschrift 1896: 172.
Reitter E. 1896e: Uebersicht der mir bekannten palaearktischen, mit der Coleopteren-Gattung Serica verwandten Gattungen und Arten. Wiener Entomologische Zeitung 15: 180–188.
Reitter E. 1896f: Beitrag zur Kenntnis der Arten und Varietäten der Coleopteren-Gattung Cetonia L.
Entomologische Nachrichten 22: 241–246.
Reitter E. 1896g: Achter Beitrag zur Coleoptere-Fauna von Europa und den angrenzenden Ländern. Wiener
Entomologische Zeitung 15: 265–272.
Reitter E. 1896h: Vierzehnter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische
Zeitung 15: 285–291.
Reitter E. 1896i: Sicardia nov. gen. Aphodinidarum. Deutsche Entomologische Zeitschrift 1896: 318.
Reitter E. 1897a: Neunter Beitrag zur Coleopteren-Fauna von Europa und den angrenzenden Ländern. Wiener
Entomologische Zeitung 16: 45–47.
Reitter E. 1897b: Coleopterologische Notizen. LIX. Wiener Entomologische Zeitung 16: 48–49.
Reitter E. 1897c: Aphodius Deubeli n. sp. Deutsche Entomologische Zeitschrift 1897: 76.
Reitter E. 1897d: Coleopterologische Notizen. Wiener Entomologische Zeitung 16: 77–80.
Reitter E. 1897e: Fünfzehter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische
Zeitung 16: 121–127.
Reitter E. 1897f: Coleopterologische Notizen. Wiener Entomologische Zeitung 16: 148–149.
Reitter E. 1897g: Zehnter Beitrag zur Coleopteren-Fauna von Europa und den angrenzenden Ländern. Wiener
Entomologische Zeitung 16: 203–206.
Reitter E. 1897h: Dreissig neue Coleopteren aus russisch Asien und der Mongolei. Deutsche Entomologische
Zeitschrift 1897: 209–228.
Reitter E. 1897i: Coleopterologische Notizen. Wiener Entomologische Zeitung 16: 217–220.
Reitter E. 1897j: Ueber die mit Pachydema Lap. verwandten Coleopteren-Gattungen und Uebersicht der
Hemictenius-Arten. Wiener Entomologische Zeitung 16: 221–225.
Reitter E. 1897k: Lethrus (Microlethrus) inermis Reitter n. sp. Deutsche Entomologische Zeitschrift 1897:
235–236.
Reitter E. 1897l: Coleopterologische Notizen. LXII. Wiener Entomologische Zeitung 16: 241–242.
Reitter E. 1897m: Abbildungen mit Beschreibungen acht neuer Coleopteren aus der palaearctischen Fauna.
Wiener Entomologische Zeitung 16: 246–252, pl. 3.
Reitter E. 1898a: Eine Decade neuer Coleopteren aus der Buchara. Wiener Entomologische Zeitung 17: 10–16.
Reitter E. 1898b: Siebzehnter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische
Zeitung 17: 109–127.
Reitter E. 1898c: Neue Coleopteren aus Europa und den angrenzenden Ländern. Deutsche Entomologische
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Trichiini. Verhandlungen des Naturforschenden Vereins in Brünn 37 [1898]: 21–111.
Reitter E. 1899b: Elfter Beitrag zur Coleopteren-Fauna von Europa und den angrezenden Ländern. Wiener
Entomologische Zeitung 18: 155–161.
Reitter E. 1899c: Beitrag zur Coleopteren-Fauna des russichen Reiches und der angrenzenden Länder.
Deutsche Entomologische Zeitschrift 1899: 193–209.
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Reitter E. 1900b: Beitrag zur Coleopteren-Fauna des russischen Reiches. Deutsche Entomologische Zeitschrift
1900: 49–59.
Reitter E. 1900c: Beitrag zur Coleopteren-Fauna von Europa und den angrenzenden Ländern. Deutsche
Entomologische Zeitschrift 1900: 81–88.
Reitter E. 1900d: Coleoptera gesammelt in Jahre 1898 in Chin. Central-Asien von Dr. Holderer in Lahr. Wiener
Entomologische Zeitung 29: 153–166.
Reitter E. 1900e: Beschreibung and Abbildung von neue neuen Coleopteren der palaearcischen Fauna.
Wiener Etomologishe Zeitung 19: 225–232.
Reitter E. 1901a: Weitere Beiträge zur Coleopteren-Fauna des russischen Reiches. Deutsche Entomologische
Zeitschrift 1901: 65–84.
Reitter E. 1901b: Dreizehnter Beitrag zur Coleopteren-Fauna von Europa und den angrenzenden Ländern.
Wiener Entomologische Zeitung 20: 99–101.
Reitter E. 1901c: Eine Serie neuer Coleopteren aus dem russischen Reiche. Deutsche Entomologische Zeitschrift
1901: 177–186.
Reitter E. 1902a: Drei neue Coleopteren aus Italien und Persien. Wiener Entomologische Zeitung 21: 81–82.
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Zilioli M. 2000a: A new species of Lucanus Scopoli, 1763, from Myanmar: Lucanus prossi sp. nov. (Coleoptera:
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Index to family-group names
Ablaberini C.É. Blanchard, 1850 210
Acmaeoderini Kerremans, 1893 438
Actenodini Gistel, 1848 494
Adoretina Burmeister, 1844 317
Adoretini Burmeister, 1844 317
Adorrhinyptiina Arrow, 1917 320
Aegialiinae Laporte, 1840 97
Aegini H. Huang & C.-C. Chen, 2013 60
Aesalinae W.S. Macleay, 1819 58
Aesalini W.S. Macleay, 1819 58
Agrilina Laporte, 1835 524
Agrilinae Laporte, 1835 524
Agrilini Laporte, 1835 524
Amorphosomatina Majer, 2000 554
Amphicominae C.É. Blanchard, 1845 87
Anchytarsinae G.C. Champion, 1897 621
Ancyronychini Ganglbauer, 1904 592
Anisopliina Burmeister, 1844 321
Anomalina Streubel, 1839 327
Anomalini Streubel, 1839 321
Anthaxiini Gory & Laporte, 1839 495
Aphanisticina Jacquelin du Val, 1859 550
Aphanisticini Jacquelin du Val, 1859 550
Aphodiina Leach, 1815 98
Aphodiinae Leach, 1815 98
Aphodiini Leach, 1815 98
Ateuchini Perty, 1830 166
Athyreinae Lynch Arribálzaga, 1878 33
Augylini Pacheco, 1964 610
Aulacocyclinae Kaup, 1868 52
Aulonocneminae A. Janssens, 1946 166
Bolbelasmini Nikolajev, 1996 33
Bolboceratinae Mulsant, 1842 33
Bolboceratini Mulsant, 1842 34
Bolbochromini Nikolajev, 1970 34
Bubastini Obenberger, 1920 513
buprestidae Leach, 1815 432
Buprestinae Leach, 1815 494
Buprestini Leach, 1815 513
buprestoidea Leach, 1815 432
byrrhidae Latreille, 1804 574
Byrrhinae Latreille, 1804 574
Byrrhini Latreille, 1804 581
byrrhoidea Latreille, 1804 574

callirhipidae Emden, 1924 624
Calyptomerinae Crowson, 1955 426
Canthonini van Lansberge, 1875 167
Cephalobyrrhinae G.C. Champion, 1925 609
Ceratocanthinae Martínez, 1968 85
Ceruchini LeConte, 1861 59
Cetoniina Leach, 1815 367
Cetoniinae Leach, 1815 367
Cetoniini Leach, 1815 367
Chalcophorini Lacordaire, 1857 458
Chalcotheina Mikšić, 1976 401
Chasmatopterini Lacordaire, 1856 211
chelonariidae C.É. Blanchard, 1845 623
Chironinae C.É. Blanchard, 1845 166
Chrysobothrini Gory & Laporte, 1938 517
Chrysochroinae Laporte, 1835 456
Chrysochroini Laporte, 1835 456
Cinnabariini Pic, 1914 428
Cisseina Majer, 2000 555
Cladotominae Pic, 1914 621
clambidae Fischer von Waldheim, 1821 426
Clambinae Fischer von Waldheim, 1821 426
Clematina Majer, 2000 555
Coenochilina Burmeister, 1842 390
Coomaniellini Bílý, 1974 521
Coprini Leach, 1815 167
Coraebina Bedel, 1921 556
Coraebini Bedel, 1921 554
Coryphocerina Burmeister, 1842 392
Corythoderini A. Schmidt, 1910 155
Cremastocheilina Burmeister & Schaum, 1841 390
Cremastocheilini Burmeister & Schaum, 1841 390
Cyclocephalini Laporte, 1840 358
Cyclommatini H. Huang & C.-C. Chen, 2013 61
Cylindromorphina Portevin, 1931 552
Cymophorina Krikken, 1984 391
dascillidae Guérin-Méneville, 1843 (1834) 428
Dascillinae Guérin-Méneville, 1843 428
Dascillini Guérin-Méneville, 1843 428
dascilloidea Guérin-Méneville, 1843 428
decliniidae Nikitsky, Lawrence, Kirejtshuk &
Gratshev, 1994 425
Dicercini Gistel, 1848 461
Dicronocephalina Krikken, 1984 398

963

Index to family-group names
Didactyliina Pittino, 1985 155
Diphycerini S.I. Medvedev, 1952 211
Diplognathini Burmeister, 1842 392
Diplotaxini Kirby, 1837 212
Dorcini Parry, 1864 61
Drepanocerina van Lansberge, 1875 174
dryopidae Billberg, 1820 (1817) 603
Dynamopodinae Arrow, 1911 208
Dynamopodini Arrow, 1911 208
Dynastinae W.S. Macleay, 1819 358
Dynastini W.S. Macleay, 1819 358

Heterocerini W.S. Macleay, 1825 613
Hopliini Latreille, 1829 216
hybosoridae Erichson, 1847 85
Hybosorinae Erichson, 1847 86
Hyphalinae Britton, 1971 609

elmidae Curtis, 1830 591
Elmina Curtis, 1830 592
Elminae Curtis, 1830 592
Elmini Curtis, 1830 592
Emmitini Escalera, 1914 430
Enoplotrupini Paulian, 1945 36
Eremazinae Iablokoff-Khnzorian, 1977
Eubolbitini Nikolajev, 1970 35
Eubrianacinae Jakobson, 1913 616
Eubriinae Lacordaire, 1857 617
Euchirini Hope, 1840 214
eucinetidae Lacordaire, 1857 425
eulichadidae Crowson, 1973 623
Eupariini A. Schmidt, 1910 156
Evidini Tôyama, 1987 467
Exomellini Casey, 1914 578

Larainae LeConte, 1861 591
Lethrinae Oken, 1843 47
Leucocelina Kraatz, 1882 388
Leucopholini Burmeister, 1855 224
limnichidae Erichson, 1846 607
Limnichinae Erichson, 1846 607
Lomapterina Burmeister, 1842 400
lucanidae Latreille, 1804 58
Lucaninae Latreille, 1804 60
Lucanini Latreille, 1804 73

Figulini Burmeister, 1847

Julodinae Lacordaire, 1857

432

Karumiinae Escalera, 1913 430
Karumiini Escalera, 1913 431
Kisanthobiini Richter, 1949 521

98

72

Galbellinae Reitter, 1911 455
Genecerini Pic, 1914 430
geotrupidae Latreille, 1802 33
Geotrupinae Latreille, 1802 36
Geotrupini Latreille, 1802 39
glaphyridae W.S. Macleay, 1819 87
Glaphyrinae W.S. Macleay, 1819 95
glaresidae H.J. Kolbe, 1905 58
Goliathini Latreille, 1829 392
Goliathopsidina Krikken, 1984 391
Gymnetina Kirby, 1827 399
Gymnetini Kirby, 1827 399
Gymnopleurini Lacordaire, 1856 171
Haplostethini LeConte, 1861 452
Heptophyllini S.I. Medvedev, 1951 215
heteroceridae W.S. Macleay, 1825 610
Heterocerinae W.S. Macleay, 1825 610

Macromina Burmeister & Schaum, 1840
Macronychini Gistel, 1848 601
Melanophilini Bedel, 1921 521
Meliboeina Majer, 2000 562
Melolonthinae Leach, 1819 210
Melolonthini Leach, 1819 226
Micilini Pacheco, 1964 616
Morychini El Moursy, 1961 578

391

Nicagini LeConte, 1861 59
Nipponocyphoninae Lawrence &
Yoshitomi, 2007 412
Noseolucanini H. Huang & C.-C. Chen, 2013
Nothochodaeini Nikolajev, 2015 85
ochodaeidae Mulsant & Rey, 1871 84
Ochodaeinae Mulsant & Rey, 1871 84
Ochodaeini Mulsant & Rey, 1871 84
Odochilini Rakovič, 1987 158
Odonteini Shokhin, 2007 35
Odontolabini Parry, 1870 81
Omorginae Nikolajev, 2005 53
Oniticellina H.J. Kolbe, 1905 175
Oniticellini H.J. Kolbe, 1905 174
Onitini Laporte, 1840 177
Onthophagini Burmeister, 1846 180

79

964
Orphninae Erichson, 1847 209
Orphnini Erichson, 1847 209
Oryctini Mulsant, 1842 359
Osmodermatini Schenkling, 1922

Index to family-group names

399

Pachydemini Burmeister, 1855 236
Pachyplectrinae Ocampo, 2006 87
Pachypodini Erichson, 1840 249
Pachyrhinadoretina Ohaus, 1912 321
Pachyschelina Böving & Craighead, 1931 567
Paratassini Bílý & Volkovitsh, 1996 467
Paratrachyini Cobos, 1980 453
passalidae Leach, 1815 52
Passalinae Leach, 1815 52
Pedilophorini Casey, 1912 579
Pentodontini Mulsant, 1842 361
Phaedimini Schoch, 1894 400
Philanthaxiina Hołyński, 1988 523
Phileurini Burmeister, 1847 366
Pilinurgina Krikken, 1984 392
Platycerini Mulsant, 1842 79
Poecilonotini Jakobson, 1913 467
Polycestina Lacordaire, 1857 453
Polycestinae Lacordaire, 1857 438
Polycestini Lacordaire, 1857 453
Polyctesini Cobos, 1955 454
Popilliina Ohaus, 1918 350
Potamophilini Mulsant & Rey, 1872 591
Prodoretina Ohaus, 1912 321
Psammodiina Mulsant, 1842 158
Psammodiini Mulsant, 1842 158
psephenidae Lacordaire, 1854 616
Psepheninae Lacordaire, 1854 620
Psephenoidinae Bollow, 1938 620
ptilodactylidae Laporte, 1838 621
Ptilodactylinae Laporte, 1838 622
Ptosimini Kerremans, 1903 455
rhipiceridae Latreille, 1834 431
Rhizotrogini Burmeister, 1855 249
Rhyparini A. Schmidt, 1910 165

Rhyssemina Pittino & Mariani, 1986
Rutelinae W.S. Macleay, 1819 317
Rutelini W.S. Macleay, 1819 356

161

Sandalinae Jakobson, 1913 431
scarabaeidae Latreille, 1802 97
Scarabaeinae Latreille, 1802 166
Scarabaeini Latreille, 1802 204
scarabaeoidea Latreille, 1802 33
Schizonychini Burmeister, 1855 280
Schizorhinini Burmeister, 1842 400
scirtidae Fleming, 1821 412
Scirtinae Fleming, 1821 412
scirtoidea Fleming, 1821 412
Sericini Kirby, 1837 281
Simplocariini Mulsant & Rey, 1869 574
Sinodendrini LeConte, 1861 60
Sisyphini Mulsant, 1842 207
Sphenopterini Lacordaire, 1857 470
Stenelmina Mulsant & Rey, 1872 599
Syncalyptinae Mulsant & Rey, 1869 588
Syncalyptini Mulsant & Rey, 1869 588
Syndesinae W.S. Macleay, 1819 59
Taenioderina Mikšić, 1976 401
Taenioderini Mikšić, 1976 401
Thaumastodinae G.C. Champion, 1924 610
Thinorycterini Semenov & Reichardt, 1925 208
Thomassetiini Bellamy, 1987 523
Toxoscelina Majer, 2000 566
Tracheina Laporte, 1835 568
Tracheini Laporte, 1835 567
Trichiina Fleming, 1821 404
Trichiini Fleming, 1821 404
Trigonostomusina Ohaus, 1912 321
trogidae W.S. Macleay, 1819 53
Troginae W.S. Macleay, 1819 55
Valgini Mulsant, 1842

409

Xenopseina Volkovitsh, 2008

454

Index to genus-group names
Synonyms are in Italics
Aaata Semenov, 1906 432
Ablaberoides C.É. Blanchard, 1850 281
Ablotemus Paulsen & Smith, 2003 274
Abothris Semenov & Richter, 1934 517
Abrognathus Jakovlev, 1890 47
Acanthobodilus G. Dellacasa, 1983 98
Acanthocephalon Théry, 1955 566
Acanthurus Kirby, 1827 411
Acarina Baraud, 1965 304
Acassidiocesta Mandl, 1972 454
Aceraius Kaup, 1868 52
Achranoxia Kraatz, 1888 226
Acindrocephalus Keith & Delpont, 2004 398
Acmaeodera Eschscholtz, 1829 438
Acmaeoderella Cobos, 1955 445
Acmaeotethya Volkovitsh, 1979 440
Acontosceles G.C. Champion, 1924 610
Acrossus Mulsant, 1842 99
Actinophorus Creutzer, 1799 206
Adoretops Kraatz, 1883 214
Adoretosoma C.É. Blanchard, 1851 327
Adoretus Dejean, 1833 317
Adorrhinyptia Arrow, 1917 320
Aegialia Latreille, 1806 97
Aegomorphus Houlbert, 1914 61
Aegus W.S. Macleay, 1819 60
Aeneobyrrhus Pütz, 1998 581
Aesalus Fabricius, 1801 58
Aethiaratrogus Decelle, 1982 249
Aethiessa Burmeister, 1842 367
Afrelia Hołyński, 2009 456
Afrocera Hołyński, 2014 438
Afroglaresis Petrovitz, 1968 58
Afroharoldius A. Janssens, 1949 183
Afromorgus Scholtz, 1986 53
Afropsephenoides Basilewsky, 1959 620
Aganocrossus Reitter, 1895 101
Agelia Laporte & Gory, 1835 456
Agestrata Eschscholtz, 1829 400
Aglaura Gistel, 1848 517
Agnorimus Miyake & Iwase, 1991 404
Agoliinus A. Schmidt, 1913 102
Agolius Mulsant & Rey, 1870 103
Agrilinus Mulsant & Rey, 1870 103

Agrilomorpha Théry, 1909 524
Agrilomorphus Portevin, 1931 553
Agrilophotus Bellamy & Westcott, 1992 524
Agrilus Curtis, 1825 524
Ahaggaria Bollow, 1938 603
Ahermes Reitter, 1891 121
Ahermodontus Báguena, 1930 105
Akiyamaia Kurosawa, 1988 513
Alabameubria H. P. Brown, 1980 617
Alaia Petrovitz, 1980 236
Alcidosoma Laporte, 1867 358
Aleucolomus Reitter, 1902 259
Aleucoschema Semenov, 1935 259
Aleurostictus Stephens, 1839 404
Alissonotum Arrow, 1908 361
Allogymnopleurus A. Janssens, 1940 171
Allotrypes François, 1904 39
Alocoderus A. Schmidt, 1913 105
Altonthophagus Kabakov, 1990 184
Amadotrogus Reitter, 1902 249
Amaladera Reitter, 1896 290
Amalus Mulsant & Rey, 1870 143
Amblis Gistel, 1834 517
Amblomala Reitter, 1903 344
Amidorinus W. Koshantschikov, 1912 141
Amidorus Mulsant & Rey, 1870 106
Amiserica Nomura, 1974 282
Ammanisoplia S.I. Medvedev, 1949 321
Ammoecius Mulsant, 1842 106
Ammogenia Semenov, 1896 350
Amorphosoma Laporte, 1835 554
Amphicoma Latreille, 1806 87
Amphimallina Reitter, 1905 250
Amphimallon Latreille, 1825 250
Amphimallum Agassiz, 1846 250
Amphimallus Mulsant, 1842 250
Amphionthophagus Martín Piera &
Zunino, 1983 184
Amphistoros Gory & Percheron, 1833 394
Anaglyptes Gistel, 1848 458
Anambus C.G. Thomson, 1864 524
Anatochilus Péringuey, 1907 390
Anatona Burmeister, 1842 368
Anchycteis Horn, 1880 621

966
Ancylocheira Eschscholtz, 1829 514
Ancylochira Eschscholtz, 1829 514
Ancylonycha Dejean, 1833 267
Ancyronyx Erichson, 1847 592
Andromeda Gistel, 1834 438
Anepsiovalgus H.J. Kolbe, 1904 409
Anisonchus Dejean, 1833 343
Anisoplia Schönherr, 1817 321
Ankareus Kerremans, 1894 452
Anoctus Sharp, 1875 180
Anomala Samouelle, 1819 328
Anomalaegialia W.J. Brown, 1931 97
Anomalepta Casey, 1915 328
Anomalocera Westwood, 1842 393
Anomalophylla Reitter, 1887 282
Anomalopides E. Strand, 1928 328
Anomalopus Casey, 1915 328
Anomius Mulsant & Rey, 1870 107
Anoplis Kirby, 1837 514
Anoplocnemus Hope, 1843 83
Anoplotrupes Jekel, 1866 39
Anoxia Laporte, 1832 226
Anoxiella Reitter, 1902 256
Anoxoides Petrovitz, 1971 228
Anthaxia Eschscholtz, 1829 495
Anthaxiola Richter, 1945 495
Anthoplia S.I. Medvedev, 1949 325
Anthracophora Burmeister, 1842 392
Anthypna Eschscholtz, 1818 89
Aparammoecius Petrovitz, 1958 108
Apatura Laporte & Gory, 1837 521
Apeosina Broun, 1881 425
Aphanisticus Latreille, 1810 550
Aphodaulacus W. Koshantschikov, 1911 109
Aphodiellus A. Schmidt, 1913 109
Aphodius Hellwig, 1798 109
Aphodobius Péringuey, 1901 111
Aphodosugrames Endrődi, 1964 134
Apleopertha Semenov, 1903 342
Aplidia Hope, 1837 254
Apogonia Kirby, 1819 212
Apotriodonta Baraud, 1962 283
Apropyga S.I. Medvedev, 1951 230
Aprosterna Hope, 1836 328
Apsteiniella A. Schmidt, 1916 111
Apterogyna Reitter, 1902 263
Aptyktophallus Steffan, 1958 598
Archeohomaloplia Nikolajev, 1982 283
Archianoxia Mikšič, 1958 227
Arctobyrrhus Munster, 1902 581

Index to genus-group names
Argante Gistel, 1834 463
Aridagrilus Alexeev, 1998 524
Armideus A. Villa & G.B. Villa, 1833 38
Arquagrilus Alexeev, 1998 524
Arrhaphipterus Schaum, 1862 431
Arrowia Obenberger, 1923 549
Artia Rambur, 1843 241
Articephala Moser, 1920 258
Asactopholis Brenske, 1894 224
Aserica Lewis, 1895 291
Asiactenius Nikolajev, 2000 236
Asiatobyrrhus Paulus, 1971 581
Asiopertha Machatschke, 1957 340
Askeptonycha Kraatz, 1883 270
Asophrops Petrovitz, 1961 255
Ataenia Schoch, 1894 403
Ataeniomorphus Balthasar, 1953 129
Ataenius Harold, 1867 156
Atanyproctus Petrovitz, 1954 237
Ateuchetus Bedel, 1892 204
Ateuchus Fabricius, 1801 206
Atlantopsis Franz, 1967 589
Atopa Paykull, 1799 428
Atropinota Heller, 1923 392
Augyles Schiödte, 1866 610
Aulacostethus C.O. Waterhouse, 1869 61
Aurigena Laporte & Gory, 1836 465
Auritulus Zang, 1905 52
Austragrilus Alexeev, 1998 524
Autanisoplia S.I. Medvedev, 1949 325
Autolethrus Semenov, 1892 47
Autoserica Brenske, 1897 283
Awadoronus Kôno, 1934 603
Babalimnichus Satô, 1994 610
Balthasarianus R. Paulian, 1934 114
Baraudia López-Colón, 1996 39
Barymorpha Guérin-Méneville, 1843 357
Belionota Eschscholtz, 1829 494
Belionota Eschscholtz, 1829 494
Bellamyina Bílý, 1994 454
Bellamyola Hołyński, 1993 462
Betelmis Matsumura, 1915 620
Bielliana Baraud, 1980 281
Bietia Fairmaire, 1898 392
Bifurcanomala J.I. Kim, 1998 328
Bipammosus Normand, 1938 160
Biraloides Balthasar, 1945 123
Biralus Mulsant & Rey, 1870 111
Bisaya Reitter, 1884 425

967

Index To Genus-group Names
Blabephorus Fairmaire, 1898 359
Blitopertha Reitter, 1903 341
Bodiloides M. Dellacasa & G. Dellacasa, 2005 111
Bodilopsis Ádám, 1994 112
Bodilus Mulsant & Rey, 1870 113
Bolbaffroides Nikolajev, 1979 34
Bolbelasmus Boucomont, 1911 33
Bolboceras Kirby, 1819 34
Bolboceratops Krikken, 1978 35
Bolbocerodema Nikolajev, 1973 34
Bolbochromus Boucomont, 1909 34
Bolbogonium Boucomont, 1911 34
Bolbohamatum Krikken, 1980 35
Bolbotrypes Olsoufieff, 1907 34
Bombodes Westwood, 1848 401
Bonsiella Ruter, 1965 395
Bootrupes Boucomont, 1911 43
Bordatius Pittino & Mariani, 1986 161
Bothriophorus Mulsant & Rey, 1852 607
Boucomontiellus Balthasar, 1932 114
Bourgoinia Obenberger, 1926 562
Brachelytrium Obenberger, 1923 513
Brachiaphodius W. Koshantschikov, 1913 114
Brachycisseis Théry, 1931 549
Brachycoraebus Kerremans, 1903 556
Brachydema Fairmaire, 1881 246
Brachyllus Brenske, 1896 255
Brachyphylla Mulsant, 1842 303
Brachyserica Brenske, 1897 292
Brachytricha Reiche, 1871 367
Brahmina C.É. Blanchard, 1851 256
Brahminella S.I. Medvedev, 1951 256
Brahminula S.I. Medvedev, 1951 270
Brancoataenius R. Paulian, 1979 157
Brancoplia Baraud, 1986 325
Brenskea Reitter, 1891 87
Brenskiella Berg, 1898 237
Brindalus Landin, 1960 158
Brontodorcus Didier, 1937 71
Bubagrilus Curletti, 1993 524
Bubalonthophagus Ádám, 1994 190
Bubas Dejean, 1833 177
Bubastoides Kerremans, 1909 513
Buettikeria Sabatinelli & Pontuale, 1998 237
Bulbocerus Acharius, 1781 50
Bunbunius Nomura, 1970 258
Buprestis Linnaeus, 1758 513
Buprestis Linnaeus, 1758 515
Buprestodes Carter, 1924 464
Butozania Mikšič 1955 258

Byrrhininus Pic, 1922 607
Byrrhinus Motschulsky, 1858 607
Byrrhobolus G. Fiori, 1965 578
Byrrhocaulus Fairmaire, 1901 582
Byrrhochomus Fabbri, 2003 578
Byrrhus Linnaeus, 1767 581
Byrrhus Linnaeus, 1767 582
Caccobius C.G. Thomson, 1859 180
Caccophilus Jekel, 1872 181
Caccothryptus Sharp, 1902 607
Caelius Lewis, 1895 97
Caelodera Dejean, 1833 249
Caenaphodius Balthasar, 1936 145
Calamosternus Motschulsky, 1860 114
Calaphodius Reitter, 1892 103
Calicnemis Laporte, 1832 361
Callanthaxia Richter, 1945 495
Callirhipis Latreille, 1829 624
Callistethus C.É. Blanchard, 1851 341
Callistopopillia Ohaus, 1903 350
Calloserica Brenske, 1894 283
Callynomes Westwood, 1873 390
Caloglycyphana Mikšić, 1968 373
Calopopillia H.J. Kolbe, 1894 352
Calopotosia Reitter, 1899 374
Caloptera Gistel, 1834 461
Calvarium Pic, 1918 412
Calyptomerus L. Redtenbacher, 1847 426
Camelonotus Fairmaire, 1883 362
Campsiura Hope, 1831 391
Canthothrupes Jekel, 1866 41
Cantonius Théry, 1929 552
Canudema M. Lacroix, 1994 237
Canuschiza M. Lacroix, 1999 280
Capnodis Eschscholtz, 1829 461
Cardiaspis Saunders, 1866 462
Carinaulus Tesař, 1945 116
Carininota Volkovitsh, 1979 446
Carolina J. Thomson, 1880 403
Carpathobyrrhulus Ganglbauer, 1902 579
Cassolus Sharp, 1875 167
Castalia Gory & Laporte, 1839 454
Castalia Laporte & Gory, 1836 467
Catalasis Dejean, 1833 226
Catharsius Hope, 1837 167
Catoxantha Dejean, 1833 456
Catoxantha Solier, 1833 458
Celaenochrous Kuijten, 1984 86
Celidotella Reitter, 1909 373

968
Centrasiobia Semenov, 1902 234
Cephalobyrrhus Pic, 1923 609
Cephalocosmus Kraatz, 1895 400
Cephaloserica Brenske, 1900 291
Ceracupes Kaup, 1871 52
Ceradryops Hinton, 1937 603
Ceramida Baraud, 1987 237
Ceratodirus Fischer von Waldheim, 1845 48
Ceratophyus Fischer von Waldheim, 1824 39
Ceruchus W.S. Macleay, 1819 59
Cetonia Fabricius, 1775 368
Cetonischema Reitter, 1899 375
Chaetadoretus Ohaus, 1914 319
Chaetonyx Schaum, 1862 209
Chaetophora Kirby, 1823 588
Chaetopisthes Westwood, 1847 155
Chaetopteroplia S.I. Medvedev, 1949 326
Chaetoserica Brenske, 1897 286
Chakriia Tongyai, 1935 562
Chalcogenia C.G. Thomson, 1879 513
Chalcogenia E. Saunders, 1871 513
Chalcophora Dejean, 1833 458
Chalcophorella Kerremans, 1903 459
Chalcosoma Hope, 1837 358
Chamoerrhipes Latreille, 1834 431
Charitovalgus H.J. Kolbe, 1904 409
Charytonia Gistel, 1834 437
Charytonia Gistel, 1848 457
Chasmatopterus Dejean, 1821 211
Cheironitis van Lansberge, 1875 177
Cheirotonus Hope, 1840 214
Chejuanomala J.I. Kim, 1998 328
Chelonarium Fabricius, 1801 623
Chelotrupes Jekel, 1866 36
Chesas Burmeister, 1876 54
Chibidoronus Satô, 1966 608
Chiloblemma Obenberger, 1926 471
Chiloloba Burmeister, 1842 373
Chilostetha Jakovlev, 1889 471
Chilostethus Nelson, 1982 471
Chilothorax Motschulsky, 1860 117
Chilotrogus Reitter, 1905 258
Chioneosoma Kraatz, 1891 259
Chionotrogus Reitter, 1902 259
Chiron W.S. Macleay, 1819 166
Chironitis A. Janssens, 1937 177
Chittius Tarasov, 2008 121
Chloresthia Fairmaire, 1905 392
Chlorobyrrhulus Fabbri, 2002 578
Chlorophorella Descarpentries, 1973 460

Index to genus-group names
Cholerastoma Mohnike, 1871 390
Choumala Kobayashi, 2008 342
Chromogeotrupes Bovo & Zunino, 1983 37
Chromovalgus H.J. Kolbe, 1897 409
Chrooxantha Hołyński, 2009 457
Chrysanthaxia Richter, 1945 503
Chrysoblemma Jakovlev, 1889 474
Chrysobothris Eschscholtz, 1829 517
Chrysobyrrhulus Reitter, 1911 580
Chrysochroa Dejean, 1833 457
Chrysodema Laporte & Gory, 1835 460
Chrysodora Gistel, 1834 484
Chrysoliocola Alexis & Delpont, 1998 375
Chrysophana LeConte, 1860 454
Chrysoplethisa Reitter, 1903 328
Chrysopotosia Mikšić, 1966 375
Chrysoserica Brenske, 1897 284
Chrysosimplocaria Paulus, 1982 574
Chrysoxantha Hołyński, 2009 457
Cinnabarium Fairmaire, 1895 428
Cisseis Gory & Laporte, 1839 555
Citellonthophagus Ádám, 1994 190
Cladognathinus Didier & Séguy, 1952 68
Cladognathus Burmeister, 1847 68
Cladophyllus Houlbert, 1912 73
Clambus Fischer von Waldheim, 1821 426
Clema Semenov, 1900 555
Cleptocaccobius Cambefort, 1984 182
Clerota Burmeister, 1842 401
Clinteria Burmeister, 1842 399
Clinterocera Motschulsky, 1858 390
Clipadoretus Ohaus, 1941 321
Clunipes Hochentwarth, 1785 50
Clypeolontha C.-L. Li & P.-S. Yang, 1999 228
Cnemargulus Semenov, 1903 121
Cnemargus Harold, 1866 134
Cnemargus Motschulsky, 1845 121
Cnematoplatys A. Schmidt, 1913 157
Cnemisus Motschulsky, 1868 121
Cobosiella Volkovitsh, 1979 442
Codocera Eschscholtz, 1821 84
Coenochilus Schaum, 1841 390
Coilodera Hope, 1831 401
Colobonthophagus Balthasar, 1963 184
Colobopterus Mulsant, 1842 121
Colorhinus Erichson, 1841 361
Comazus Fairmaire & Laboulbène, 1854 426
Coniotrogus Brenske, 1903 256
Contacyphon Gozis, 1886 412
Convexagrilus Alexeev, 1998 524

969

Index To Genus-group Names
Coomaniella Bourgoin, 1924 521
Coomaniellina Bílý, 1974 521
Coomanius R. Paulian, 1932 197
Cophaesthetus C.O. Waterhouse, 1880 619
Coprimorphus Mulsant, 1842 122
Copriodes Gistel, 1857 204
Copris Geoffroy, 1762 168
Coptocera A. Murray, 1868 428
Coptochiroides Balthasar, 1938 123
Coptognathus Burmeister, 1847 362
Coraebus Gory & Laporte, 1839 556
Coraegrilus Fairmaire, 1889 556
Coroebina Obenberger, 1923 555
Coroebus Agassiz, 1846 556
Coryphe W.S. Macleay, 1838 393
Coryphocera Burmeister, 1842 393
Cosmiomorpha W.W. Saunders, 1852 392
Cosmiomorpha W.W. Saunders, 1852 392
Cosmiomorphomima Mikšić, 1976 396
Cosmiorrhina Bourgoin, 1917 395
Cosmocephalus Schoch, 1896 400
Costapopillia Benderitter, 1922 352
Cratomerella Richter, 1949 503
Cratomerus Solier, 1833 502
Crator Semenov, 1889 366
Cryphaeobius Kraatz, 1882 273
Cryptanthaxia Richter, 1945 503
Cryptocratomerus Richter, 1949 503
Cryptodactylus Deyrolle, 1864 566
Cryptomera Broun, 1893 425
Cryptotrogus Kraatz, 1888 229
Curimopsis Ganglbauer, 1902 589
Curimus Erichson, 1846 587
Cuspidevia Jäch & Boukal, 1995 601
Cyalithoides Fisher, 1922 460
Cyclanthaxia Richter, 1949 495
Cyclolimnius Nomura, 1958 595
Cyclommatellus Nagel, 1936 61
Cyclommatinus Didier, 1927 61
Cyclommatus Parry, 1863 61
Cyclophthalmus Hope & Westwood, 1845 73
Cyclophthorus Arrow, 1950 61
Cyclorasis J. Thomson, 1862 73
Cycloserica Reitter, 1896 292
Cyclotrogus Wasmann, 1918 183
Cyclotropus R. Oberthür & Houlbert, 1913 68
Cylindrocaulus Fairmaire, 1880 52
Cylindromorphus Kiesenwetter, 1857 553
Cylindropterus Motschulsky, 1864 157
Cymophorus Kirby, 1827 391

Cyphelytra C.O. Waterhouse, 1875 356
Cyphochilus C.O. Waterhouse, 1867 224
Cyphon Paykull, 1799 416
Cyphonicus Sharp, 1902 607
Cyphonotus Fischer von Waldheim, 1824 229
Cyphonoxia Reitter, 1889 229
Cyphosoma Mannerheim, 1837 463
Cyphostetha Jakovlev, 1893 478
Cyphostetha Jakovlev, 1900 471
Cypriacis Casey, 1909 515
Cyprotrogus Nikolajev, 2004 260
Cyriopertha Reitter, 1903 342
Cyriopertha Reitter, 1903 342
Cyrtocamenta Brenske, 1897 210
Cytilus Erichson, 1847 587
Cytoderhinus Seabra, 1909 109
Dactylopopillia Arrow, 1917 350
Dascillus Latreille, 1797 428
Dasylepida Moser, 1913 225
Dasysterna Rambur, 1843 241
Dasytrogus Reitter, 1902 260
Dasytrogus Reitter, 1902 261
Dasyvalgoides Endrődi, 1952 409
Dasyvalgus H.J. Kolbe, 1904 409
Decamera Mulsant, 1842 217
Declinia Nikitsky, Lawrence, Kirejtshuk &
Gratshev, 1994 425
Dedalopterus Sabatinelli & Pontuale, 1998 225
Dejeania C.É. Blanchard, 1850 211
Delopleurus Erichson, 1847 166
Demochroa White, 1859 457
Demoxantha Hołyński, 2009 457
Dendrochariessa Gistel, 1848 467
Dentagrilus Alexeev, 1998 524
Depsaria Gistel, 1856 484
Dermestocyphon Pic, 1918 416
Descarpentriesina Leraut, 1983 470
Dessumia Descarpentries & Villiers, 1966 566
Deudora Jakovlev, 1899 478
Diastictus Mulsant, 1842 158
Dicaulocephalus Gestro, 1888 357
Dicerca Eschscholtz, 1829 463
Dichelomorpha Burmeister, 1855 211
Dicheros Gory & Percheron, 1833 393
Dichodontus Burmeister, 1847 359
Dichomala Reitter, 1903 328
Dicranobia Reitter, 1900 376
Dicranocephalus Hope, 1831 398
Dicranoceps S.I. Medvedev, 1972 398

970
Dicranophorus Reitter, 1899 376
Dicranoplia Reitter, 1903 350
Dicranopselaphus Guérin-Méneville, 1861 617
Dicronocephalus Hope, 1831 398
Didactylia d’Orbigny, 1896 155
Didrepanephorus Wood-Mason, 1878 357
Digitonthophagus Balthasar, 1959 182
Digonophorus C.O. Waterhouse, 1895 62
Digonorhinus Seabra, 1909 106
Dimalia Mulsant & Rey, 1870 97
Dipelicus Hope, 1845 362
Diphucrania Dejean, 1833 555
Diphycerus Fairmaire, 1878 211
Diphydactylus J. Thomson, 1858 217
Diphydactylus J. Thomson, 1858 219
Diphyllomorpha Hope, 1843 393
Diplolophotus Abeille de Perrin, 1897 524
Diplomala Reitter, 1903 328
Discoderoides Théry, 1936 566
Ditomoderus Parry, 1864 62
Dorcus W.S. Macleay, 1819 62
Dorochoviella Jendek, 2006 549
Drepanognathus van Lansberge, 1886 210
Drepanopodus A. Janssens, 1940 206
Drilocephalus Pic, 1918 431
Drupeubria Nakane, 1952 619
Drupeus Lewis, 1895 621
Dryopidius Grouvelle, 1896 606
Dryopomorphus Hinton, 1936 591
Dryops A.G. Olivier, 1791 603
Dupophilus Mulsant & Rey, 1872 592
Durelius Houlbert, 1915 62
Duttus Curletti, 1993 524
Dynamopus Semenov, 1896 208
Dynodorcus Didier, 1937 62
Eccoptaphodius Iablokoff-Khnzorian, 1964 146
Echinoaesalus Zelenka, 1993 59
Echmatophorus C.O. Waterhouse, 1895 357
Ectinohoplia L. Redtenbacher, 1868 216
Ectopria LeConte, 1854 618
Elaphocera Gené, 1836 238
Elaphocerida Reitter, 1902 238
Eligmodontus Houlbert, 1915 78
Elminus Rafinesque, 1815 592
Elmis Latreille, 1802 592
Elmomorphellus Chûjô & Satô, 1964 606
Elmomorphus Sharp, 1888 606
Elodes Latreille, 1797 416
Elsianus Sharp, 1882 596

Index to genus-group names
Elytrobrachium Bílý & Bellamy, 2000 513
Emadiellus A. Schmidt, 1913 123
Emadus Mulsant & Rey, 1870 145
Emma Escalera, 1913 430
Emmita Escalera, 1914 430
Empecamenta Brenske, 1895 210
Empectida Moser, 1914 224
Emphalena Reitter, 1903 328
Endebius van Lansberge, 1880 359
Endelus Deyrolle, 1864 552
Endrodia Mikšić, 1958 376
Engycera Saunders, 1866 523
Ennometes Pascoe, 1866 624
Ennometidium Emden, 1929 624
Enoplotarsus P.H. Lucas, 1859 388
Enoplotrupes P.H. Lucas, 1869 36
Enoplotrypes Fairmaire, 1886 36
Enoplus Reiche, 1860 359
Eoanthaxites Haupt, 1950 495
Eodrepanus Barbero, Palestrini &
Roggero, 2009 174
Eogeotrupes Bovo & Zunino, 1983 37
Eoglaphyrus Semenov, 1926 95
Eoglaresis Semenov & S.I. Medvedev, 1932 58
Eolucanus Kurosawa, 1970 73
Eonychius Jäch & Boukal, 1996 602
Eonychus Jäch & Boukal, 1995 602
Eophileurus Arrow, 1908 366
Eotrichia S.I. Medvedev, 1951 261
Epacmene Gistel, 1848 456
Epadoretus Reitter, 1903 321
Ephillopus Reitter, 1892 204
Epicometis Burmeister, 1842 387
Epidorcus Séguy, 1954 66
Epigeotrupes Bovo & Zunino, 1983 41
Epilichas White, 1859 621
Epitrichius Tagawa, 1941 404
Eremadoretus Semenov, 1912 320
Eremazus Mulsant, 1851 98
Eremoglaresis Semenov & S.I. Medvedev, 1932 58
Eremonthophagus Zunino, 1979 185
Eremoryctes Semenov & S.I. Medvedev, 1932 359
Eretusa Reitter, 1902 278
Erichia Reitter, 1895 610
Eriomela Reitter, 1903 344
Eriotrogus Reitter, 1902 265
Erytrotrogus Iablokoff-Khnzorian, 1955 250
Erytus Mulsant & Rey, 1870 123
Escalerina Bolívar y Pieltain, 1926 431
Escarabaeus Zídek & Pokorný, 2011 205

971

Index To Genus-group Names
Esimaphodius Reitter, 1892 131
Esolus Mulsant & Rey, 1872 593
Estenomenus Faldermann, 1835 391
Esymoides Balthasar, 1945 111
Esymus Mulsant & Rey, 1870 123
Ethon Gory & Laporte, 1839 555
Euacmaeoderella Volkovitsh, 1979 447
Euanthaxia Richter, 1945 495
Eubolbitus Reitter, 1892 35
Eubria Latreille, 1829 618
Eubrianax Kiesenwetter, 1874 616
Eucampsiura Mikšić, 1987 391
Eucetonia Schoch, 1894 370
Euchlora W.S. Macleay, 1819 328
Euchloris Billberg, 1820 457
Euchloropus Arrow, 1907 394
Euchromoplia S.I. Medvedev, 1952 218
Euchronomala Reitter, 1903 328
Euchrysinda Reitter, 1903 327
Eucilodes Vít, 1985 425
Eucinetella Nikitsky, 1996 425
Eucinetus Germar, 1818 425
Eucirrus Melly, 1832 225
Eucteis Guérin-Méneville, 1861 622
Eudolus Mulsant & Rey, 1870 125
Eudora Laporte, 1840 73
Euheptaulacus G. Dellacasa, 1983 125
Euiridotaenia Hołyński, 1997 458
Eulasia Truqui, 1848 89
Eulichas Jakobson, 1913 623
Eulichas Jakobson, 1913 623
Eulucanus Didier, 1927 68
Eumaladera Nomura, 1967 284
Eumimela Kraatz, 1881 368
Eumimimetica Kraatz, 1881 376
Euocys Gistel, 1856 517
Euoniticellus A. Janssens, 1953 175
Euonthophagus Balthasar, 1959 182
Euorodalus G. Dellacasa, 1983 126
Eupachnoda H.J. Kolbe, 1892 374
Eupatorus Burmeister, 1847 358
Eupleurus Mulsant, 1842 126
Eupopillia H.J. Kolbe, 1894 352
Euporochlora Reitter, 1903 328
Euporomala Reitter, 1903 328
Eupotosia Mikšić, 1954 376
Euranoxia Semenov, 1890 230
Eurea LeConte, 1854 618
Eurhipis Laporte, 1834 431
Europtron Marseul, 1867 240

Eurydorcus Didier, 1937 62
Eurythyrea Dejean, 1833 516
Eurytrachellelus Didier, 1937 62
Eurytrachelus J. Thomson, 1862 62
Euselates J. Thomson, 1880 402
Euserica Reitter, 1896 284
Eusericula Reitter, 1902 292
Eutyctus Semenov, 1889 362
Evides Dejean, 1833 467
Excisivalgus Endrődi, 1952 410
Exochomoscirtes Pic, 1916 419
Exolontha Reitter, 1902 229
Exoma Casey, 1908 578
Exomala Reitter, 1903 342
Exomella Casey, 1914 578
Exonthophagus Kabakov, 2006 185
Fairmairiana Alexis & Delpont, 1998 377
Falcicornis Planet, 1894 67
Falsodaemon Pic, 1913 622
Falsotoclinius Keith & Montreuil, 2004 240
Figulus W.S. Macleay, 1819 72
Firminus Coca-Abia, 2003 262
Flatipalpus Fairmaire, 1873 241
Flavohelodes Klausnitzer, 1980 423
Formicdubius Philips & Scholtz, 2000 183
Foveopotosia Mikšić, 1959 377
Freyiella Bollow, 1938 592
Furcilethrus Nikolajev, 1968 48
Furcipalpus Guérin-Méneville, 1861 617
Furconthophagus Zunino, 1979 185
Galbella Westwood, 1848 455
Gametis Burmeister, 1842 371
Garreta A. Janssens, 1940 172
Gassneria Obenberger, 1919 562
Gastromaladera Nomura, 1973 285
Gastroserica Brenske, 1897 285
Gemadoretus Reitter, 1903 320
Genecerus Walker, 1871 430
Genestia Théry, 1923 470
Geobius Brullé, 1832 209
Geotrogus Guérin-Méneville, 1842 263
Geotrupes Latreille, 1797 40
Geotrypes Agassiz, 1846 40
Gibbonthophagus Balthasar, 1963 186
Gilletianus Balthasar, 1933 127
Glacilitarsus Matsumoto, 2014 265
Glaphyrus Latreille, 1802 95
Glaresis Erichson, 1848 58

972
Glenopopillia Lin, 1980 350
Glycetonia Reitter, 1891 373
Glycosia Schoch, 1896 372
Glycyphana Burmeister, 1842 373
Glycyphaniola Mikšić, 1968 373
Glyphana Gistel, 1848 458
Glyptogeotrupes Nikolajev, 1979 41
Gnaphaloryx Burmeister, 1847 67
Gnaphalostetha Reiche & Saulcy, 1856 265
Gnathocera Kirby, 1825 394
Gnathocerida Péringuey, 1907 394
Gnorimotrichius Krajčík, 2009 404
Gnorimus Le Peletier & Audinet-Serville, 1828 404
Godschama Reitter, 1903 352
Goliathopsis Janson, 1881 391
Gonaphodius Reitter, 1892 127
Goniodorcus Didier, 1937 62
Goniolethrus Semenov, 1894 50
Goniophthalma Chevrolat, 1838 550
Gonometopus Houlbert, 1915 78
Grammeubria Kiesenwetter, 1874 618
Grananoxia Brenske, 1890 234
Grandinaphodius Ziani, 2002 127
Granida Motschulsky, 1862 234
Granuleubria Jäch & Lee, 1994 618
Granulopsammodius Rakovič, 1981 158
Graphelmis Delève, 1968 594
Gridelliana Bollow, 1939 591
Grouvelleus Zaitzev, 1908 594
Grouvellinus G.C. Champion, 1923 594
Guanyinaphodius Masumoto & Kiuchi, 2001 127
Gymnodus Kirby, 1837 399
Gymnopleurus Illiger, 1803 172
Gymnota Gistel, 1834 514
Gynaecoplotrupes Boucomont, 1912 37
Gynaecoserica Brenske, 1897 286
Gynexophylla S.I. Medvedev, 1951 234
Gynoplotrupes R. Oberthür, 1883 37
Habroloma C.G. Thomson, 1864 568
Hadroginidius Kraatz, 1896 73
Hadropopillia Kraatz, 1892 341
Hamaxobium Duftschmid, 1825 425
Haplanthaxia Reitter, 1911 503
Haplidia Harold, 1869 254
Haplonthophagus Ádám, 1994 190
Haplosoma Semenov, 1889 362
Haplostethus LeConte, 1860 453
Haroldius Boucomont, 1914 183
Hecatomnus Fairmaire, 1891 271

Index to genus-group names
Heibrianax Lee, Satô & P.-S. Yang, 1999 616
Helichus Erichson, 1847 606
Heliocantharus W.S. Macleay, 1821 206
Heliocopris Hope, 1837 170
Hellaserica Baraud & Nicolas, 1966 287
Helmis Bedel, 1878 592
Helodes Agassiz, 1846 416
Hemichaetoplia Baraud, 1986 327
Hemicoryphocera Mikšić, 1974 393
Hemictenius Reitter, 1897 240
Hemicyclorhinus Seabra, 1909 99
Hemidicerca Richter, 1952 463
Hemiglaphyrus Champenois, 1903 96
Hemiheterorrhina Mikšić, 1974 394
Hemiopta Bedel, 1896 388
Hemiprigenia Mikšić, 1974 400
Hemiserica Brenske, 1894 287
Hemisodorcus J. Thomson, 1862 62
Hemispilota Casey, 1915 328
Heptaulaculus G. Dellacasa & Baraud, 1978 127
Heptaulacus Mulsant, 1842 127
Heptophylla Motschulsky, 1858 215
Herculaisia Seillière, 1910 394
Herthania Klausnitzer, 2006 419
Hetamius Fairmaire, 1893 287
Heterlimnius Hinton, 1935 595
Heterocerus Fabricius, 1792 613
Heterocnemis Albers, 1853 373
Heteronychus Dejean, 1833 362
Heteroplia Burmeister, 1844 328
Heteroplistodus Jakovlev, 1890 49
Heteroprotaetia Mikšić, 1963 386
Heterorhina Westwood, 1842 394
Heterorrhiniolla Mikšić, 1974 395
Hexalus Mulsant & Rey, 1870 156
Hexaphyllus Mulsant, 1839 74
Hexarthrius Hope, 1842 67
Hexataenius Fairmaire, 1891 215
Hilyotrogus Fairmaire, 1886 215
Himacyphon Ruta, Yoshitomi &
Klausnitzer, 2013 416
Himalayoligus Fabbri, 2002 575
Himalhoplia Sabatinelli, 1983 217
Himaloaesalus H. Huang & C.-C. Chen, 2013 59
Himalobuprestis Kurosawa, 1988 513
Hippotrichia Arrow, 1948 271
Histeridium Motschulsky, 1860 180
Hoggarobothris Théry, 1935 520
Holochelus Reitter, 1889 265
Holoschiza van Lansberge, 1886 281

973

Index To Genus-group Names
Holotrichia Hope, 1837 267
Holotrochus Brenske, 1894 271
Homalanthaxia Richter, 1945 495
Homalocopris Solsky, 1871 171
Homaloplia Erichson, 1847 303
Homoderinus Kriesche, 1926 68
Homoeogenus C.O. Waterhouse, 1880 618
Homoeorhipis Fairmaire, 1887 624
Homothyrea H.J. Kolbe, 1895 388
Homovalgus H.J. Kolbe, 1897 411
Hontoriella Escalera, 1914 343
Hoplandrocneme Semenov, 1899 482
Hoplia Illiger, 1803 217
Hoplia Illiger, 1803 219
Hoplistura Jakovlev, 1889 482
Hoplitocranum Jakovlev, 1896 68
Hoplomaladera Nomura, 1974 287
Hoplopus Laporte, 1832 343
Hoploryctoderus Prell, 1933 359
Hoplosternus Guérin-Méneville, 1838 230
Horatocera Lewis, 1895 624
Horiella Takizawa, 1983 575
Horonotus Burmeister, 1847 362
Hovoryctes Prell, 1934 360
Humerimopsis Fabbri, 2003 589
Hybalonomala Reitter, 1903 328
Hybalus Dejean, 1833 209
Hybosorus W.S. Macleay, 1819 86
Hybovalgus H.J. Kolbe, 1904 410
Hydera Latreille, 1816 592
Hydrocyphon L. Redtenbacher, 1858 419
Hymenochelus Reitter, 1890 287
Hymenoplia Eschscholtz, 1830 287
Hyperis Dejean, 1833 222
Hyperius Fairmaire, 1878 212
Hyphalus Britton, 1971 609
Hypochrus Fairmaire, 1891 215
Hypolorus Mulsant & Rey, 1869 578
Hypolorus Mulsant & Rey, 1869 580
Hypselothorax Kirsch, 1866 622
Ianthe Marseul, 1865 455
Idiocnema Faldermann, 1835 328
Idiocnemina Reitter, 1903 328
Idionycha Arrow, 1932 280
Iliola Semenov & S.I. Medvedev, 1949
Indachorius Balthasar, 1941 188
Indiadactylus Bellamy, 1992 566
Indobolbus Nikolajev, 1979 34
Indocetonia Mikšić, 1965 371

328

Indognorimus Krikken, 2009 405
Indonthophagus Kabakov, 2006 188
Indoprotaetia Mikšić, 1968 377
Indosolus Bollow, 1940 602
Indotrichius Krikken, 2009 405
Indovalgus Ricchiardi, 2013 411
Ingrisma Fairmaire, 1893 395
Insulicopris Tsukamoto, 1974 168
Iranotrogus Nikolajev, 2006 270
Iridomroczkowskia Hołyński, 1997 458
Iridotaenia Deyrolle, 1864 458
Iridotoenia Deyrolle, 1864 458
Ischnopopillia Kraatz, 1892 351
Isocamenta H.J. Kolbe, 1914 210
Isochirus Reitter, 1892 154
Ixorida J. Thomson, 1880 402
Jacobsonia W. Koshantschikov, 1912 111
Jaechanax Lee, Satô & P.-S. Yang, 2000 617
Jaechobyrrhinus Pütz, 1991 610
Jalalabadia Balthasar, 1967 241
Jalodis Eschscholtz, 1829 433
Janthe Marseul, 1866 455
Jekelius López-Colón, 1989 41
Jilanzhunychus Jäch & Boukal, 1995 602
Jinbrianax Lee, Satô & P.-S. Yang, 1999 617
Julodella Semenov, 1893 432
Julodis Eschscholtz, 1829 433
Jumnos W.W. Saunders, 1839 395
Kamosia Kerremans, 1898 562
Karumia Escalera, 1913 431
Kheper A. Janssens, 1940 205
Kibakoganea Nagai, 1984 357
Kirchnerius Schenk, 2009 68
Kisanthobia Marseul, 1865 521
Kisonthophagus Ádám, 1994 190
Knowltonia Fisher, 1935 517
Kocheridia Cobos, 1958 450
Kolbeus Boucomont, 1911 33
Koreoxyomus J.I. Kim, 1996 128
Koshantschikovius A. Schmidt, 1913 128
Kryzhanovskia Nikolajev & Kabakov, 1977 241
Kubaniellus Kalashian, 1997 552
Kurosawaia Tôyama & Ohmomo, 1985 523
Labarrus Mulsant & Rey, 1870 128
Lachnota Reitter, 1889 270
Lagopelas Burmeister, 1876 54
Lakhonia Descarpentries & Villiers, 1967

562

974
Lamoana Casey, 1915 328
Lampetis Dejean, 1833 464
Lampra Dejean, 1833 467
Lamprobyrrhulus Ganglbauer, 1902 580
Lamprocheila E. Saunders, 1871 517
Lamprodila Motschulsky, 1860 467
Laorina Jäch, 1997 591
Lareynia Jacquelin du Val, 1859 592
Lasiexis Semenov & S.I. Medvedev, 1936 270
Lasiobrahmina S.I. Medvedev, 1951 270
Lasiodorcus Didier, 1937 71
Lasiomorychus Ganglbauer, 1902 576
Lasioplia S.I. Medvedev, 1949 321
Lasiopsis Erichson, 1847 270
Lasioserica Brenske, 1896 289
Lasiotrichius Reitter, 1899 406
Lasiotropus Reitter, 1913 271
Latelmis Reitter, 1883 595
Lathelmis Zaitzev, 1908 595
Latipalpis Solier, 1833 465
Leiopsammodius Rakovič, 1981 159
Lepadoretus Reitter, 1903 320
Lephthelmis Zaitzev, 1910 599
Lepidiota Kirby, 1828 225
Leptaegialia W.J. Brown, 1931 97
Leptaulacides Zang, 1905 53
Leptaulax Kaup, 1868 53
Leptelmis Sharp, 1888 599
Leptochristina Baraud & Branco, 1991 241
Lethrotrypes Jakobson, 1892 46
Lethrulus Semenov, 1892 49
Lethrus Scopoli, 1777 47
Lethrus Scopoli, 1777 50
Leucocelidia Olsoufieff, 1916 389
Leucolasium Semenov, 1935 260
Leucopholis Dejean, 1833 226
Leucophorus Brenske, 1894 226
Leucoschema Semenov, 1935 260
Leucoschesis Semenov, 1935 259
Leucoserica Reitter, 1896 292
Leucotrogus Nikolajev, 2004 260
Leuroserica Arrow, 1946 290
Liatongus Reitter, 1892 176
Lichas Westwood, 1853 623
Ligniperda Fabricius, 1790 60
Limaroides Balthasar, 1932 144
Limarus Mulsant & Rey, 1870 129
Limnichomorphus Pic, 1922 608
Limnichus Dejean, 1821 608
Limnichus Latreille, 1829 608

Index to genus-group names
Limnius Illiger, 1802 595
Lindbergianus Petrovitz, 1961 129
Liocola C.G. Thomson, 1859 377
Liogastria Volkovitsh, 1979 450
Liogenys Guérin-Méneville, 1831 214
Lioglyptoxenus Pittino, 2006 165
Liothorax Motschulsky, 1860 129
Lisposcelis Volkovitsh, 1979 442
Lissadoretus Arrow, 1917 320
Litocopris C.O. Waterhouse, 1891 168
Littorimus Gozis, 1885 610
Loboparius A. Schmidt, 1913 130
Loepotethya Volkovitsh, 1979 442
Loraphodius Reitter, 1892 130
Loraspis Mulsant & Rey, 1870 131
Loricaster Mulsant & Rey, 1861 428
Lucanus Scopoli, 1763 74
Lucidonthophagus Ádám, 1994 190
Ludibrius Gozis, 1886 230
Lunaphodius Balthasar, 1964 131
Lycaste Gistel, 1834 453
Macrator Reitter, 1895 362
Macrelmis Motschulsky, 1860 596
Macrobyrrhinus Pic, 1922 607
Macrodorcas Motschulsky, 1862 62
Macrodorcinus Maes, 1990 68
Macroeubria Pic, 1916 618
Macrognathus Hope & Westwood, 1845 68
Macrolinus Kaup, 1868 53
Macroliocola Alexis & Delpont, 1998 378
Macronotops Krikken, 1977 403
Macronoxia Crotch, 1873 235
Macronthophagus Ochi, 2003 188
Macronychus P.W.J. Müller, 1806 602
Macropropus Agassiz, 1846 214
Macroprotaetia Mikšić, 1965 378
Macroretrus Péringuey, 1908 131
Macroryctes Semenov & S.I. Medvedev, 1932 360
Macroserica S.I. Medvedev, 1952 292
Madateuchus R. Paulian, 1953 206
Madotrogus Reitter, 1902 261
Madrasostes R. Paulian, 1975 85
Malacopsephenoides Jeng & Satô, 2006 620
Maladera Mulsant & Rey, 1871 290
Maladera Mulsant & Rey, 1871 293
Malaiseianus Bollow, 1940 607
Malaisius Arrow, 1941 226
Maraxes Lewis, 1895 167
Marcsikiella Hołyński, 2014 460

975

Index To Genus-group Names
Mastogenius Solier, 1851 453
Mataeopsephus C.O. Waterhouse, 1876 620
Matashia Matsumura, 1938 188
Mawenzhena Alexis & Delpont, 2001 395
Mecynodes Mulsant & Rey, 1870 131
Medeotrogus Keith, 2001 271
Megalisus Mulsant & Rey, 1870 114
Megaloprepes J. Thomson, 1862 61
Megalotrox Preudhomme de Borre, 1886 54
Megaloxantha Kerremans, 1903 456
Meganoxia Reitter, 1902 229
Megapertha Reitter, 1903 343
Megarhipis Laporte, 1834 431
Megatelus Reitter, 1892 132
Megistophylla Burmesiter, 1855 271
Melanomaladera Miyake & Yamaya, 2001 283
Melanophila Eschscholtz, 1829 521
Melanopopillia Lin, 1980 343
Melanosa Mulsant & Rey, 1871 378
Melanotrupes F. Blanchard, 1888 39
Melanthaxia Richter, 1949 510
Melaphodius Reitter, 1892 113
Melaserica Brenske, 1897 282
Meliacanthus Théry, 1942 562
Meliboeoides Théry, 1942 562
Meliboeus Deyrolle, 1864 562
Melichrus Brenske, 1892 215
Melicoraebus Théry, 1932 562
Melinopterus Mulsant, 1842 132
Melinospila Kraatz, 1890 403
Melixes Schaefer, 1949 562
Melolontha Fabricius, 1775 230
Mendidaphodius Reitter, 1901 134
Mendidius Harold, 1868 134
Meraphodius D. Koshantschikov, 1894 131
Meretricula Özdikmen, 2008 393
Merocratus Bílý, 1989 513
Meroloba J. Thomson, 1880 403
Mesanoxia S.I. Medvedev, 1951 227
Mesanthaxia Richter, 1945 503
Mescirtes Motschulsky, 1863 421
Mesolethrus Nikolajev, 2003 50
Mesontoplatys Motschulsky, 1864 136
Mesopolyphylla S.I. Medvedev, 1951 235
Mesoscarabaeus Zídek & Pokorný, 2008 205
Mesoserica Matsumura, 1911 312
Mesotrypes Jakobson, 1892 38
Messyrhus Rakovič & Král, 1997 161
Metabolus Fairmaire, 1887 274
Metacatharsius Montreuil, 1998 170

Metaceraspis Frey, 1962 212
Metagymnopleurus Kabakov, 2006 174
Metallactulus Ritsema, 1885 68
Metapopillia H.J. Kolbe, 1894 352
Metasambus Kerremans, 1903 555
Metatoxoscelus Hattori, 1990 566
Metonius Say, 1833 567
Metopodontus Hope & Westwood, 1845 68
Metopotropus R. Oberthür & Houlbert, 1913 68
Micilus Mulsant & Rey, 1872 616
Micragrilus Alexeev, 1998 524
Micramphimallon S.I. Medvedev, 1951 261
Micreubrianax Pic, 1922 620
Microacmaeodera Cobos, 1966 451
Microacmaeodera Cobos, 1966 451
Microcamenta Brenske, 1898 210
Microcara C.G. Thomson, 1859 421
Microcopris Balthasar, 1958 171
Microcosmiomorpha Mikšić, 1974 393
Microdes Motschulsky, 1860 594
Microdonta Hope, 1837 250
Microdrupeus Nakane, 1993 622
Microeubria Lee & P.-S. Yang, 1994 619
Microgenestia Bellamy, 1986 470
Microlethrus Semenov, 1894 47
Microlontha Petrovitz, 1967 344
Micromicilus J.R. Sahlberg, 1900 616
Micronthophagus Balthasar, 1963 189
Micropertha Baraud, 1991 344
Microphylla Kraatz, 1890 234
Microrhipis Guérin-Méneville, 1831 431
Microryctes Arrow, 1908 363
Microserica Brenske, 1894 298
Microtrichia Brenske, 1900 278
Miksicoprotaetia Legrand & Chew
Kea Foo, 2010 378
Millingenia Sharp, 1874 98
Miltotrogus Reitter, 1902 266
Mimela Kirby, 1823 344
Minotaurus Mulsant & Godart, 1855 38
Miridiba Reitter, 1902 271
Mitshekia Hołyński, 1994 460
Miwanus H. Huang & C.-C. Chen, 2013 68
Mnematidium Ritsema, 1889 206
Mnematium W.S. Macleay, 1821 206
Monotropus Erichson, 1847 272
Morphobyrrhulus Pütz, 2007 578
Morychus Erichson, 1847 578
Moseriana Ruter, 1965 395
Mothon Semenov & S.I. Medvedev, 1927 136

976
Mozartius Nomura & Nakane, 1951
Mundaria Kerremans, 1894 562
Musurgus Vauloger, 1898 363
Mycteristes Laporte, 1840 400
Myrhessus Balthasar, 1956 161

Index to genus-group names
136

Nalanda Théry, 1904 562
Nannovalgus H.J. Kolbe, 1904 409
Neagolius W. Koshantschikov, 1912 136
Neateuchus Gillet, 1911 206
Negreia Cobos, 1962 556
Nematopsephus Jäch & Jeng, 1995 621
Neoblitopertha Baraud, 1991 342
Neobodilus Hollande & Thérond, 1999 113
Neocalaphodius Bordat, 1990 137
Neochaetopisthes Wasmann, 1918 155
Neocoraebus Kerremans, 1903 556
Neoctodon Bedel, 1892 206
Neodipelicus Ritsema, 1887 362
Neodontocnema Arrow, 1948 271
Neolethrus Nikolajev, 1987 50
Neolucanus J. Thomson, 1862 81
Neomaladera Baraud, 1965 299
Neomnematium A. Janssens, 1938 206
Neophaedimus P.H. Lucas, 1870 395
Neoptosima Théry, 1905 453
Neoriohelmis Nomura, 1958 596
Neosambus Fisher, 1921 562
Neosaprosites Endrődi, 1951 157
Neoserica Brenske, 1894 299
Neosericania Miyake & Yamaya, 1994 301
Neosisyphus J. Müller, 1942 207
Neotemnorrhynchus A. Janssens, 1949 366
Neotoxoscelus Fisher, 1921 566
Neotrichiorhyssemus Rakovič & Král, 1997 161
Neovalgus Miyake, 1985 411
Nepaloserica Frey, 1965 301
Netocia A. Costa, 1852 378
Nialaphodius H.J. Kolbe, 1908 138
Nialosternus Hollande & Thérond, 1999 138
Nialus Mulsant & Rey, 1870 138
Nicagus LeConte, 1861 59
Niehuisia Curletti, 1995 555
Nigidionus Kriesche, 1926 73
Nigidius W.S. Macleay, 1819 73
Nimbus Mulsant & Rey, 1870 138
Niponoprotaetia Mikšić, 1966 381
Nipponaphodius Nakane, 1959 139
Nipponeubria Lee & Satô, 1996 619
Nipponoagoliinus Ochi & Kawahara, 2001 139

Nipponobuprestis Tôyama, 1986 460
Nipponobuprestisia Peng, 1995 461
Nipponocyphon Lawrence & Yoshitomi, 2007 412
Nipponodorcus Nomura & Kurosawa, 1960 62
Nipponoserica Nomura, 1973 302
Nipponovalgus Sawada, 1941 411
Noanda Reitter, 1902 278
Nobiellus M. Dellacasa & G. Dellacasa, 2005 139
Nobius Mulsant & Rey, 1870 140
Nomuraelmis Satô, 1964 596
Nongoma Péringuey, 1902 328
Normandia Pic, 1900 598
Norosus Mulsant & Rey, 1869 587
Noseolucanus Araya & Tanaka, 1998 79
Nothochodaeus Nikolajev, 2005 85
Notiocyphon Blackburn, 1896 607
Nycteus Latreille, 1829 425
Nyholmia Klausnitzer, 2013 421
Obenbergerietta E. Strand, 1942 452
Ochicanthon Vaz de Mello, 2003 167
Ochodaeus Dejean, 1821 84
Ochranoxia Kraatz, 1888 226
Octodon van Lansberge, 1875 206
Odeles Klausnitzer, 2004 422
Odochilus Harold, 1877 158
Odontacrossus G. Dellacasa, Král,
M. Dellacasa & Bordat, 2014 140
Odontaeus Dejean, 1821 35
Odontanax Lee, Satô & P.-S. Yang, 2000 617
Odonteus Samouelle, 1819 35
Odontiellus Keith & Montreuil, 2004 246
Odontolabis Hope, 1842 83
Odontomus Kirby, 1837 517
Odontonyx Guérin-Méneville, 1843 622
Odontotrupes Boucomont, 1905 43
Odontotrypes Fairmaire, 1887 43
Ohkubous Sawada, 1938 357
Oligophylla Kraatz, 1894 234
Oloperus Mulsant & Rey, 1870 146
Omaladera Reitter, 1896 294
Omaloplia Schönherr, 1817 303
Omorgus Erichson, 1847 54
Omphalothorax Volkovitsh, 1979 451
Oniticellus Dejean, 1821 177
Oniticellus Le Peletier de St. Fardeau &
Audinet-Serville, 1828 177
Onitis Fabricius, 1798 178
Onthophagiellus Balthasar, 1935 189
Onthophagus Latreille, 1802 184

977

Index To Genus-group Names
Onthophagus Latreille, 1802 189
Onthosphaenus Motschulsky, 1860 176
Onthotrogus Motschulsky, 1860 199
Onychosophrops Frey, 1972 272
Onychothecus Boucomont, 1912 166
Ootoma C.É. Blanchard, 1850 241
Ophrygonius Zang, 1904 53
Ophthalmoserica Brenske, 1897 307
Oplistura Jakovlev, 1889 482
Oplosternus Guérin-Méneville, 1838 230
Optioservus Sanderson, 1953 596
Ora Clark, 1865 423
Orchiserica Miyake & Yamaya, 2001 289
Ordobrevia Sanderson, 1953 599
Oreocyphon Klausnitzer, 2008 416
Oreoderus Burmeister, 1842 411
Oreopotosia S.I. Medvedev, 1964 381
Orientagrilus Alexeev, 1998 524
Orientelmis Shepard, 1998 597
Ornatobyrrhus Fabbri, 2000 585
Orodaliscus Reitter, 1900 140
Orodalus Mulsant & Rey, 1870 123
Oromia López-Colón, 1989 241
Oromiella López-Colón, 1992 241
Oromus Mulsant & Rey, 1870 140
Orphnomala Reitter, 1903 328
Orphnus W.S. Macleay, 1819 210
Orthocheira Richter, 1952 516
Orubesa Reitter, 1895 208
Oryctes Hellwig, 1798 359
Oryx Guérin-Méneville, 1838 365
Osmanius Branco & Baraud, 1988 141
Osmoderma LePeletier &
Audinet-Serville, 1828 399
Otoclinius Brenske, 1897 241
Otophorus Mulsant, 1842 141
Oulimnius Gozis, 1886 597
Oxyadoretus Arrow, 1917 321
Oxyaphodius Balthasar, 1965 141
Oxycetonia Arrow, 1910 371
Oxycorythus Solsky, 1876 155
Oxyomus Dejean, 1833 141
Oxypteris Kirby, 1837 521
Oxyserica Brenske, 1900 305
Oxythyrea Mulsant, 1842 389
Pachnoda Burmeister, 1842 374
Pachnodula Schoch, 1898 374
Pachnotosia Reitter, 1891 377
Pachychaeta Brenske, 1897 237

Pachydema Laporte, 1832 241
Pachydemocera Reitter, 1902 245
Pachymerus Faldermann, 1835 95
Pachyparnus Fairmaire, 1889 606
Pachyprotaetia Mikšić, 1965 381
Pachypus Audinet-Serville, 1825 249
Pachypus Dejean, 1821 249
Pachyrhinadoretus Ohaus, 1912 321
Pachyschelus Solier, 1833 567
Pachyserica Brenske, 1897 305
Pachystethus C.É. Blanchard, 1851 341
Pakistanotrogus Keith, 2006 267
Palaeonthophagus Zunino, 1979 190
Palaeotethya Volkovitsh, 1979 442
Paleira Reiche, 1871 390
Palmar Schaefer, 1949 469
Palmerstonia Blackburn, 1888 362
Palnia Stebnicka, 1985 166
Palpilatis Bílý, 1980 465
Panelus Lewis, 1895 167
Panotrogus Reitter, 1902 272
Papuana Arrow, 1911 363
Parabodilus Hollande & Thérond, 1999 142
Parachorius Harold, 1873 166
Paracmaeodera Théry, 1946 443
Paracopris Balthasar, 1939 171
Paracoptochirus Balthasar, 1963 142
Paracrossidius Balthasar, 1932 142
Paracrusis Newman, 1839 344
Paracryptodactylus Alexeev, 1975 566
Paracylindromorphus Théry, 1930 553
Paradeudora Obenberger, 1924 484
Paradidactylia Balthasar, 1937 142
Paragematis Reitter, 1903 328
Paragolius Balthasar, 1945 145
Paragymnopleurus Shipp, 1897 174
Paragynaecoserica Khan & Ghai, 1982 286
Parahabroloma Kurosawa, 1959 568
Parahelichus Löbl & Smetana, 2006 606
Paraixodina Roggero, Barbero &
Palestrini, 2015 174
Paralethrus Nikolajev, 2003 51
Paralichas White, 1859 622
Paralophotus Curletti, 1998 524
Paramacronychus Nomura, 1958 602
Paramaladera Nomura, 1974 306
Paramelinopterus Rakovič, 1984 113
Paramimela Ohaus, 1915 344
Parammoecius Seidlitz, 1888 143
Paranomala Casey, 1915 328

978
Paranthaxia Gozis, 1886 495
Paraphanaeomorphus Balthasar, 1959 197
Paraphytus Harold, 1877 167
Parapilinurgus Arrow, 1910 392
Pararhinyptia Machatschke, 1957 327
Pararhyssemus Balthasar, 1956 161
Parasambus Descarpentries & Villiers, 1966 549
Parascatonomus R. Paulian, 1932 197
Paraserica Reitter, 1896 306
Parastasia Westwood, 1842 357
Parataenius Balthasar, 1961 157
Paratassa Marseul, 1882 467
Paratethya Bílý, 1983 452
Parateuchus Shipp, 1895 206
Paratrachys E. Saunders, 1873 453
Paratrichius Janson, 1881 406
Paratriodonta Baraud, 1962 306
Paremadus Nakane, 1967 108
Parentius Zunino, 1979 198
Parexolontha Chang, 1965 229
Parnus Fabricius, 1792 603
Parodochilus Rakovič, 1997 158
Paroniticellus Balthasar, 1963 177
Parvulomaladera D. Ahrens, 1995 305
Passalaphodius Martinez, 1953 157
Paulianellus Balthasar, 1938 143
Pectinichelus Ballion, 1871 273
Pedilophorus Steffahny, 1843 580
Pelochares Mulsant & Rey, 1869 609
Pelocherops Pic, 1923 607
Peltonotus Burmeister, 1847 358
Penichrolucanus Deyrolle, 1863 73
Pentodon Hope, 1837 363
Peperonota Westwood, 1847 358
Peradoretus Reitter, 1903 320
Periphanesthes Kraatz, 1880 395
Peritryssus Reitter, 1918 214
Perotis Dejean, 1833 465
Personatus Curletti, 1993 524
Petalon Schönherr, 1833 430
Petrovitzia López-Colón, 1996 42
Petrovitzia Mikšić, 1965 395
Phacosoma Boucomont, 1914 167
Phaeadoretus Reitter, 1903 321
Phaeaphodius Reitter, 1892 144
Phaenops Dejean, 1833 522
Phaeochroops Candèze, 1876 86
Phaeochrous Laporte, 1840 86
Phaeocroides Péringuey, 1908 87
Phalacronothus Motschulsky, 1860 145
Phalangonyx Reitter, 1889 245

Index to genus-group names
Phalantha Gistel, 1834 458
Phalops Erichson, 1848 204
Phanaeomorphus Balthasar, 1963 199
Pharaonus C.É. Blanchard, 1851 352
Pharaphodius Reitter, 1892 146
Phelix Marseul, 1865 466
Phelotrupes Jekel, 1866 37
Philanthaxia Deyrolle, 1864 523
Philhelena J. Thomson, 1880 381
Philistina W.S. Macleay, 1838 400
Philocteanus Deyrolle, 1864 458
Philydrus Duftschmid, 1805 592
Phlexis Erichson, 1841 241
Phoberus W.S. Macleay, 1819 57
Photyna Brenske, 1897 212
Phygotoxeuma Brenske, 1897 241
Phyllis Gistel, 1834 433
Phyllognathus Eschscholtz, 1830 365
Phyllopertha Stephens, 1830 348
Phyrites Schiödte, 1866 613
Physopalpus Heyden, 1900 242
Phytotera Gistel, 1856 569
Pilleriana Baraud, 1991 325
Pittinius Rakovič & Král, 1997 161
Placonycha Horn, 1880 616
Plagiogonus Mulsant, 1842 146
Planolinellus M. Dellacasa &
G. Dellacasa, 2005 147
Planolinoides M. Dellacasa &
G. Dellacasa, 2005 148
Planolinus Mulsant & Rey, 1870 148
Platycephalus Cuvier, 1797 109
Platycerus Geoffroy, 1762 79
Platycerus Gory, 1833 73
Platycerus Latreille, 1806 59
Platyderides A. Schmidt, 1916 148
Platyderus A. Schmidt, 1913 148
Platypelochares G.C. Champion, 1923 609
Platyprosopus Hope & Westwood, 1845 71
Platysodes Westwood, 1873 391
Platytomus Mulsant, 1842 161
Pledina Reitter, 1902 272
Pleopertha Reitter, 1903 342
Pleuraphodius A. Schmidt, 1913 148
Pleurophorus Mulsant, 1842 162
Plocovalgus H.J. Kolbe, 1904 409
Podalgus Burmeister, 1847 366
Podoserica Breit, 1912 307
Poecilonota Eschscholtz, 1829 470
Poecilophilides Kraatz, 1898 392
Poecilosticta Kraatz, 1892 341

Index To Genus-group Names
Pogonodorcus Séguy, 1954 67
Pollaplonyx C.O. Waterhouse, 1875 274
Polycesta Dejean, 1833 453
Polyctesis Marseul, 1865 455
Polydomia J. Thomson, 1880 403
Polylamina A. R. Hardy, 1974 235
Polymoechus LeConte, 1856 357
Polyonychus Chevrolat, 1838 566
Polyphylla Harris, 1841 234
Polyphylla Harris, 1841 235
Polytrichius Heer, 1834 407
Pomatinus Sturm, 1853 606
Ponerotrogus Silvestri, 1924 183
Popillia Dejean, 1821 352
Porcinolus Mulsant & Rey, 1869 588
Porropus Laporte, 1840 214
Potamodytes Grouvelle, 1896 591
Potamophilinus Grouvelle, 1896 592
Potamophilus Germar, 1811 592
Potosia Mulsant & Rey, 1871 382
Praehelichus Löbl & Smetana, 2006 606
Prionadoretus Ohaus, 1914 317
Prionocyphon L. Redtenbacher, 1858 423
Prismognathus Motschulsky, 1860 78
Pristadoretus Reitter, 1903 320
Proagoderus van Lansberge, 1883 199
Proagopertha Reitter, 1903 349
Procantonius Kalashian, 2004 553
Proeuzkus Vít, 2000 426
Progymnopleurus Garreta, 1914 174
Propomacrus Newman, 1837 214
Prosopocoilus Hope & Westwood, 1845 68
Protaetia Burmeister, 1842 374
Protaetia Burmeister, 1842 386
Protaetiola Mikšić, 1963 386
Protoctenius S.I. Medvedev, 1952 245
Psammobius Heer, 1841 159
Psammodius Fallén, 1807 159
Psammoporus C.G. Thomson, 1859 97
Psephenoides Gahan, 1914 620
Psephus Kirby & Spence, 1826 84
Pseudacrossus Reitter, 1892 148
Pseudadoretus Semenov, 1889 320
Pseudagenius Heller, 1923 407
Pseudagrilus Laporte, 1835 549
Pseudammoecius A. Schmidt, 1913 156
Pseudamophilus Bollow, 1940 598
Pseudanthracophora Kraatz, 1898 376
Pseuderytus Hollande & Thérond, 1999 149
Pseudesymus d’Orbigny, 1896 150
Pseudianthe Fairmaire, 1901 452

979
Pseudoahermodontus G. Dellacasa & M. Dellacasa,
1997 150
Pseudoapterogyna Escalera, 1914 263
Pseudoathyreus Howden & Martínez, 1963 33
Pseudoblitopertha Keith, 2000 356
Pseudocalamosternus Balthasar, 1936 128
Pseudocalopotosia Mikšić, 1966 386
Pseudocastalia Kraatz, 1896 453
Pseudochelonarium Pic, 1916 623
Pseudocladotoma Pic, 1918 622
Pseudocymophorus Schein, 1954 391
Pseudodiceros Mikšić, 1974 396
Pseudoepilichas Armstrong & Nakane, 1956 621
Pseudoeuselates Mikšić, 1974 402
Pseudoheptaulacus M. Dellacasa &
G. Dellacasa, 2000 150
Pseudohoplia Reitter, 1890 222
Pseudolichas Fairmaire, 1878 430
Pseudolimarus Balthasar, 1931 103
Pseudolontha Fairmaire, 1897 256
Pseudolucanus Hope & Westwood, 1845 74
Pseudomaladera Nikolajev, 1980 302
Pseudomaladera Nomura, 1974 307
Pseudomothon Pittino, 1984 155
Pseudonetocia S.I. Medvedev, 1964 386
Pseudoniticellus Kraatz, 1895 177
Pseudonthophagus Balthasar, 1959 197
Pseudooxythyrea Baraud, 1985 390
Pseudopachydema Balthasar, 1930 245
Pseudopanotrogus Petrovitz, 1969 274
Pseudopopillia H.J. Kolbe, 1894 352
Pseudoquercagrilus Alexeev, 1998 524
Pseudorhaetus Planet, 1899 71
Pseudorhomborrhina Mikšić, 1956 396
Pseudosericania Kobayashi, 1980 307
Pseudosinghala Heller, 1891 349
Pseudosugrames Kiseritzky, 1928 152
Pseudosymmachia Dalla Torre, 1913 274
Pseudoteuchestes G. Dellacasa, 1986 150
Pseudothryptus Hernando & Ribera, 2005 609
Pseudotocama Keith, 2008 236
Pseudotorynorrhina Mikšić, 1967 396
Pseudotrematodes Jacquelin du Val, 1860 275
Pseudotrox Robinson, 1948 55
Pseudotrypocopris Mikšić, 1954 46
Pseudyamina Richter, 1952 516
Psilodema C.É. Blanchard, 1845 89
Pterorthochaetes Gestro, 1898 86
Ptilodactyla Illiger, 1807 622
Ptosima Dejean, 1833 455
Ptychomus Marseul, 1865 444

980

Index to genus-group names

Ptychophorus Schaum, 1841 391
Ptyocerus C. Thunberg, 1806 431
Pubinus Mulsant & Rey, 1870 150
Pusilloderes Obenberger, 1916 555
Pygopleurus Motschulsky, 1860 92
Pyranthe Gistel, 1834 457
Pyropotosia Reitter, 1899 386
Qingaphodius Král, 1997 150
Quercuagrilus Alexeev, 1998 524
Rakovicius Pittino, 2006 160
Redotus Fairmaire, 1860 281
Reitterius López-Colón, 1996 42
Relictonthophagus Kabakov, 1979 198
Renaudia López-Colón, 1996 44
Renaudius Balthasar, 1951 150
Renaudtrupes López-Colón, 2006 44
Rhabdotis Burmeister, 1842 387
Rhaetulus Westwood, 1871 71
Rhaetus Parry, 1864 71
Rhampadoretus Ohaus, 1912 320
Rhaphidochila Jakovlev, 1900 478
Rhinyptia Burmeister, 1844 327
Rhizocolax Motschulsky, 1860 256
Rhizocolax Motschulsky, 1860 256
Rhizoplatys Westwood, 1841 367
Rhizotrogus Latreille, 1825 275
Rhodaphodius Ádám, 1994 150
Rhombonyx Hope, 1837 344
Rhomborhina Hope, 1837 396
Rhynchosymmela Frey, 1974 287
Rhyparus Agassiz, 1846 165
Rhysothorax Bedel, 1911 97
Rhyssemodes Reitter, 1892 162
Rhyssemus Mulsant, 1842 163
Riolus Mulsant & Rey, 1872 598
Robertius Théry, 1947 524
Rosagrilus Alexeev, 1998 524
Rossiella Obenberger, 1942 460
Rotundobyrrhus Fabbri, 2000 585
Rudeulasia Baraud, 1990 90
Rudolfpetrovitzia A. Rey & López-Colón, 2003
Rugacmaeodera Holm, 1978 444
Rykanes Minck, 1919 360
Rykanoryctes Minck, 1919 360
Ryparus Westwood, 1845 165
Saccosoma Motschulsky, 1859
Sacodes LeConte, 1854 423

433

42

Sahlbergianus M. Dellacasa & G. Dellacasa,
2002 151
Samboides Kerremans, 1900 524
Sambus Deyrolle, 1864 549
Sandalus Knoch, 1801 431
Sapaia Bílý, 1994 461
Saprosites L. Redtenbacher, 1858 157
Sarawakita Obenberger, 1924 524
Scarabaeus Linnaeus, 1758 206
Scelolethrus Semenov, 1892 51
Schinostethus C.O. Waterhouse, 1880 619
Schinostethus C.O. Waterhouse, 1880 619
Schismatocera Gautier des Cottes, 1872 214
Schistocometa Brenske, 1903 256
Schizonycha Dejean, 1833 281
Schoenherria Burmeister, 1855 228
Schoutedenia Obenberger, 1924 455
Schoutedeniastes Burgeon, 1941 455
Scintillatrix Obenberger, 1956 467
Scirtes Illiger, 1807 424
Sebasteos Westwood, 1847 206
Seleucosorus Kuijten, 1983 87
Seminolus Mulsant & Rey, 1869 586
Serica W.S. Macleay, 1819 307
Sericania Motschulsky, 1860 312
Sericotrupes Zunino, 1984 45
Serraphodius Kabakov, 1996 151
Serrognathus Motschulsky, 1862 71
Serrophorus Balthasar, 1963 199
Setylaides Stebnicka, 1994 158
Shangaia R. Lucas, 1920 271
Shanna Pütz, 2007 576
Sicardia Reitter, 1896 160
Sicardia Théry, 1909 452
Sigorus Mulsant & Rey, 1870 151
Silluvia Landin, 1949 98
Silotrupes Mulsant & Rey, 1871 46
Silphodes Westwood, 1841 86
Silphotrupes Jekel, 1866 45
Simianus C.É. Blanchard, 1853 624
Simplocaria Stephens, 1829 576
Simplocaria Stephens, 1829 576
Sinagrilus Alexeev, 1998 524
Sinaphodius Červenka, 1994 151
Sinelmis Satô & Kishimoto, 2001 599
Sinobyrrhus Fabbri, 2000 586
Sinocaulus Fairmaire, 1878 430
Sinochelus Fairmaire, 1900 211
Sinocopris Ochi, Kon & Bai, 2009 170
Sinodendron Hellwig, 1792 60

981

Index To Genus-group Names
Sinodiapterna G. Dellacasa, 1986 151
Sinodorcus H. Huang & C.-C. Chen, 2013 72
Sinodrepanus Simonis, 1985 174
Sinogeotrupes Bovo & Zunino, 1983 38
Sinokele Bílý, 1989 517
Sinonthophagus Kabakov, 2006 200
Sinonychus Jäch & Boukal, 1995 602
Sinoplia S.I. Medvedev, 1952 222
Sinopsephenoides J. Yang, 1993 621
Sinopsephenus Nakane, 1964 620
Sinorychomus Fabbri, 2003 579
Sinoserica Miyake & Yamaya, 2001 316
Sinuatiagrilus Alexeev, 1998 524
Sissantobius Ritsema, 1889 210
Sisyphus Latreille, 1806 207
Sitiphus Fairmaire, 1894 152
Socotrabolbus Cambefort, 1998 35
Socotraproctus Král, Sehnal & Bezděk, 2012 246
Solskiola Semenov, 1903 91
Sophrops Fairmaire, 1887 278
Sostea Pascoe, 1860 607
Sparophysa Burmeister, 1855 242
Spermohybosorus Pic, 1922 87
Sphaerobothris Semenov & Richter, 1934 520
Sphaerotrochalus Brenske, 1900 314
Sphenoptera Dejean, 1833 471
Sphenoptera Dejean, 1833 484
Sphenopterella Volkovitsh & Kalashian, 1994 493
Sphodroxia Kraatz, 1890 236
Spileuchlora Ohaus, 1903 341
Spilopopillia Kraatz, 1892 356
Spilota Burmeister, 1844 341
Spinanomala Ohaus, 1910 350
Spineubria Nakane, 1952 617
Spinigymnopleurus Shipp, 1897 172
Spinohoplia Sabatinelli, 1997 216
Spinthoptera Casey, 1909 464
Spiragrilus Alexeev, 1998 524
Sponsor Gory & Laporte, 1839 453
Sponsor Gory & Laporte, 1839 453
Stalagmopygus Kraatz, 1882 387
Stalagmosoma Burmeister, 1842 387
Steffahnyellus Portevin, 1931 578
Stenactyla Fairmaire, 1896 622
Stenelmis Dufour, 1835 599
Stenobronchus A. Schmidt, 1916 152
Stenomystax Kodada, Jäch & Čiampor, 2003 607
Stenonota Fairmaire, 1889 403
Stenosophrops Nomura, 1977 279
Stenothorax A. Schmidt, 1916 152

Stenotothorax A. Schmidt, 1913 152
Stenuris Kirby, 1837 463
Steraspis Dejean, 1833 458
Stereopyge A. Costa, 1847 41
Sternocera Eschscholtz, 1829 437
Sternoplidius Moser, 1908 372
Sternotrupes Jekel, 1866 46
Sternoxus Billberg, 1820 437
Sternuchus LeConte, 1850 426
Stichadoretus Reitter, 1903 320
Stigmatophorella Tôyama, 1986 460
Stomphax Fischer von Wladheim, 1823
Strandius Balthasar, 1963 200
Streopuge A. Costa, 1847 41
Strigoptera Dejean, 1833 454
Strigopteroides Cobos, 1981 454
Subrinus Mulsant & Rey, 1870 152
Subulistomella M. Sakai, 1980 426
Sugrames Reitter, 1894 152
Sulcodrepanus Krikken, 2009 175
Sundodrupeus Pic, 1916 619
Sunenaga Ochi, 2003 200
Svatacesta Zabransky, 2004 453
Sybacodes Fairmaire, 1896 166
Symmachia Brenske, 1892 274
Synapsis Bates, 1868 171
Syncalypta Dillwyn, 1829 588
Syngonovalgus H.J. Kolbe, 1904 409
Systenocerus J. Weise, 1883 79

84

Taenheterocerus Kuwert, 1890 610
Taeniocerus Kaup, 1871 52
Taeniodera Burmeister, 1842 403
Taiwanoserica Nomura, 1974 310
Taiwanotrichia Kobayashi, 1990 216
Tamamushia Miwa & Chûjô, 1935 461
Tamerlanius Keith, Sabatinelli & Uliana, 2015
Tamina Gistel, 1848 517
Tanyproctocera Reitter, 1902 246
Tanyproctoides Petrovitz, 1971 246
Tanyproctus Ménétriés, 1832 246
Tanyproctus Ménétriés, 1832 247
Tarandus Dejean, 1833 59
Taxipertha Baraud, 1991 343
Tecinoa A. Costa, 1852 368
Telodorcus Didier, 1937 62
Temnorhynchus Hope, 1837 366
Teratolethrus Semenov, 1894 52
Teres Harris, 1829 524
Termitopisthes Wasman, 1918 155

352

982
Tesarius Rakovič, 1981 160
Tetragonus Gory & Percheron, 1833 400
Tetraproctus Iablokoff-Khnzorian, 1953 249
Tetrarthrius Didier, 1926 79
Tetraserica D. Ahrens, 2004 314
Tetrodontus Y.-W. Zhang, 1991 367
Teuchestes Mulsant, 1842 153
Thaiaphodius Masumoto, 1991 148
Thailandia Bílý, 1990 513
Thaumastopeus Kraatz, 1885 401
Theotimius Huchet, 2000 166
Therilus Mulsant & Rey, 1870 113
Theriomorphus Pic, 1913 622
Theryola Nelson, 1997 454
Therysambus Descarpentries & Villiers, 1967 524
Thinorycter Semenov & Reichardt, 1925 208
Thorectes Mulsant, 1842 45
Thorectomimus Nikolajev, 2009 44
Thurntaxisia Schatzmayr, 1929 454
Thyreogonia Reitter, 1899 387
Tiberioides Gravely, 1913 53
Tibiodrepanus Krikken, 2009 175
Tibiotrichius Miyake, 1994 407
Tinclirea J. Thomson, 1880 399
Tiniocellus Péringuey, 1902 177
Tocama Reitter, 1902 236
Tohlezkus Vít, 1977 426
Tokaranodicerca Hattori, 2004 466
Tolisus Sharp, 1874 98
Tomentoprotaetia Mikšić, 1987 387
Tonkinula Obenberger, 1923 562
Torynorrhina Arrow, 1907 397
Tosevskiana Pavićević, 1985 280
Touzalinia Théry, 1922 466
Toxoscelus Deyrolle, 1864 567
Toxospathius Fairmaire, 1878 216
Trachypteris Kirby, 1837 523
Trachys Fabricius, 1801 569
Trematodes Faldermann, 1835 280
Tribopertha Reitter, 1903 356
Trichanomala Arrow, 1917 350
Trichaphodius A. Schmidt, 1913 153
Trichiorhyssemus Clouët des Pesruches, 1901 165
Trichius Fabricius, 1775 407
Trichobyrrhulus Ganglbauer, 1902 578
Trichocratomerus Richter, 1945 495
Trichogomphus Burmeister, 1847 361
Tricholontha Nomura, 1952 236
Trichomaladera Nomura, 1974 315
Trichonotulus Bedel, 1911 153
Trichonotus Mulsant, 1842 153

Index to genus-group names
Trichonthophagus Zunino, 1979 200
Trichopertha Reitter, 1903 343
Trichopleurus Motschulsky, 1860 91
Trichopopillia Ohaus, 1897 356
Trichoserica Reitter, 1896 307
Trichovalgus H.J. Kolbe, 1904 409
Triclirea Schoch, 1895 399
Trigonocnemis Kraatz, 1894 350
Trigonophorinus Pouillaude, 1913 397
Trigonophorus Hope, 1831 397
Trigonoscelus Petrovitz, 1963 155
Trimela Ohaus, 1924 344
Trinaria Mulsant & Rey, 1869 577
Triodonta Mulsant, 1842 315
Triodontella Reitter, 1919 315
Trioserica Moser, 1922 316
Trirania Mroczkowski, 1983 577
Tristaphodius Balthasar, 1932 154
Tristria Hołyński, 2001 466
Trochaloschema Reitter, 1896 316
Trochaloserica Brenske, 1900 281
Troglonthophagus Ádám, 1994 190
Tropeopeltis Jakovlev, 1902 493
Tropinota Mulsant, 1842 387
Tropinota Mulsant, 1842 388
Tropiorhynchus C.É. Blanchard, 1851 327
Trox Fabricius, 1775 55
Trypocopris Motschulsky, 1860 46
Trypocopris Motschulsky, 1860 46
Trypoxylus Minck, 1920 358
Tuberniella Jendek & Kalashian, 1999 521
Turanella Semenov, 1905 154
Typhaeus Leach, 1815 38
Uenodryops Satô, 1981 603
Uposlotus A. Costa, 1853 177
Uragrilus Semenov, 1935 524
Uralohelmis Roubal, 1940 596
Urleta Westwood, 1875 357
Urumaelmis Satô, 1963 602
Urumaeubria Satô, 1968 618
Valgus Scriba, 1790 411
Vertumnus Reiche, 1859 366
Victoria Miyahara, 1941 218
Vietelmis Delève, 1968 602
Vietnamoprotaetia Mikšić, 1971 387
Vittateulasia Baraud, 1990 91
Vladimirellus M. Dellacasa, G. Dellacasa &
Bordat, 2002 154
Volinus Mulsant & Rey, 1870 154

983

Index To Genus-group Names
Wallaceilus Hołyński, 2003 524
Weinreichius J.P. Lacroix, 1978 71
Weyersiella Théry, 1930 549
Wooldridgeus Spangler, 1999 609
Xanthelaeus Fairmaire, 1897 208
Xantheremia Volkovitsh, 1979 452
Xanthotrogus Reitter, 1902 280
Xanthotrogus Reitter, 1902 280
Xenoceraspis Arrow, 1920 212
Xenogenius H.J. Kolbe, 1892 390
Xenoplia S.I. Medvedev, 1952 222
Xenopopillia H.J. Kolbe, 1910 352
Xenoserica D. Ahrens, 2005 317
Xeragrilus Alexeev, 1998 524
Xerasiobia S.I. Medvedev, 1951 235
Xestotrogus Reitter, 1902 280
Xizangia Y.-W. Zhang, 1988 73
Xorema Reitter, 1902 282

Xyloscaptes Prell, 1934 358
Xylotrupes Hope, 1837 359
Xynophron Harold, 1880 167
Yamina Kerremans, 1903 517
Yanovalgus Nomura, 1952 412
Yoshitomia Klausnitzer, 2013 424
Ypsiloncyphon Klausnitzer, 2009 424
Yrdops Steffan, 1961 603
Zaitzevia G.C. Champion, 1923 603
Zaitzeviaria Nomura, 1959 603
Zantheumia Stephens, 1829 250
Zarudniola Semenov & Martynov, 1925
Zaspilota Casey, 1915 341
Zhangia Bunalski, 2002 230
Zoolrecordia Hołyński, 2006 466
Zuninoeus López-Colón, 1989 47

431